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<document id="BFCAB12B2D63B1D5938E51D04AD2717C" ID-CLB-Dataset="172790" ID-DOI="10.11646/zootaxa.5198.1.1" ID-GBIF-Dataset="17babbe7-1645-47c8-ac5f-2299f1ad436c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7251602" ID-ZooBank="019FE488-4263-4BC2-8606-446E599E226A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666769542164" checkinUser="plazi" docAuthor="Hou, Yanmeng, Gao, Zhizhong &amp; Zhang, Feng" docDate="2022" docId="8A6487A5FFF9A917FF11FE30FDF0FC4A" docLanguage="en" docName="zootaxa.5198.1.1.pdf" docOrigin="Zootaxa 5198 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5198.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Lagynochthonius yaowangguensis Hou &amp; Gao &amp; Zhang 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="61" masterDocId="765DFFDDFFCEA92BFF86FFEAFFEBFFCB" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="65" masterPageNumber="1" pageNumber="56" updateTime="1701997268366" updateUser="plazi">
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<mods:title id="E66ED5586C728AD569D52920CD29D7E4">Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="93C3068AB7247EDC1C09C3B5A39FAE70">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="DB7C2DD49CF71365A6525B0B1D2D0216">Gao, Zhizhong</mods:namePart>
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<mods:namePart id="A8F03E12670432E4B0973BE6963E23CD">Zhang, Feng</mods:namePart>
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<mods:title id="D2CE945AB862B8CAE74DB7BE6D8640AE">Zootaxa</mods:title>
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<mods:date id="207729E67BA7921DF05CCC30D2619750">2022</mods:date>
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<mods:number id="93B193486D4E762E913D59867ABCA851">2022-10-25</mods:number>
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<mods:number id="FA4A6170E57B90F1AB0EABEDB6A825D8">5198</mods:number>
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<treatment id="8A6487A5FFF9A917FF11FE30FDF0FC4A" ID-DOI="http://doi.org/10.5281/zenodo.7255285" ID-GBIF-Taxon="203411083" ID-Zenodo-Dep="7255285" LSID="urn:lsid:plazi:treatment:8A6487A5FFF9A917FF11FE30FDF0FC4A" httpUri="http://treatment.plazi.org/id/8A6487A5FFF9A917FF11FE30FDF0FC4A" lastPageId="60" lastPageNumber="61" pageId="55" pageNumber="56">
<subSubSection id="4AD76538FFF9A91CFF11FE30FC82FE3F" box="[151,873,474,501]" pageId="55" pageNumber="56" type="nomenclature">
<paragraph id="027236B3FFF9A91CFF11FE30FC82FE3F" blockId="55.[151,873,474,537]" box="[151,873,474,501]" pageId="55" pageNumber="56">
<heading id="593A81DFFFF9A91CFF11FE30FC82FE3F" bold="true" box="[151,873,474,501]" fontSize="11" level="2" pageId="55" pageNumber="56" reason="4">
<emphasis id="30B9EAA1FFF9A91CFF11FE30FC82FE3F" bold="true" box="[151,873,474,501]" pageId="55" pageNumber="56">
<taxonomicName id="C5CD4D30FFF9A91CFF11FE30FDD2FE3E" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[151,569,474,501]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF9A91CFF11FE30FDD2FE3E" bold="true" box="[151,569,474,501]" italics="true" pageId="55" pageNumber="56">Lagynochthonius yaowangguensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B8A57DAFFF9A91CFDC7FE31FD75FE3E" box="[577,670,475,501]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
(ŪƗỖIJ伪oi)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFF9A91CFF11FE15FEFFFDD2" box="[151,276,510,537]" pageId="55" pageNumber="56" type="description">
<paragraph id="027236B3FFF9A91CFF11FE15FEFFFDD2" blockId="55.[151,873,474,537]" box="[151,276,510,537]" pageId="55" pageNumber="56">
<figureCitation id="9AF62A36FFF9A91CFF11FE15FEFFFDD2" box="[151,276,510,537]" captionStart-0="FIGURE 42" captionStart-1="FIGURE 43" captionStart-2="FIGURE 44" captionStart-3="FIGURE 45" captionStartId-0="56.[151,250,1890,1914]" captionStartId-1="57.[151,250,847,871]" captionStartId-2="58.[151,250,1899,1923]" captionStartId-3="59.[151,250,1875,1899]" captionTargetBox-0="[151,1436,525,1867]" captionTargetBox-1="[162,1436,181,824]" captionTargetBox-2="[151,1436,267,1876]" captionTargetBox-3="[168,1396,189,1847]" captionTargetId-0="figure-205@56.[151,1436,525,1867]" captionTargetId-1="figure-19@57.[151,1436,181,824]" captionTargetId-2="figure-33@58.[151,1436,237,1876]" captionTargetId-3="figure-19@59.[168,1420,181,1851]" captionTargetPageId-0="56" captionTargetPageId-1="57" captionTargetPageId-2="58" captionTargetPageId-3="59" captionText-0="FIGURE 42. Unnamed Cave 4, type locality of Lagynochthonius yaowangguensis sp. nov., A. Entrance; B. Inside the cave entrance; C–D. Peculiar landscapes inside the cave; E. Area where L. yaowangguensis sp. nov. specimens were collected; F–G. Live individuals of L. yaowangguensis sp. nov. (F: male, G: female) in their natural environment." captionText-1="FIGURE 43. Lagynochthonius yaowangguensis sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." captionText-2="FIGURE 44. Lagynochthonius yaowangguensis sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." captionText-3="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi-0="http://doi.org/10.5281/zenodo.7251732" figureDoi-1="http://doi.org/10.5281/zenodo.7251734" figureDoi-2="http://doi.org/10.5281/zenodo.7251736" figureDoi-3="http://doi.org/10.5281/zenodo.7251738" httpUri-0="https://zenodo.org/record/7251732/files/figure.png" httpUri-1="https://zenodo.org/record/7251734/files/figure.png" httpUri-2="https://zenodo.org/record/7251736/files/figure.png" httpUri-3="https://zenodo.org/record/7251738/files/figure.png" pageId="55" pageNumber="56">Figs 42–45</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFF9A91CFF11FDADFB73FD3A" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="027236B3FFF9A91CFF11FDADFB73FD3A" blockId="55.[151,1437,582,2013]" pageId="55" pageNumber="56">
<emphasis id="30B9EAA1FFF9A91CFF11FDADFEA7FDAB" bold="true" box="[151,332,582,609]" pageId="55" pageNumber="56">Type material.</emphasis>
<materialsCitation id="B2A53CEEFFF9A91CFEDDFDADFBA1FD62" ID-GBIF-Occurrence="3946861307" collectingDate="2021-04-19" collectionCode="MHBU-HBUARA" collectorName="Zegang Feng" country="China" county="Malipo County" elevation="1221" latitude="23.010584" location="Laoshanyaowanggu" longLatPrecision="1" longitude="104.82155" municipality="Tianbao Town" pageId="55" pageNumber="56" specimenCode="MHBU-HBUARA#2021-27901" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="DD768811FFF9A91CFEDDFDADFE28FDAA" box="[347,451,583,609]" pageId="55" pageNumber="56" type="holotype">Holotype</typeStatus>
<specimenCount id="14CBFD3AFFF9A91CFE57FDADFE0DFDAA" box="[465,486,583,609]" pageId="55" pageNumber="56" type="male">♂</specimenCount>
(Ps.-
<specimenCode id="526B9EC8FFF9A91CFDA4FDADFC7CFDAA" box="[546,919,583,609]" collectionCode="MHBU-HBUARA" pageId="55" pageNumber="56">MHBU-HBUARA#2021-27901</specimenCode>
):
<collectingCountry id="7ADA7623FFF9A91CFC37FDADFC1CFDAA" box="[945,1015,583,609]" name="China" pageId="55" pageNumber="56">China</collectingCountry>
,
<collectingRegion id="C009F851FFF9A91CFB8EFDADFB39FDAA" box="[1032,1234,583,609]" country="China" name="Yunnan" pageId="55" pageNumber="56">Yunnan Province</collectingRegion>
,
<collectingCounty id="EB134E3FFFF9A91CFB63FDADFA72FDAA" box="[1253,1433,583,609]" pageId="55" pageNumber="56">Malipo County</collectingCounty>
