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<document id="D26F3921449EB193F93285AE8393D223" ID-CLB-Dataset="63559" ID-DOI="10.11646/zootaxa.5150.1.1" ID-GBIF-Dataset="cb0065b0-bad7-4fb5-b6cb-4920f80293cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="6609961" ID-ZooBank="0A7818EA-FBCE-4597-9557-28C208CAC97A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1654244705367" checkinUser="plazi" docAuthor="Souma, Jun &amp; Ishikawa, Tadashi" docDate="2022" docId="A202791FFFA2FFEE69F7FDBDFE7FF974" docLanguage="en" docName="zootaxa.5150.1.1.pdf" docOrigin="Zootaxa 5150 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Physatocheila euthenia Souma &amp; Ishikawa 2022, comb. nov." docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="5E3B0167FFB0FFFD6960FFD7FFE7FFFB" masterDocTitle="Taxonomic review of the tingine genera Cysteochila, Hurdchila, Physatocheila, and Xynotingis from Japan, with description of a new genus (Hemiptera: Heteroptera: Tingidae)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="19" updateTime="1699338851664" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F5BC31DEE3887EBBAAC0CE22030AB60E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DCEB55E86E70EF54B5C4F92B1126EC52">
<mods:title id="E0E41ED78F6E1C26E375CFFB27F397B4">Taxonomic review of the tingine genera Cysteochila, Hurdchila, Physatocheila, and Xynotingis from Japan, with description of a new genus (Hemiptera: Heteroptera: Tingidae)</mods:title>
</mods:titleInfo>
<mods:name id="E703E7F4652486359CDECFD390AF56B3" type="personal">
<mods:role id="ACFA45505FE2450684659C25CA5F9DD4">
<mods:roleTerm id="62A805A50DC8E01774A50253ECEC631B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D32160EA80D7147C20BDDDCAA00A8F1">Souma, Jun</mods:namePart>
<mods:nameIdentifier id="14E487FCA3AFF90726C588DAC110360E" type="ORCID">0000-0002-2238-5015</mods:nameIdentifier>
<mods:affiliation id="0D8EEAF4C020408252F24A1F38442F20">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, Japan. kodokusignal @ gmail. com; https: // orcid. org / 0000 - 0002 - 2238 - 5015 &amp; Research Fellowship for Young Scientists (DC 1), Japan Society for the Promotion of Science, Tokyo, Japan</mods:affiliation>
<mods:nameIdentifier id="BF6E05040EC94133B5365F5F70375650" type="email">kodokusignal@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="54E1B5F2366ECC7EB207D72334B16BD4" type="personal">
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<mods:roleTerm id="F47C96D5C0D5F0B1E4D5F3D549BA5FE3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A0C665EB4F16E5025C1D0F0856AD786">Ishikawa, Tadashi</mods:namePart>
<mods:affiliation id="E67BB4DC875AABB186A7EC79D31F16BB">Laboratory of Entomology, Faculty of Agriculture, Tokyo University of Agriculture, Kanagawa, Japan.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1110485AE2E239F1E5AD81791A89BB5D">text</mods:typeOfResource>
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<mods:titleInfo id="D667CC7DCBDDEC3BEFD41596C4D9EE5D">
<mods:title id="FA5D8725DC66F662DE30346C79489B29">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="29B718CBAD0703D2B8E33A9F36594AE1">
<mods:date id="EE260F512B50AF32B3C3B367F68DD055">2022</mods:date>
<mods:detail id="27C0FCB738585998ECA34387C4FDA9C6" type="pubDate">
<mods:number id="BB375A7E4D439999C08CC2696885D64E">2022-06-03</mods:number>
</mods:detail>
<mods:detail id="34EAFEDD6A4FE4C8377FA9331B3DCAE5" type="volume">
<mods:number id="84A3A9688F0B43CDE48BDD1AA3F6EFC6">5150</mods:number>
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<mods:detail id="EBF786077A70FC85B266875F6BA69E4F" type="issue">
<mods:number id="B790C64FBA0E60CC36ECC2B5B4F8CE91">1</mods:number>
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<mods:start id="4C00E5668BCDEB810F62022D43EC8946">1</mods:start>
<mods:end id="4FC8FD27A0866D4AFAC3C7D1C029516C">42</mods:end>
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<mods:classification id="0C43B10A4EA78BCE5C6A1B696BFE7E18">journal article</mods:classification>
<mods:identifier id="F42A8FA86B99A28419318806403742F3" type="CLB-Dataset">63559</mods:identifier>
<mods:identifier id="D4061ED86CE4A2954766A28F6C24D7E0" type="DOI">10.11646/zootaxa.5150.1.1</mods:identifier>
<mods:identifier id="BC104DA82B61DDFEE226AE6DBE9711C2" type="GBIF-Dataset">cb0065b0-bad7-4fb5-b6cb-4920f80293cc</mods:identifier>
<mods:identifier id="F808EA8B16FEA1B267AD1269DBA3AA57" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A202791FFFA2FFEE69F7FDBDFE7FF974" ID-DOI="http://doi.org/10.5281/zenodo.6611344" ID-GBIF-Taxon="195729302" ID-Zenodo-Dep="6611344" LSID="urn:lsid:plazi:treatment:A202791FFFA2FFEE69F7FDBDFE7FF974" httpUri="http://treatment.plazi.org/id/A202791FFFA2FFEE69F7FDBDFE7FF974" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="62B19B82FFA2FFEF69F7FDBDFC9FFD7E" box="[151,888,618,645]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="2A14C809FFA2FFEF69F7FDBDFC9FFD7E" blockId="18.[151,888,618,681]" box="[151,888,618,645]" pageId="18" pageNumber="19">
<heading id="715C7F65FFA2FFEF69F7FDBDFC9FFD7E" bold="true" box="[151,888,618,645]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="18DF141BFFA2FFEF69F7FDBDFC9FFD7E" bold="true" box="[151,888,618,645]" pageId="18" pageNumber="19">
<taxonomicName id="EDABB38AFFA2FFEF69F7FDBDFD03FD7F" authority="(Drake &amp; Ruhoff, 1965)" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake &amp; Ruhoff" baseAuthorityYear="1965" box="[151,740,618,645]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euthenia" status="comb. nov.">
<emphasis id="18DF141BFFA2FFEF69F7FDBDFE55FD7E" bold="true" box="[151,434,618,645]" italics="true" pageId="18" pageNumber="19">Physatocheila euthenia</emphasis>
(Drake &amp; Ruhoff, 1965)
</taxonomicName>
,
<taxonomicNameLabel id="03ECA960FFA2FFEF6B92FDBCFC9FFD7E" box="[754,888,619,645]" pageId="18" pageNumber="19" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA2FFEF69F7FD58FDC1FD52" box="[151,550,655,681]" pageId="18" pageNumber="19" type="description">
<paragraph id="2A14C809FFA2FFEF69F7FD58FDC1FD52" blockId="18.[151,888,618,681]" box="[151,550,655,681]" pageId="18" pageNumber="19">
(
