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<document id="807B077F393144F076ECCC7A4B09A558" ID-CLB-Dataset="63559" ID-DOI="10.11646/zootaxa.5150.1.1" ID-GBIF-Dataset="cb0065b0-bad7-4fb5-b6cb-4920f80293cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="6609961" ID-ZooBank="0A7818EA-FBCE-4597-9557-28C208CAC97A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1654244705367" checkinUser="plazi" docAuthor="Souma, Jun &amp; Ishikawa, Tadashi" docDate="2022" docId="A202791FFFBFFFEC69F7F9A4FB5DFF49" docLanguage="en" docName="zootaxa.5150.1.1.pdf" docOrigin="Zootaxa 5150 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Physatocheila consueta Souma &amp; Ishikawa 2022, comb. nov." docType="treatment" docVersion="5" lastPageNumber="17" masterDocId="5E3B0167FFB0FFFD6960FFD7FFE7FFFB" masterDocTitle="Taxonomic review of the tingine genera Cysteochila, Hurdchila, Physatocheila, and Xynotingis from Japan, with description of a new genus (Hemiptera: Heteroptera: Tingidae)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="16" updateTime="1699338851664" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="3CFBBF9F07CCBE089580AFE45460EF69" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ED77F4FD72A942188BB38C429E05E0AF">
<mods:title id="76295A1415CC58ACF643257720833C29">Taxonomic review of the tingine genera Cysteochila, Hurdchila, Physatocheila, and Xynotingis from Japan, with description of a new genus (Hemiptera: Heteroptera: Tingidae)</mods:title>
</mods:titleInfo>
<mods:name id="652371F4C4A143E4ADF9DFA013077E0B" type="personal">
<mods:role id="7E80B3547327EE43CE07EF79366FBD01">
<mods:roleTerm id="80ED60D41F4C5B65E2C0F07694BCF414">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4B36BCEC061DF4E2F3A016E7E7C636FB">Souma, Jun</mods:namePart>
<mods:nameIdentifier id="21E7B97B907E47B9D5B2018EBDF001ED" type="ORCID">0000-0002-2238-5015</mods:nameIdentifier>
<mods:affiliation id="F235A3F34B9E29F1F5C56367A03C2D2C">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, Japan. kodokusignal @ gmail. com; https: // orcid. org / 0000 - 0002 - 2238 - 5015 &amp; Research Fellowship for Young Scientists (DC 1), Japan Society for the Promotion of Science, Tokyo, Japan</mods:affiliation>
<mods:nameIdentifier id="15C6989FF159CB58940F5BA1F1100BE5" type="email">kodokusignal@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="4903B87AFD33E7923D1356F5F774E520" type="personal">
<mods:role id="3C06368B8AB12AD02604C2E98EAD44D8">
<mods:roleTerm id="646052D34224DDF7F4F0B0A33E6C9A77">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7C354229E93E19FA6764C2551405394B">Ishikawa, Tadashi</mods:namePart>
<mods:affiliation id="697DD3A6C4F315B53C5E6374715F16B9">Laboratory of Entomology, Faculty of Agriculture, Tokyo University of Agriculture, Kanagawa, Japan.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1E5B5D695A909C9EC0C359402A8C9137">text</mods:typeOfResource>
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<mods:title id="02C8198084A799E01BDC41B6D78A83FB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="149B053F02F74F55DF0AFDC20A6F8653">
<mods:date id="9991460B973E67622D7CA16A4FEE82CB">2022</mods:date>
<mods:detail id="68F64ADA6C9770EF5065941931E92EBE" type="pubDate">
<mods:number id="8D3E34180BD14BADAA9E2CFFF2192136">2022-06-03</mods:number>
</mods:detail>
<mods:detail id="0F9D4434F553428E579C30FC6D01239B" type="volume">
<mods:number id="270B887B69C84315EA497E5F334984B7">5150</mods:number>
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<mods:number id="9271FC59EDF94424F9920A07964A3FE1">1</mods:number>
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</mods:extent>
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<mods:classification id="452DFEE1EF85DF2531A01F6CCE5FFCA1">journal article</mods:classification>
<mods:identifier id="295F56B62FECF6EB5C4B83F5C5F3F3E4" type="CLB-Dataset">63559</mods:identifier>
<mods:identifier id="EE5CF79CF1AA17B3B788DAE10E1A220F" type="DOI">10.11646/zootaxa.5150.1.1</mods:identifier>
<mods:identifier id="1D65F6DE639E11070A0FA8DA4D431752" type="GBIF-Dataset">cb0065b0-bad7-4fb5-b6cb-4920f80293cc</mods:identifier>
<mods:identifier id="427A75DF4387B9D422D29333E2947A57" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A202791FFFBFFFEC69F7F9A4FB5DFF49" ID-DOI="http://doi.org/10.5281/zenodo.6611330" ID-GBIF-Taxon="195729295" ID-Zenodo-Dep="6611330" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49" httpUri="http://treatment.plazi.org/id/A202791FFFBFFFEC69F7F9A4FB5DFF49" lastPageId="17" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="62B19B82FFBFFFF269F7F9A4FD1CF975" box="[151,763,1651,1678]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="2A14C809FFBFFFF269F7F9A4FD1CF975" blockId="15.[151,763,1651,1714]" box="[151,763,1651,1678]" pageId="15" pageNumber="16">
<heading id="715C7F65FFBFFFF269F7F9A4FD1CF975" bold="true" box="[151,763,1651,1678]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="18DF141BFFBFFFF269F7F9A4FD1CF975" bold="true" box="[151,763,1651,1678]" pageId="15" pageNumber="16">
<taxonomicName id="EDABB38AFFBFFFF269F7F9A4FD80F976" authority="(Drake, 1948)" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[151,615,1651,1678]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFBFFFF269F7F9A4FE53F975" bold="true" box="[151,436,1651,1678]" italics="true" pageId="15" pageNumber="16">Physatocheila consueta</emphasis>
(Drake, 1948)
</taxonomicName>
,
<taxonomicNameLabel id="03ECA960FFBFFFF26B15F9A3FD1CF975" box="[629,763,1652,1678]" pageId="15" pageNumber="16" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFBFFFF269F7F940FD6CF949" box="[151,651,1687,1714]" pageId="15" pageNumber="16" type="description">
<paragraph id="2A14C809FFBFFFF269F7F940FD6CF949" blockId="15.[151,763,1651,1714]" box="[151,651,1687,1714]" pageId="15" pageNumber="16">
(
<figureCitation id="B290D48CFFBFFFF269FFF940FEE5F949" box="[159,258,1687,1714]" captionStart="FIGURE 8" captionStartId="13.[151,250,1851,1875]" captionTargetBox="[157,1428,996,1820]" captionTargetId="figure-410@13.[152,1436,993,1828]" captionTargetPageId="13" captionText="FIGURE 8. Nine Physatocheila species from Japan. Dried specimens, dorsal view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi from Japan proper (E); P. fieberi from the Ryukyu Islands (F); P. miyatakei (G); P. nigrintegerrima (H); P. orientis (I); P. veteris (J). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6609989" httpUri="https://zenodo.org/record/6609989/files/figure.png" pageId="15" pageNumber="16">Figs. 8B</figureCitation>
,
<figureCitation id="B290D48CFFBFFFF2686DF94FFED7F949" box="[269,304,1688,1714]" captionStart="FIGURE 9" captionStartId="14.[151,250,1804,1828]" captionTargetBox="[151,1435,489,1780]" captionTargetId="figure-184@14.[151,1435,489,1780]" captionTargetPageId="14" captionText="FIGURE 9. Rostra and sternal laminae of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609991" httpUri="https://zenodo.org/record/6609991/files/figure.png" pageId="15" pageNumber="16">9B</figureCitation>
,
<figureCitation id="B290D48CFFBFFFF2685CF94FFE8AF949" box="[316,365,1688,1714]" captionStart="FIGURE 10" captionStartId="19.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1198]" captionTargetId="figure-17@19.[151,1436,181,1198]" captionTargetPageId="19" captionText="FIGURE 10. Male terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609995" httpUri="https://zenodo.org/record/6609995/files/figure.png" pageId="15" pageNumber="16">10B</figureCitation>
,
<figureCitation id="B290D48CFFBFFFF26819F94FFE4FF949" box="[377,424,1688,1714]" captionStart="FIGURE 11" captionStartId="21.[151,250,1579,1603]" captionTargetBox="[151,1436,525,1555]" captionTargetId="figure-182@21.[151,1436,525,1555]" captionTargetPageId="21" captionText="FIGURE 11. Female terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6610129" httpUri="https://zenodo.org/record/6610129/files/figure.png" pageId="15" pageNumber="16">11B</figureCitation>
,
<figureCitation id="B290D48CFFBFFFF268D4F94FFDEFF949" box="[436,520,1688,1714]" captionStart="FIGURE 13" captionStartId="25.[151,250,526,550]" captionTargetBox="[151,1436,181,502]" captionTargetId="figure-17@25.[151,1436,181,502]" captionTargetPageId="25" captionText="FIGURE 13. A living adult (A) and a living fifth instar nymph (B) of Physatocheila consueta comb. nov. from Ishigaki Island, the Ryukyu Islands, Japan (photographed indoor). Host plant (Ampelopsis glandulosa (Wall.) Momiy. var. hancei (Planch.) Momiy.) (C) of P. consueta comb. nov. on Ishigaki Island, the Ryukyu Islands, Japan." figureDoi="http://doi.org/10.5281/zenodo.6610008" httpUri="https://zenodo.org/record/6610008/files/figure.png" pageId="15" pageNumber="16">13A, B</figureCitation>