,
<collectingMunicipality id="E216ACC9FFF9A91CFF11FD81FED1FD4E" box="[151,314,619,645]" pageId="55" pageNumber="56">Tianbao Town</collectingMunicipality>
,
<location id="07126068FFF9A91CFEC0FD81FDC2FD4E" LSID="urn:lsid:plazi:treatment:8A6487A5FFF9A917FF11FE30FDF0FC4A:07126068FFF9A91CFEC0FD81FDC2FD4E" box="[326,553,619,645]" country="China" county="Malipo County" latitude="23.010584" longLatPrecision="1" longitude="104.82155" municipality="Tianbao Town" name="Laoshanyaowanggu" pageId="55" pageNumber="56" stateProvince="Yunnan">Laoshanyaowanggu</location>
, cave without name (
<location id="07126068FFF9A91CFC93FD81FC2EFD4E" LSID="urn:lsid:plazi:treatment:8A6487A5FFF9A917FF11FE30FDF0FC4A:07126068FFF9A91CFC93FD81FC2EFD4E" box="[789,965,619,645]" country="China" county="Malipo County" latitude="23.010584" longLatPrecision="1" longitude="104.82155" municipality="Tianbao Town" name="Unnamed Cave" pageId="55" pageNumber="56" stateProvince="Yunnan">Unnamed Cave</location>
4), under stones and detritus in deep zone [
<geoCoordinate id="67F95074FFF9A91CFF18FD65FEA1FD62" box="[158,330,655,681]" degrees="23" direction="north" minutes="00" orientation="latitude" pageId="55" pageNumber="56" precision="1" seconds="38.10" value="23.010584">23°00'38.10&quot;N</geoCoordinate>
,
<geoCoordinate id="67F95074FFF9A91CFEDFFD64FDE4FD62" box="[345,527,654,681]" degrees="104" direction="east" minutes="49" orientation="longitude" pageId="55" pageNumber="56" precision="1" seconds="17.57" value="104.82155">104°49'17.57&quot;E</geoCoordinate>
],
<elevation id="89E0D180FFF9A91CFDA3FD65FD56FD62" box="[549,701,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.2209999999999999" pageId="55" pageNumber="56" unit="m" value="1221.0">
<quantity id="C5359B56FFF9A91CFDA3FD65FD96FD62" box="[549,637,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.2209999999999999" pageId="55" pageNumber="56" unit="m" value="1221.0">1221 m</quantity>
a.s.l.
</elevation>
,
<date id="76731073FFF9A91CFD4BFD65FC99FD62" box="[717,882,655,681]" pageId="55" pageNumber="56" value="2021-04-19">
<collectingDate id="6637E99BFFF9A91CFD4BFD65FC99FD62" box="[717,882,655,681]" pageId="55" pageNumber="56" value="2021-04-19">19 April 2021</collectingDate>
</date>
,
<collectorName id="AF385365FFF9A91CFC06FD65FBFDFD62" box="[896,1046,655,681]" pageId="55" pageNumber="56">Zegang Feng</collectorName>
leg.
</materialsCitation>
<materialsCitation id="B2A53CEEFFF9A91CFBD2FD65FCC2FD06" ID-GBIF-Occurrence="3946861314" collectingDate="2021-04-19" collectorName="Zegang Feng" country="China" county="Malipo County" elevation="1221" latitude="23.010584" location="Laoshanyaowanggu" longLatPrecision="1" longitude="104.82155" municipality="Tianbao Town" pageId="55" pageNumber="56" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="DD768811FFF9A91CFBD2FD65FB2EFD62" box="[1108,1221,655,681]" pageId="55" pageNumber="56" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="14CBFD3AFFF9A91CFB55FD65FAEBFD62" box="[1235,1280,655,681]" pageId="55" pageNumber="56" type="female">1 ♀</specimenCount>
(Ps.-MHBU-HBUARA#2021-27902), the same data as the holotype
</materialsCitation>
;
<materialsCitation id="B2A53CEEFFF9A91CFCBCFD59FB73FD3A" ID-GBIF-Occurrence="3946861322" collectingDate="2021-07-22" collectionCode="MSWU-HBUARA" collectorName="Zegang Feng &amp; Hongru Xu &amp; Liu Fu &amp; Nana Zhan" country="China" county="Malipo County" elevation="1221" latitude="23.010584" location="Laoshanyaowanggu" longLatPrecision="1" longitude="104.82155" municipality="Tianbao Town" pageId="55" pageNumber="56" specimenCode="MSWU-HBUARA#2021-18001" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="14CBFD3AFFF9A91CFCBCFD59FC81FD06" box="[826,874,691,717]" pageId="55" pageNumber="56" type="female">1 ♀</specimenCount>
(Ps.-
<specimenCode id="526B9EC8FFF9A91CFC22FD59FAF0FD06" box="[932,1307,691,717]" collectionCode="MSWU-HBUARA" pageId="55" pageNumber="56">MSWU-HBUARA#2021-18001</specimenCode>
), the same location as the holotype,
<date id="76731073FFF9A91CFE36FD3DFDAFFD3A" box="[432,580,727,753]" pageId="55" pageNumber="56" value="2021-07-22">
<collectingDate id="6637E99BFFF9A91CFE36FD3DFDAFFD3A" box="[432,580,727,753]" pageId="55" pageNumber="56" value="2021-07-22">22 July 2021</collectingDate>
</date>
,
<collectorName id="AF385365FFF9A91CFDC9FD3DFD0EFD3A" box="[591,741,727,753]" pageId="55" pageNumber="56">Zegang Feng</collectorName>
,
<collectorName id="AF385365FFF9A91CFD76FD3DFC9AFD3A" box="[752,881,727,753]" pageId="55" pageNumber="56">Hongru Xu</collectorName>
,
<collectorName id="AF385365FFF9A91CFCFAFD3DFC23FD3A" box="[892,968,727,753]" pageId="55" pageNumber="56">Liu Fu</collectorName>
&amp;
<collectorName id="AF385365FFF9A91CFC6AFD3DFB83FD3A" box="[1004,1128,727,753]" pageId="55" pageNumber="56">Nana Zhan</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFF9A91CFF41FD11FBB2FCDE" box="[199,1113,763,789]" pageId="55" pageNumber="56" type="etymology">
<paragraph id="027236B3FFF9A91CFF41FD11FBB2FCDE" blockId="55.[151,1437,582,2013]" box="[199,1113,763,789]" pageId="55" pageNumber="56">
<emphasis id="30B9EAA1FFF9A91CFF41FD11FEA5FCDE" bold="true" box="[199,334,763,789]" pageId="55" pageNumber="56">Etymology.</emphasis>
Named after the valley of Laoshanyaowanggu, near the
<typeStatus id="DD768811FFF9A91CFC4EFD11FC13FCDE" box="[968,1016,763,789]" pageId="55" pageNumber="56">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFF9A91CFF41FCF4FD86FC26" pageId="55" pageNumber="56" type="diagnosis">
<paragraph id="027236B3FFF9A91CFF41FCF4FD86FC26" blockId="55.[151,1437,582,2013]" pageId="55" pageNumber="56">
<emphasis id="30B9EAA1FFF9A91CFF41FCF4FE6FFCF7" bold="true" box="[199,388,798,828]" pageId="55" pageNumber="56">
Diagnosis (
<specimenCount id="14CBFD3AFFF9A91CFECCFCCAFEB4FCF7" box="[330,351,800,828]" pageId="55" pageNumber="56" type="male">♂</specimenCount>
<specimenCount id="14CBFD3AFFF9A91CFED9FCCAFE9FFCF7" box="[351,372,800,828]" pageId="55" pageNumber="56" type="female">♀</specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with 2 setae; tergites I–VII each with 4 setae. Pedipalps slender, femur 10.46 (
<specimenCount id="14CBFD3AFFF9A91CFBF3FC8DFB67FC4A" box="[1141,1164,871,897]" pageId="55" pageNumber="56" type="male">♂</specimenCount>
), 10.07–10.38 (
<specimenCount id="14CBFD3AFFF9A91CFABBFC8DFAA4FC4A" box="[1341,1359,871,897]" pageId="55" pageNumber="56" type="female">♀</specimenCount>
) times longer than broad; chela 8.76 (
<specimenCount id="14CBFD3AFFF9A91CFE68FC61FDEEFC6E" box="[494,517,907,933]" pageId="55" pageNumber="56" type="male">♂</specimenCount>
), 8.30–8.32 (
<specimenCount id="14CBFD3AFFF9A91CFD1AFC61FD45FC6E" box="[668,686,907,933]" pageId="55" pageNumber="56" type="female">♀</specimenCount>
) times longer than broad; both chelal fingers with intercalary teeth and fixed chelal finger with a modified accessory tooth (
<emphasis id="30B9EAA1FFF9A91CFC81FC45FCF6FC02" box="[775,797,943,969]" italics="true" pageId="55" pageNumber="56">td</emphasis>
) on dorso-antiaxial face; chemosensory setae (
<emphasis id="30B9EAA1FFF9A91CFAA3FC45FAD7FC03" box="[1317,1340,943,968]" italics="true" pageId="55" pageNumber="56">sc</emphasis>
) present on dorsum of chelal hand; sensilla absent.