<figureCitation id="B290D48CFFA2FFEF69FFFD58FEE4FD52" box="[159,259,655,681]" captionStart="FIGURE 8" captionStartId="13.[151,250,1851,1875]" captionTargetBox="[157,1428,996,1820]" captionTargetId="figure-410@13.[152,1436,993,1828]" captionTargetPageId="13" captionText="FIGURE 8. Nine Physatocheila species from Japan. Dried specimens, dorsal view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi from Japan proper (E); P. fieberi from the Ryukyu Islands (F); P. miyatakei (G); P. nigrintegerrima (H); P. orientis (I); P. veteris (J). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6609989" httpUri="https://zenodo.org/record/6609989/files/figure.png" pageId="18" pageNumber="19">Figs. 8D</figureCitation>
,
<figureCitation id="B290D48CFFA2FFEF686FFD58FED4FD52" box="[271,307,655,681]" captionStart="FIGURE 9" captionStartId="14.[151,250,1804,1828]" captionTargetBox="[151,1435,489,1780]" captionTargetId="figure-184@14.[151,1435,489,1780]" captionTargetPageId="14" captionText="FIGURE 9. Rostra and sternal laminae of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609991" httpUri="https://zenodo.org/record/6609991/files/figure.png" pageId="18" pageNumber="19">9D</figureCitation>
,
<figureCitation id="B290D48CFFA2FFEF685FFD58FE95FD52" box="[319,370,655,681]" captionStart="FIGURE 10" captionStartId="19.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1198]" captionTargetId="figure-17@19.[151,1436,181,1198]" captionTargetPageId="19" captionText="FIGURE 10. Male terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609995" httpUri="https://zenodo.org/record/6609995/files/figure.png" pageId="18" pageNumber="19">10D</figureCitation>
,
<figureCitation id="B290D48CFFA2FFEF681DFD58FE48FD52" box="[381,431,655,681]" captionStart="FIGURE 11" captionStartId="21.[151,250,1579,1603]" captionTargetBox="[151,1436,525,1555]" captionTargetId="figure-182@21.[151,1436,525,1555]" captionTargetPageId="21" captionText="FIGURE 11. Female terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6610129" httpUri="https://zenodo.org/record/6610129/files/figure.png" pageId="18" pageNumber="19">11D</figureCitation>
,
<figureCitation id="B290D48CFFA2FFEF68DBFD58FE01FD52" box="[443,486,655,681]" captionStart="FIGURE 17" captionStartId="28.[151,250,1870,1894]" captionTargetBox="[156,1434,888,1845]" captionTargetId="figure-335@28.[151,1436,885,1847]" captionTargetPageId="28" captionText="FIGURE 17. Pygophores of eleven species of Cysteochila and its close relatives from Japan, dorsal view: C. abundantis (A); Limnostatua lewisi comb. nov. (B); Physatocheila chiniana comb. nov. (C); P. consueta comb. nov. (D); P. distinguenda (E); P. euthenia comb. nov. (F); P. fieberi from Japan proper (G); P. fieberi from the Ryukyu Islands (H); P. miyatakei (I); P. nigrintegerrima (J); P. orientis (K); P. veteris (L). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6610019" httpUri="https://zenodo.org/record/6610019/files/figure.png" pageId="18" pageNumber="19">17F</figureCitation>
,
<figureCitation id="B290D48CFFA2FFEF6892FD58FDF9FD52" box="[498,542,655,681]" captionStart="FIGURE 18" captionStartId="29.[151,250,1878,1902]" captionTargetBox="[154,1433,1025,1851]" captionTargetId="figure-333@29.[151,1436,1022,1854]" captionTargetPageId="29" captionText="FIGURE 18. Parameres of eleven species of Cysteochila and its close relatives from Japan, dorsal view: C. abundantis (A); Limnostatua lewisi comb. nov. (B); Physatocheila chiniana comb. nov. (C); P. consueta comb. nov. (D); P. distinguenda (E); P. euthenia comb. nov. (F); P. fieberi from Japan proper (G); P. fieberi from the Ryukyu Islands (H); P. miyatakei (I); P. nigrintegerrima (J); P. orientis (K); P. veteris (L). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6610022" httpUri="https://zenodo.org/record/6610022/files/figure.png" pageId="18" pageNumber="19">18F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA2FFEF69F7FD04FB6DFC07" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="2A14C809FFA2FFEF69F7FD04FEF2FCF2" blockId="18.[151,1437,723,777]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="0ABBEF27FFA2FFEF69F7FD04FEF2FCF2" pageId="18" pageNumber="19">
<treatmentCitation id="AB0AEE18FFA2FFEF69F7FD04FD56FD10" author="Drake, C. J. &amp; Ruhoff, F. A." box="[151,689,723,747]" page="254" pageId="18" pageNumber="19" year="1965">
<taxonomicName id="EDABB38AFFA2FFEF69F7FD04FD56FD10" ID-CoL="33PBG" authority="Drake &amp; Ruhoff, 1965 b: 254" authorityName="Drake &amp; Ruhoff" authorityPageNumber="254" authorityYear="1965" box="[151,689,723,747]" class="Insecta" family="Tingidae" genus="Cysteochila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euthenia">
<emphasis id="18DF141BFFA2FFEF69F7FD04FE97FD10" box="[151,368,723,747]" italics="true" pageId="18" pageNumber="19">Cysteochila euthenia</emphasis>
<materialsCitation id="9AC3C254FFA2FFEF681AFD04FD56FD10" ID-GBIF-Occurrence="3802857310" box="[378,689,723,747]" collectorName="N. Borneo" country="Malaysia" location="Sabah" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<bibRefCitation id="4E3AB5F8FFA2FFEF681AFD04FD56FD10" author="Drake, C. J. &amp; Ruhoff, F. A." box="[378,689,723,747]" pageId="18" pageNumber="19" pagination="243 - 290" refId="ref36581" refString="Drake, C. J. &amp; Ruhoff, F. A. (1965 b) Lacebugs from New Guinea, Borneo, Solomons, and other islands of the South Pacific and Indian oceans (Hemiptera: Tingidae). Pacific Insects, 7 (2), 243 - 290." type="journal article" year="1965">Drake &amp; Ruhoff, 1965b: 254</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="F51076ABFFA2FFEF6BDEFD04FCC7FD10" box="[702,800,723,747]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
: macropterous
<specimenCount id="3CAD0380FFA2FFEF6AA2FD03FC30FD10" box="[962,983,724,747]" pageId="18" pageNumber="19" type="male">♂</specimenCount>
,
<collectingCountry id="52BC8899FFA2FFEF6A85FD04FBA1FD10" box="[997,1094,723,747]" name="Malaysia" pageId="18" pageNumber="19">Malaysia</collectingCountry>
:
<collectingRegion id="E86F06EBFFA2FFEF6D33FD04FB76FD10" box="[1107,1169,723,747]" country="Malaysia" name="Sabah" pageId="18" pageNumber="19">Sabah</collectingRegion>
(
<collectorName id="875EADDFFFA2FFEF6DC3FD04FAF3FD10" box="[1187,1300,723,747]" pageId="18" pageNumber="19">N. Borneo</collectorName>
), Tenompok; BPBM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2A14C809FFA2FFEF69F7FCF8FBA4FCBC" blockId="18.[151,1091,815,839]" box="[151,1091,815,839]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="0ABBEF27FFA2FFEF69F7FCF8FBA4FCBC" box="[151,1091,815,839]" pageId="18" pageNumber="19">
