289 lines
37 KiB
XML
289 lines
37 KiB
XML
<document id="836CAA3D679EE9ADAAF615E1234DEA11" ID-CLB-Dataset="7165" ID-DOI="10.11646/zootaxa.4958.1.11" ID-GBIF-Dataset="ef663e33-b442-492e-8d5c-a373429ef059" ID-ISSN="1175-5326" ID-Zenodo-Dep="4691515" ID-ZooBank="ADD5204B-A342-4A85-8F10-778241D70E9E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618457602964" checkinUser="plazi" docAuthor="Galindo-Malagón, Ximena Alejandra, Morales, Irina & Moreira, Felipe Ferraz Figueiredo" docDate="2021" docId="E7216366DF1E5A79FF35B9C0FDAD688B" docLanguage="en" docName="zootaxa.4958.1.11.pdf" docOrigin="Zootaxa 4958 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhagovelia graziae Galindo-Malagón & Morales & Moreira 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="199" masterDocId="1B181B1EDF015A59FFA2B957FFCD6926" masterDocTitle="Revision of the Rhagovelia angustipes complex (Insecta: Hemiptera: Veliidae from Colombia" masterLastPageNumber="225" masterPageNumber="167" pageNumber="198" updateTime="1698936821219" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="DB4A3D1BF0693FCE86F2279CFF7394A7">
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<mods:title id="7E30B64DA71C3E0B7DBAC5E60012EFC5">Revision of the Rhagovelia angustipes complex (Insecta: Hemiptera: Veliidae from Colombia</mods:title>
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<mods:name id="8D458DF99D4C31B83EDC0A2DE9DBAAA8" type="personal">
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<mods:roleTerm id="F058C68AE90D07F8AF170C358731DA72">Author</mods:roleTerm>
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<mods:namePart id="5FDF0EC09E42E4441F46CBE634FD2041">Galindo-Malagón, Ximena Alejandra</mods:namePart>
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<mods:nameIdentifier id="17586D0C8DB79010DF0C7BC37EF3AA55" type="ORCID">0000-0003-0617-9403</mods:nameIdentifier>
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<mods:affiliation id="CD661B523B281B882A846335840DBCF8">Grupo de Investigación Sistemática Biológica-SisBio, Laboratorio de Entomología, Universidad Pedagógica y Tecnológica de Colombia, Tunja, BY, Colombia. & ximena. galindo @ uptc. edu. co; https: // orcid. org / 0000 - 0003 - 0617 - 9403</mods:affiliation>
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<mods:nameIdentifier id="A27A064EB3D535A682C31F52B54524A2" type="email">ximena.galindo@uptc.edu.co</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="96EFC8A06729AE73AD995C516610FABB">Author</mods:roleTerm>
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<mods:namePart id="CA6877A346964A4AB17F3E49CD6A5C3D">Morales, Irina</mods:namePart>
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<mods:nameIdentifier id="BA94ADF14F4C6BA6CE6A5139E7E979CB" type="ORCID">0000-0003-2456-5674</mods:nameIdentifier>
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<mods:affiliation id="4D0AA4CEC243D70A8C7D894ECD4C2EF6">Grupo de Investigación Sistemática Biológica-SisBio, Laboratorio de Entomología, Universidad Pedagógica y Tecnológica de Colombia, Tunja, BY, Colombia. & irina. morales @ uptc. edu. co; https: // orcid. org / 0000 - 0003 - 2456 - 5674</mods:affiliation>
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<mods:nameIdentifier id="6244A5425BA6A018888693539FF3468D" type="email">irina.morales@uptc.edu.co</mods:nameIdentifier>
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<mods:roleTerm id="E63724E8090A37EE59D2AA9DF54066A5">Author</mods:roleTerm>
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<mods:namePart id="3D00EF7FDF316BDC82008FACECDE078D">Moreira, Felipe Ferraz Figueiredo</mods:namePart>
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<mods:affiliation id="70A5CDF8E429FF609E22A1466020A1E3">Laboratório de Biodiversidade Entomológica, Instituto Oswaldo Cruz, Fundação Oswaldo Cruz, Rio de Janeiro, RJ, Brasil.</mods:affiliation>
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<mods:title id="562DB9B9EB459A8F7A6F28C1AE2C0826">Zootaxa</mods:title>
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<mods:part id="E35D4030634D575D6A88285D2B5EA8EA">
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<mods:date id="DC6A8D2A2781A8A18954D47CA96D343C">2021</mods:date>
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<mods:number id="B602E313A9269E50F1EC25CA32AFB6A7">2021-04-14</mods:number>
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<mods:number id="3FFCD4132B174CF51BB698D9EE2A633C">4958</mods:number>
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<mods:identifier id="20191ABB3DCA9AEFDD204BE0BF8A2BED" type="DOI">10.11646/zootaxa.4958.1.11</mods:identifier>