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFF9A911FF41FC1CFC8CFF1E" lastPageId="58" lastPageNumber="59" pageId="55" pageNumber="56" type="description">
<paragraph id="027236B3FFF9A91CFF41FC1CFBC3FBDA" blockId="55.[151,1437,582,2013]" box="[199,1064,1014,1041]" pageId="55" pageNumber="56">
<emphasis id="30B9EAA1FFF9A91CFF41FC1CFEB2FBDB" bold="true" box="[199,345,1014,1040]" pageId="55" pageNumber="56">Description.</emphasis>
Male (
<typeStatus id="DD768811FFF9A91CFE2EFC1DFDFBFBDA" box="[424,528,1015,1041]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
), female unknown (
<figureCitation id="9AF62A36FFF9A91CFD6BFC1DFCB9FBDA" box="[749,850,1015,1041]" captionStart="FIGURE 42" captionStartId="56.[151,250,1890,1914]" captionTargetBox="[151,1436,525,1867]" captionTargetId="figure-205@56.[151,1436,525,1867]" captionTargetPageId="56" captionText="FIGURE 42. Unnamed Cave 4, type locality of Lagynochthonius yaowangguensis sp. nov., A. Entrance; B. Inside the cave entrance; C–D. Peculiar landscapes inside the cave; E. Area where L. yaowangguensis sp. nov. specimens were collected; F–G. Live individuals of L. yaowangguensis sp. nov. (F: male, G: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251732" httpUri="https://zenodo.org/record/7251732/files/figure.png" pageId="55" pageNumber="56">Figs 42F</figureCitation>
,
<figureCitation id="9AF62A36FFF9A91CFCDBFC1DFC7BFBDA" box="[861,912,1015,1041]" captionStart="FIGURE 43" captionStartId="57.[151,250,847,871]" captionTargetBox="[162,1436,181,824]" captionTargetId="figure-19@57.[151,1436,181,824]" captionTargetPageId="57" captionText="FIGURE 43. Lagynochthonius yaowangguensis sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251734" httpUri="https://zenodo.org/record/7251734/files/figure.png" pageId="55" pageNumber="56">43A</figureCitation>
,
<figureCitation id="9AF62A36FFF9A91CFC1DFC1DFC1BFBDA" box="[923,1008,1015,1041]" captionStart="FIGURE 44" captionStartId="58.[151,250,1899,1923]" captionTargetBox="[151,1436,267,1876]" captionTargetId="figure-33@58.[151,1436,237,1876]" captionTargetPageId="58" captionText="FIGURE 44. Lagynochthonius yaowangguensis sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251736" httpUri="https://zenodo.org/record/7251736/files/figure.png" pageId="55" pageNumber="56">44A–D</figureCitation>
,
<figureCitation id="9AF62A36FFF9A91CFC7AFC1CFBF7FBDA" box="[1020,1052,1014,1041]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="55" pageNumber="56">45</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFF9A91CFF41FBF1FEE0FB92" blockId="55.[151,1437,582,2013]" pageId="55" pageNumber="56">
Colour (
<figureCitation id="9AF62A36FFF9A91CFEA3FBF1FE60FBFE" box="[293,395,1051,1077]" captionStart="FIGURE 42" captionStartId="56.[151,250,1890,1914]" captionTargetBox="[151,1436,525,1867]" captionTargetId="figure-205@56.[151,1436,525,1867]" captionTargetPageId="56" captionText="FIGURE 42. Unnamed Cave 4, type locality of Lagynochthonius yaowangguensis sp. nov., A. Entrance; B. Inside the cave entrance; C–D. Peculiar landscapes inside the cave; E. Area where L. yaowangguensis sp. nov. specimens were collected; F–G. Live individuals of L. yaowangguensis sp. nov. (F: male, G: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251732" httpUri="https://zenodo.org/record/7251732/files/figure.png" pageId="55" pageNumber="56">Figs 42F</figureCitation>
,
<figureCitation id="9AF62A36FFF9A91CFE11FBF1FE21FBFE" box="[407,458,1051,1077]" captionStart="FIGURE 43" captionStartId="57.[151,250,847,871]" captionTargetBox="[162,1436,181,824]" captionTargetId="figure-19@57.[151,1436,181,824]" captionTargetPageId="57" captionText="FIGURE 43. Lagynochthonius yaowangguensis sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251734" httpUri="https://zenodo.org/record/7251734/files/figure.png" pageId="55" pageNumber="56">43A</figureCitation>
,
<figureCitation id="9AF62A36FFF9A91CFE50FBF1FDC4FBFE" box="[470,559,1051,1077]" captionStart="FIGURE 44" captionStartId="58.[151,250,1899,1923]" captionTargetBox="[151,1436,267,1876]" captionTargetId="figure-33@58.[151,1436,237,1876]" captionTargetPageId="58" captionText="FIGURE 44. Lagynochthonius yaowangguensis sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251736" httpUri="https://zenodo.org/record/7251736/files/figure.png" pageId="55" pageNumber="56">44A–D</figureCitation>
): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
</paragraph>
<paragraph id="027236B3FFF9A91CFF41FB89FEC6FA0A" blockId="55.[151,1437,582,2013]" pageId="55" pageNumber="56">
Cephalothorax (
<figureCitation id="9AF62A36FFF9A91CFEFCFB89FE09FBB6" box="[378,482,1123,1149]" captionStart="FIGURE 44" captionStartId="58.[151,250,1899,1923]" captionTargetBox="[151,1436,267,1876]" captionTargetId="figure-33@58.[151,1436,237,1876]" captionTargetPageId="58" captionText="FIGURE 44. Lagynochthonius yaowangguensis sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251736" httpUri="https://zenodo.org/record/7251736/files/figure.png" pageId="55" pageNumber="56">Figs 44B</figureCitation>
,
<figureCitation id="9AF62A36FFF9A91CFE6BFB88FDC9FBB6" box="[493,546,1122,1149]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="55" pageNumber="56">45A</figureCitation>
): carapace 0.93 times longer than broad, markedly narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome very pointed and small, triangular, with 2 setae flanking base; with 16 setae arranged 4: 4: 4: 2: 2, preocular setae missing, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3–4, III 4–5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 9–11 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="9AF62A36FFF9A91CFC24FA69FBE3FA56" box="[930,1032,1410,1437]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="55" pageNumber="56">Fig. 45C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<paragraph id="027236B3FFF9A91CFF41FA21FE7CF92A" blockId="55.[151,1437,582,2013]" pageId="55" pageNumber="56">
Chelicera (
<figureCitation id="9AF62A36FFF9A91CFEC0FA21FE5FFA2E" box="[326,436,1483,1509]" captionStart="FIGURE 44" captionStartId="58.[151,250,1899,1923]" captionTargetBox="[151,1436,267,1876]" captionTargetId="figure-33@58.[151,1436,237,1876]" captionTargetPageId="58" captionText="FIGURE 44. Lagynochthonius yaowangguensis sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251736" httpUri="https://zenodo.org/record/7251736/files/figure.png" pageId="55" pageNumber="56">Figs 44C</figureCitation>
,
<figureCitation id="9AF62A36FFF9A91CFE42FA20FE1CFA2E" box="[452,503,1482,1509]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="55" pageNumber="56">45B</figureCitation>
): large, about as long as carapace, 2.39 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12 teeth, distal one largest; movable finger with 11 retrorse contiguous small teeth; galea completely vestigial (