,
<figureCitation id="B290D48CFFBFFFF26B73F94FFDA2F949" box="[531,581,1688,1714]" captionStart="FIGURE 17" captionStartId="28.[151,250,1870,1894]" captionTargetBox="[156,1434,888,1845]" captionTargetId="figure-335@28.[151,1436,885,1847]" captionTargetPageId="28" captionText="FIGURE 17. Pygophores of eleven species of Cysteochila and its close relatives from Japan, dorsal view: C. abundantis (A); Limnostatua lewisi comb. nov. (B); Physatocheila chiniana comb. nov. (C); P. consueta comb. nov. (D); P. distinguenda (E); P. euthenia comb. nov. (F); P. fieberi from Japan proper (G); P. fieberi from the Ryukyu Islands (H); P. miyatakei (I); P. nigrintegerrima (J); P. orientis (K); P. veteris (L). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6610019" httpUri="https://zenodo.org/record/6610019/files/figure.png" pageId="15" pageNumber="16">17D</figureCitation>
,
<figureCitation id="B290D48CFFBFFFF26B32F94FFD64F949" box="[594,643,1688,1714]" captionStart="FIGURE 18" captionStartId="29.[151,250,1878,1902]" captionTargetBox="[154,1433,1025,1851]" captionTargetId="figure-333@29.[151,1436,1022,1854]" captionTargetPageId="29" captionText="FIGURE 18. Parameres of eleven species of Cysteochila and its close relatives from Japan, dorsal view: C. abundantis (A); Limnostatua lewisi comb. nov. (B); Physatocheila chiniana comb. nov. (C); P. consueta comb. nov. (D); P. distinguenda (E); P. euthenia comb. nov. (F); P. fieberi from Japan proper (G); P. fieberi from the Ryukyu Islands (H); P. miyatakei (I); P. nigrintegerrima (J); P. orientis (K); P. veteris (L). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6610022" httpUri="https://zenodo.org/record/6610022/files/figure.png" pageId="15" pageNumber="16">18D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFBFFFF269F7F900FD8CF892" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="2A14C809FFBFFFF269F7F900FC0EF914" blockId="15.[151,1001,1750,1775]" box="[151,1001,1750,1775]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="0ABBEF27FFBFFFF269F7F900FC0EF914" box="[151,1001,1750,1775]" pageId="15" pageNumber="16">
<treatmentCitation id="AB0AEE18FFBFFFF269F7F900FDFBF915" author="Drake, C. J." box="[151,540,1750,1775]" page="5" pageId="15" pageNumber="16" year="1948">
<taxonomicName id="EDABB38AFFBFFFF269F7F900FDFBF915" ID-CoL="33PB4" authority="Drake, 1948 b: 5" authorityName="Drake" authorityPageNumber="5" authorityYear="1948" box="[151,540,1750,1775]" class="Insecta" family="Tingidae" genus="Cysteochila" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="consueta">
<emphasis id="18DF141BFFBFFFF269F7F900FE88F915" box="[151,367,1751,1775]" italics="true" pageId="15" pageNumber="16">Cysteochila consueta</emphasis>
<bibRefCitation id="4E3AB5F8FFBFFFF26816F900FDFBF915" author="Drake, C. J." box="[374,540,1750,1775]" pageId="15" pageNumber="16" pagination="1 - 9" refId="ref36172" refString="Drake, C. J. (1948 b) Some Tingidae (Hemiptera) from China, Japan and India. Notes d'Entomologie Chinoise, 12 (1), 1 - 9." type="journal article" year="1948">Drake, 1948b: 5</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="F51076ABFFBFFFF26B46F900FD6FF914" box="[550,648,1751,1775]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
: macropterous
<specimenCount id="3CAD0380FFBFFFF26A42F900FCD0F915" box="[802,823,1751,1774]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
,
<collectingCountry id="52BC8899FFBFFFF26A21F900FC68F914" box="[833,911,1751,1775]" name="Taiwan" pageId="15" pageNumber="16">Taiwan</collectingCountry>
; USNM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2A14C809FFBFFFF269F7F8C3FD8CF892" blockId="15.[151,1436,1812,1897]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="0ABBEF27FFBFFFF269F7F8C3FD8CF892" pageId="15" pageNumber="16">
<taxonomicName id="EDABB38AFFBFFFF269F7F8C3FDE1F8D7" ID-CoL="33PB4" authority="Drake, 1948 b" authorityName="Drake" authorityYear="1948" box="[151,518,1812,1836]" class="Insecta" family="Tingidae" genus="Cysteochila" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="consueta">
<emphasis id="18DF141BFFBFFFF269F7F8C3FE97F8D7" box="[151,368,1812,1836]" italics="true" pageId="15" pageNumber="16">Cysteochila consueta</emphasis>
<bibRefCitation id="4E3AB5F8FFBFFFF26817F8C3FDE1F8D7" author="Drake, C. J." box="[375,518,1812,1836]" pageId="15" pageNumber="16" pagination="1 - 9" refId="ref36172" refString="Drake, C. J. (1948 b) Some Tingidae (Hemiptera) from China, Japan and India. Notes d'Entomologie Chinoise, 12 (1), 1 - 9." type="journal article" year="1948">Drake, 1948b</bibRefCitation>
</taxonomicName>
: Takeya (1951: 21) (checklist: eastern Asia);
<treatmentCitation id="AB0AEE18FFBFFFF26AB5F8C3FAE1F8D7" author="Drake, C. J. &amp; Ruhoff, F. A." box="[981,1286,1812,1836]" page="166" pageId="15" pageNumber="16" year="1965">
<bibRefCitation id="4E3AB5F8FFBFFFF26AB5F8C3FAE1F8D7" author="Drake, C. J. &amp; Ruhoff, F. A." box="[981,1286,1812,1836]" pageId="15" pageNumber="16" pagination="1 - 634" refId="ref36512" refString="Drake, C. J. &amp; Ruhoff, F. A. (1965 a) Lacebugs of the world, a catalog (Hemiptera: Tingidae). Bulletin of the United States National Museum, 243, i - viii + 1 - 634, pls. frontis piece + 1 - 56. https: // doi. org / 10.5479 / si. 03629236.243.1" type="journal article" year="1965">Drake &amp; Ruhoff (1965a: 166)</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="AB0AEE18FFBFFFF26C16F8C3FEDAF8B0" author="Lee, C. E." page="107" pageId="15" pageNumber="16" year="1967">
<bibRefCitation id="4E3AB5F8FFBFFFF26C16F8C3FEDAF8B0" author="Lee, C. E." pageId="15" pageNumber="16" pagination="91 - 111" refId="ref38587" refString="Lee, C. E. (1967) Tingidae from Korea (Heteroptera). Mushi, 41, 91 - 111." type="journal article" year="1967">Lee (1967: 107)</bibRefCitation>
</treatmentCitation>
(distribution);
<treatmentCitation id="AB0AEE18FFBFFFF268B5F8E4FD16F8B0" author="Pericart, J. &amp; Golub, V. B." box="[469,753,1843,1867]" page="26" pageId="15" pageNumber="16" year="1996">
<bibRefCitation id="4E3AB5F8FFBFFFF268B5F8E4FD16F8B0" author="Pericart, J. &amp; Golub, V. B." box="[469,753,1843,1867]" pageId="15" pageNumber="16" pagination="3 - 78" refId="ref40107" refString="Pericart, J. &amp; Golub, V. B. (1996) Superfamily Tingoidea Laporte, 1832. In: Aukema, B. &amp; Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 2, Cimicomorpha I. The Netherlands Entomological Society, Amsterdam, pp. 3 - 78." type="book chapter" year="1996">Péricart &amp; Golub (1996: 26)</bibRefCitation>
</treatmentCitation>
(checklist: Palaearctic);
<treatmentCitation id="AB0AEE18FFBFFFF26A89F8E4FB3DF8B0" author="Kwon, Y. J. &amp; Suh, S. J. &amp; Kim, J. A." box="[1001,1242,1843,1867]" page="188" pageId="15" pageNumber="16" year="2001">
<bibRefCitation id="4E3AB5F8FFBFFFF26A89F8E4FB3DF8B0" author="Kwon, Y. J. &amp; Suh, S. J. &amp; Kim, J. A." box="[1001,1242,1843,1867]" pageId="15" pageNumber="16" pagination="1 - 512" refId="ref38530" refString="Kwon, Y. J., Suh, S. J. &amp; Kim, J. A. (2001) Hemiptera. In: Economic insects of Korea. Vol. 18. Insecta Koreana. Supplement 25. National Institute of Agricultural Science and Technology, Suwon, pp. 1 - 512" type="book chapter" year="2001">
Kwon
<emphasis id="18DF141BFFBFFFF26D4DF8E4FBBEF8B0" box="[1069,1113,1843,1867]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. (2001: 188)
</bibRefCitation>
</treatmentCitation>
(checklist:
<collectingCountry id="52BC8899FFBFFFF26C2FF8E4FA76F8B0" box="[1359,1425,1843,1867]" name="South Korea" pageId="15" pageNumber="16">Korea</collectingCountry>
);
<treatmentCitation id="AB0AEE18FFBFFFF269A7F886FE4FF892" author="Cho, G. &amp; Kim, W. &amp; Kwon, Y. S. &amp; Lee, S." box="[199,424,1873,1897]" page="738" pageId="15" pageNumber="16" year="2020">
<bibRefCitation id="4E3AB5F8FFBFFFF269A7F886FE4FF892" author="Cho, G. &amp; Kim, W. &amp; Kwon, Y. S. &amp; Lee, S." box="[199,424,1873,1897]" pageId="15" pageNumber="16" pagination="736 - 745" refId="ref35840" refString="Cho, G., Kim, W., Kwon, Y. S. &amp; Lee, S. (2020) Check list of lace bugs (Hemiptera: Tingidae) of Korea. Journal of Asia-Pacific Entomology, 23, 736 - 745. https: // doi. org / 10.1016 / j. aspen. 2020.04.005" type="journal article" year="2020">
Cho
<emphasis id="18DF141BFFBFFFF26998F885FEC3F892" box="[248,292,1873,1897]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. (2020: 738)