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<treatment id="E7216366DF1E5A79FF35B9C0FDAD688B" ID-DOI="http://doi.org/10.5281/zenodo.4691763" ID-GBIF-Taxon="181653115" ID-Zenodo-Dep="4691763" LSID="urn:lsid:plazi:treatment:E7216366DF1E5A79FF35B9C0FDAD688B" httpUri="http://treatment.plazi.org/id/E7216366DF1E5A79FF35B9C0FDAD688B" lastPageId="32" lastPageNumber="199" pageId="31" pageNumber="198">
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<subSubSection id="279281FBDF1E5A46FF35B9C0FE296997" box="[151,484,151,178]" pageId="31" pageNumber="198" type="nomenclature">
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<paragraph id="6F37D270DF1E5A46FF35B9C0FE296997" blockId="31.[151,624,151,213]" box="[151,484,151,178]" pageId="31" pageNumber="198">
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<heading id="347F651CDF1E5A46FF35B9C0FE296997" bold="true" box="[151,484,151,178]" fontSize="11" level="1" pageId="31" pageNumber="198" reason="1">
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<emphasis id="5DFC0E62DF1E5A46FF35B9C0FE296997" bold="true" box="[151,484,151,178]" pageId="31" pageNumber="198">
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<taxonomicName id="A888A9F3DF1E5A46FF35B9C0FEB26997" authority="Galindo-Malagón & Morales & Moreira, 2021" authorityName="Galindo-Malagón & Morales & Moreira" authorityYear="2021" box="[151,383,151,178]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
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<emphasis id="5DFC0E62DF1E5A46FF35B9C0FEB26997" bold="true" box="[151,383,151,178]" italics="true" pageId="31" pageNumber="198">Rhagovelia graziae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="46CFB319DF1E5A46FE25B9C0FE296997" box="[391,484,151,177]" pageId="31" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="279281FBDF1E5A46FF35B9ECFDBD69F3" box="[151,624,187,213]" pageId="31" pageNumber="198" type="description">
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<paragraph id="6F37D270DF1E5A46FF35B9ECFDBD69F3" blockId="31.[151,624,151,213]" box="[151,624,187,213]" pageId="31" pageNumber="198">
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(
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<figureCitation id="F7B3CEF5DF1E5A46FF3DB9ECFECD69F3" box="[159,256,187,213]" captionStart="FIGURE 9" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[280,1306,182,1877]" captionTargetId="figure-26@26.[280,1307,181,1887]" captionTargetPageId="26" captionText="FIGURE 9. Males of the Rhagovelia bisignata and hambletoni groups recorded from Colombia, dorsal view: A. R. caunapi (holotype); B. R. cimarrona (holotype); C. R. gastrotricha. D. R. grandis (holotype); E. R. graziae sp. nov. (holotype); F. R. longipes." figureDoi="http://doi.org/10.5281/zenodo.4691545" httpUri="https://zenodo.org/record/4691545/files/figure.png" pageId="31" pageNumber="198">Figs. 9E</figureCitation>
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,
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||
<figureCitation id="F7B3CEF5DF1E5A46FEAEB9ECFEF669F3" box="[268,315,187,213]" captionStart="FIGURE 10" captionStartId="27.[151,250,1936,1960]" captionTargetBox="[261,1322,186,1907]" captionTargetId="figure-17@27.[261,1327,181,1912]" captionTargetPageId="27" captionText="FIGURE 10. Males of the Rhagovelia bisignata and hambletoni groups recorded from Colombia, ventral view: A. R. caunapi (holotype); B. R. cimarrona (holotype); C. R. gastrotricha. D. R. grandis (holotype); E. R. graziae sp. nov. (holotype); F. R. longipes." figureDoi="http://doi.org/10.5281/zenodo.4691549" httpUri="https://zenodo.org/record/4691549/files/figure.png" pageId="31" pageNumber="198">10E</figureCitation>
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,
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<figureCitation id="F7B3CEF5DF1E5A46FEE5B9ECFEBB69F3" box="[327,374,187,213]" captionStart="FIGURE 11" captionStartId="28.[151,250,1913,1937]" captionTargetBox="[286,1296,185,1881]" captionTargetId="figure-26@28.[286,1301,181,1890]" captionTargetPageId="28" captionText="FIGURE 11. Females of the Rhagovelia bisignata and hambletoni groups recorded from Colombia, dorsal view: A. R. caunapi (holotype); B. R. cimarrona (paratype); C. R. gastrotricha. D. R. grandis (paratype); E. R. graziae sp. nov. (paratype); F. R. longipes." figureDoi="http://doi.org/10.5281/zenodo.4691553" httpUri="https://zenodo.org/record/4691553/files/figure.png" pageId="31" pageNumber="198">11E</figureCitation>
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,