<treatmentCitation id="AB0AEE18FFA2FFEF69F7FCF8FDA3FCBC" author="Pericart, J." box="[151,580,815,839]" page=")" pageId="18" pageNumber="19" year="1985">
<taxonomicName id="EDABB38AFFA2FFEF69F7FCF8FDA3FCBC" ID-CoL="33PBG" authority=": Pericart (1985 a: 48)" authorityName="Pericart" authorityPageNumber="48" authorityYear="1985" box="[151,580,815,839]" class="Insecta" family="Tingidae" genus="Cysteochila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euthenia">
<emphasis id="18DF141BFFA2FFEF69F7FCF8FE8BFCBC" box="[151,364,815,839]" italics="true" pageId="18" pageNumber="19">Cysteochila euthenia</emphasis>
:
<bibRefCitation id="4E3AB5F8FFA2FFEF681AFCF8FDA3FCBC" author="Pericart, J." box="[378,580,815,839]" pageId="18" pageNumber="19" pagination="27 - 62" refId="ref39935" refString="Pericart, J. (1985 a) Tingidae nouveaux ou interessants du Nord du sous-continent indien (Hemiptera). II. Entomologica Basiliensia, 10, 27 - 62. [in French, English summary]" type="journal article" year="1985">Péricart (1985a: 48)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(distribution);
<treatmentCitation id="AB0AEE18FFA2FFEF6BBEFCF8FC48FCBC" ID-DOI="http://doi.org/10.5281/zenodo.6113567" ID-Zenodo-Dep="6113567" author="Guilbert, E." box="[734,943,815,839]" httpUri="http://treatment.plazi.org/id/191287AE891CFFFFFF41FE1E1E8F062F" page="536" pageId="18" pageNumber="19" year="2015">
<bibRefCitation id="4E3AB5F8FFA2FFEF6BBEFCF8FC48FCBC" author="Guilbert, E." box="[734,943,815,839]" pageId="18" pageNumber="19" pagination="531 - 546" refId="ref37512" refString="Guilbert, E. (2015) New species and new records of Tingidae (Hemiptera: Heteroptera) from Vietnam. Zootaxa, 3956 (4), 531 - 546. https: // doi. org / 10.11646 / zootaxa. 3956.4.5" type="journal article" year="2015">Guilbert (2015: 536)</bibRefCitation>
</treatmentCitation>
(distribution).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2A14C809FFA2FFEF69F7FCA2FB6DFC07" blockId="18.[151,1437,885,2027]" pageId="18" pageNumber="19">
<emphasis id="18DF141BFFA2FFEF69F7FCA2FE71FC74" bold="true" box="[151,406,885,911]" pageId="18" pageNumber="19">Specimens examined.</emphasis>
<materialsCitation id="9AC3C254FFA2FFEF68FCFCA1FD7DFC6B" ID-GBIF-Occurrence="3802857343" box="[412,666,885,912]" location="Non-types" pageId="18" pageNumber="19" specimenCount="5" specimenCount-female="1" specimenCount-male="4">
<location id="2F749ED2FFA2FFEF68FCFCA1FDF6FC6B" LSID="urn:lsid:plazi:treatment:A202791FFFA2FFEE69F7FDBDFE7FF974:2F749ED2FFA2FFEF68FCFCA1FDF6FC6B" box="[412,529,886,912]" name="Non-types" pageId="18" pageNumber="19">Non-types</location>
(
<specimenCount id="3CAD0380FFA2FFEF6B40FCA2FDB8FC6B" box="[544,607,885,912]" pageId="18" pageNumber="19" type="male">4 ♂♂</specimenCount>
<specimenCount id="3CAD0380FFA2FFEF6B06FCA2FD6AFC6B" box="[614,653,885,912]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="9AC3C254FFA2FFEF6BC6FCA2FDE5FC4F" ID-GBIF-Occurrence="3802857383" collectingDate="1960-11-23" collectionCode="ELKU" collectorName="K. Yasumatsu" country="Japan" location="Yaeyama Islands" municipality="Ryukyu Islands" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<emphasis id="18DF141BFFA2FFEF6BC6FCA2FB43FC74" bold="true" box="[678,1188,885,912]" pageId="18" pageNumber="19">
<collectingCountry id="52BC8899FFA2FFEF6BC6FCA2FCE3FC74" box="[678,772,885,911]" name="Japan" pageId="18" pageNumber="19">JAPAN</collectingCountry>
:
<collectingMunicipality id="CA705273FFA2FFEF6A70FCA2FC29FC74" box="[784,974,885,911]" pageId="18" pageNumber="19">Ryukyu Islands</collectingMunicipality>
:
<emphasis id="18DF141BFFA2FFEF6AB9FCA1FB7CFC6B" bold="true" box="[985,1179,885,912]" italics="true" pageId="18" pageNumber="19">
<location id="2F749ED2FFA2FFEF6AB9FCA1FB7CFC6B" LSID="urn:lsid:plazi:treatment:A202791FFFA2FFEE69F7FDBDFE7FF974:2F749ED2FFA2FFEF6AB9FCA1FB7CFC6B" box="[985,1179,885,912]" country="Japan" municipality="Ryukyu Islands" name="Yaeyama Islands" pageId="18" pageNumber="19" stateProvince="Okinawa">Yaeyama Islands</location>
</emphasis>
:
</emphasis>
<collectingRegion id="E86F06EBFFA2FFEF6DCBFCA2FAF4FC74" box="[1195,1299,885,911]" country="Japan" name="Okinawa" pageId="18" pageNumber="19">Okinawa</collectingRegion>
,
<date id="5E15EEC9FFA2FFEF6C7DFCA2FA70FC74" box="[1309,1431,885,911]" pageId="18" pageNumber="19" value="1960-11-23">
<collectingDate id="4E511721FFA2FFEF6C7DFCA2FA70FC74" box="[1309,1431,885,911]" pageId="18" pageNumber="19" value="1960-11-23">23.xi.1960</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA2FFEF69A7FC4DFE85FC4F" box="[199,354,922,948]" pageId="18" pageNumber="19">K. Yasumatsu</collectorName>
(
<specimenCount id="3CAD0380FFA2FFEF6814FC4EFE79FC4F" box="[372,414,921,948]" pageId="18" pageNumber="19" type="male">1 ♂</specimenCount>
,
<collectionCode id="4CBA50CCFFA2FFEF68CAFC4DFE1DFC4F" box="[426,506,922,948]" pageId="18" pageNumber="19">ELKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA2FFEF6B6DFC4EFB52FC4F" ID-GBIF-Occurrence="3802857439" box="[525,1205,921,948]" collectingDate="1963-10-08" collectionCode="KUM" collectorName="K. Morimoto" country="Japan" location="Sonai" municipality="Iriomote Is." pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa">
<collectingMunicipality id="CA705273FFA2FFEF6B6DFC4EFD76FC4F" box="[525,657,921,948]" pageId="18" pageNumber="19">Iriomote Is.</collectingMunicipality>
,
<location id="2F749ED2FFA2FFEF6BFDFC4EFD07FC48" LSID="urn:lsid:plazi:treatment:A202791FFFA2FFEE69F7FDBDFE7FF974:2F749ED2FFA2FFEF6BFDFC4EFD07FC48" box="[669,736,921,947]" country="Japan" municipality="Iriomote Is." name="Sonai" pageId="18" pageNumber="19" stateProvince="Okinawa">Sonai</location>
,
<date id="5E15EEC9FFA2FFEF6B8BFC4EFCA8FC48" box="[747,847,921,947]" pageId="18" pageNumber="19" value="1963-10-08">
<collectingDate id="4E511721FFA2FFEF6B8BFC4EFCA8FC48" box="[747,847,921,947]" pageId="18" pageNumber="19" value="1963-10-08">8.x.1963</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA2FFEF6AEBFC4DFBF8FC48" box="[907,1055,921,948]" pageId="18" pageNumber="19">K. Morimoto</collectorName>
(
<specimenCount id="3CAD0380FFA2FFEF6D50FC4EFBBFFC4F" box="[1072,1112,921,948]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