</bibRefCitation>
</treatmentCitation>
(checklist:
<collectingCountry id="52BC8899FFBFFFF26B7EF886FD87F892" box="[542,608,1873,1897]" name="South Korea" pageId="15" pageNumber="16">Korea</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFBFFFED69F7F840FAB1FD9A" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="2A14C809FFBFFFED69F7F840FAB1FD9A" blockId="15.[151,1436,1943,2042]" lastBlockId="16.[151,1437,150,2013]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="18DF141BFFBFFFF269F7F840FE7EF84A" bold="true" box="[151,409,1943,1969]" pageId="15" pageNumber="16">Specimens examined.</emphasis>
<materialsCitation id="9AC3C254FFBFFFF268C1F84FFC11F82D" ID-GBIF-Occurrence="3802857391" collectingDate="1996-04-27" collectionCode="TUA" collectorName="Non-types &amp; T. Nishimura" country="Japan" county="Hirakubo" location="Ryukyu Islands" municipality="Ishigaki-shi" pageId="15" pageNumber="16" specimenCount="34" specimenCount-female="16" specimenCount-male="18">
<collectorName id="875EADDFFFBFFFF268C1F84FFDF1F849" box="[417,534,1944,1970]" pageId="15" pageNumber="16">Non-types</collectorName>
(
<specimenCount id="3CAD0380FFBFFFF26B48F84FFD9FF849" box="[552,632,1944,1970]" pageId="15" pageNumber="16" type="male">17 ♂♂</specimenCount>
<specimenCount id="3CAD0380FFBFFFF26BE1F840FD37F849" box="[641,720,1943,1970]" pageId="15" pageNumber="16" type="female">16 ♀♀</specimenCount>
1 nymph),
<emphasis id="18DF141BFFBFFFF26A37F840FAB8F849" bold="true" box="[855,1375,1943,1970]" pageId="15" pageNumber="16">
<collectingCountry id="52BC8899FFBFFFF26A37F840FC52F84A" box="[855,949,1943,1969]" name="Japan" pageId="15" pageNumber="16">JAPAN</collectingCountry>
:
<location id="2F749ED2FFBFFFF26AA3F84FFB63F849" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFBFFFF26AA3F84FFB63F849" box="[963,1156,1944,1970]" country="Japan" county="Hirakubo" municipality="Ishigaki-shi" name="Ryukyu Islands" pageId="15" pageNumber="16">Ryukyu Islands</location>
:
<emphasis id="18DF141BFFBFFFF26DF2F84FFAB1F849" bold="true" box="[1170,1366,1943,1970]" italics="true" pageId="15" pageNumber="16">
<location id="2F749ED2FFBFFFF26DF2F84FFAB1F849" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFBFFFF26DF2F84FFAB1F849" box="[1170,1366,1943,1970]" country="Japan" county="Hirakubo" municipality="Ishigaki-shi" name="Yaeyama Islands" pageId="15" pageNumber="16">Yaeyama Islands</location>
</emphasis>
:
</emphasis>
<location id="2F749ED2FFBFFFF26C09F840FEE0F82E" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFBFFFF26C09F840FEE0F82E" country="Japan" county="Hirakubo" municipality="Ishigaki-shi" name="Okinawa-ken" pageId="15" pageNumber="16">Okinawa-ken</location>
,
<collectingMunicipality id="CA705273FFBFFFF26873F86CFE7DF82E" box="[275,410,1979,2005]" pageId="15" pageNumber="16">Ishigaki-shi</collectingMunicipality>
,
<collectingCounty id="C375B085FFBFFFF268C5F86CFDF5F82E" box="[421,530,1979,2005]" pageId="15" pageNumber="16">Hirakubo</collectingCounty>
,
<date id="5E15EEC9FFBFFFF26B7EF86CFD71F82E" box="[542,662,1979,2005]" pageId="15" pageNumber="16" value="1996-04-27">
<collectingDate id="4E511721FFBFFFF26B7EF86CFD71F82E" box="[542,662,1979,2005]" pageId="15" pageNumber="16" value="1996-04-27">27.iv.1996</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFBFFFF26BB3F86BFC81F82E" box="[723,870,1979,2006]" pageId="15" pageNumber="16">T. Nishimura</collectorName>
(
<specimenCount id="3CAD0380FFBFFFF26A19F86BFC43F82D" box="[889,932,1980,2006]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
,
<collectionCode id="4CBA50CCFFBFFFF26AD0F86BFC09F82D" box="[944,1006,1980,2006]" pageId="15" pageNumber="16">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFBFFFF26D62F86CFEF3F801" ID-GBIF-Occurrence="3802857407" collectingDate="1995-05" collectionCode="NSMT" collectorName="M. Takai" country="Japan" location="Ishigaki Is." pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1">
<location id="2F749ED2FFBFFFF26D62F86CFB66F82D" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFBFFFF26D62F86CFB66F82D" box="[1026,1153,1979,2006]" country="Japan" name="Ishigaki Is." pageId="15" pageNumber="16">Ishigaki Is.</location>
,
<date id="5E15EEC9FFBFFFF26DEEF86CFB3CF82D" box="[1166,1243,1979,2006]" pageId="15" pageNumber="16" value="1995-05">
<collectingDate id="4E511721FFBFFFF26DEEF86CFB3CF82D" box="[1166,1243,1979,2006]" pageId="15" pageNumber="16" value="1995-05">v.1995</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFBFFFF26C78F86BFA9AF82E" box="[1304,1405,1979,2006]" pageId="15" pageNumber="16">M. Takai</collectorName>
(
<specimenCount id="3CAD0380FFBFFFF26CF0F86BFF49F801" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
,
<collectionCode id="4CBA50CCFFBFFFF269D9F837FEEBF801" box="[185,268,2016,2042]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="15" pageNumber="16">NSMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFBFFFF2687EF808FBD5F801" ID-GBIF-Occurrence="3802857311" box="[286,1074,2015,2042]" collectingDate="1998-03-16" collectionCode="NSMT" collectorName="Ishigaki-jima Is. &amp; K. Takahashi" country="Japan" location="Hoshino" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1">
<collectorName id="875EADDFFFBFFFF2687EF808FE32F801" box="[286,469,2015,2042]" pageId="15" pageNumber="16">Ishigaki-jima Is.</collectorName>
,
<location id="2F749ED2FFBFFFF26881F808FDA4F802" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFBFFFF26881F808FDA4F802" box="[481,579,2015,2041]" country="Japan" name="Hoshino" pageId="15" pageNumber="16">Hoshino</location>
,
<date id="5E15EEC9FFBFFFF26B2CF808FD2FF802" box="[588,712,2015,2041]" pageId="15" pageNumber="16" value="1998-03-16">
<collectingDate id="4E511721FFBFFFF26B2CF808FD2FF802" box="[588,712,2015,2041]" pageId="15" pageNumber="16" value="1998-03-16">16.iii.1998</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFBFFFF26A61F837FC74F802" box="[769,915,2015,2042]" pageId="15" pageNumber="16">K. Takahashi</collectorName>
(
<specimenCount id="3CAD0380FFBFFFF26AC4F837FC2AF801" box="[932,973,2016,2042]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
,
<collectionCode id="4CBA50CCFFBFFFF26AB8F837FBCCF801" box="[984,1067,2016,2042]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="15" pageNumber="16">NSMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFBFFFED6D5BF837FF25FF4A" ID-GBIF-Occurrence="3802857487" collectorName="Ishigaki Is." country="Japan" lastPageId="16" lastPageNumber="17" location="Yaeyama Isls." pageId="15" pageNumber="16" specimenCount="1">
<location id="2F749ED2FFBFFFF26D5BF837FB3FF802" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFBFFFF26D5BF837FB3FF802" box="[1083,1240,2015,2042]" country="Japan" name="Yaeyama Isls." pageId="15" pageNumber="16">Yaeyama Isls.</location>
,
<collectorName id="875EADDFFFBFFFF26D83F808FAB8F801" box="[1251,1375,2015,2042]" pageId="15" pageNumber="16">Ishigaki Is.</collectorName>
,
<materialsCitation id="9AC3C254FFBFFFED6C0AF808FF25FF4A" ID-GBIF-Occurrence="3802857379" collectingDate="1996-04-03" collectionCode="TUA" collectorName="M. Hayashi" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1">Kainan</materialsCitation>
</materialsCitation>
,
<date id="5E15EEC9FFA0FFED69ADFF40FEDFFF4A" box="[205,312,151,177]" pageId="16" pageNumber="17" value="1996-04-03">
<collectingDate id="4E511721FFA0FFED69ADFF40FEDFFF4A" box="[205,312,151,177]" pageId="16" pageNumber="17" value="1996-04-03">3.iv.1996</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED6812FF40FE13FF4A" box="[370,500,151,177]" pageId="16" pageNumber="17">M. Hayashi</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6B65FF40FDCAFF4A" box="[517,557,151,177]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6B5AFF40FD9FFF4A" box="[570,632,151,177]" pageId="16" pageNumber="17">TUA</collectionCode>
);
<materialsCitation id="9AC3C254FFA0FFED6BEBFF40FACBFF4A" ID-GBIF-Occurrence="3802857449" box="[651,1324,150,177]" collectingDate="2018-11-15" collectionCode="TUA" collectorName="J. Souma" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1">
as above but
<date id="5E15EEC9FFA0FFED6A7CFF41FC71FF4A" box="[796,918,150,177]" pageId="16" pageNumber="17" value="2018-11-15">
<collectingDate id="4E511721FFA0FFED6A7CFF41FC71FF4A" box="[796,918,150,177]" pageId="16" pageNumber="17" value="2018-11-15">15.xi.2018</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED6AB1FF40FBD0FF4A" box="[977,1079,151,177]" pageId="16" pageNumber="17">J. Souma</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6D28FF40FB97FF4A" box="[1096,1136,151,177]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