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<figureCitation id="F7B3CEF5DF1E5A46FE23B9ECFE7D69F3" box="[385,432,187,213]" captionStart="FIGURE 12" captionStartId="29.[151,250,1925,1949]" captionTargetBox="[243,1345,183,1892]" captionTargetId="figure-17@29.[239,1347,181,1901]" captionTargetPageId="29" captionText="FIGURE 12. Females of the Rhagovelia bisignata and hambletoni groups recorded from Colombia, ventral view:A. R. caunapi (holotype); B. R. cimarrona (paratype); C. R. gastrotricha. D. R. grandis (paratype); E. R. graziae sp. nov. (paratype); F. R. longipes." figureDoi="http://doi.org/10.5281/zenodo.4691557" httpUri="https://zenodo.org/record/4691557/files/figure.png" pageId="31" pageNumber="198">12E</figureCitation>
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,
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<figureCitation id="F7B3CEF5DF1E5A46FE1EB9ECFE2269F3" box="[444,495,187,213]" captionStart="FIGURE 19" captionStartId="45.[151,250,1854,1878]" captionTargetBox="[190,1403,190,1822]" captionTargetId="figure-17@45.[184,1404,181,1830]" captionTargetPageId="45" captionText="FIGURE 19. Parameres of species of the Rhagovelia angustipes complex recorded from Colombia: A. R. arcuata; B. R. colombiana; C. R. nuqui; D. R. plumbea; E. R. tintipan; F. R. angustipes; G. R. boyacensis sp. nov. (paratype); H. R. calceola; I. R. calopa; J. R. cardia (paratype); K. R. cimarrona (paratype); L. R. gastrotricha; M. R. grandis (paratype); N. R. graziae sp. nov. (paratype); O. R. longipes; P. R. molanoi sp. nov. (paratype); Q. R. rosensis (paratype); R. R. santanderi; S. R. spinosa; T. R. tantilla; U. R. tenuipes V. R. guachiconoense (modified from: Padilla-Gil, 2019b)." figureDoi="http://doi.org/10.5281/zenodo.4691583" httpUri="https://zenodo.org/record/4691583/files/figure.png" pageId="31" pageNumber="198">19N</figureCitation>
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,
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<figureCitation id="F7B3CEF5DF1E5A46FE58B9ECFDE369F3" box="[506,558,187,213]" captionStart="FIGURE 20" captionStartId="46.[151,250,1862,1886]" captionTargetBox="[420,1169,189,1826]" captionTargetId="figure-26@46.[408,1179,181,1837]" captionTargetPageId="46" captionText="FIGURE 20. Male proctiger of species of the Rhagovelia angustipes complex recorded from Colombia: A. R. arcuata; B. R. colombiana; C. R. nuqui; D. R. plumbea; E. R. tintipan; F. R. angustipes; G. R. boyacensis sp. nov. (paratype); H. R. calceola; I. R. calopa; J. R. cardia (paratype); K. R. cimarrona (paratype); L. R. gastrotricha; M. R. grandis (paratype); N. R. graziae sp. nov. (paratype); O. R. longipes; P. R. molanoi sp. nov. (paratype); Q. R. rosensis (paratype); R. R. santanderi; S. R. spinosa; T. R. tantilla; U. R. tenuipes; V. R. guachiconoense (modified from: Padilla-Gil, 2019b)." figureDoi="http://doi.org/10.5281/zenodo.4691587" httpUri="https://zenodo.org/record/4691587/files/figure.png" pageId="31" pageNumber="198">20N</figureCitation>
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,
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<figureCitation id="F7B3CEF5DF1E5A46FD9BB9ECFDA569F3" box="[569,616,187,213]" captionStart="FIGURE 23" captionStartId="51.[151,250,1923,1947]" captionTargetBox="[290,1296,189,1891]" captionTargetId="figure-17@51.[286,1301,181,1899]" captionTargetPageId="51" captionText="FIGURE 23. Geographic distribution of species of the Rhagovelia bisignata and hambletoni groups in Colombia: A. R. barbacoensis, R. caunapi, R. cimarrona, R. guachiconoense; B. R. grandis, R. penta, R. rosensis; C. R. boyacensis sp. nov., R. graziae sp. nov., R. molanoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4691597" httpUri="https://zenodo.org/record/4691597/files/figure.png" pageId="31" pageNumber="198">23C</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="279281FBDF1E5A46FF35B854FD646F9B" pageId="31" pageNumber="198" type="materials_examined">
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<paragraph id="6F37D270DF1E5A46FF35B854FDC968AF" blockId="31.[151,1437,259,2014]" pageId="31" pageNumber="198">
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<materialsCitation id="DFE0D82DDF1E5A46FF35B854FDC968AF" ID-GBIF-Occurrence="3091613369" collectionCode="III, PL, PW" pageId="31" pageNumber="198" specimenCount="1" typeStatus="holotype">
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<emphasis id="5DFC0E62DF1E5A46FF35B854FE70683B" bold="true" box="[151,445,259,285]" pageId="31" pageNumber="198">
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<typeStatus id="B0336CD2DF1E5A46FF35B854FEC9683B" box="[151,260,259,285]" pageId="31" pageNumber="198" type="holotype">Holotype</typeStatus>
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apterous male.