<figureCitation id="9AF62A36FFF9A91CFBBDF9B1FB49F9BE" box="[1083,1186,1626,1653]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="55" pageNumber="56">Fig. 45B</figureCitation>
). Serrula exterior with 25 and serrula interior with 14 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="9AF62A36FFF9A91CFEA7F92DFE61F92A" box="[289,394,1734,1761]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="55" pageNumber="56">Fig. 45D</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFF9A913FF41F901FBE2FE3E" blockId="55.[151,1437,582,2013]" lastBlockId="56.[151,1437,151,501]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
Pedipalp (
<figureCitation id="9AF62A36FFF9A91CFEB1F901FE49F8CE" box="[311,418,1771,1797]" captionStart="FIGURE 44" captionStartId="58.[151,250,1899,1923]" captionTargetBox="[151,1436,267,1876]" captionTargetId="figure-33@58.[151,1436,237,1876]" captionTargetPageId="58" captionText="FIGURE 44. Lagynochthonius yaowangguensis sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251736" httpUri="https://zenodo.org/record/7251736/files/figure.png" pageId="55" pageNumber="56">Figs 44A</figureCitation>
,
<figureCitation id="9AF62A36FFF9A91CFE2AF900FDEAF8CE" box="[428,513,1770,1797]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="55" pageNumber="56">45E–G</figureCitation>
): long and slender, trochanter 1.81, femur 10.46, patella 2.82, chela 8.76, hand 3.33 times longer than broad; femur 2.83 times longer than patella; movable chelal finger 1.64 times longer than hand and 0.63 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
<figureCitation id="9AF62A36FFF9A91CFAE9F8D9FF22F8BA" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="55" pageNumber="56">Fig. 45E</figureCitation>
). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
<emphasis id="30B9EAA1FFF9A91CFD14F875FD43F872" box="[658,680,1951,1977]" italics="true" pageId="55" pageNumber="56">ib</emphasis>
and
<emphasis id="30B9EAA1FFF9A91CFD63F875FCEDF872" box="[741,774,1951,1977]" italics="true" pageId="55" pageNumber="56">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis id="30B9EAA1FFF9A91CFF58F829FF13F816" box="[222,248,1987,2013]" italics="true" pageId="55" pageNumber="56">eb</emphasis>
,
<emphasis id="30B9EAA1FFF9A91CFE81F829FEC7F816" box="[263,300,1987,2013]" italics="true" pageId="55" pageNumber="56">esb</emphasis>
and
<emphasis id="30B9EAA1FFF9A91CFEE3F829FE94F817" box="[357,383,1987,2012]" italics="true" pageId="55" pageNumber="56">ist</emphasis>
forming a straight oblique row at base of fixed chelal finger;
<emphasis id="30B9EAA1FFF9A91CFBC6F829FBBBF817" box="[1088,1104,1987,2012]" italics="true" pageId="55" pageNumber="56">it</emphasis>
slightly distal to
<emphasis id="30B9EAA1FFF9A91CFA91F829FADDF817" box="[1303,1334,1987,2012]" italics="true" pageId="55" pageNumber="56">est</emphasis>
, situated subdistally;
<emphasis id="30B9EAA1FFF6A913FE99FF7DFED8FF7B" box="[287,307,151,176]" italics="true" pageId="56" pageNumber="57">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis id="30B9EAA1FFF6A913FBA3FF7DFBD4FF7A" box="[1061,1087,151,177]" italics="true" pageId="56" pageNumber="57">dx</emphasis>
situated distal to
<emphasis id="30B9EAA1FFF6A913FA85FF7DFAFCFF7B" box="[1283,1303,151,176]" italics="true" pageId="56" pageNumber="57">et</emphasis>
;
<emphasis id="30B9EAA1FFF6A913FAA0FF7DFAD4FF7A" box="[1318,1343,151,177]" italics="true" pageId="56" pageNumber="57">sb</emphasis>
situated midway between
<emphasis id="30B9EAA1FFF6A913FEE7FF51FE84FF1E" box="[353,367,187,213]" italics="true" pageId="56" pageNumber="57">b</emphasis>
and
<emphasis id="30B9EAA1FFF6A913FE2CFF51FE56FF1F" box="[426,445,187,212]" italics="true" pageId="56" pageNumber="57">st</emphasis>
;
<emphasis id="30B9EAA1FFF6A913FE48FF51FE37FF1E" box="[462,476,187,213]" italics="true" pageId="56" pageNumber="57">b</emphasis>
and
<emphasis id="30B9EAA1FFF6A913FD91FF51FDF4FF1F" box="[535,543,187,212]" italics="true" pageId="56" pageNumber="57">t</emphasis>
situated subdistally,
<emphasis id="30B9EAA1FFF6A913FC96FF51FCF5FF1E" box="[784,798,187,213]" italics="true" pageId="56" pageNumber="57">b</emphasis>
situated at same level as
<emphasis id="30B9EAA1FFF6A913FBC1FF51FB8DFF1F" box="[1095,1126,187,212]" italics="true" pageId="56" pageNumber="57">est</emphasis>
;
<emphasis id="30B9EAA1FFF6A913FBFEFF51FB6BFF1F" box="[1144,1152,187,212]" italics="true" pageId="56" pageNumber="57">t</emphasis>
situated distal to
<emphasis id="30B9EAA1FFF6A913FAC8FF51FAB5FF1F" box="[1358,1374,187,212]" italics="true" pageId="56" pageNumber="57">it</emphasis>
(
<figureCitation id="9AF62A36FFF6A913FAE9FF51FF2CFF32" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="56" pageNumber="57">Fig. 45F</figureCitation>
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis id="30B9EAA1FFF6A913FB08FF35FB43FF33" box="[1166,1192,223,248]" italics="true" pageId="56" pageNumber="57">ist</emphasis>
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 34 macrodenticles, slightly retrorse and pointed, plus
<specimenCount id="14CBFD3AFFF6A913FD9CFEA1FD5EFEAE" box="[538,693,331,357]" pageId="56" pageNumber="57" type="intercalary">21 intercalary</specimenCount>
microdenticles and a modified accessory tooth on dorso-antiaxial face (
<emphasis id="30B9EAA1FFF6A913FF50FE85FF07FE42" box="[214,236,367,393]" italics="true" pageId="56" pageNumber="57">td</emphasis>
, slightly distal to
<emphasis id="30B9EAA1FFF6A913FE30FE85FE3BFE42" box="[438,464,367,393]" italics="true" pageId="56" pageNumber="57">dx</emphasis>
),
<quantity id="C5359B56FFF6A913FE6EFE84FDCAFE42" box="[488,545,366,393]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="56" pageNumber="57" unit="in" value="56.0">56 in</quantity>
total; movable chelal finger with 23 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus
<specimenCount id="14CBFD3AFFF6A913FCB5FE78FC3AFE66" box="[819,977,402,429]" pageId="56" pageNumber="57" type="intercalary">15 intercalary</specimenCount>
microdenticles and 8 vestigial, rounded and contiguous basal teeth (
<figureCitation id="9AF62A36FFF6A913FE5EFE5DFDD5FE1A" box="[472,574,438,465]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="56" pageNumber="57">Fig. 45F</figureCitation>
),
<quantity id="C5359B56FFF6A913FDD2FE5DFD64FE1A" box="[596,655,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.1684" pageId="56" pageNumber="57" unit="in" value="46.0">46 in</quantity>
total. Sensilla absent. Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="9AF62A36FFF6A913FC12FE31FC16FE3E" box="[916,1021,474,501]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="56" pageNumber="57">Fig. 45G</figureCitation>
).