1 nymph,
<collectionCode id="4CBA50CCFFA0FFED6D85FF40FAC4FF4A" box="[1253,1315,151,177]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6C55FF40FC75FF2E" ID-GBIF-Occurrence="3802857328" collectingDate="1996-06-18" collectionCode="TUA" collectorName="Ishigaki Is. &amp; M. Hayashi" location="Omoto" municipality="Yaeyama Isls." pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="CA705273FFA0FFED6C55FF40FF20FF2E" pageId="16" pageNumber="17">Yaeyama Isls.</collectingMunicipality>
,
<collectorName id="875EADDFFFA0FFED69B5FF6CFEB3FF2E" box="[213,340,187,213]" pageId="16" pageNumber="17">Ishigaki Is.</collectorName>
,
<location id="2F749ED2FFA0FFED6803FF6CFE53FF2E" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED6803FF6CFE53FF2E" box="[355,436,187,213]" municipality="Yaeyama Isls." name="Omoto" pageId="16" pageNumber="17">Omoto</location>
,
<date id="5E15EEC9FFA0FFED68A0FF6CFDDDFF2E" box="[448,570,187,213]" pageId="16" pageNumber="17" value="1996-06-18">
<collectingDate id="4E511721FFA0FFED68A0FF6CFDDDFF2E" box="[448,570,187,213]" pageId="16" pageNumber="17" value="1996-06-18">18.vi.1996</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED6B1AFF6CFD18FF2E" box="[634,767,187,213]" pageId="16" pageNumber="17">M. Hayashi</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6A72FF6CFCDBFF2E" box="[786,828,187,213]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6A2BFF6CFC6EFF2E" box="[843,905,187,213]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6AFFFF6CFE48FF02" ID-GBIF-Occurrence="3802857431" collectingDate="1997-04-06" collectionCode="TUA" collectorName="Ishigaki-jima Is. &amp; T. Ishikawa" location="Banna Park" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1">
<collectorName id="875EADDFFFA0FFED6AFFFF6CFBBDFF2E" box="[927,1114,187,213]" pageId="16" pageNumber="17">Ishigaki-jima Is.</collectorName>
,
<location id="2F749ED2FFA0FFED6D08FF6CFB09FF2E" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED6D08FF6CFB09FF2E" box="[1128,1262,187,213]" name="Banna Park" pageId="16" pageNumber="17">Banna Park</location>
,
<date id="5E15EEC9FFA0FFED6D9BFF6CFA81FF2E" box="[1275,1382,187,213]" pageId="16" pageNumber="17" value="1997-04-06">
<collectingDate id="4E511721FFA0FFED6D9BFF6CFA81FF2E" box="[1275,1382,187,213]" pageId="16" pageNumber="17" value="1997-04-06">6.iv.1997</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED69F7FF08FEFCFF02" box="[151,283,223,249]" pageId="16" pageNumber="17">T. Ishikawa</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED684FFF08FEBBFF02" box="[303,348,223,249]" pageId="16" pageNumber="17" type="male">1 ♂</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6809FF08FE40FF02" box="[361,423,223,249]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED68DDFF08FCB5FF02" ID-GBIF-Occurrence="3802857351" box="[445,850,223,249]" collectingDate="1997-04-07" collectionCode="TUA" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1">
as above but
<date id="5E15EEC9FFA0FFED6B35FF08FD5AFF02" box="[597,701,223,249]" pageId="16" pageNumber="17" value="1997-04-07">
<collectingDate id="4E511721FFA0FFED6B35FF08FD5AFF02" box="[597,701,223,249]" pageId="16" pageNumber="17" value="1997-04-07">7.iv.1997</collectingDate>
</date>
(
<specimenCount id="3CAD0380FFA0FFED6BB1FF08FD1BFF02" box="[721,764,223,249]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6A6BFF08FCAEFF02" box="[779,841,223,249]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6A3FFF08FB1DFF02" ID-GBIF-Occurrence="3802857308" box="[863,1274,223,249]" collectingDate="2000-05-17" collectionCode="TUA" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1">
as above but
<date id="5E15EEC9FFA0FFED6A98FF08FB81FF02" box="[1016,1126,223,249]" pageId="16" pageNumber="17" value="2000-05-17">
<collectingDate id="4E511721FFA0FFED6A98FF08FB81FF02" box="[1016,1126,223,249]" pageId="16" pageNumber="17" value="2000-05-17">17.v.2000</collectingDate>
</date>
(
<specimenCount id="3CAD0380FFA0FFED6D1AFF08FB43FF02" box="[1146,1188,223,249]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6DD3FF08FB16FF02" box="[1203,1265,223,249]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6C67FF08FD06FEE6" ID-GBIF-Occurrence="3802857376" collectingDate="1999-06-04" collectionCode="TUA" collectorName="Ishigaki-jima Is. &amp; H. Mizushima" location="Itona" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1">
<collectorName id="875EADDFFFA0FFED6C67FF08FF53FEE6" pageId="16" pageNumber="17">Ishigaki-jima Is.</collectorName>
,
<location id="2F749ED2FFA0FFED69A1FED4FF1AFEE6" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED69A1FED4FF1AFEE6" box="[193,253,259,285]" name="Itona" pageId="16" pageNumber="17">Itona</location>
,
<date id="5E15EEC9FFA0FFED6868FED4FE92FEE6" box="[264,373,259,285]" pageId="16" pageNumber="17" value="1999-06-04">
<collectingDate id="4E511721FFA0FFED6868FED4FE92FEE6" box="[264,373,259,285]" pageId="16" pageNumber="17" value="1999-06-04">4.vi.1999</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED68D1FED4FDB6FEE6" box="[433,593,259,285]" pageId="16" pageNumber="17">H. Mizushima</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6B04FED4FD6BFEE6" box="[612,652,259,285]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6BFAFED4FD3FFEE6" box="[666,728,259,285]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6B8CFED4FF10FEBA" ID-GBIF-Occurrence="3802857320" collectingDate="2008-06-17" collectionCode="NIAES" collectorName="Ishigaki Is. &amp; Y. Nakatani" location="Nosoko-rindo" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1">
<collectorName id="875EADDFFFA0FFED6B8CFED4FC8DFEE6" box="[748,874,259,285]" pageId="16" pageNumber="17">Ishigaki Is.</collectorName>
,
<location id="2F749ED2FFA0FFED6A17FED4FBF1FEE6" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED6A17FED4FBF1FEE6" box="[887,1046,259,285]" name="Nosoko-rindo" pageId="16" pageNumber="17">Nosoko-rindo</location>
,
<date id="5E15EEC9FFA0FFED6D41FED4FB7CFEE6" box="[1057,1179,259,285]" pageId="16" pageNumber="17" value="2008-06-17">
<collectingDate id="4E511721FFA0FFED6D41FED4FB7CFEE6" box="[1057,1179,259,285]" pageId="16" pageNumber="17" value="2008-06-17">17.vi.2008</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED6DB6FED4FABDFEE6" box="[1238,1370,259,285]" pageId="16" pageNumber="17">Y. Nakatani</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6C0DFED4FA72FEE6" box="[1389,1429,259,285]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED69F7FEF0FF09FEBA" box="[151,238,295,321]" country="Japan" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="16" pageNumber="17">NIAES</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6862FEF0FC95FEBA" ID-GBIF-Occurrence="3802857408" box="[258,882,295,321]" collectingDate="2018-11-14" collectionCode="TUA" collectorName="J. Souma" pageId="16" pageNumber="17" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
as above but
<date id="5E15EEC9FFA0FFED68F5FEF0FDF7FEBA" box="[405,528,295,321]" pageId="16" pageNumber="17" value="2018-11-14">
<collectingDate id="4E511721FFA0FFED68F5FEF0FDF7FEBA" box="[405,528,295,321]" pageId="16" pageNumber="17" value="2018-11-14">14.xi.2018</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED6B2BFEF0FD55FEBA" box="[587,690,295,321]" pageId="16" pageNumber="17">J. Souma</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6BA4FEF0FD0AFEBA" box="[708,749,295,321]" pageId="16" pageNumber="17" type="male">1 ♂</specimenCount>
<specimenCount id="3CAD0380FFA0FFED6B94FEF0FCF9FEBA" box="[756,798,295,321]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6A4CFEF0FC8DFEBA" box="[812,874,295,321]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6A1CFEF0FCDBFE9E" ID-GBIF-Occurrence="3802857375" accessionNumber="N 12409" collectingDate="2018-11-11" collectionCode="TUA" collectorName="Ishigaki Is. &amp; J. Souma" location="Okawa" municipality="Yaeyama Isls." pageId="16" pageNumber="17" specimenCount="7" specimenCount-female="3" specimenCount-male="4">
<collectingMunicipality id="CA705273FFA0FFED6A1CFEF0FBFCFEBA" box="[892,1051,295,321]" pageId="16" pageNumber="17">Yaeyama Isls.</collectingMunicipality>
,
<collectorName id="875EADDFFFA0FFED6D46FEF0FB43FEBA" box="[1062,1188,295,321]" pageId="16" pageNumber="17">Ishigaki Is.</collectorName>
,
<location id="2F749ED2FFA0FFED6DD1FEF0FAE4FEBA" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED6DD1FEF0FAE4FEBA" box="[1201,1283,295,321]" municipality="Yaeyama Isls." name="Okawa" pageId="16" pageNumber="17">Okawa</location>