,
<collectionCode id="4CBA50CCFFA2FFEF6D06FC4DFB4BFC4F" box="[1126,1196,922,948]" pageId="18" pageNumber="19">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA2FFEF6DA0FC4EFD1AFC2C" ID-GBIF-Occurrence="3802857495" collectingDate="1991-03-27" collectionCode="NSMT" collectorName="M. Hayashi" country="Japan" location="Iriomote Is." pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<location id="2F749ED2FFA2FFEF6DA0FC4EFAA3FC4F" LSID="urn:lsid:plazi:treatment:A202791FFFA2FFEE69F7FDBDFE7FF974:2F749ED2FFA2FFEF6DA0FC4EFAA3FC4F" box="[1216,1348,921,948]" country="Japan" name="Iriomote Is." pageId="18" pageNumber="19" stateProvince="Okinawa">Iriomote Is.</location>
, Hoshidate,
<date id="5E15EEC9FFA2FFEF69B5FC6AFEB6FC2C" box="[213,337,957,983]" pageId="18" pageNumber="19" value="1991-03-27">
<collectingDate id="4E511721FFA2FFEF69B5FC6AFEB6FC2C" box="[213,337,957,983]" pageId="18" pageNumber="19" value="1991-03-27">27.iii.1991</collectingDate>
</date>
, leg.
<collectorName id="875EADDFFFA2FFEF68F0FC69FDF3FC2C" box="[400,532,957,984]" pageId="18" pageNumber="19">M. Hayashi</collectorName>
<emphasis id="18DF141BFFA2FFEF6B7CFC69FDA8FC2C" box="[540,591,957,983]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (
<specimenCount id="3CAD0380FFA2FFEF6B09FC6AFD72FC23" box="[617,661,957,984]" pageId="18" pageNumber="19" type="male">1 ♂</specimenCount>
,
<collectionCode id="4CBA50CCFFA2FFEF6BC2FC6AFD12FC2C" box="[674,757,957,983]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="18" pageNumber="19">NSMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9AC3C254FFA2FFEF6A6AFC69FB25FC23" ID-GBIF-Occurrence="3802857441" box="[778,1218,957,984]" collectingDate="1996-03-30" collectionCode="TUA" country="Japan" pageId="18" pageNumber="19" specimenCount="2" specimenCount-male="2" stateProvince="Okinawa">
as above but
<date id="5E15EEC9FFA2FFEF6AC1FC6AFBFDFC2C" box="[929,1050,957,983]" pageId="18" pageNumber="19" value="1996-03-30">
<collectingDate id="4E511721FFA2FFEF6AC1FC6AFBFDFC2C" box="[929,1050,957,983]" pageId="18" pageNumber="19" value="1996-03-30">30.iii.1996</collectingDate>
</date>
(
<specimenCount id="3CAD0380FFA2FFEF6D4BFC6AFB89FC23" box="[1067,1134,957,984]" pageId="18" pageNumber="19" type="male">2 ♂♂</specimenCount>
,
<collectionCode id="4CBA50CCFFA2FFEF6D1BFC69FB5DFC23" box="[1147,1210,958,984]" pageId="18" pageNumber="19">TUA</collectionCode>
)
</materialsCitation>
. A single specimen deposited in ELKU is labeled “
<taxonomicName id="EDABB38AFFA2FFEF6896FC36FC79FC00" authority="Takeya, 1962" authorityName="Takeya" authorityYear="1962" box="[502,926,993,1019]" class="Insecta" family="Tingidae" genus="Cysteochila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="yasumatsui">
<emphasis id="18DF141BFFA2FFEF6896FC36FD18FC00" box="[502,767,993,1019]" italics="true" pageId="18" pageNumber="19">Cysteochila yasumatsui</emphasis>
Takeya, 1962
</taxonomicName>
” (manuscript name).
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA2FFEF69A7FBD2FB45FAC4" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="2A14C809FFA2FFEF69A7FBD2FB45FAC4" blockId="18.[151,1437,885,2027]" pageId="18" pageNumber="19">
<emphasis id="18DF141BFFA2FFEF69A7FBD2FEA5FBE4" bold="true" box="[199,322,1029,1055]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
Recognized among other species of
<taxonomicName id="EDABB38AFFA2FFEF6B8AFBD2FC6EFBE4" authorityName="Fieber" authorityYear="1844" box="[746,905,1029,1055]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="18DF141BFFA2FFEF6B8AFBD2FC6EFBE4" box="[746,905,1029,1055]" italics="true" pageId="18" pageNumber="19">Physatocheila</emphasis>
</taxonomicName>
by a combination of the following characters: general color ashen (
<figureCitation id="B290D48CFFA2FFEF68E2FBFEFE01FBB8" box="[386,486,1065,1091]" captionStart="FIGURE 8" captionStartId="13.[151,250,1851,1875]" captionTargetBox="[157,1428,996,1820]" captionTargetId="figure-410@13.[152,1436,993,1828]" captionTargetPageId="13" captionText="FIGURE 8. Nine Physatocheila species from Japan. Dried specimens, dorsal view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi from Japan proper (E); P. fieberi from the Ryukyu Islands (F); P. miyatakei (G); P. nigrintegerrima (H); P. orientis (I); P. veteris (J). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6609989" httpUri="https://zenodo.org/record/6609989/files/figure.png" pageId="18" pageNumber="19">Figs. 8D</figureCitation>
,
<figureCitation id="B290D48CFFA2FFEF6893FBFEFDF0FBB8" box="[499,535,1065,1091]" captionStart="FIGURE 9" captionStartId="14.[151,250,1804,1828]" captionTargetBox="[151,1435,489,1780]" captionTargetId="figure-184@14.[151,1435,489,1780]" captionTargetPageId="14" captionText="FIGURE 9. Rostra and sternal laminae of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609991" httpUri="https://zenodo.org/record/6609991/files/figure.png" pageId="18" pageNumber="19">9D</figureCitation>
,
<figureCitation id="B290D48CFFA2FFEF6B44FBFEFDBFFBB8" box="[548,600,1065,1091]" captionStart="FIGURE 10" captionStartId="19.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1198]" captionTargetId="figure-17@19.[151,1436,181,1198]" captionTargetPageId="19" captionText="FIGURE 10. Male terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609995" httpUri="https://zenodo.org/record/6609995/files/figure.png" pageId="18" pageNumber="19">10D</figureCitation>
); median spine on head extending beyond bases of frontal spines; rostrum reaching middle part of metasternum; pubescence on pronotum more than 0.5 times as long as diameter of compound eye; lateral carinae of pronotum close to each other in anterior part, concealed by paranotum in anterior part; paranota completely covering pronotal disc in anterior part, not touching each other, not concealing median carina of pronotum, widened posteriorly, bulged upward in posterior part, forming a cyst; cyst of paranotum concave in posterior part; outer margin of paranotum gently curved outward in its entire length; costal area of hemelytron wider than subcostal area at widest part, more than 0.5 times as wide as discoidal area at widest part, with 2 rows of areolae in its entire length; and subcostal area subvertical, with 2 rows of areolae in its entire length.