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</emphasis>
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BL 2.72; HL 0.33; HW 0.70; INT 0.16; ANT I 0.75, ANT II 0.38, ANT
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<collectionCode id="09994AB5DF1E5A46FB5DB853FAD66838" box="[1279,1307,260,286]" pageId="31" pageNumber="198">III</collectionCode>
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0.46, ANT IV 0.47; EYE 0.27;
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<collectionCode id="09994AB5DF1E5A46FEDEB87FFE516864" box="[380,412,296,322]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="31" pageNumber="198" type="Museum">PL</collectionCode>
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0.19;
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<collectionCode id="09994AB5DF1E5A46FE46B87FFDC06864" box="[484,525,296,322]" country="Thailand" name="Paleontological Collections" pageId="31" pageNumber="198">PW</collectionCode>
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||
0.75; FORELEG: FEM 0.86; TIB 0.91; TAR I 0.02; TAR II 0.01; TAR
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||
<collectionCode id="09994AB5DF1E5A46FAE2B87FFA916864" box="[1344,1372,296,322]" pageId="31" pageNumber="198">III</collectionCode>
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0.21; MIDLEG: FEM 1.53; TIB 1.01; TAR I 0.07; TAR II 0.52; TAR
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<collectionCode id="09994AB5DF1E5A46FCCDB81BFC466840" box="[879,907,332,358]" pageId="31" pageNumber="198">III</collectionCode>
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0.67; HINDLEG: FEM 1.12; TIB 1.17; TAR I 0.04; TAR II 0.09; TAR
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<collectionCode id="09994AB5DF1E5A46FE0BB827FE0868AC" box="[425,453,368,394]" pageId="31" pageNumber="198">III</collectionCode>
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0.27.
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</materialsCitation>
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<paragraph id="6F37D270DF1E5A46FF65B8C4FC8B6A1C" blockId="31.[151,1437,259,2014]" pageId="31" pageNumber="198">
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Head dorsally black, covered with golden pubescence; longitudinal midline and a pair of oblique indentations at base impressed and shiny.Venter of head black. Buccula yellow. Labium brown. Eye dark red.Antenniferous tubercle brown. Antennomere I yellow for almost basal half; apex of I and rest of antenna brown. Pronotum dark orange between eyes behind vertex of head; dark brown laterally and posteriorly. Meso- and metanota black, covered with golden pubescence. Pro-, meso- and metapleura black, covered with greyish pubescence. Pro, meso- and metasterna black, covered with greyish pubescence. Pro-, meso- and metacetabula yellow. Coxae and trochanters yellow. Fore femur with yellow base, then black; middle femur black; hind femur with short yellow base, then black. Tibiae and tarsi black. Abdominal mediotergites black, covered with golden pubescence; VII with shiny black central spot; tergum VIII shiny black, covered with short golden setae. Abdominal laterotergites black, covered with golden pubescence, with lateral margins shiny black. Abdominal sterna black, covered with greyish pubescence, with a median comb of long golden setae on segments II–
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<collectionCode id="09994AB5DF1E5A46FD71BBABFD3F6A30" box="[723,754,764,790]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="31" pageNumber="198">VI</collectionCode>
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; VII shiny black medially, with weak carina posteriorly; VIII light brown, with central carina and long lateral golden setae.
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</paragraph>
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<paragraph id="6F37D270DF1E5A46FF65BA14FDE66C73" blockId="31.[151,1437,259,2014]" pageId="31" pageNumber="198">
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Head short, covered with short setae; frons with longer setae. Antenna covered with brown setae, denser on antennomere IV; antennomere I with at least six long, thick brown setae. Antennomeres I–
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||
<collectionCode id="09994AB5DF1E5A46FB73BA3FFB206AA4" box="[1233,1261,872,898]" pageId="31" pageNumber="198">III</collectionCode>
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cylindrical; IV fusiform; I and IV wider at the middle than II–
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<collectionCode id="09994AB5DF1E5A46FD02BADBFD706A80" box="[672,701,908,934]" pageId="31" pageNumber="198">III</collectionCode>
|
||
, which are subequal in width. Labium short. Ocular setae present. Pronotum short, not covering mesonotum, covered with short golden setae, denser laterally, posterior margin slightly concave. Mesonotum covered with short golden setae, denser on posterior margin; posterior margin convex centrally. Metanotum short; posterior margin straight centrally. Metasternum covered with long golden setae. Sides of thorax with long golden setae. Legs covered with short setae, with rows of longer thicker setae on femora and tibiae. Trochanters without spines. Fore tibia slightly curved, with weak preapical depression; grasping comb extending slightly beyond apex. Hind femur slightly surpassing apex of abdomen, wider than middle femur, with posterior margin sinuous; distal half with row of about 7 spines decreasing in size towards apex. Hind tibia straight, apically with 4 subequal short denticles at posterior surface, apex with straight spur. Abdominal mediotergites subrectangular. Abdominal laterotergites raised, but not vertical, with short golden setae. Abdominal sterna II–
|
||
<collectionCode id="09994AB5DF1E5A46FA2ABD87FA516DCC" box="[1416,1436,1232,1258]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="198" type="Museum">V</collectionCode>
|
||
covered with long golden setae medially. Proctiger subtriangular, dorsum with thick, long brown setae; basal lobes strong, short; apex rounded, densely covered with setae. Paramere elongated, basal part oval with long setae on the upper margin, apex acute, elongated.
|
||
</paragraph>
|
||
<paragraph id="6F37D270DF1E5A46FF65BC08FD9C6CC3" blockId="31.[151,1437,259,2014]" pageId="31" pageNumber="198">
|
||
<emphasis id="5DFC0E62DF1E5A46FF65BC08FDCE6C5F" bold="true" box="[199,515,1375,1401]" pageId="31" pageNumber="198">
|
||
<typeStatus id="B0336CD2DF1E5A46FF65BC08FEF96C5F" box="[199,308,1375,1401]" pageId="31" pageNumber="198" type="paratype">Paratype</typeStatus>
|
||
apterous female.