</paragraph>
<caption id="56B2663BFFF6A913FF11F888FB9DF809" ID-DOI="http://doi.org/10.5281/zenodo.7251732" ID-Zenodo-Dep="7251732" httpUri="https://zenodo.org/record/7251732/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,1890,1914]" targetBox="[151,1436,525,1867]" targetPageId="56">
<paragraph id="027236B3FFF6A913FF11F888FB9DF809" blockId="56.[151,1437,1890,1986]" pageId="56" pageNumber="57">
<emphasis id="30B9EAA1FFF6A913FF11F888FEC9F8B1" bold="true" box="[151,290,1890,1914]" pageId="56" pageNumber="57">FIGURE 42.</emphasis>
Unnamed Cave 4, type locality of
<taxonomicName id="C5CD4D30FFF6A913FD17F888FC04F8B1" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[657,1007,1890,1914]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF6A913FD17F888FC04F8B1" box="[657,1007,1890,1914]" italics="true" pageId="56" pageNumber="57">Lagynochthonius yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF6A913FC71F888FBA1F8B1" bold="true" box="[1015,1098,1890,1914]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="2B8A57DAFFF6A913FC71F888FBA1F8B1" box="[1015,1098,1890,1914]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; B. Inside the cave entrance; C–D. Peculiar landscapes inside the cave; E. Area where
<taxonomicName id="C5CD4D30FFF6A913FCBDF86DFC16F855" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[827,1021,1927,1950]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF6A913FCBDF86DFC16F855" box="[827,1021,1927,1950]" italics="true" pageId="56" pageNumber="57">L. yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF6A913FB85F86CFBB8F855" bold="true" box="[1027,1107,1926,1950]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="2B8A57DAFFF6A913FB85F86CFBB8F855" box="[1027,1107,1926,1950]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected; F–G. Live individuals of
<taxonomicName id="C5CD4D30FFF6A913FED8F841FDCBF809" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[350,544,1963,1986]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF6A913FED8F841FDCBF809" box="[350,544,1963,1986]" italics="true" pageId="56" pageNumber="57">L. yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF6A913FDA1F840FD9CF809" bold="true" box="[551,631,1962,1986]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="2B8A57DAFFF6A913FDA1F840FD9CF809" box="[551,631,1962,1986]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(F: male, G: female) in their natural environment.
</paragraph>
</caption>
<caption id="56B2663BFFF7A912FF11FCA5FECEFC40" ID-DOI="http://doi.org/10.5281/zenodo.7251734" ID-Zenodo-Dep="7251734" httpUri="https://zenodo.org/record/7251734/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,847,871]" targetBox="[162,1436,181,824]" targetPageId="57">
<paragraph id="027236B3FFF7A912FF11FCA5FECEFC40" blockId="57.[151,1436,847,907]" pageId="57" pageNumber="58">
<emphasis id="30B9EAA1FFF7A912FF11FCA5FECBFCAC" bold="true" box="[151,288,847,871]" pageId="57" pageNumber="58">FIGURE 43.</emphasis>
<taxonomicName id="C5CD4D30FFF7A912FEA3FCA5FD6AFCAC" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[293,641,847,871]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF7A912FEA3FCA5FD6AFCAC" box="[293,641,847,871]" italics="true" pageId="57" pageNumber="58">Lagynochthonius yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF7A912FD01FCBAFD33FCA3" bold="true" box="[647,728,848,872]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="2B8A57DAFFF7A912FD01FCBAFD33FCA3" box="[647,728,848,872]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view).
</paragraph>
</caption>
<paragraph id="027236B3FFF7A912FF41FC50FB59FB8B" blockId="57.[151,1437,954,2025]" pageId="57" pageNumber="58">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 5: 5: 4: T2T: 0; tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 11: 9: 8: 8: 7: 8: 7: 9: 0: 2. Anterior genital operculum with 9 setae, genital opening slit-like, with 14–15 marginal setae on each side,
<quantity id="C5359B56FFF7A912FC3AFBCCFC1EFB8B" box="[956,1013,1062,1088]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" pageId="57" pageNumber="58" unit="in" value="38.0">38 in</quantity>
total (
<figureCitation id="9AF62A36FFF7A912FBBBFBCCFB4CFB8B" box="[1085,1191,1062,1088]" captionStart="FIGURE 44" captionStartId="58.[151,250,1899,1923]" captionTargetBox="[151,1436,267,1876]" captionTargetId="figure-33@58.[151,1436,237,1876]" captionTargetPageId="58" captionText="FIGURE 44. Lagynochthonius yaowangguensis sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251736" httpUri="https://zenodo.org/record/7251736/files/figure.png" pageId="57" pageNumber="58">Fig. 44D</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFF7A912FF41FBA0FEDBFAF7" blockId="57.[151,1437,954,2025]" pageId="57" pageNumber="58">
Legs (
<figureCitation id="9AF62A36FFF7A912FE8BFBA0FE60FBAF" box="[269,395,1098,1125]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="57" pageNumber="58">Fig. 45H, I</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur I, IV and patella I, IV. Femur of leg I 2.05 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.34 times longer than tibia. Femoropatella of leg IV 5.09 times longer than deep; tibia 7.40 times longer than deep; with basal tactile setae on both tarsal segments: basitarsus 4.00 times longer than deep (TS= 0.22), telotarsus 15.17 times longer than deep and 2.84 times longer than basitarsus (TS= 0.24). Setae of leg I (trochanter to tibia) 3: 11: 11: 15, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 9: 11. Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="027236B3FFF7A912FF41FAACFEB2F9F2" blockId="57.[151,1437,954,2025]" pageId="57" pageNumber="58">
Adult females (
<typeStatus id="DD768811FFF7A912FEF0FAACFE0EFAAB" box="[374,485,1350,1376]" pageId="57" pageNumber="58" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="9AF62A36FFF7A912FE7AFAACFD82FAAB" box="[508,617,1350,1376]" captionStart="FIGURE 42" captionStartId="56.[151,250,1890,1914]" captionTargetBox="[151,1436,525,1867]" captionTargetId="figure-205@56.[151,1436,525,1867]" captionTargetPageId="56" captionText="FIGURE 42. Unnamed Cave 4, type locality of Lagynochthonius yaowangguensis sp. nov., A. Entrance; B. Inside the cave entrance; C–D. Peculiar landscapes inside the cave; E. Area where L. yaowangguensis sp. nov. specimens were collected; F–G. Live individuals of L. yaowangguensis sp. nov. (F: male, G: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251732" httpUri="https://zenodo.org/record/7251732/files/figure.png" pageId="57" pageNumber="58">Figs 42G</figureCitation>
,
<figureCitation id="9AF62A36FFF7A912FDF3FAACFD4DFAAB" box="[629,678,1350,1376]" captionStart="FIGURE 43" captionStartId="57.[151,250,847,871]" captionTargetBox="[162,1436,181,824]" captionTargetId="figure-19@57.[151,1436,181,824]" captionTargetPageId="57" captionText="FIGURE 43. Lagynochthonius yaowangguensis sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251734" httpUri="https://zenodo.org/record/7251734/files/figure.png" pageId="57" pageNumber="58">43B</figureCitation>
,
<figureCitation id="9AF62A36FFF7A912FD35FAACFD0EFAAB" box="[691,741,1350,1376]" captionStart="FIGURE 44" captionStartId="58.[151,250,1899,1923]" captionTargetBox="[151,1436,267,1876]" captionTargetId="figure-33@58.[151,1436,237,1876]" captionTargetPageId="58" captionText="FIGURE 44. Lagynochthonius yaowangguensis sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251736" httpUri="https://zenodo.org/record/7251736/files/figure.png" pageId="57" pageNumber="58">44E</figureCitation>
). Mostly same as male, but a little larger; carapace 0.95–0.98 times longer than broad, gently narrowed posteriorly,
<specimenCount id="14CBFD3AFFF7A912FD73FA80FC9AFA4F" box="[757,881,1386,1412]" pageId="57" pageNumber="58" type="female">one female</specimenCount>
has a pair of preocular microsetae; chaetotaxy of coxae: P 3, I 3, II 4,
<specimenCode id="526B9EC8FFF7A912FEF3FA64FE4DFA62" box="[373,422,1422,1449]" collectionCode="III" pageId="57" pageNumber="58">III 5</specimenCode>
,
<specimenCode id="526B9EC8FFF7A912FE37FA64FE0EFA62" box="[433,485,1422,1449]" collectionCode="IV" pageId="57" pageNumber="58">IV 5</specimenCode>
; tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 4: 5: 4:
<specimenCode id="526B9EC8FFF7A912FB9FFA64FBD1FA63" box="[1049,1082,1422,1448]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="57" pageNumber="58">T2</specimenCode>
T: 0; sternal chaetotaxy IV–XII: 10: 8: 7: 7–8: 7–9: 8–9: 8–9: 0: 2; anterior genital operculum with 9 setae, posterior margin with 10–11 marginal setae,
<quantity id="C5359B56FFF7A912FF5BFA3CFEABFA3B" box="[221,320,1494,1520]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.08" metricValueMin="4.826" pageId="57" pageNumber="58" unit="in" value="19.5" valueMax="20.0" valueMin="19.0">19–20 in</quantity>
total; leg IV with a long tactile seta on both tarsal segments: basitarsus 4.29–4.75 times longer than deep (TS= 0.20–0.21), telotarsus 16.00–18.60 times longer than deep and 2.53–3.10 times longer than basitarsus (TS= 0.25–0.29).