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA2FFEF69A7FA9EFEBEFA08" pageId="18" pageNumber="19" type="description">
<paragraph id="2A14C809FFA2FFEF69A7FA9EFEBEFA08" blockId="18.[151,1437,885,2027]" pageId="18" pageNumber="19">
<emphasis id="18DF141BFFA2FFEF69A7FA9EFE04FA98" bold="true" box="[199,483,1353,1379]" pageId="18" pageNumber="19">Description of genitalia.</emphasis>
Pygophore (
<figureCitation id="B290D48CFFA2FFEF6B11FA9EFD05FA98" box="[625,738,1353,1379]" captionStart="FIGURE 10" captionStartId="19.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1198]" captionTargetId="figure-17@19.[151,1436,181,1198]" captionTargetPageId="19" captionText="FIGURE 10. Male terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609995" httpUri="https://zenodo.org/record/6609995/files/figure.png" pageId="18" pageNumber="19">Figs. 10D</figureCitation>
,
<figureCitation id="B290D48CFFA2FFEF6B8DFA9EFCFEFA98" box="[749,793,1353,1379]" captionStart="FIGURE 17" captionStartId="28.[151,250,1870,1894]" captionTargetBox="[156,1434,888,1845]" captionTargetId="figure-335@28.[151,1436,885,1847]" captionTargetPageId="28" captionText="FIGURE 17. Pygophores of eleven species of Cysteochila and its close relatives from Japan, dorsal view: C. abundantis (A); Limnostatua lewisi comb. nov. (B); Physatocheila chiniana comb. nov. (C); P. consueta comb. nov. (D); P. distinguenda (E); P. euthenia comb. nov. (F); P. fieberi from Japan proper (G); P. fieberi from the Ryukyu Islands (H); P. miyatakei (I); P. nigrintegerrima (J); P. orientis (K); P. veteris (L). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6610019" httpUri="https://zenodo.org/record/6610019/files/figure.png" pageId="18" pageNumber="19">17F</figureCitation>
) compressed dorsoventrally, semicircular in ventral view, strongly concave at anterior margin of dorsum, elevated at center of venter, smooth on surface, irregularly punctate in middle part of dorsum. Paramere (
<figureCitation id="B290D48CFFA2FFEF6B4DFA46FD6BFA50" box="[557,652,1425,1451]" captionStart="FIGURE 18" captionStartId="29.[151,250,1878,1902]" captionTargetBox="[154,1433,1025,1851]" captionTargetId="figure-333@29.[151,1436,1022,1854]" captionTargetPageId="29" captionText="FIGURE 18. Parameres of eleven species of Cysteochila and its close relatives from Japan, dorsal view: C. abundantis (A); Limnostatua lewisi comb. nov. (B); Physatocheila chiniana comb. nov. (C); P. consueta comb. nov. (D); P. distinguenda (E); P. euthenia comb. nov. (F); P. fieberi from Japan proper (G); P. fieberi from the Ryukyu Islands (H); P. miyatakei (I); P. nigrintegerrima (J); P. orientis (K); P. veteris (L). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6610022" httpUri="https://zenodo.org/record/6610022/files/figure.png" pageId="18" pageNumber="19">Fig. 18F</figureCitation>
) expanded in middle part, curved inward in apical part; outer and inner margins covered with pubescence in middle part. Female terminalia (
<figureCitation id="B290D48CFFA2FFEF6AFBFA62FC18FA34" box="[923,1023,1461,1487]" captionStart="FIGURE 11" captionStartId="21.[151,250,1579,1603]" captionTargetBox="[151,1436,525,1555]" captionTargetId="figure-182@21.[151,1436,525,1555]" captionTargetPageId="21" captionText="FIGURE 11. Female terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6610129" httpUri="https://zenodo.org/record/6610129/files/figure.png" pageId="18" pageNumber="19">Fig. 11D</figureCitation>
) pentagonal in ventral view, covered with pubescence.
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA2FFEF69A7FA2AFDB6F85F" pageId="18" pageNumber="19" type="discussion">
<paragraph id="2A14C809FFA2FFEF69A7FA2AFDB4F9A4" blockId="18.[151,1437,885,2027]" pageId="18" pageNumber="19">
<emphasis id="18DF141BFFA2FFEF69A7FA2AFEDCF9EC" bold="true" box="[199,315,1533,1559]" pageId="18" pageNumber="19">Remarks.</emphasis>
This species belongs to
<taxonomicName id="EDABB38AFFA2FFEF6B32FA2AFD16F9EC" authorityName="Fieber" authorityYear="1844" box="[594,753,1533,1559]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="18DF141BFFA2FFEF6B32FA2AFD16F9EC" box="[594,753,1533,1559]" italics="true" pageId="18" pageNumber="19">Physatocheila</emphasis>
</taxonomicName>
without doubt because of the presence of generic characteristics, as described in the Diagnosis section. Therefore, the new combination,
<taxonomicName id="EDABB38AFFA2FFEF6D7CF9F6FED2F99B" authority="(Drake &amp; Ruhoff, 1965)" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake &amp; Ruhoff" baseAuthorityYear="1965" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euthenia" status="comb. nov.">
<emphasis id="18DF141BFFA2FFEF6D7CF9F6FAC2F9C0" box="[1052,1317,1569,1595]" italics="true" pageId="18" pageNumber="19">Physatocheila euthenia</emphasis>
(Drake &amp; Ruhoff, 1965)
</taxonomicName>
<emphasis id="18DF141BFFA2FFEF685CF992FE5AF9A4" bold="true" box="[316,445,1605,1631]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="03ECA960FFA2FFEF685CF992FE5AF9A4" box="[316,445,1605,1631]" pageId="18" pageNumber="19" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, is proposed.