|
||
</emphasis>
|
||
BL 3.00; HL 0.39; HW 0.75; INT 0.16; ANT I 0.74, ANT II 0.38, ANT
|
||
<collectionCode id="09994AB5DF1E5A46FAE2BC37FA916C5C" box="[1344,1372,1376,1402]" pageId="31" pageNumber="198">III</collectionCode>
|
||
0.44, ANT IV 0.41; EYE 0.29;
|
||
<collectionCode id="09994AB5DF1E5A46FE1FBCD3FE106CB8" box="[445,477,1412,1438]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="31" pageNumber="198" type="Museum">PL</collectionCode>
|
||
0.20;
|
||
<collectionCode id="09994AB5DF1E5A46FD87BCD3FD836CB8" box="[549,590,1412,1438]" country="Thailand" name="Paleontological Collections" pageId="31" pageNumber="198">PW</collectionCode>
|
||
0.83; FORELEG: FEM 0.82; TIB 0.83; TAR I 0.05; TAR II 0.02; TAR
|
||
<collectionCode id="09994AB5DF1E5A46FA22BCD3FA516CB8" box="[1408,1436,1412,1438]" pageId="31" pageNumber="198">III</collectionCode>
|
||
0.24; MIDLEG: FEM 1.52; TIB 1.06; TAR I 0.09; TAR II 0.49; TAR
|
||
<collectionCode id="09994AB5DF1E5A46FC1ABCFFFC196CE4" box="[952,980,1448,1474]" pageId="31" pageNumber="198">III</collectionCode>
|
||
0.63; HINDLEG: FEM 1.13; TIB 1.20; TAR I 0.05; TAR II 0.09; TAR
|
||
<collectionCode id="09994AB5DF1E5A46FE54BC9BFDDF6CC0" box="[502,530,1484,1510]" pageId="31" pageNumber="198">III</collectionCode>
|
||
0.27.
|
||
</paragraph>
|
||
<paragraph id="6F37D270DF1E5A46FF65BCB8FD646F9B" blockId="31.[151,1437,259,2014]" pageId="31" pageNumber="198">
|
||
Similar to apterous male in structure and color. Hind femur slightly thinner than in male, with 5–6 spines on distal half. Abdominal mediotergites with abundant short silvery setae; I raised posteriorly; II not raised, in the same plane as
|
||
<collectionCode id="09994AB5DF1E5A46FF5EBF6FFED46F74" box="[252,281,1592,1618]" pageId="31" pageNumber="198">III</collectionCode>
|
||
–
|
||
<collectionCode id="09994AB5DF1E5A46FE8BBF6FFE856F74" box="[297,328,1592,1618]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="31" pageNumber="198">VI</collectionCode>
|
||
; dorsum of segments VII–VIII shiny black centrally. Abdominal laterotergites covered with short setae, slightly sloping downward and merging with apex of tergum VIII; lateral margins of
|
||
<collectionCode id="09994AB5DF1E5A46FB21BF0BFB526F50" box="[1155,1183,1628,1654]" pageId="31" pageNumber="198">III</collectionCode>
|
||
–
|
||
<collectionCode id="09994AB5DF1E5A46FB0FBF0BFB0C6F50" box="[1197,1217,1628,1654]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="198" type="Museum">V</collectionCode>
|
||
with abundant long golden setae; IV–
|
||
<collectionCode id="09994AB5DF1E5A46FEF8BFD7FEBA6FBC" box="[346,375,1664,1690]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="31" pageNumber="198">VI</collectionCode>
|
||
broader than anterior and posterior segments.Abdominal sterna without carina or comb of setae medially; VII with shiny brown mark medially.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="279281FBDF1E5A46FF65BF90FD6B6FC7" box="[199,678,1735,1762]" pageId="31" pageNumber="198" type="distribution">
|
||
<paragraph id="6F37D270DF1E5A46FF65BF90FD6B6FC7" blockId="31.[151,1437,259,2014]" box="[199,678,1735,1762]" pageId="31" pageNumber="198">
|
||
<emphasis id="5DFC0E62DF1E5A46FF65BF90FE946FC7" bold="true" box="[199,345,1735,1761]" pageId="31" pageNumber="198">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="179F92E0DF1E5A46FEC5BF90FE176FC7" box="[359,474,1735,1761]" name="Colombia" pageId="31" pageNumber="198">Colombia</collectingCountry>
|
||
:
|
||
<emphasis id="5DFC0E62DF1E5A46FE47BF90FDEE6FC7" bold="true" box="[485,547,1735,1761]" pageId="31" pageNumber="198">
|
||
<collectingRegion id="AD4C1C92DF1E5A46FE47BF90FDEE6FC7" box="[485,547,1735,1761]" country="Colombia" name="Meta" pageId="31" pageNumber="198">Meta</collectingRegion>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="F7B3CEF5DF1E5A46FD90BF90FD546FC7" box="[562,665,1735,1761]" captionStart="FIGURE 23" captionStartId="51.[151,250,1923,1947]" captionTargetBox="[290,1296,189,1891]" captionTargetId="figure-17@51.[286,1301,181,1899]" captionTargetPageId="51" captionText="FIGURE 23. Geographic distribution of species of the Rhagovelia bisignata and hambletoni groups in Colombia: A. R. barbacoensis, R. caunapi, R. cimarrona, R. guachiconoense; B. R. grandis, R. penta, R. rosensis; C. R. boyacensis sp. nov., R. graziae sp. nov., R. molanoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4691597" httpUri="https://zenodo.org/record/4691597/files/figure.png" pageId="31" pageNumber="198">Fig. 23C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="279281FBDF1E5A79FF65BFBCFD9B69F3" lastPageId="32" lastPageNumber="199" pageId="31" pageNumber="198" type="discussion">
|
||
<paragraph id="6F37D270DF1E5A79FF65BFBCFD9B69F3" blockId="31.[151,1437,259,2014]" lastBlockId="32.[151,1437,151,429]" lastPageId="32" lastPageNumber="199" pageId="31" pageNumber="198">
|
||
<emphasis id="5DFC0E62DF1E5A46FF65BFBCFE856E23" bold="true" box="[199,328,1771,1797]" pageId="31" pageNumber="198">Comments</emphasis>
|
||
.