</paragraph>
<paragraph id="027236B3FFF7A912FF41F9A8FBEDF8DA" blockId="57.[151,1437,954,2025]" pageId="57" pageNumber="58">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Male: body length 1.63. Pedipalps: trochanter 0.29/0.16 (1.81), femur 1.36/0.13 (10.46), patella 0.48/0.17 (2.82), chela 1.84/0.21 (8.76), hand 0.70/0.21 (3.33), movable chelal finger length 1.15. Chelicera 0.67/0.28 (2.39), movable finger length 0.37. Carapace 0.55/0.59 (0.93). Leg I: trochanter 0.20/0.12 (1.67), femur 0.78/0.08 (9.75), patella 0.38/0.08 (4.75), tibia 0.35/0.07 (5.00), tarsus 0.82/0.06 (13.67). Leg IV: trochanter 0.27/0.16 (1.69), femoropatella 1.12/0.22 (5.09), tibia 0.74/0.10 (7.40), basitarsus 0.32/0.08 (4.00), telotarsus 0.91/0.06 (15.17).</paragraph>
<paragraph id="027236B3FFF7A911FF41F8F0FC8CFF1E" blockId="57.[151,1437,954,2025]" lastBlockId="58.[151,1436,150,213]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">Females: body length 1.81–2.20. Pedipalps: trochanter 0.31–0.35/0.17–0.19 (1.82–1.84), femur 1.35– 1.41/0.13–0.14 (10.07–10.38), patella 0.49–0.52/0.18 (2.72–2.89), chela 1.83–1.91/0.22–0.23 (8.30–8.32), hand 0.68–0.69/0.22–0.23 (2.96–3.14), movable chelal finger length 1.15–1.19. Chelicera 0.70–0.74/0.29–0.30 (2.41– 2.47), movable finger length 0.39–0.40. Carapace 0.57–0.61/0.60–0.62 (0.95–0.98). Leg I: trochanter 0.19/0.13– 0.14 (1.36–1.46), femur 0.76–0.79/0.08–0.09 (8.44–9.88), patella 0.36–0.41/0.08 (4.50–5.13), tibia 0.32–0.38/0.05– 0.07 (5.43–6.40), tarsus 0.80–0.83/0.05 (16.00–16.60). Leg IV: trochanter 0.25–0.33/0.15–0.19 (1.67–1.74), femoropatella 1.09–1.17/0.19–0.21 (5.19–6.16), tibia 0.69–0.74/0.09–0.10 (7.40–7.67), basitarsus 0.30–0.38/0.07– 0.08 (4.29–4.75), telotarsus 0.93–0.96/0.05–0.06 (16.00–18.60).</paragraph>
</subSubSection>
<caption id="56B2663BFFF4A911FF11F881FEC6F800" ID-DOI="http://doi.org/10.5281/zenodo.7251736" ID-Zenodo-Dep="7251736" httpUri="https://zenodo.org/record/7251736/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,250,1899,1923]" targetBox="[151,1436,267,1876]" targetPageId="58">
<paragraph id="027236B3FFF4A911FF11F881FEC6F800" blockId="58.[151,1436,1899,1995]" pageId="58" pageNumber="59">
<emphasis id="30B9EAA1FFF4A911FF11F881FEC8F848" bold="true" box="[151,291,1899,1923]" pageId="58" pageNumber="59">FIGURE 44.</emphasis>
<taxonomicName id="C5CD4D30FFF4A911FEADF881FD61F848" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[299,650,1899,1923]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF4A911FEADF881FD61F848" box="[299,650,1899,1923]" italics="true" pageId="58" pageNumber="59">Lagynochthonius yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF4A911FD15F881FD0CF848" bold="true" box="[659,743,1899,1923]" pageId="58" pageNumber="59">
<taxonomicNameLabel id="2B8A57DAFFF4A911FD15F881FD0CF848" box="[659,743,1899,1923]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view).
</paragraph>
</caption>
<caption id="56B2663BFFF5A910FF11F8B9FA68F830" ID-DOI="http://doi.org/10.5281/zenodo.7251738" ID-Zenodo-Dep="7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="59" pageNumber="60" startId="59.[151,250,1875,1899]" targetBox="[168,1396,189,1847]" targetPageId="59">
<paragraph id="027236B3FFF5A910FF11F8B9FA68F830" blockId="59.[151,1437,1875,2043]" pageId="59" pageNumber="60">
<emphasis id="30B9EAA1FFF5A910FF11F8B9FECAF8A0" bold="true" box="[151,289,1875,1899]" pageId="59" pageNumber="60">FIGURE 45.</emphasis>
<taxonomicName id="C5CD4D30FFF5A910FEAFF8B9FD6DF8A0" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[297,646,1875,1899]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF5A910FEAFF8B9FD6DF8A0" box="[297,646,1875,1899]" italics="true" pageId="59" pageNumber="60">Lagynochthonius yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF5A910FD08F8B9FD0BF8A0" bold="true" box="[654,736,1875,1899]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="2B8A57DAFFF5A910FD08F8B9FD0BF8A0" box="[654,736,1875,1899]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D).