</paragraph>
<paragraph id="2A14C809FFA2FFEF69A7F9BEFA93F8E8" blockId="18.[151,1437,885,2027]" pageId="18" pageNumber="19">
The aforementioned specimens match the illustration (http://n
<quantity id="ED5365ECFFA2FFEF6AF4F9BEFC4DF97F" box="[916,938,1641,1668]" metricMagnitude="3" metricUnit="kg" metricValue="2.0" pageId="18" pageNumber="19" unit="t" value="2.0">2t</quantity>
.net/ark:/65665/369f879a9-f75d-467b-b4c9- 6269c9c8361f) and the original description (
<bibRefCitation id="4E3AB5F8FFA2FFEF6BE6F95AFC6CF953" author="Drake, C. J. &amp; Ruhoff, F. A." box="[646,907,1677,1704]" pageId="18" pageNumber="19" pagination="243 - 290" refId="ref36581" refString="Drake, C. J. &amp; Ruhoff, F. A. (1965 b) Lacebugs from New Guinea, Borneo, Solomons, and other islands of the South Pacific and Indian oceans (Hemiptera: Tingidae). Pacific Insects, 7 (2), 243 - 290." type="journal article" year="1965">Drake &amp; Ruhoff 1965b</bibRefCitation>
) of
<taxonomicName id="EDABB38AFFA2FFEF6AD7F959FBD5F95C" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake &amp; Ruhoff" baseAuthorityYear="1965" box="[951,1074,1677,1703]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euthenia" status="comb. nov.">
<emphasis id="18DF141BFFA2FFEF6AD7F959FBD5F95C" box="[951,1074,1677,1703]" italics="true" pageId="18" pageNumber="19">P. euthenia</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA2FFEF6D59F95AFB5EF95C" bold="true" box="[1081,1209,1677,1703]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="03ECA960FFA2FFEF6D59F95AFB5EF95C" box="[1081,1209,1677,1703]" pageId="18" pageNumber="19" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
described from
<collectingCountry id="52BC8899FFA2FFEF6C0EF959FF3CF930" name="Malaysia" pageId="18" pageNumber="19">Malaysia</collectingCountry>
, with the exception of the rostrum reaching the middle part of the metasternum. Because it is currently unclear whether the difference in the length of the rostrum indicates a difference at the species level or reflects infraspecific variation within the same species, the Japanese population is provisionally identified as
<taxonomicName id="EDABB38AFFA2FFEF6D10F92DFB0BF8E8" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake &amp; Ruhoff" baseAuthorityYear="1965" box="[1136,1260,1785,1811]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euthenia" status="comb. nov.">
<emphasis id="18DF141BFFA2FFEF6D10F92DFB0BF8E8" box="[1136,1260,1785,1811]" italics="true" pageId="18" pageNumber="19">P. euthenia</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA2FFEF6D93F92EFA93F8E8" bold="true" box="[1267,1396,1785,1811]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="03ECA960FFA2FFEF6D93F92EFA93F8E8" box="[1267,1396,1785,1811]" pageId="18" pageNumber="19" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="2A14C809FFA2FFEF69A7F8CAFDB6F85F" blockId="18.[151,1437,885,2027]" pageId="18" pageNumber="19">
<taxonomicName id="EDABB38AFFA2FFEF69A7F8CAFE28F8CC" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake &amp; Ruhoff" baseAuthorityYear="1965" box="[199,463,1821,1847]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euthenia" status="comb. nov.">
<emphasis id="18DF141BFFA2FFEF69A7F8CAFE28F8CC" box="[199,463,1821,1847]" italics="true" pageId="18" pageNumber="19">Physatocheila euthenia</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA2FFEF68B9F8CAFDBBF8CC" bold="true" box="[473,604,1821,1847]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="03ECA960FFA2FFEF68B9F8CAFDBBF8CC" box="[473,604,1821,1847]" pageId="18" pageNumber="19" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="EDABB38AFFA2FFEF6BBFF8C9FCAEF8CC" baseAuthorityName="Scott" baseAuthorityYear="1874" box="[735,841,1821,1848]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fieberi">
<emphasis id="18DF141BFFA2FFEF6BBFF8C9FD17F8CC" box="[735,752,1822,1847]" italics="true" pageId="18" pageNumber="19">P</emphasis>
.
<emphasis id="18DF141BFFA2FFEF6A60F8CAFCAEF8CC" box="[768,841,1821,1847]" italics="true" pageId="18" pageNumber="19">fieberi</emphasis>
</taxonomicName>
in the shape of paranotum but can be easily distinguished from the latter by the following characteristics: general color ashen (
<figureCitation id="B290D48CFFA2FFEF6D68F896FB8AF8A0" box="[1032,1133,1857,1883]" captionStart="FIGURE 8" captionStartId="13.[151,250,1851,1875]" captionTargetBox="[157,1428,996,1820]" captionTargetId="figure-410@13.[152,1436,993,1828]" captionTargetPageId="13" captionText="FIGURE 8. Nine Physatocheila species from Japan. Dried specimens, dorsal view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi from Japan proper (E); P. fieberi from the Ryukyu Islands (F); P. miyatakei (G); P. nigrintegerrima (H); P. orientis (I); P. veteris (J). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6609989" httpUri="https://zenodo.org/record/6609989/files/figure.png" pageId="18" pageNumber="19">Figs. 8D</figureCitation>
,
<figureCitation id="B290D48CFFA2FFEF6D1AF896FB78F8A0" box="[1146,1183,1857,1883]" captionStart="FIGURE 9" captionStartId="14.[151,250,1804,1828]" captionTargetBox="[151,1435,489,1780]" captionTargetId="figure-184@14.[151,1435,489,1780]" captionTargetPageId="14" captionText="FIGURE 9. Rostra and sternal laminae of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609991" httpUri="https://zenodo.org/record/6609991/files/figure.png" pageId="18" pageNumber="19">9D</figureCitation>
,
<figureCitation id="B290D48CFFA2FFEF6DCCF896FB06F8A0" box="[1196,1249,1857,1883]" captionStart="FIGURE 10" captionStartId="19.[151,250,1222,1246]" captionTargetBox="[151,1436,181,1198]" captionTargetId="figure-17@19.[151,1436,181,1198]" captionTargetPageId="19" captionText="FIGURE 10. Male terminalia of nine Physatocheila species from Japan, ventral view: P. chiniana comb. nov. (A); P. consueta comb. nov. (B); P. distinguenda (C); P. euthenia comb. nov. (D); P. fieberi (E); P. miyatakei (F); P. nigrintegerrima (G); P. orientis (H); P. veteris (I). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6609995" httpUri="https://zenodo.org/record/6609995/files/figure.png" pageId="18" pageNumber="19">10D</figureCitation>
); pubescence on pronotum more than 0.5 times as long as diameter of compound eye; costal area of hemelytron more than 0.5 times as wide as discoidal area at widest part.