|
||
<taxonomicName id="A888A9F3DF1E5A46FEF5BFBCFDFF6E23" authority="Galindo-Malagón & Morales & Moreira, 2021" authorityName="Galindo-Malagón & Morales & Moreira" authorityYear="2021" box="[343,562,1771,1797]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
|
||
<emphasis id="5DFC0E62DF1E5A46FEF5BFBCFDFF6E23" box="[343,562,1771,1797]" italics="true" pageId="31" pageNumber="198">Rhagovelia graziae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DFC0E62DF1E5A46FD98BFBCFD596E23" bold="true" box="[570,660,1771,1797]" pageId="31" pageNumber="198">
|
||
<taxonomicNameLabel id="46CFB319DF1E5A46FD98BFBCFD596E23" box="[570,660,1771,1797]" pageId="31" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a relatively small species, with body length ~
|
||
<quantity id="A8707F95DF1E5A46FB16BFBCFAC96E23" box="[1204,1284,1771,1798]" metricMagnitude="-2" metricUnit="m" metricValue="6.985" pageId="31" pageNumber="198" unit="in" value="2.75">2.75 in</quantity>
|
||
the male and ~3.00 in the female. Other species within this size range are
|
||
<taxonomicName id="A888A9F3DF1E5A46FCF6BE47FC1C6E0F" authority="Galindo-Malagón & Morales & Moreira, 2021" authorityName="Galindo-Malagón & Morales & Moreira" authorityYear="2021" box="[852,977,1807,1833]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="molanoi" status="sp. nov.">
|
||
<emphasis id="5DFC0E62DF1E5A46FCF6BE47FC1C6E0F" box="[852,977,1807,1833]" italics="true" pageId="31" pageNumber="198">R. molanoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DFC0E62DF1E5A46FC78BE58FBF86E0F" bold="true" box="[986,1077,1807,1833]" pageId="31" pageNumber="198">
|
||
<taxonomicNameLabel id="46CFB319DF1E5A46FC78BE58FBF86E0F" box="[986,1077,1807,1833]" pageId="31" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="A888A9F3DF1E5A46FBE7BE47FB0F6E0F" authorityName="Padilla-Gil" authorityYear="2011" box="[1093,1218,1808,1833]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="rosensis">
|
||
<emphasis id="5DFC0E62DF1E5A46FBE7BE47FB0F6E0F" box="[1093,1218,1808,1833]" italics="true" pageId="31" pageNumber="198">R. rosensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A888A9F3DF1E5A46FB70BE47FAA66E0F" authorityName="Padilla-Gil" authorityYear="2015" box="[1234,1387,1807,1833]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="santanderi">
|
||
<emphasis id="5DFC0E62DF1E5A46FB70BE47FAA66E0F" box="[1234,1387,1807,1833]" italics="true" pageId="31" pageNumber="198">R. santanderi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A888A9F3DF1E5A46FF35BE63FEC46E6B" authorityName="Drake & Harris" authorityYear="1933" box="[151,265,1843,1869]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="tantilla">
|
||
<emphasis id="5DFC0E62DF1E5A46FF35BE63FEC46E6B" box="[151,265,1843,1869]" italics="true" pageId="31" pageNumber="198">R. tantilla</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="A888A9F3DF1E5A46FEBBBE64FE3F6E6B" authorityName="Drake & Harris" authorityYear="1933" box="[281,498,1843,1869]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="tantilla">
|
||
<emphasis id="5DFC0E62DF1E5A46FEBBBE64FE3F6E6B" box="[281,498,1843,1869]" italics="true" pageId="31" pageNumber="198">Rhagovelia tantilla</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A888A9F3DF1E5A46FD8EBE63FD646E6B" authority="Galindo-Malagón & Morales & Moreira, 2021" authorityName="Galindo-Malagón & Morales & Moreira" authorityYear="2021" box="[556,681,1843,1869]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="molanoi" status="sp. nov.">
|
||
<emphasis id="5DFC0E62DF1E5A46FD8EBE63FD646E6B" box="[556,681,1843,1869]" italics="true" pageId="31" pageNumber="198">R. molanoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DFC0E62DF1E5A46FD10BE64FCC06E6B" bold="true" box="[690,781,1843,1869]" pageId="31" pageNumber="198">
|
||
<taxonomicNameLabel id="46CFB319DF1E5A46FD10BE64FCC06E6B" box="[690,781,1843,1869]" pageId="31" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be readily distinguished from the others by the male hind tibia unarmed, without small spines throughout length or apical spur.