, 2421'42.2&quot;
<accessionNumber id="35F855EAFFA0FFED6CE7FEF0FF06FE9E" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/N12409" pageId="16" pageNumber="17" type="EnaNcbi">N 12409</accessionNumber>
'52.5&quot;E,
<date id="5E15EEC9FFA0FFED685DFE9CFE50FE9E" box="[317,439,331,357]" pageId="16" pageNumber="17" value="2018-11-11">
<collectingDate id="4E511721FFA0FFED685DFE9CFE50FE9E" box="[317,439,331,357]" pageId="16" pageNumber="17" value="2018-11-11">11.xi.2018</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED6891FE9CFDB0FE9E" box="[497,599,331,357]" pageId="16" pageNumber="17">J. Souma</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6B05FE9CFD42FE9E" box="[613,677,331,357]" pageId="16" pageNumber="17" type="male">4 ♂♂</specimenCount>
<specimenCount id="3CAD0380FFA0FFED6BCBFE9CFD0DFE9E" box="[683,746,331,357]" pageId="16" pageNumber="17" type="female">3 ♀♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6B96FE9CFCD3FE9E" box="[758,820,331,357]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6A27FE9CFB16FE9E" ID-GBIF-Occurrence="3802857435" box="[839,1265,331,357]" collectingDate="2018-11-16" collectionCode="TUA" pageId="16" pageNumber="17" specimenCount="4" specimenCount-male="4">
as above but
<date id="5E15EEC9FFA0FFED6AB8FE9CFBB7FE9E" box="[984,1104,331,357]" pageId="16" pageNumber="17" value="2018-11-16">
<collectingDate id="4E511721FFA0FFED6AB8FE9CFBB7FE9E" box="[984,1104,331,357]" pageId="16" pageNumber="17" value="2018-11-16">16.xi.2018</collectingDate>
</date>
(
<specimenCount id="3CAD0380FFA0FFED6D3EFE9CFB78FE9E" box="[1118,1183,331,357]" pageId="16" pageNumber="17" type="male">4 ♂♂</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6DCAFE9CFB0FFE9E" box="[1194,1256,331,357]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6D9AFE9CFB77FE72" ID-GBIF-Occurrence="3802857397" accessionNumber="N 12412" collectingDate="2018-11-17" collectionCode="TUA" collectorName="Ishigaki Is. &amp; J. Souma" location="Hirae" municipality="Yaeyama Isls." pageId="16" pageNumber="17" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<collectingMunicipality id="CA705273FFA0FFED6D9AFE9CFA7FFE9E" box="[1274,1432,331,357]" pageId="16" pageNumber="17">Yaeyama Isls.</collectingMunicipality>
,
<collectorName id="875EADDFFFA0FFED69F7FEB8FEF3FE72" box="[151,276,367,393]" pageId="16" pageNumber="17">Ishigaki Is.</collectorName>
,
<location id="2F749ED2FFA0FFED6840FEB8FE86FE72" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED6840FEB8FE86FE72" box="[288,353,367,393]" municipality="Yaeyama Isls." name="Hirae" pageId="16" pageNumber="17">Hirae</location>
, 2423'28.6&quot;
<accessionNumber id="35F855EAFFA0FFED6884FEB8FDAEFE72" box="[484,585,367,393]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/N12412" pageId="16" pageNumber="17" type="EnaNcbi">N 12412</accessionNumber>
'00.7&quot;E,
<date id="5E15EEC9FFA0FFED6BC5FEB8FCF8FE72" box="[677,799,367,393]" pageId="16" pageNumber="17" value="2018-11-17">
<collectingDate id="4E511721FFA0FFED6BC5FEB8FCF8FE72" box="[677,799,367,393]" pageId="16" pageNumber="17" value="2018-11-17">17.xi.2018</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED6A3AFEB8FC27FE72" box="[858,960,367,393]" pageId="16" pageNumber="17">J. Souma</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6AB1FEB8FC1EFE72" box="[977,1017,367,393]" pageId="16" pageNumber="17" type="male">1 ♂</specimenCount>
<specimenCount id="3CAD0380FFA0FFED6A9FFEB8FBD9FE72" box="[1023,1086,367,393]" pageId="16" pageNumber="17" type="female">2 ♀♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6D2AFEB8FB6FFE72" box="[1098,1160,367,393]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6DFAFEB8FBBAFE56" ID-GBIF-Occurrence="3802857494" accessionNumber="N 12409" collectingDate="2018-11-18" collectionCode="TUA" collectorName="Ishigaki Is. &amp; J. Souma" location="Misaki-cho" municipality="Yaeyama Isls." pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1">
<collectingMunicipality id="CA705273FFA0FFED6DFAFEB8FAD0FE72" box="[1178,1335,367,393]" pageId="16" pageNumber="17">Yaeyama Isls.</collectingMunicipality>
,
<collectorName id="875EADDFFFA0FFED6C22FEB8FF53FE56" pageId="16" pageNumber="17">Ishigaki Is.</collectorName>
,
<location id="2F749ED2FFA0FFED69A4FE44FEA1FE56" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED69A4FE44FEA1FE56" box="[196,326,403,429]" municipality="Yaeyama Isls." name="Misaki-cho" pageId="16" pageNumber="17">Misaki-cho</location>
, 2420'13.0&quot;
<accessionNumber id="35F855EAFFA0FFED68AEFE44FDDDFE56" box="[462,570,403,429]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/N12409" pageId="16" pageNumber="17" type="EnaNcbi">N 12409</accessionNumber>
'19.9&quot;E,
<date id="5E15EEC9FFA0FFED6BF9FE44FCF4FE56" box="[665,787,403,429]" pageId="16" pageNumber="17" value="2018-11-18">
<collectingDate id="4E511721FFA0FFED6BF9FE44FCF4FE56" box="[665,787,403,429]" pageId="16" pageNumber="17" value="2018-11-18">18.xi.2018</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED6A37FE44FC25FE56" box="[855,962,403,429]" pageId="16" pageNumber="17">J. Souma</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6AB8FE44FBE0FE56" box="[984,1031,403,429]" pageId="16" pageNumber="17" type="male">1 ♂</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6D76FE44FBB3FE56" box="[1046,1108,403,429]" pageId="16" pageNumber="17">TUA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6D0CFE44FC83FE2A" ID-GBIF-Occurrence="3802857425" collectingDate="2021-05-28" collectingDateMax="2021-05-31" collectingDateMin="2021-05-28" collectionCode="ELKU" collectorName="S. Imada &amp; N. Tsuji" location="Torogawa" municipality="Okinawa-ken" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="CA705273FFA0FFED6D0CFE44FAE1FE56" box="[1132,1286,403,429]" pageId="16" pageNumber="17">Okinawa-ken</collectingMunicipality>
,
<collectorName id="875EADDFFFA0FFED6C75FE44FA70FE56" box="[1301,1431,403,429]" pageId="16" pageNumber="17">Ishigaki Is.</collectorName>
,
<location id="2F749ED2FFA0FFED69F7FE60FEECFE2A" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED69F7FE60FEECFE2A" box="[151,267,439,465]" municipality="Okinawa-ken" name="Torogawa" pageId="16" pageNumber="17">Tôrogawa</location>
,
<date id="5E15EEC9FFA0FFED6876FE60FE56FE2A" box="[278,433,439,465]" pageId="16" pageNumber="17" value="2021-05-28" valueMax="2021-05-31" valueMin="2021-05-28">
<collectingDate id="4E511721FFA0FFED6876FE60FE56FE2A" box="[278,433,439,465]" pageId="16" pageNumber="17" value="2021-05-28" valueMax="2021-05-31" valueMin="2021-05-28">28–31.v.2021</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED688AFE60FDAAFE2A" box="[490,589,439,465]" pageId="16" pageNumber="17">S. Imada</collectorName>
&amp;
<collectorName id="875EADDFFFA0FFED6B0FFE60FD20FE2A" box="[623,711,439,465]" pageId="16" pageNumber="17">N. Tsuji</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6BB8FE60FD18FE2A" box="[728,767,439,465]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6A6DFE60FCBAFE2A" box="[781,861,439,465]" pageId="16" pageNumber="17">ELKU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="9AC3C254FFA0FFED6A0DFE61FE13FE0E" ID-GBIF-Occurrence="3802857365" collectingDate="1938-06-19" collectingDateMax="1938-06-20" collectingDateMin="1938-06-19" collectionCode="ELKU" collectorName="S. Nomura" country="Taiwan" location="Mt. Rara" municipality="Bunzan-gun" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1">
<emphasis id="18DF141BFFA0FFED6A0DFE61FC0FFE2B" bold="true" box="[877,1000,438,464]" pageId="16" pageNumber="17">
<collectingCountry id="52BC8899FFA0FFED6A0DFE61FC04FE2B" box="[877,995,438,464]" name="Taiwan" pageId="16" pageNumber="17">TAIWAN</collectingCountry>
:
</emphasis>
<collectingMunicipality id="CA705273FFA0FFED6A8EFE60FB9DFE2A" box="[1006,1146,439,465]" pageId="16" pageNumber="17">Bunzan-gun</collectingMunicipality>
,
<location id="2F749ED2FFA0FFED6DE4FE60FB0DFE2A" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED6DE4FE60FB0DFE2A" box="[1156,1258,439,465]" country="Taiwan" municipality="Bunzan-gun" name="Mt. Rara" pageId="16" pageNumber="17">Mt. Rara</location>
,
<date id="5E15EEC9FFA0FFED6D93FE60FA70FE2A" box="[1267,1431,439,465]" pageId="16" pageNumber="17" value="1938-06-19" valueMax="1938-06-20" valueMin="1938-06-19">