</paragraph>
</caption>
<subSubSection id="4AD76538FFF2A917FF41FF7DFD07FD62" pageId="60" pageNumber="61" type="discussion">
<paragraph id="027236B3FFF2A917FF41FF7DFE65FED6" blockId="60.[151,1437,151,897]" pageId="60" pageNumber="61">
<emphasis id="30B9EAA1FFF2A917FF41FF7DFED0FF7A" bold="true" box="[199,315,151,177]" pageId="60" pageNumber="61">Remarks.</emphasis>
<taxonomicName id="C5CD4D30FFF2A917FEC4FF7DFD2FFF7B" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[322,708,151,177]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF2A917FEC4FF7DFD2FFF7B" box="[322,708,151,177]" italics="true" pageId="60" pageNumber="61">Lagynochthonius yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF2A917FD4DFF7DFCCEFF7A" bold="true" box="[715,805,151,177]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="2B8A57DAFFF2A917FD4DFF7DFCCEFF7A" box="[715,805,151,177]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="C5CD4D30FFF2A917FC3EFF7DFBE5FF7A" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[952,1038,151,177]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="fengi" status="sp. nov.">
<emphasis id="30B9EAA1FFF2A917FC3EFF7DFBE5FF7A" box="[952,1038,151,177]" italics="true" pageId="60" pageNumber="61">L. fengi</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF2A917FB93FF7DFB84FF7A" bold="true" box="[1045,1135,151,177]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="2B8A57DAFFF2A917FB93FF7DFB84FF7A" box="[1045,1135,151,177]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having intercalary teeth on both chelal fingers and a pointed and triangular epistome, but differs by the longer chela (e.g. chela length 1.84
<emphasis id="30B9EAA1FFF2A917FF11FF35FF5DFF33" box="[151,182,223,248]" italics="true" pageId="60" pageNumber="61">vs.</emphasis>
<quantity id="C5359B56FFF2A917FF38FF34FE8FFF32" box="[190,356,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.535" metricValueMax="1.54" metricValueMin="1.53" pageId="60" pageNumber="61" unit="mm" value="1.535" valueMax="1.54" valueMin="1.53">1.53–1.54 mm</quantity>
, movable chelal finger length 1.15
<emphasis id="30B9EAA1FFF2A917FD71FF35FCFDFF33" box="[759,790,223,248]" italics="true" pageId="60" pageNumber="61">vs.</emphasis>
<quantity id="C5359B56FFF2A917FC98FF35FC2EFF32" box="[798,965,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="8.95" metricValueMax="9.1" metricValueMin="8.8" pageId="60" pageNumber="61" unit="mm" value="0.895" valueMax="0.91" valueMin="0.88">0.88–0.91 mm</quantity>
; all in males) and the number of setae on tergites I–IV (4
<emphasis id="30B9EAA1FFF2A917FECCFEE9FE82FED7" box="[330,361,259,284]" italics="true" pageId="60" pageNumber="61">vs.</emphasis>
2).
</paragraph>
<paragraph id="027236B3FFF2A917FF41FECDFD5CFE42" blockId="60.[151,1437,151,897]" pageId="60" pageNumber="61">
<taxonomicName id="C5CD4D30FFF2A917FF41FECDFDA1FE8B" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[199,586,295,321]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF2A917FF41FECDFDA1FE8B" box="[199,586,295,321]" italics="true" pageId="60" pageNumber="61">Lagynochthonius yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF2A917FDD4FECDFD45FE8A" bold="true" box="[594,686,295,321]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="2B8A57DAFFF2A917FDD4FECDFD45FE8A" box="[594,686,295,321]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="C5CD4D30FFF2A917FCC0FECDFBFCFE8A" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[838,1047,295,321]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="bailongtanensis">
<emphasis id="30B9EAA1FFF2A917FCC0FECDFBFCFE8A" box="[838,1047,295,321]" italics="true" pageId="60" pageNumber="61">L. bailongtanensis</emphasis>
</taxonomicName>
in the shape of epistome (pointed and triangular
<emphasis id="30B9EAA1FFF2A917FEBAFEA1FEB8FEAF" box="[316,339,331,356]" italics="true" pageId="60" pageNumber="61">vs</emphasis>
. rounded and obtuse), and the pattern of dentation on the chelal fingers (with intercalary teeth on both chelal fingers
<emphasis id="30B9EAA1FFF2A917FEE9FE85FE6DFE43" box="[367,390,367,392]" italics="true" pageId="60" pageNumber="61">vs</emphasis>
. without intercalary teeth).
</paragraph>
<paragraph id="027236B3FFF2A917FF41FE79FEBDFE3E" blockId="60.[151,1437,151,897]" pageId="60" pageNumber="61">
<taxonomicName id="C5CD4D30FFF2A917FF41FE79FDA1FE67" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[199,586,403,429]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF2A917FF41FE79FDA1FE67" box="[199,586,403,429]" italics="true" pageId="60" pageNumber="61">Lagynochthonius yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF2A917FDD4FE79FD46FE66" bold="true" box="[594,685,403,429]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="2B8A57DAFFF2A917FDD4FE79FD46FE66" box="[594,685,403,429]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="C5CD4D30FFF2A917FCC0FE79FBFBFE66" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[838,1040,403,429]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="laoxueyanensis">
<emphasis id="30B9EAA1FFF2A917FCC0FE79FBFBFE66" box="[838,1040,403,429]" italics="true" pageId="60" pageNumber="61">L. laoxueyanensis</emphasis>
</taxonomicName>
by the number of setae on tergites I–II (4 and 4
<emphasis id="30B9EAA1FFF2A917FEABFE5DFEAFFE1B" box="[301,324,439,464]" italics="true" pageId="60" pageNumber="61">vs</emphasis>
. 2 and 2, respectively), and the presence of intercalary teeth on both chelal fingers (
<bibRefCitation id="665C4B42FFF2A917FB7CFE5DFA7CFE1A" author="Li, Y. C. &amp; Liu, H. &amp; Shi, A. M." box="[1274,1431,439,465]" pageId="60" pageNumber="61" pagination="28 - 34" refId="ref39131" refString="Li, Y. C., Liu, H. &amp; Shi, A. M. (2019) A new cave-dwelling species of Lagynochthonius (Arachnida: Pseudoscorpiones: Chthoniidae) from Yunnan Province, China. Zootaxa, 4571 (1), 28 - 34. https: // doi. org / 10.11646 / zootaxa. 4571.1.2" type="journal article" year="2019">
Li
<emphasis id="30B9EAA1FFF2A917FA9AFE5DFAA5FE1A" box="[1308,1358,439,465]" italics="true" pageId="60" pageNumber="61">et al</emphasis>
. 2019
</bibRefCitation>
;
<bibRefCitation id="665C4B42FFF2A917FF11FE31FEA1FE3E" author="Hou, Y. M. &amp; Gao, Z. Z. &amp; Zhang, F." box="[151,330,475,501]" pageId="60" pageNumber="61" pagination="65 - 83" refId="ref38684" refString="Hou, Y. M., Gao, Z. Z. &amp; Zhang, F. (2022) Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China. ZooKeys, 1097, 65 - 83. https: // doi. org / 10.3897 / zookeys. 1097.82527" type="journal article" year="2022">
Hou
<emphasis id="30B9EAA1FFF2A917FF48FE31FEEBFE3E" box="[206,256,475,501]" italics="true" pageId="60" pageNumber="61">et al</emphasis>
. 2022
</bibRefCitation>
).