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA2FFEE69A7F87AFB31FA05" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="distribution">
<paragraph id="2A14C809FFA2FFEF69A7F87AFF19F810" blockId="18.[151,1437,885,2027]" pageId="18" pageNumber="19">
<emphasis id="18DF141BFFA2FFEF69A7F87AFE86F83C" bold="true" box="[199,353,1965,1991]" pageId="18" pageNumber="19">Distribution.</emphasis>
<collectingCountry id="52BC8899FFA2FFEF680BF879FE4CF833" box="[363,427,1966,1992]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
(Ryukyu Islands:
<collectingRegion id="E86F06EBFFA2FFEF6BE1F87AFD01F83C" box="[641,742,1965,1991]" country="Japan" name="Okinawa" pageId="18" pageNumber="19">Okinawa</collectingRegion>
Island?, Iriomote Island),
<collectingCountry id="52BC8899FFA2FFEF6D7CF87AFB64F83C" box="[1052,1155,1965,1991]" name="Malaysia" pageId="18" pageNumber="19">Malaysia</collectingCountry>
(Borneo Island),
<collectingCountry id="52BC8899FFA2FFEF6C32F87AFA7FF83C" box="[1362,1432,1965,1991]" name="Nepal" pageId="18" pageNumber="19">Nepal</collectingCountry>
,
<collectingCountry id="52BC8899FFA2FFEF69F7F806FF1DF810" box="[151,250,2001,2027]" name="Vietnam" pageId="18" pageNumber="19">Vietnam</collectingCountry>
.
</paragraph>
<caption id="7ED49881FFA3FFEE69F7FB11FD96FADE" ID-DOI="http://doi.org/10.5281/zenodo.6609995" ID-Zenodo-Dep="6609995" httpUri="https://zenodo.org/record/6609995/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1222,1246]" targetBox="[151,1436,181,1198]" targetPageId="19">
<paragraph id="2A14C809FFA3FFEE69F7FB11FD96FADE" blockId="19.[151,1436,1222,1318]" pageId="19" pageNumber="20">
<emphasis id="18DF141BFFA3FFEE69F7FB11FEF8FB25" bold="true" box="[151,287,1222,1246]" pageId="19" pageNumber="20">FIGURE 10.</emphasis>
Male terminalia of nine
<taxonomicName id="EDABB38AFFA3FFEE6B78FB11FD11FB25" box="[536,758,1222,1246]" pageId="19" pageNumber="20">
<emphasis id="18DF141BFFA3FFEE6B78FB11FD4FFB25" box="[536,680,1222,1246]" italics="true" pageId="19" pageNumber="20">Physatocheila</emphasis>
species
</taxonomicName>
from Japan, ventral view:
<taxonomicName id="EDABB38AFFA3FFEE6D63FB10FB9FFB25" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1942" box="[1027,1144,1222,1246]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="chiniana" status="comb. nov.">
<emphasis id="18DF141BFFA3FFEE6D63FB10FBF5FB25" box="[1027,1042,1223,1246]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="18DF141BFFA3FFEE6D7FFB11FB9FFB25" box="[1055,1144,1222,1246]" italics="true" pageId="19" pageNumber="20">chiniana</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA3FFEE6D1DFB11FB17FB25" bold="true" box="[1149,1264,1222,1246]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="03ECA960FFA3FFEE6D1DFB11FB17FB25" box="[1149,1264,1222,1246]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(A);
<taxonomicName id="EDABB38AFFA3FFEE6C47FB10FA7BFB25" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake" baseAuthorityYear="1948" box="[1319,1436,1222,1246]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="consueta" status="comb. nov.">
<emphasis id="18DF141BFFA3FFEE6C47FB10FAD1FB25" box="[1319,1334,1223,1246]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="18DF141BFFA3FFEE6C22FB10FA7BFB25" box="[1346,1436,1223,1246]" italics="true" pageId="19" pageNumber="20">consueta</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA3FFEE69F7FB3DFEEBFAF9" bold="true" box="[151,268,1258,1282]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="03ECA960FFA3FFEE69F7FB3DFEEBFAF9" box="[151,268,1258,1282]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(B);
<taxonomicName id="EDABB38AFFA3FFEE6824FB3CFE03FAF9" authorityName=": Horvath" authorityYear="1906" box="[324,484,1258,1282]" class="Insecta" family="Tingidae" genus="Physatochila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="distinguenda">
<emphasis id="18DF141BFFA3FFEE6824FB3CFEB4FAF9" box="[324,339,1259,1282]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="18DF141BFFA3FFEE6801FB3DFE03FAF9" box="[353,484,1258,1282]" italics="true" pageId="19" pageNumber="20">distinguenda</emphasis>
</taxonomicName>
(C);
<taxonomicName id="EDABB38AFFA3FFEE6B7BFB3CFD68FAF9" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake &amp; Ruhoff" baseAuthorityYear="1965" box="[539,655,1258,1282]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="euthenia" status="comb. nov.">
<emphasis id="18DF141BFFA3FFEE6B7BFB3CFDCDFAF9" box="[539,554,1259,1282]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="18DF141BFFA3FFEE6B58FB3DFD68FAF9" box="[568,655,1258,1282]" italics="true" pageId="19" pageNumber="20">euthenia</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA3FFEE6BF7FB3DFCEBFAF9" bold="true" box="[663,780,1258,1282]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="03ECA960FFA3FFEE6BF7FB3DFCEBFAF9" box="[663,780,1258,1282]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(D);
<taxonomicName id="EDABB38AFFA3FFEE6A25FB3CFC43FAF9" baseAuthorityName="Scott" baseAuthorityYear="1874" box="[837,932,1258,1282]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="fieberi">
<emphasis id="18DF141BFFA3FFEE6A25FB3CFCB3FAF9" box="[837,852,1259,1282]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="18DF141BFFA3FFEE6A02FB3DFC43FAF9" box="[866,932,1258,1282]" italics="true" pageId="19" pageNumber="20">fieberi</emphasis>
</taxonomicName>
(E);
<taxonomicName id="EDABB38AFFA3FFEE6ABAFB3CFBBEFAF9" authorityName="Miyamoto" authorityYear="1964" box="[986,1113,1258,1282]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="miyatakei">
<emphasis id="18DF141BFFA3FFEE6ABAFB3CFC0EFAF9" box="[986,1001,1259,1282]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="18DF141BFFA3FFEE6A97FB3DFBBEFAF9" box="[1015,1113,1258,1282]" italics="true" pageId="19" pageNumber="20">miyatakei</emphasis>
</taxonomicName>
(F);
<taxonomicName id="EDABB38AFFA3FFEE6DEFFB3CFAA9FAF9" authorityName="Souma" authorityYear="2019" box="[1167,1358,1258,1282]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrintegerrima">
<emphasis id="18DF141BFFA3FFEE6DEFFB3CFB79FAF9" box="[1167,1182,1259,1282]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="18DF141BFFA3FFEE6DCCFB3CFAA9FAF9" box="[1196,1358,1259,1282]" italics="true" pageId="19" pageNumber="20">nigrintegerrima</emphasis>
</taxonomicName>
(G);