<collectingDate id="4E511721FFA0FFED6D93FE60FA70FE2A" box="[1267,1431,439,465]" pageId="16" pageNumber="17" value="1938-06-19" valueMax="1938-06-20" valueMin="1938-06-19">19–20.vi.1938</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED69ABFE0CFEAEFE0E" box="[203,329,475,501]" pageId="16" pageNumber="17">S. Nomura</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED683EFE0CFE6BFE0E" box="[350,396,475,501]" pageId="16" pageNumber="17" type="male">1 ♂</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED68FCFE0CFE0BFE0E" box="[412,492,475,501]" pageId="16" pageNumber="17">ELKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED6B62FE0CFA7FFE0E" ID-GBIF-Occurrence="3802857371" box="[514,1432,475,501]" collectingDate="1932-07-12" collectionCode="ELKU" collectorName="T. Esaki" country="Taiwan" county="Taich-sh" location="Hassenzan" municipality="Kahodai-Reimei" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1">
<collectingCounty id="C375B085FFA0FFED6B62FE0CFD82FE0E" box="[514,613,475,501]" pageId="16" pageNumber="17">Taich-sh</collectingCounty>
,
<collectingMunicipality id="CA705273FFA0FFED6B14FE0CFCD7FE0E" box="[628,816,475,501]" pageId="16" pageNumber="17">Kahodai-Reimei</collectingMunicipality>
,
<location id="2F749ED2FFA0FFED6A5FFE0CFC5DFE0E" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED6A5FFE0CFC5DFE0E" box="[831,954,475,501]" country="Taiwan" county="Taich-sh" municipality="Kahodai-Reimei" name="Hassenzan" pageId="16" pageNumber="17">Hassenzan</location>
,
<date id="5E15EEC9FFA0FFED6AA9FE0CFBADFE0E" box="[969,1098,475,501]" pageId="16" pageNumber="17" value="1932-07-12">
<collectingDate id="4E511721FFA0FFED6AA9FE0CFBADFE0E" box="[969,1098,475,501]" pageId="16" pageNumber="17" value="1932-07-12">12.vii.1932</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED6DEEFE0CFB0BFE0E" box="[1166,1260,475,501]" pageId="16" pageNumber="17">T. Esaki</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6C63FE0CFAC8FE0E" box="[1283,1327,475,501]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED6C20FE0CFA77FE0E" box="[1344,1424,475,501]" pageId="16" pageNumber="17">ELKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA0FFED69F7FE28FE01FDC6" ID-GBIF-Occurrence="3802857461" accessionNumber="N 12109" collectingDate="2016-05-19" collectionCode="NMNS" collectorName="Y. Tamadera" country="Taiwan" county="Ren'ai Township" location="Lixing Industry Road" municipality="Tattaka" pageId="16" pageNumber="17" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nantou">
<collectingRegion id="E86F06EBFFA0FFED69F7FE28FEAFFDE2" box="[151,328,511,537]" country="Taiwan" name="Nantou" pageId="16" pageNumber="17">Nantou County</collectingRegion>
,
<collectingCounty id="C375B085FFA0FFED6836FE28FDF0FDE2" box="[342,535,511,537]" pageId="16" pageNumber="17">Ren'ai Township</collectingCounty>
,
<collectingMunicipality id="CA705273FFA0FFED6B45FE28FD9DFDE2" box="[549,634,511,537]" pageId="16" pageNumber="17">Tattaka</collectingMunicipality>
,
<location id="2F749ED2FFA0FFED6BE8FE28FC64FDE2" LSID="urn:lsid:plazi:treatment:A202791FFFBFFFEC69F7F9A4FB5DFF49:2F749ED2FFA0FFED6BE8FE28FC64FDE2" box="[648,899,511,537]" country="Taiwan" county="Ren'ai Township" municipality="Tattaka" name="Lixing Industry Road" pageId="16" pageNumber="17" stateProvince="Nantou">Lixing Industry Road</location>
, 2404'25.4&quot;
<accessionNumber id="35F855EAFFA0FFED6D6AFE29FB95FDE2" box="[1034,1138,510,537]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/N12109" pageId="16" pageNumber="17" type="EnaNcbi">N 12109</accessionNumber>
'32.2&quot;E,
<date id="5E15EEC9FFA0FFED6DB3FE28FAA4FDE2" box="[1235,1347,511,537]" pageId="16" pageNumber="17" value="2016-05-19">
<collectingDate id="4E511721FFA0FFED6DB3FE28FAA4FDE2" box="[1235,1347,511,537]" pageId="16" pageNumber="17" value="2016-05-19">19.v.2016</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA0FFED6CE5FE28FEE2FDC6" pageId="16" pageNumber="17">Y. Tamadera</collectorName>
(
<specimenCount id="3CAD0380FFA0FFED6879FDF4FEA4FDC6" box="[281,323,547,573]" pageId="16" pageNumber="17" type="male">1 ♂</specimenCount>
<specimenCount id="3CAD0380FFA0FFED682CFDF4FE9FFDC6" box="[332,376,547,573]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA0FFED68E8FDF4FE39FDC6" box="[392,478,547,573]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="16" pageNumber="17">NMNS</collectionCode>
)
</materialsCitation>
. Two specimens deposited in ELKU are labeled “
<taxonomicName id="EDABB38AFFA0FFED6D41FDF4FA75FDC6" authority="Takeya, 1962" authorityName="Takeya" authorityYear="1962" box="[1057,1426,547,573]" class="Insecta" family="Tingidae" genus="Cysteochila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="esakii">
<emphasis id="18DF141BFFA0FFED6D41FDF4FB08FDC6" box="[1057,1263,547,573]" italics="true" pageId="16" pageNumber="17">Cysteochila esakii</emphasis>
Takeya, 1962
</taxonomicName>
” (manuscript name). A single nymph is in a poor condition and, therefore, not described in the present article.
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA0FFED69A7FDBDFE07FC32" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="2A14C809FFA0FFED69A7FDBDFE07FC32" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">
<emphasis id="18DF141BFFA0FFED69A7FDBDFEA5FD7F" bold="true" box="[199,322,618,644]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
Recognized among other species of
<taxonomicName id="EDABB38AFFA0FFED6B8AFDBCFC6EFD7E" authorityName="Fieber" authorityYear="1844" box="[746,905,619,645]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="18DF141BFFA0FFED6B8AFDBCFC6EFD7E" box="[746,905,619,645]" italics="true" pageId="16" pageNumber="17">Physatocheila</emphasis>
</taxonomicName>
by a combination of the following characters: general color light brown (
<figureCitation id="B290D48CFFA0FFED68DFFD58FDC7FD52" box="[447,544,655,681]" captionStart="FIGURE 8" captionStartId="13.[151,250,1851,1875]" captionTargetBox="[157,1428,996,1820]" captionTargetId="figure-410@13.[152,1436,993,1828]" captionTargetPageId="13" captionText="FIGURE 8. Nine Physatocheila species from Japan. Dried specimens, dorsal view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi from Japan proper (E); P. fieberi from the Ryukyu Islands (F); P. miyatakei (G); P. nigrintegerrima (H); P. orientis (I); P. veteris (J). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6609989" httpUri="https://zenodo.org/record/6609989/files/figure.png" pageId="16" pageNumber="17">Figs. 8B</figureCitation>
,
<figureCitation id="B290D48CFFA0FFED6B4BFD58FDA9FD52" box="[555,590,655,681]" captionStart="FIGURE 9" captionStartId="14.[151,250,1804,1828]" captionTargetBox="[151,1435,489,1780]" captionTargetId="figure-184@14.[151,1435,489,1780]" captionTargetPageId="14" captionText="FIGURE 9. Rostra and sternal laminae of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609991" httpUri="https://zenodo.org/record/6609991/files/figure.png" pageId="16" pageNumber="17">9B</figureCitation>
,
<figureCitation id="B290D48CFFA0FFED6B39FD58FD6DFD52" box="[601,650,655,681]" captionStart="FIGURE 10" captionStartId="19.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1198]" captionTargetId="figure-17@19.[151,1436,181,1198]" captionTargetPageId="19" captionText="FIGURE 10. Male terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609995" httpUri="https://zenodo.org/record/6609995/files/figure.png" pageId="16" pageNumber="17">10B</figureCitation>
,
<figureCitation id="B290D48CFFA0FFED6BF5FD58FD21FD52" box="[661,710,655,681]" captionStart="FIGURE 11" captionStartId="21.[151,250,1579,1603]" captionTargetBox="[151,1436,525,1555]" captionTargetId="figure-182@21.[151,1436,525,1555]" captionTargetPageId="21" captionText="FIGURE 11. Female terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6610129" httpUri="https://zenodo.org/record/6610129/files/figure.png" pageId="16" pageNumber="17">11B</figureCitation>
); median spine on head extending beyond bases of frontal spines; rostrum reaching posterior margin of mesosternum; pubescence on pronotum less than 0.5 times as long as diameter of compound eye; lateral carinae of pronotum close to each other in anterior part, concealed by paranotum in anterior part; paranota incompletely covering pronotal disc in anterior part, not touching each other, not concealing median carina of pronotum, widened posteriorly, bulged upward in posterior part, forming a cyst; cyst of paranotum not concave in posterior part; outer margin of paranotum gently curved outward in its entire length; costal area of hemelytron wider than subcostal area at widest part, less than 0.5 times as wide as discoidal area at widest part, with a single row of areolae in apical part and 2 rows in remaining parts; and subcostal area subhorizontal, with 2 rows of areolae in its entire length.