</paragraph>
<paragraph id="027236B3FFF2A917FF41FE15FD07FD62" blockId="60.[151,1437,151,897]" pageId="60" pageNumber="61">
In
<taxonomicName id="C5CD4D30FFF2A917FF62FE15FE52FDD3" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[228,441,511,536]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="yaowangguensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF2A917FF62FE15FE52FDD3" box="[228,441,511,536]" italics="true" pageId="60" pageNumber="61">L. yaowangguensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF2A917FE39FE15FDFCFDD2" bold="true" box="[447,535,511,537]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="2B8A57DAFFF2A917FE39FE15FDFCFDD2" box="[447,535,511,537]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the hand of the chela is slightly narrowed and the apodeme of the movable chelal finger is weakly sclerotized, but the presence of a modified accessory tooth (
<emphasis id="30B9EAA1FFF2A917FC69FDC9FBEEFDF6" box="[1007,1029,547,573]" italics="true" pageId="60" pageNumber="61">td</emphasis>
) on the dorso-antiaxial face of fixed chelal finger (
<figureCitation id="9AF62A36FFF2A917FEBEFDADFE2FFDAA" box="[312,452,582,609]" captionStart="FIGURE 45" captionStartId="59.[151,250,1875,1899]" captionTargetBox="[168,1396,189,1847]" captionTargetId="figure-19@59.[168,1420,181,1851]" captionTargetPageId="59" captionText="FIGURE 45. Lagynochthonius yaowangguensis sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips and modified tooth (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251738" httpUri="https://zenodo.org/record/7251738/files/figure.png" pageId="60" pageNumber="61">Figs 45F–G</figureCitation>
,
<figureCitation id="9AF62A36FFF2A917FE54FDADFDEAFDAA" box="[466,513,583,609]" captionStart="FIGURE 46" captionStartId="62.[151,250,1614,1638]" captionTargetBox="[151,1436,202,1590]" captionTargetId="figure-20@62.[151,1436,181,1590]" captionTargetPageId="62" captionText="FIGURE 46. Comparison of the chela of some cave-dwelling species of Lagynochthonius and Tyrannochthonius from Yunnan (black dotted bordered rectangles refer to the narrowed chelal hands and the sclerotized apodeme of movable chelal fingers). A. Left chela of L. crassus sp. nov. (lateral view); B. Left chela of L. fengi sp. nov. (lateral view); C. Left chela of L. magnidentatus sp. nov. (lateral view); D. Left chela of L. minimus sp. nov. (lateral view); E. Left chela of L. retrorsus sp. nov. (lateral view); F. Left chela of L. serratus sp. nov. (lateral view); G. Left chela of L. spinulentus sp. nov. (lateral view); H. Left chela of L. xiaolinensis sp. nov. (lateral view); I. Left chela of L. xibaiensis sp. nov. (lateral view); J. Left chela of L. xinjiaoensis sp. nov. (lateral view); K. Left chela of L. yaowangguensis sp. nov. (lateral view); L. A modified accessory tooth (td) present on dorso-antiaxial face of fixed chelal finger of L. yaowangguensis sp. nov. (red arrow, dorsal view); M–N. Left chela of Tyrannochthonius sp. nov. (lateral view). Scale bars: 0.25 mm (A–C, E–K, M–N); 0.10 mm (D, L)." figureDoi="http://doi.org/10.5281/zenodo.7251740" httpUri="https://zenodo.org/record/7251740/files/figure.png" pageId="60" pageNumber="61">46L</figureCitation>
) is a typical character of the genus
<taxonomicName id="C5CD4D30FFF2A917FC25FDADFB8FFDAA" authorityName="Beier" authorityYear="1951" box="[931,1124,583,609]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="30B9EAA1FFF2A917FC25FDADFB8FFDAA" box="[931,1124,583,609]" italics="true" pageId="60" pageNumber="61">Lagynochthonius</emphasis>
</taxonomicName>
(
<bibRefCitation id="665C4B42FFF2A917FBF3FDADFAABFDAA" author="Chamberlin, J. C." box="[1141,1344,583,609]" pageId="60" pageNumber="61" pagination="303 - 352" refId="ref38231" refString="Chamberlin, J. C. (1962) New and little-known false scorpions, principally from caves, belonging to the families Chthoniidae and Neobisiidae (Arachnida, Chelonethida). Bulletin of the American Museum of Natural History, 123, 303 - 352." type="journal article" year="1962">Chamberlin 1962</bibRefCitation>
;
<bibRefCitation id="665C4B42FFF2A917FAC8FDADFF3AFD4E" author="Judson, M. L. I." pageId="60" pageNumber="61" pagination="53 - 68" refId="ref38953" refString="Judson, M. L. I. (2007) A new and endangered species of the pseudoscorpion genus Lagynochthonius from a cave in Vietnam, with notes on chelal morphology and the composition of the Tyrannochthoniini (Arachnida, Chelonethi, Chthoniidae). Zootaxa, 1627, 53 - 68. https: // doi. org / 10.11646 / zootaxa. 1627.1.4" type="journal article" year="2007">Judson 2007</bibRefCitation>
) and maintained in this species. Therefore, considering all the characteristic factors, it is most appropriate to place this new species in the genus
<taxonomicName id="C5CD4D30FFF2A917FDA3FD65FD0DFD62" authorityName="Beier" authorityYear="1951" box="[549,742,655,681]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="30B9EAA1FFF2A917FDA3FD65FD0DFD62" box="[549,742,655,681]" italics="true" pageId="60" pageNumber="61">Lagynochthonius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFF2A917FF41FD58FDF0FC4A" pageId="60" pageNumber="61" type="distribution">
<paragraph id="027236B3FFF2A917FF41FD58FDF0FC4A" blockId="60.[151,1437,151,897]" pageId="60" pageNumber="61">
<emphasis id="30B9EAA1FFF2A917FF41FD58FE12FD07" bold="true" box="[199,505,690,717]" pageId="60" pageNumber="61">Distribution and habitat.</emphasis>
This species is only known from the
<typeStatus id="DD768811FFF2A917FC3DFD59FC00FD06" box="[955,1003,691,717]" pageId="60" pageNumber="61">type</typeStatus>
locality, Unnamed Cave 4 (
<figureCitation id="9AF62A36FFF2A917FAB0FD59FA7CFD06" box="[1334,1431,691,717]" captionStart="FIGURE 1" captionStartId="3.[151,250,1542,1566]" captionTargetBox="[152,1433,567,1514]" captionTargetId="figure-169@3.[151,1436,561,1519]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of the type locality of troglomorphic Lagynochthonius species in Yunnan (China). A. L. retrorsus sp. nov. (Qinglong Cave); B. L. fengi sp. nov. (Unnamed Cave 2); C. L. xiaolinensis sp. nov. (Dayan Cave); D. L. xibaiensis sp. nov. (Hama Cave); E. L. spinulentus sp. nov. (Yingshan Cave); F. L. xinjiaoensis sp. nov. (Yueliang Cave); G. L. magnidentatus sp. nov. (Xiao Cave); H. L. bailongtanensis sp. nov. (Qianfo Cave); I. L. laoxueyanensis sp. nov. (Laoxueyan Cave); J. L. serratus sp. nov. (Unnamed Cave 3); K. L. yaowangguensis sp. nov. (Unnamed Cave 4); L. L. crassus sp. nov. (Unnamed Cave 1); M–N. L. minimus sp. nov. (M: Yanfen Cave, N: Cizhu Cave)." figureDoi="http://doi.org/10.5281/zenodo.7251604" httpUri="https://zenodo.org/record/7251604/files/figure.png" pageId="60" pageNumber="61">Figs 1K</figureCitation>
,
<figureCitation id="9AF62A36FFF2A917FF11FD3DFF00FD3A" box="[151,235,727,753]" captionStart="FIGURE 42" captionStartId="56.[151,250,1890,1914]" captionTargetBox="[151,1436,525,1867]" captionTargetId="figure-205@56.[151,1436,525,1867]" captionTargetPageId="56" captionText="FIGURE 42. Unnamed Cave 4, type locality of Lagynochthonius yaowangguensis sp. nov., A. Entrance; B. Inside the cave entrance; C–D. Peculiar landscapes inside the cave; E. Area where L. yaowangguensis sp. nov. specimens were collected; F–G. Live individuals of L. yaowangguensis sp. nov. (F: male, G: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251732" httpUri="https://zenodo.org/record/7251732/files/figure.png" pageId="60" pageNumber="61">42A–E</figureCitation>
), which is located about
<quantity id="C5359B56FFF2A917FD82FD3DFDB8FD3A" box="[516,595,727,753]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="60" pageNumber="61" unit="km" value="0.8">0.8 km</quantity>
northwest of Laoshanyaowanggu (a valley near the border between
<collectingCountry id="7ADA7623FFF2A917FADFFD3DFA77FD3A" box="[1369,1436,727,753]" name="China" pageId="60" pageNumber="61">China</collectingCountry>
and
<collectingCountry id="7ADA7623FFF2A917FF43FD11FEC2FCDE" box="[197,297,763,789]" name="Vietnam" pageId="60" pageNumber="61">Vietnam</collectingCountry>
). The cave was found near the top of a mountain. Several boulders were scattered around the triangular entrance (about 2 meters high and 3 meters wide). The interior space is large, and extends horizontally. The total length of the cave is unknown. All specimens were collected under stones and detritus in the deep zone (temperature around 11°C, humidity over 85%).
</paragraph>
</subSubSection>
</treatment>
</document>
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