<taxonomicName id="EDABB38AFFA3FFEE6CE6FB3CFF03FADD" authorityName="Drake" authorityYear="1942" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="orientis">
<emphasis id="18DF141BFFA3FFEE6CE6FB3CFA72FAF9" box="[1414,1429,1259,1282]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="18DF141BFFA3FFEE69F7FAD8FF03FADD" box="[151,228,1295,1318]" italics="true" pageId="19" pageNumber="20">orientis</emphasis>
</taxonomicName>
(H);
<taxonomicName id="EDABB38AFFA3FFEE687BFAD8FE9BFADD" authorityName="Drake" authorityYear="1942" box="[283,380,1294,1318]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="veteris">
<emphasis id="18DF141BFFA3FFEE687BFAD8FECDFADD" box="[283,298,1295,1318]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="18DF141BFFA3FFEE6858FAD8FE9BFADD" box="[312,380,1295,1318]" italics="true" pageId="19" pageNumber="20">veteris</emphasis>
</taxonomicName>
(I). Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="2A14C809FFA3FFEE69A7FA83FB5FFA69" blockId="19.[151,1437,1364,1679]" pageId="19" pageNumber="20">
<taxonomicName id="EDABB38AFFA3FFEE69A7FA83FE2BFA95" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake &amp; Ruhoff" baseAuthorityYear="1965" box="[199,460,1364,1390]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="euthenia" status="comb. nov.">
<emphasis id="18DF141BFFA3FFEE69A7FA83FE2BFA95" box="[199,460,1364,1390]" italics="true" pageId="19" pageNumber="20">Physatocheila euthenia</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA3FFEE68B2FA83FDB6FA95" bold="true" box="[466,593,1364,1390]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="03ECA960FFA3FFEE68B2FA83FDB6FA95" box="[466,593,1364,1390]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
was recorded in
<collectingCountry id="52BC8899FFA3FFEE6A6CFA82FCABFA94" box="[780,844,1365,1391]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
for the first time. Discovery of the
<taxonomicName id="EDABB38AFFA3FFEE6DB0FA82FAA9FA95" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake &amp; Ruhoff" baseAuthorityYear="1965" box="[1232,1358,1364,1391]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="euthenia" status="comb. nov.">
<emphasis id="18DF141BFFA3FFEE6DB0FA82FB06FA95" box="[1232,1249,1365,1390]" italics="true" pageId="19" pageNumber="20">P</emphasis>
.
<emphasis id="18DF141BFFA3FFEE6D8EFA83FAA9FA95" box="[1262,1358,1364,1390]" italics="true" pageId="19" pageNumber="20">euthenia</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA3FFEE6C34FA83FF2FFA69" bold="true" pageId="19" pageNumber="20">
<taxonomicNameLabel id="03ECA960FFA3FFEE6C34FA83FF2FFA69" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
from the Iriomote Island represents the northernmost distributional record of this species.
</paragraph>
<paragraph id="2A14C809FFA3FFEE69A7FA4BFB31FA05" blockId="19.[151,1437,1364,1679]" pageId="19" pageNumber="20">
The notation “Okinawa” in the label of a single specimen deposited in ELKU likely refers to the Okinawa “Prefecture” (an administrative area), which includes a number of islands, and not the Okinawa “
<collectingCountry id="52BC8899FFA3FFEE6DA6FA17FAE8FA21" box="[1222,1295,1472,1498]" name="Iceland" pageId="19" pageNumber="20">Island</collectingCountry>
” (an island). Therefore, whether
<taxonomicName id="EDABB38AFFA3FFEE6815FA32FE17FA05" authorityName="Souma &amp; Ishikawa" authorityYear="2022" baseAuthorityName="Drake &amp; Ruhoff" baseAuthorityYear="1965" box="[373,496,1508,1534]" class="Insecta" family="Tingidae" genus="Physatocheila" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="euthenia" status="comb. nov.">
<emphasis id="18DF141BFFA3FFEE6815FA32FE17FA05" box="[373,496,1508,1534]" italics="true" pageId="19" pageNumber="20">P. euthenia</emphasis>
</taxonomicName>
<emphasis id="18DF141BFFA3FFEE6898FA33FD9FFA05" bold="true" box="[504,632,1508,1534]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="03ECA960FFA3FFEE6898FA33FD9FFA05" box="[504,632,1508,1534]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
was recorded on the Okinawa Island remains unclear.
</paragraph>
</subSubSection>
<subSubSection id="62B19B82FFA3FFEE69A7F9DFFE7FF974" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="2A14C809FFA3FFEE69A7F9DFFE2EF9D9" blockId="19.[151,1437,1364,1679]" box="[199,457,1544,1570]" pageId="19" pageNumber="20">
<emphasis id="18DF141BFFA3FFEE69A7F9DFFEACF9D9" bold="true" box="[199,331,1544,1570]" pageId="19" pageNumber="20">Host plant.</emphasis>
Unknown.
</paragraph>
<paragraph id="2A14C809FFA3FFEE69A7F9FBFE7FF974" blockId="19.[151,1437,1364,1679]" pageId="19" pageNumber="20">
<emphasis id="18DF141BFFA3FFEE69A7F9FBFEC0F9BD" bold="true" box="[199,295,1580,1606]" pageId="19" pageNumber="20">Biology.</emphasis>
In
<collectingCountry id="52BC8899FFA3FFEE6832F9FAFE72F9BC" box="[338,405,1581,1607]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
, adults were observed in October, November, and March (the present study). Conversely, adults were collected from September to February in other distribution areas (
<bibRefCitation id="4E3AB5F8FFA3FFEE6D43F987FAD1F990" author="Drake, C. J. &amp; Ruhoff, F. A." box="[1059,1334,1616,1643]" pageId="19" pageNumber="20" pagination="243 - 290" refId="ref36581" refString="Drake, C. J. &amp; Ruhoff, F. A. (1965 b) Lacebugs from New Guinea, Borneo, Solomons, and other islands of the South Pacific and Indian oceans (Hemiptera: Tingidae). Pacific Insects, 7 (2), 243 - 290." type="journal article" year="1965">Drake &amp; Ruhoff 1965b</bibRefCitation>
;
<bibRefCitation id="4E3AB5F8FFA3FFEE6C25F987FF38F974" author="Pericart, J." pageId="19" pageNumber="20" pagination="27 - 62" refId="ref39935" refString="Pericart, J. (1985 a) Tingidae nouveaux ou interessants du Nord du sous-continent indien (Hemiptera). II. Entomologica Basiliensia, 10, 27 - 62. [in French, English summary]" type="journal article" year="1985">Péricart 1985a</bibRefCitation>
;
<bibRefCitation id="4E3AB5F8FFA3FFEE698AF9A3FE6BF974" author="Guilbert, E." box="[234,396,1652,1679]" pageId="19" pageNumber="20" pagination="531 - 546" refId="ref37512" refString="Guilbert, E. (2015) New species and new records of Tingidae (Hemiptera: Heteroptera) from Vietnam. Zootaxa, 3956 (4), 531 - 546. https: // doi. org / 10.11646 / zootaxa. 3956.4.5" type="journal article" year="2015">Guilbert 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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