|
||
<taxonomicName id="A888A9F3DF1E5A46FBA2BE00FB136E57" authority="Galindo-Malagón & Morales & Moreira, 2021" authorityName="Galindo-Malagón & Morales & Moreira" authorityYear="2021" box="[1024,1246,1879,1905]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
|
||
<emphasis id="5DFC0E62DF1E5A46FBA2BE00FB136E57" box="[1024,1246,1879,1905]" italics="true" pageId="31" pageNumber="198">Rhagovelia graziae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DFC0E62DF1E5A46FB4BBE00FA8B6E57" bold="true" box="[1257,1350,1879,1905]" pageId="31" pageNumber="198">
|
||
<taxonomicNameLabel id="46CFB319DF1E5A46FB4BBE00FA8B6E57" box="[1257,1350,1879,1905]" pageId="31" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="A888A9F3DF1E5A46FECDBE2BFE256EB3" authorityName="Padilla-Gil" authorityYear="2011" box="[367,488,1916,1941]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="rosensis">
|
||
<emphasis id="5DFC0E62DF1E5A46FECDBE2BFE256EB3" box="[367,488,1916,1941]" italics="true" pageId="31" pageNumber="198">R. rosensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A888A9F3DF1E5A46FDBEBE2BFD7F6EB3" authorityName="Padilla-Gil" authorityYear="2015" box="[540,690,1915,1941]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="198" phylum="Arthropoda" rank="species" species="santanderi">
|
||
<emphasis id="5DFC0E62DF1E5A46FDBEBE2BFD7F6EB3" box="[540,690,1915,1941]" italics="true" pageId="31" pageNumber="198">R. santanderi</emphasis>
|
||
</taxonomicName>
|
||
by the coxae and trochanters yellow (vs. dark brown to black in the other two), male abdominal sterna II–V with a median comb of long golden setae (vs. only with short setae), male abdominal segment VIII large and robust (vs. small), the shape of the paramere (
|
||
<figureCitation id="F7B3CEF5DF1E5A46FB8EBE94FB586EFB" box="[1068,1173,1987,2013]" captionStart="FIGURE 19" captionStartId="45.[151,250,1854,1878]" captionTargetBox="[190,1403,190,1822]" captionTargetId="figure-17@45.[184,1404,181,1830]" captionTargetPageId="45" captionText="FIGURE 19. Parameres of species of the Rhagovelia angustipes complex recorded from Colombia: A. R. arcuata; B. R. colombiana; C. R. nuqui; D. R. plumbea; E. R. tintipan; F. R. angustipes; G. R. boyacensis sp. nov. (paratype); H. R. calceola; I. R. calopa; J. R. cardia (paratype); K. R. cimarrona (paratype); L. R. gastrotricha; M. R. grandis (paratype); N. R. graziae sp. nov. (paratype); O. R. longipes; P. R. molanoi sp. nov. (paratype); Q. R. rosensis (paratype); R. R. santanderi; S. R. spinosa; T. R. tantilla; U. R. tenuipes V. R. guachiconoense (modified from: Padilla-Gil, 2019b)." figureDoi="http://doi.org/10.5281/zenodo.4691583" httpUri="https://zenodo.org/record/4691583/files/figure.png" pageId="31" pageNumber="198">Fig. 19N</figureCitation>
|
||
), the lateral margins of female abdomen sinuous (vs. evenly tapering or bowed), and the posterior margin of female abdominal tergum VIII with long setae (vs. without long setae).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="279281FBDF215A79FF65B988FDF86867" pageId="32" pageNumber="199" type="etymology">
|
||
<paragraph id="6F37D270DF215A79FF65B988FDF86867" blockId="32.[151,1437,151,429]" pageId="32" pageNumber="199">
|
||
<emphasis id="5DFC0E62DF215A79FF65B988FE8569DF" bold="true" box="[199,328,223,249]" pageId="32" pageNumber="199">Etymology</emphasis>
|
||
. This species is named in honor of the Brazilian researcher Dr. Jocélia Grazia (Universidade Federal do
|
||
<collectingRegion id="AD4C1C92DF215A79FF1CB854FE526838" box="[190,415,259,286]" country="Brazil" name="Rio Grande do Sul" pageId="32" pageNumber="199">Rio Grande do Sul</collectingRegion>
|
||
), who has made innumerable contributions to the knowledge of
|
||
<taxonomicName id="A888A9F3DF215A79FB34B854FAD0683B" box="[1174,1309,259,285]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="32" pageNumber="199" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
|
||