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA0FFED69A7FC05FE5FFB86" pageId="16" pageNumber="17" type="description">
<paragraph id="2A14C809FFA0FFED69A7FC05FE5FFB86" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">
<emphasis id="18DF141BFFA0FFED69A7FC05FE00FC17" bold="true" box="[199,487,978,1004]" pageId="16" pageNumber="17">Description of genitalia.</emphasis>
Pygophore (
<figureCitation id="B290D48CFFA0FFED6B18FC04FD0DFC16" box="[632,746,979,1005]" captionStart="FIGURE 10" captionStartId="19.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1198]" captionTargetId="figure-17@19.[151,1436,181,1198]" captionTargetPageId="19" captionText="FIGURE 10. Male terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609995" httpUri="https://zenodo.org/record/6609995/files/figure.png" pageId="16" pageNumber="17">Figs. 10B</figureCitation>
,
<figureCitation id="B290D48CFFA0FFED6B96FC04FCC0FC16" box="[758,807,979,1005]" captionStart="FIGURE 17" captionStartId="28.[151,250,1870,1894]" captionTargetBox="[156,1434,888,1845]" captionTargetId="figure-335@28.[151,1436,885,1847]" captionTargetPageId="28" captionText="FIGURE 17. Pygophores of eleven species of Cysteochila and its close relatives from Japan, dorsal view: C. abundantis (A); Limnostatua lewisi comb. nov. (B); Physatocheila chiniana comb. nov. (C); P. consueta comb. nov. (D); P. distinguenda (E); P. euthenia comb. nov. (F); P. fieberi from Japan proper (G); P. fieberi from the Ryukyu Islands (H); P. miyatakei (I); P. nigrintegerrima (J); P. orientis (K); P. veteris (L). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6610019" httpUri="https://zenodo.org/record/6610019/files/figure.png" pageId="16" pageNumber="17">17D</figureCitation>
) compressed dorsoventrally, hexagonal in ventral view, strongly concave at anterior margin of dorsum, elevated at center of venter, smooth on surface, irregularly punctate in middle part of dorsum. Paramere (
<figureCitation id="B290D48CFFA0FFED6B20FBCCFD4FFBCE" box="[576,680,1051,1077]" captionStart="FIGURE 18" captionStartId="29.[151,250,1878,1902]" captionTargetBox="[154,1433,1025,1851]" captionTargetId="figure-333@29.[151,1436,1022,1854]" captionTargetPageId="29" captionText="FIGURE 18. Parameres of eleven species of Cysteochila and its close relatives from Japan, dorsal view: C. abundantis (A); Limnostatua lewisi comb. nov. (B); Physatocheila chiniana comb. nov. (C); P. consueta comb. nov. (D); P. distinguenda (E); P. euthenia comb. nov. (F); P. fieberi from Japan proper (G); P. fieberi from the Ryukyu Islands (H); P. miyatakei (I); P. nigrintegerrima (J); P. orientis (K); P. veteris (L). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6610022" httpUri="https://zenodo.org/record/6610022/files/figure.png" pageId="16" pageNumber="17">Fig. 18D</figureCitation>
) expanded in middle part, curved inward in apical part; outer and inner margins covered with pubescence in middle part. Female terminalia (
<figureCitation id="B290D48CFFA0FFED6A91FBE8FBB1FBA2" box="[1009,1110,1087,1113]" captionStart="FIGURE 11" captionStartId="21.[151,250,1579,1603]" captionTargetBox="[151,1436,525,1555]" captionTargetId="figure-182@21.[151,1436,525,1555]" captionTargetPageId="21" captionText="FIGURE 11. Female terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6610129" httpUri="https://zenodo.org/record/6610129/files/figure.png" pageId="16" pageNumber="17">Fig. 11B</figureCitation>
) pentagonal in ventral view, covered with pubescence.
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA0FFED69A7FB50FD3DF9F2" pageId="16" pageNumber="17" type="discussion">
<paragraph id="2A14C809FFA0FFED69A7FB50FEE1FB12" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">
<emphasis id="18DF141BFFA0FFED69A7FB50FEDCFB5A" bold="true" box="[199,315,1159,1185]" pageId="16" pageNumber="17">Remarks.</emphasis>
This species belongs to
<taxonomicName id="EDABB38AFFA0FFED6B29FB50FD0FFB5A" authorityName="Fieber" authorityYear="1844" box="[585,744,1159,1185]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="18DF141BFFA0FFED6B29FB50FD0FFB5A" box="[585,744,1159,1185]" italics="true" pageId="16" pageNumber="17">Physatocheila</emphasis>
</taxonomicName>
based on the presence of generic characteristics, as mentioned in the Diagnosis section. Therefore, the new combination,
<taxonomicName id="EDABB38AFFA0FFED6A59FB7CFB0AFB3E" authority="(Drake, 1948)" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[825,1261,1195,1221]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED6A59FB7CFBA2FB3F" box="[825,1093,1195,1221]" italics="true" pageId="16" pageNumber="17">Physatocheila consueta</emphasis>
(Drake, 1948)
</taxonomicName>
<emphasis id="18DF141BFFA0FFED6D97FB7CFA9EFB3E" bold="true" box="[1271,1401,1195,1221]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED6D97FB7CFA9EFB3E" box="[1271,1401,1195,1221]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, is proposed.
</paragraph>
<paragraph id="2A14C809FFA0FFED69A7FB24FB0CFAAE" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">
These specimens matched the photographs of the
<typeStatus id="F51076ABFFA0FFED6A5FFB24FC46FAF6" box="[831,929,1267,1293]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
(http://twinsecttype.nmns.edu.tw/specimen/?id=NMNS-USNM-00042; http://n
<quantity id="ED5365ECFFA0FFED6B4BFAC1FDA6FACA" box="[555,577,1302,1329]" metricMagnitude="3" metricUnit="kg" metricValue="2.0" pageId="16" pageNumber="17" unit="t" value="2.0">2t</quantity>
.net/ark:/65665/30e912f6b-9ac6-4a4c-8f15-67eddf9f871e) and the original description (Drake 1948) of
<taxonomicName id="EDABB38AFFA0FFED68B6FAECFDBEFAAF" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[470,601,1339,1365]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED68B6FAECFE00FAAF" box="[470,487,1339,1364]" italics="true" pageId="16" pageNumber="17">P</emphasis>
.
<emphasis id="18DF141BFFA0FFED6895FAECFDBEFAAF" box="[501,601,1339,1364]" italics="true" pageId="16" pageNumber="17">consueta</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED6B00FAECFD07FAAE" bold="true" box="[608,736,1339,1365]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED6B00FAECFD07FAAE" box="[608,736,1339,1365]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
described from
<collectingCountry id="52BC8899FFA0FFED6AF6FAECFC0FFAAE" box="[918,1000,1339,1365]" name="Taiwan" pageId="16" pageNumber="17">Taiwan</collectingCountry>
in general appearance.
</paragraph>
<paragraph id="2A14C809FFA0FFED69A7FA88FD3DF9F2" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">
<taxonomicName id="EDABB38AFFA0FFED69A7FA88FE37FA83" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[199,464,1375,1401]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED69A7FA88FE37FA83" box="[199,464,1375,1401]" italics="true" pageId="16" pageNumber="17">Physatocheila consueta</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED68B7FA88FDB0FA82" bold="true" box="[471,599,1375,1401]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED68B7FA88FDB0FA82" box="[471,599,1375,1401]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="EDABB38AFFA0FFED6BB4FA88FCB2FA82" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1942" box="[724,853,1375,1401]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="chiniana" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED6BB4FA88FD02FA83" box="[724,741,1375,1400]" italics="true" pageId="16" pageNumber="17">P</emphasis>
.
<emphasis id="18DF141BFFA0FFED6B93FA88FCB2FA82" box="[755,853,1375,1401]" italics="true" pageId="16" pageNumber="17">chiniana</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED6A3BFA88FC3CFA82" bold="true" box="[859,987,1375,1401]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED6A3BFA88FC3CFA82" box="[859,987,1375,1401]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
in general appearance but can be distinguished from it by the following characteristics: paranotum incompletely covering pronotal disc in anterior part, not touching median carina of pronotum (
<figureCitation id="B290D48CFFA0FFED6B5EFA70FD71FA3A" box="[574,662,1447,1473]" captionStart="FIGURE 8" captionStartId="13.[151,250,1851,1875]" captionTargetBox="[157,1428,996,1820]" captionTargetId="figure-410@13.[152,1436,993,1828]" captionTargetPageId="13" captionText="FIGURE 8. Nine Physatocheila species from Japan. Dried specimens, dorsal view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi from Japan proper (E); P. fieberi from the Ryukyu Islands (F); P. miyatakei (G); P. nigrintegerrima (H); P. orientis (I); P. veteris (J). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6609989" httpUri="https://zenodo.org/record/6609989/files/figure.png" pageId="16" pageNumber="17">Fig. 8B</figureCitation>
); costal area of hemelytron with a single row of areolae in apical part and in 2 rows in remaining parts; subcostal area of hemelytron subhorizontal: metasternum wider than mesosternum (
<figureCitation id="B290D48CFFA0FFED69C0FA38FF10F9F2" box="[160,247,1519,1545]" captionStart="FIGURE 9" captionStartId="14.[151,250,1804,1828]" captionTargetBox="[151,1435,489,1780]" captionTargetId="figure-184@14.[151,1435,489,1780]" captionTargetPageId="14" captionText="FIGURE 9. Rostra and sternal laminae of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609991" httpUri="https://zenodo.org/record/6609991/files/figure.png" pageId="16" pageNumber="17">Fig. 9B</figureCitation>
); and metasternal laminae curved outward.
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA0FFED69A7F9C5FF2DF8FE" pageId="16" pageNumber="17" type="distribution">
<paragraph id="2A14C809FFA0FFED69A7F9C5FBB2F9D6" blockId="16.[151,1437,150,2013]" box="[199,1109,1554,1582]" pageId="16" pageNumber="17">
<emphasis id="18DF141BFFA0FFED69A7F9C5FE86F9D7" bold="true" box="[199,353,1554,1580]" pageId="16" pageNumber="17">Distribution.</emphasis>
<collectingCountry id="52BC8899FFA0FFED6807F9C4FE40F9D6" box="[359,423,1555,1581]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
(Ryukyu Islands: Ishigaki Island),
<collectingCountry id="52BC8899FFA0FFED6A51F9C4FC9DF9D6" box="[817,890,1555,1581]" name="South Korea" pageId="16" pageNumber="17">Korea</collectingCountry>
?,
<collectingCountry id="52BC8899FFA0FFED6AEFF9C4FC03F9D6" box="[911,996,1555,1581]" name="Taiwan" pageId="16" pageNumber="17">Taiwan</collectingCountry>
,
<collectingCountry id="52BC8899FFA0FFED6A8FF9C4FBB5F9D6" box="[1007,1106,1555,1581]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="2A14C809FFA0FFED69A7F9E0FF2DF8FE" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">
As mentioned in the section on
<taxonomicName id="EDABB38AFFA0FFED6B49F9E0FD41F9AA" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1942" box="[553,678,1591,1617]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="chiniana" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED6B49F9E0FD41F9AA" box="[553,678,1591,1617]" italics="true" pageId="16" pageNumber="17">P. chiniana</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED6BCDF9E0FCCAF9AA" bold="true" box="[685,813,1591,1617]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED6BCDF9E0FCCAF9AA" box="[685,813,1591,1617]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, previous records of
<taxonomicName id="EDABB38AFFA0FFED6D7AF9E0FB7EF9AB" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[1050,1177,1591,1616]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED6D7AF9E0FB7EF9AB" box="[1050,1177,1591,1616]" italics="true" pageId="16" pageNumber="17">P. consueta</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED6DFFF9E0FAFFF9AA" bold="true" box="[1183,1304,1591,1617]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED6DFFF9E0FAFFF9AA" box="[1183,1304,1591,1617]" pageId="16" pageNumber="17" rank="species">comb nov.</taxonomicNameLabel>
</emphasis>
from
<collectingCountry id="52BC8899FFA0FFED6C3CF9E0FA7BF9AA" box="[1372,1436,1591,1617]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
are all doubtful. Therefore, reliable records of the “true”
<taxonomicName id="EDABB38AFFA0FFED6A4AF98CFC4AF98F" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[810,941,1627,1652]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED6A4AF98CFC4AF98F" box="[810,941,1627,1652]" italics="true" pageId="16" pageNumber="17">P. consueta</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED6AD6F98CFBDEF98E" bold="true" box="[950,1081,1627,1653]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED6AD6F98CFBDEF98E" box="[950,1081,1627,1653]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
from
<collectingCountry id="52BC8899FFA0FFED6DE2F98CFB25F98E" box="[1154,1218,1627,1653]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
are presented here for the first time. The photograph of
<taxonomicName id="EDABB38AFFA0FFED6B57F9A8FD5BF963" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[567,700,1663,1689]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="consueta">
<emphasis id="18DF141BFFA0FFED6B57F9A8FDADF962" box="[567,586,1663,1689]" italics="true" pageId="16" pageNumber="17">C</emphasis>
.
<emphasis id="18DF141BFFA0FFED6B38F9A8FD5BF963" box="[600,700,1663,1688]" italics="true" pageId="16" pageNumber="17">consueta</emphasis>
</taxonomicName>
from
<collectingCountry id="52BC8899FFA0FFED6A62F9A8FCA1F962" box="[770,838,1663,1689]" name="South Korea" pageId="16" pageNumber="17">Korea</collectingCountry>
provided by
<bibRefCitation id="4E3AB5F8FFA0FFED6ABCF9A8FB79F962" author="Cho, G. &amp; Kim, W. &amp; Kwon, Y. S. &amp; Lee, S." box="[988,1182,1663,1689]" pageId="16" pageNumber="17" pagination="736 - 745" refId="ref35840" refString="Cho, G., Kim, W., Kwon, Y. S. &amp; Lee, S. (2020) Check list of lace bugs (Hemiptera: Tingidae) of Korea. Journal of Asia-Pacific Entomology, 23, 736 - 745. https: // doi. org / 10.1016 / j. aspen. 2020.04.005" type="journal article" year="2020">
Cho
<emphasis id="18DF141BFFA0FFED6D73F9A8FBA3F962" box="[1043,1092,1663,1689]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
. (2020)
</bibRefCitation>
represents
<taxonomicName id="EDABB38AFFA0FFED6C7EF9A8FA7BF962" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1942" box="[1310,1436,1663,1689]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="chiniana" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED6C7EF9A8FA7BF962" box="[1310,1436,1663,1689]" italics="true" pageId="16" pageNumber="17">P. chiniana</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED69F7F974FEFFF946" bold="true" box="[151,280,1699,1725]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED69F7F974FEFFF946" box="[151,280,1699,1725]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, therefore the occurrence of
<taxonomicName id="EDABB38AFFA0FFED6B3FF974FD38F947" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[607,735,1699,1724]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED6B3FF974FD38F947" box="[607,735,1699,1724]" italics="true" pageId="16" pageNumber="17">P. consueta</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED6B86F974FC80F946" bold="true" box="[742,871,1699,1725]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED6B86F974FC80F946" box="[742,871,1699,1725]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
in
<collectingCountry id="52BC8899FFA0FFED6AECF974FC37F946" box="[908,976,1699,1725]" name="South Korea" pageId="16" pageNumber="17">Korea</collectingCountry>
is in need of confirmation. Discovery of
<taxonomicName id="EDABB38AFFA0FFED69F7F910FEFCF91B" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[151,283,1735,1761]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED69F7F910FF4FF91B" box="[151,168,1735,1760]" italics="true" pageId="16" pageNumber="17">P</emphasis>
.
<emphasis id="18DF141BFFA0FFED69D7F910FEFCF91B" box="[183,283,1735,1760]" italics="true" pageId="16" pageNumber="17">consueta</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED6842F910FE7BF91A" bold="true" box="[290,412,1735,1761]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED6842F910FE7BF91A" box="[290,412,1735,1761]" pageId="16" pageNumber="17" rank="species">comb nov.</taxonomicNameLabel>
</emphasis>
from Ishigaki Island represents the northernmost reliable distributional record for this species.
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA0FFEC69A7F8D8FB5DFF49" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="biology_ecology">
<paragraph id="2A14C809FFA0FFED69A7F8D8FBA4F8B6" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">
<emphasis id="18DF141BFFA0FFED69A7F8D8FEACF8D2" bold="true" box="[199,331,1807,1833]" pageId="16" pageNumber="17">Host plant.</emphasis>
The first author of the present study confirmed
<taxonomicName id="EDABB38AFFA0FFED6A03F8D8FE32F8B5" authority="(Planch.) Momiy." authorityName="Momiy." baseAuthorityName="Planch." class="Magnoliopsida" family="Vitaceae" genus="Ampelopsis" kingdom="Plantae" order="Vitales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="variety" species="glandulosa" variety="hancei">
<emphasis id="18DF141BFFA0FFED6A03F8D8FB80F8D2" box="[867,1127,1807,1833]" italics="true" pageId="16" pageNumber="17">Ampelopsis glandulosa</emphasis>
(Wall.) Momiy. var.
<emphasis id="18DF141BFFA0FFED6C32F8D8FA7AF8D2" box="[1362,1437,1807,1833]" italics="true" pageId="16" pageNumber="17">hancei</emphasis>
(Planch.) Momiy. (Vitaceae)
</taxonomicName>
(
<figureCitation id="B290D48CFFA0FFED6885F8E4FDAEF8B6" box="[485,585,1843,1869]" captionStart="FIGURE 13" captionStartId="25.[151,250,526,550]" captionTargetBox="[151,1436,181,502]" captionTargetId="figure-17@25.[151,1436,181,502]" captionTargetPageId="25" captionText="FIGURE 13. A living adult (A) and a living fifth instar nymph (B) of Physatocheila consueta comb. nov. from Ishigaki Island, the Ryukyu Islands, Japan (photographed indoor). Host plant (Ampelopsis glandulosa (Wall.) Momiy. var. hancei (Planch.) Momiy.) (C) of P. consueta comb. nov. on Ishigaki Island, the Ryukyu Islands, Japan." figureDoi="http://doi.org/10.5281/zenodo.6610008" httpUri="https://zenodo.org/record/6610008/files/figure.png" pageId="16" pageNumber="17">Fig. 13C</figureCitation>
) as the host plant for
<taxonomicName id="EDABB38AFFA0FFED6A5CF8E4FC5BF8B7" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[828,956,1843,1868]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED6A5CF8E4FC5BF8B7" box="[828,956,1843,1868]" italics="true" pageId="16" pageNumber="17">P. consueta</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED6AA2F8E4FBA4F8B6" bold="true" box="[962,1091,1843,1869]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED6AA2F8E4FBA4F8B6" box="[962,1091,1843,1869]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="2A14C809FFA0FFED69A7F881FD79F842" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">
<emphasis id="18DF141BFFA0FFED69A7F881FEC0F88B" bold="true" box="[199,295,1878,1904]" pageId="16" pageNumber="17">Biology.</emphasis>
A number of individuals of
<taxonomicName id="EDABB38AFFA0FFED6B04F880FC89F88B" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[612,878,1879,1905]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFA0FFED6B04F880FC89F88B" box="[612,878,1879,1905]" italics="true" pageId="16" pageNumber="17">Physatocheila consueta</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA0FFED6A15F880FC11F88A" bold="true" box="[885,1014,1879,1905]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="03ECA960FFA0FFED6A15F880FC11F88A" box="[885,1014,1879,1905]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
were collected from the leaves of the host plant
<taxonomicName id="EDABB38AFFA0FFED686BF8ACFD68F86E" box="[267,655,1915,1941]" class="Magnoliopsida" family="Vitaceae" genus="Ampelopsis" kingdom="Plantae" order="Vitales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="variety" species="glandulosa" variety="hancei">
<emphasis id="18DF141BFFA0FFED686BF8ACFDE9F86E" box="[267,526,1915,1941]" italics="true" pageId="16" pageNumber="17">Ampelopsis glandulosa</emphasis>
var.
<emphasis id="18DF141BFFA0FFED6B24F8ACFD68F86E" box="[580,655,1915,1941]" italics="true" pageId="16" pageNumber="17">hancei</emphasis>
</taxonomicName>
in the Ryukyu Islands, suggesting that this species is folivorous, similar to many tingids (
<bibRefCitation id="4E3AB5F8FFA0FFED6819F848FD75F842" author="Schuh, R. T. &amp; Weirauch, C." box="[377,658,1951,1977]" pageId="16" pageNumber="17" refId="ref40209" refString="Schuh, R. T. &amp; Weirauch, C. (2020) True Bugs of the World (Hemiptera: Heteroptera). Classification and natural history (Second Edition). Siri Scientific Press, Manchester, 768 pp., 32 pls." type="book" year="2020">Schuh &amp; Weirauch 2020</bibRefCitation>
).
</paragraph>
<paragraph id="2A14C809FFA0FFEC69A7F814FB5DFF49" blockId="16.[151,1437,150,2013]" lastBlockId="17.[151,1210,151,249]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
In
<collectingCountry id="52BC8899FFA0FFED6986F814FECEF826" box="[230,297,1987,2013]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
, adults were observed from April to June and in November, and nymphs were observed in November (present study). Conversely, adults were collected from May to July in
<collectingCountry id="52BC8899FFA1FFEC6AD2FF40FBE3FF4A" box="[946,1028,151,177]" name="Taiwan" pageId="17" pageNumber="18">Taiwan</collectingCountry>
(present study).
</paragraph>
</subSubSection>
</treatment>
</document>
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