, especially Pentatomoidea, from the Neotropics.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="279281FBDF215A79FF65B81CFDAD688B" pageId="32" pageNumber="199" type="materials_examined">
|
||
<paragraph id="6F37D270DF215A79FF65B81CFDAD688B" blockId="32.[151,1437,151,429]" pageId="32" pageNumber="199">
|
||
<emphasis id="5DFC0E62DF215A79FF65B81CFE226842" bold="true" box="[199,495,330,357]" pageId="32" pageNumber="199">Type material examined.</emphasis>
|
||
<materialsCitation id="DFE0D82DDF215A79FE54B81CFC2468AF" ID-GBIF-Occurrence="3091613515" collectingDate="2015-02-10" collectorName="Col & N. Torres'" country="Colombia" county="Puerto Gaitan" elevation="189" location="rio Iteviare" municipality="Puerto Trujillo" pageId="32" pageNumber="199" specimenCount="1" specimenCount-male="1" stateProvince="Meta" typeStatus="holotype">
|
||
<typeStatus id="B0336CD2DF215A79FE54B81CFD936843" box="[502,606,331,357]" pageId="32" pageNumber="199" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="798E19F9DF215A79FDC4B81CFDB66843" box="[614,635,331,357]" pageId="32" pageNumber="199" type="male">♂</specimenCount>
|
||
apterous (UPTC-In-0007): ‘
|
||
<collectingCountry id="179F92E0DF215A79FC15B81CFBE66843" box="[951,1067,331,357]" name="Colombia" pageId="32" pageNumber="199">Colombia</collectingCountry>
|
||
\
|
||
<collectingRegion id="AD4C1C92DF215A79FB98B81CFBBE6843" box="[1082,1139,331,357]" country="Colombia" name="Meta" pageId="32" pageNumber="199">Meta</collectingRegion>
|
||
\ municipio
|
||
<collectingCounty id="8656AAFCDF215A79FAA3B81CFA506843" box="[1281,1437,331,357]" pageId="32" pageNumber="199">Puerto Gaitán</collectingCounty>
|
||
\ vereda
|
||
<collectingMunicipality id="8F53480ADF215A79FF58B838FE5268AF" box="[250,415,367,393]" pageId="32" pageNumber="199">Puerto Trujillo</collectingMunicipality>
|
||
\
|
||
<location id="6A5784ABDF215A79FE15B838FDF968AF" LSID="urn:lsid:plazi:treatment:E7216366DF1E5A79FF35B9C0FDAD688B:6A5784ABDF215A79FE15B838FDF968AF" box="[439,564,367,393]" country="Colombia" county="Puerto Gaitan" municipality="Puerto Trujillo" name="rio Iteviare" pageId="32" pageNumber="199" stateProvince="Meta">río Iteviaré</location>
|
||
\
|
||
<quantity id="A8707F95DF215A79FDE9B838FD5E68AF" box="[587,659,367,393]" metricMagnitude="2" metricUnit="m" metricValue="1.89" pageId="32" pageNumber="199" unit="m" value="189.0">
|
||
<elevation id="E4A53543DF215A79FDE9B838FD5E68AF" box="[587,659,367,393]" metricMagnitude="2" metricUnit="m" metricValue="1.89" pageId="32" pageNumber="199" unit="m" value="189.0">189 m</elevation>
|
||
</quantity>
|
||
\
|
||
<date id="1B36F4B0DF215A79FD09B839FCD268AF" box="[683,799,366,393]" pageId="32" pageNumber="199" value="2015-02-10">
|
||
<collectingDate id="0B720D58DF215A79FD09B839FCD268AF" box="[683,799,366,393]" pageId="32" pageNumber="199" value="2015-02-10">10.II.2015</collectingDate>
|
||
</date>
|
||
\
|
||
<collectorName id="C27DB7A6DF215A79FC9AB838FCA868AF" box="[824,869,367,393]" pageId="32" pageNumber="199">Col</collectorName>
|
||
:
|
||
<collectorName id="C27DB7A6DF215A79FCD2B838FC2468AF" box="[880,1001,367,393]" pageId="32" pageNumber="199">N. Tórres’</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="B0336CD2DF215A79FC56B838FBAC68AF" box="[1012,1121,367,393]" pageId="32" pageNumber="199" type="paratype">Paratypes</typeStatus>
|
||
<specimenCount id="798E19F9DF215A79FBCBB838FB5968AF" box="[1129,1172,367,393]" pageId="32" pageNumber="199" type="male">1 ♂</specimenCount>
|
||
apterous,
|
||
<materialsCitation id="DFE0D82DDF215A79FAA8B838FD90688B" ID-GBIF-Occurrence="3091613409" collectingDate="2015-02-10" collectorName="Col & N. Torres'" country="Colombia" county="Puerto Gaitan" elevation="189" location="rio Iteviare" municipality="Puerto Trujillo" pageId="32" pageNumber="199" specimenCount="1" specimenCount-female="1" stateProvince="Meta" typeStatus="paratype">
|
||
<specimenCount id="798E19F9DF215A79FAA8B838FAF868AF" box="[1290,1333,367,393]" pageId="32" pageNumber="199" type="female">1 ♀</specimenCount>
|
||
apterous (UPTC-In-0008): same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |