503 lines
77 KiB
XML
503 lines
77 KiB
XML
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<mods:title id="19EAEDA422029DFF8DB611D1589BE27C">Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="504732D254E2212EF05297FFF61B3F8E">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="45AEDBF1ACB75E87749909B48A7CCAA7">Zhang, Feng</mods:namePart>
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<mods:date id="32B08DD37E47D8F2CCA47BDAFE2E62E7">2022</mods:date>
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<treatment id="8A6487A5FFEBA901FF11FAD0FD8AFBFE" ID-DOI="http://doi.org/10.5281/zenodo.7251618" ID-GBIF-Taxon="203411085" ID-Zenodo-Dep="7251618" LSID="urn:lsid:plazi:treatment:8A6487A5FFEBA901FF11FAD0FD8AFBFE" httpUri="http://treatment.plazi.org/id/8A6487A5FFEBA901FF11FAD0FD8AFBFE" lastPageId="42" lastPageNumber="43" pageId="37" pageNumber="38">
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<heading id="593A81DFFFEBA90EFF11FAD0FCF0FA98" bold="true" box="[151,795,1338,1365]" fontSize="11" level="2" pageId="37" pageNumber="38" reason="4">
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<taxonomicName id="C5CD4D30FFEBA90EFF11FAD0FE15FA9E" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[151,510,1338,1365]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="spinulentus" status="sp. nov.">
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<emphasis id="30B9EAA1FFEBA90EFF11FAD0FE15FA9E" bold="true" box="[151,510,1338,1365]" italics="true" pageId="37" pageNumber="38">Lagynochthonius spinulentus</emphasis>
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<taxonomicNameLabel id="2B8A57DAFFEBA90EFD80FAD1FD8FFA9E" box="[518,612,1339,1365]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
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(ØḦIJ伪oi)
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="4AD76538FFEBA90EFF11FAB5FEFEFAB2" box="[151,277,1375,1401]" pageId="37" pageNumber="38" type="description">
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<paragraph id="027236B3FFEBA90EFF11FAB5FEFEFAB2" blockId="37.[151,795,1338,1401]" box="[151,277,1375,1401]" pageId="37" pageNumber="38">
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<figureCitation id="9AF62A36FFEBA90EFF11FAB5FEFEFAB2" box="[151,277,1375,1401]" captionStart-0="FIGURE 27" captionStart-1="FIGURE 28" captionStart-2="FIGURE 29" captionStart-3="FIGURE 30" captionStartId-0="38.[151,250,715,739]" captionStartId-1="38.[151,250,1556,1580]" captionStartId-2="40.[151,250,1848,1872]" captionStartId-3="41.[151,250,1868,1892]" captionTargetBox-0="[151,1437,181,691]" captionTargetBox-1="[193,1436,811,1518]" captionTargetBox-2="[151,1435,181,1824]" captionTargetBox-3="[182,1405,181,1844]" captionTargetId-0="figure-20@38.[151,1437,181,691]" captionTargetId-1="figure-49@38.[151,1437,800,1533]" captionTargetId-2="figure-20@40.[151,1436,181,1824]" captionTargetId-3="figure-19@41.[182,1405,181,1844]" captionTargetPageId-0="38" captionTargetPageId-1="38" captionTargetPageId-2="40" captionTargetPageId-3="41" captionText-0="FIGURE 27. Yingshan Cave, type locality of Lagynochthonius spinulentus sp. nov., A. Entrance; B. Live male of L. spinulentus sp. nov. in its natural environment." captionText-1="FIGURE 28. Lagynochthonius spinulentus sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." captionText-2="FIGURE 29. Lagynochthonius spinulentus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." captionText-3="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi-0="http://doi.org/10.5281/zenodo.7251687" figureDoi-1="http://doi.org/10.5281/zenodo.7251691" figureDoi-2="http://doi.org/10.5281/zenodo.7251695" figureDoi-3="http://doi.org/10.5281/zenodo.7251699" httpUri-0="https://zenodo.org/record/7251687/files/figure.png" httpUri-1="https://zenodo.org/record/7251691/files/figure.png" httpUri-2="https://zenodo.org/record/7251695/files/figure.png" httpUri-3="https://zenodo.org/record/7251699/files/figure.png" pageId="37" pageNumber="38">Figs 27–30</figureCitation>
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</paragraph>
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<subSubSection id="4AD76538FFEBA90EFF11FA4DFC8DF99A" pageId="37" pageNumber="38" type="materials_examined">
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<emphasis id="30B9EAA1FFEBA90EFF11FA4DFEA3FA0B" bold="true" box="[151,328,1446,1473]" pageId="37" pageNumber="38">Type material.</emphasis>
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(Ps.-
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):
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,
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<collectingRegion id="C009F851FFEBA90EFC6BFA4DFB58FA0A" box="[1005,1203,1447,1473]" country="China" name="Yunnan" pageId="37" pageNumber="38">Yunnan Province</collectingRegion>
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,
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<collectingCounty id="EB134E3FFFEBA90EFB47FA4DFA72FA0A" box="[1217,1433,1447,1473]" pageId="37" pageNumber="38">Zhenxiong County</collectingCounty>
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,
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,
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<location id="07126068FFEBA90EFED7FA21FE03FA2E" LSID="urn:lsid:plazi:treatment:8A6487A5FFEBA901FF11FAD0FD8AFBFE:07126068FFEBA90EFED7FA21FE03FA2E" box="[337,488,1483,1509]" country="China" county="Zhenxiong County" latitude="27.360672" longLatPrecision="1" longitude="104.82131" municipality="Zhongtun Town" name="Qixin Village" pageId="37" pageNumber="38" stateProvince="Yunnan">Qixin Village</location>
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,
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<location id="07126068FFEBA90EFE69FA21FD73FA2E" LSID="urn:lsid:plazi:treatment:8A6487A5FFEBA901FF11FAD0FD8AFBFE:07126068FFEBA90EFE69FA21FD73FA2E" box="[495,664,1483,1509]" country="China" county="Zhenxiong County" latitude="27.360672" longLatPrecision="1" longitude="104.82131" municipality="Zhongtun Town" name="Yingshan Cave" pageId="37" pageNumber="38" stateProvince="Yunnan">Yingshan Cave</location>
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, under stones within
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<quantity id="C5359B56FFEBA90EFCFFFA21FC0CFA2E" box="[889,999,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="1.1" metricValueMax="2.0" metricValueMin="0.2" pageId="37" pageNumber="38" unit="m" value="110.0" valueMax="200.0" valueMin="20.0">
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</quantity>
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from the cave entrance [
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,
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],
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a.s.l.
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</elevation>
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,
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<collectingDate id="6637E99BFFEBA90EFE71FA05FD3BF9C2" box="[503,720,1519,1545]" pageId="37" pageNumber="38" value="2021-09-28">28 September 2021</collectingDate>
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</date>
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,
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<collectorName id="AF385365FFEBA90EFD5FFA05FC86F9C2" box="[729,877,1519,1545]" pageId="37" pageNumber="38">Zegang Feng</collectorName>
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,
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<collectorName id="AF385365FFEBA90EFCF3FA05FBFCF9C2" box="[885,1047,1519,1545]" pageId="37" pageNumber="38">Yanmeng Hou</collectorName>
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,
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<collectorName id="AF385365FFEBA90EFBA6FA05FB67F9C2" box="[1056,1164,1519,1545]" pageId="37" pageNumber="38">Lu Zhang</collectorName>
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&
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||
<collectorName id="AF385365FFEBA90EFB2DFA05FB1EF9C2" box="[1195,1269,1519,1545]" pageId="37" pageNumber="38">Liu Fu</collectorName>
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leg.
|
||
</materialsCitation>
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<typeStatus id="DD768811FFEBA90EFAAEFA05FA72F9C2" box="[1320,1433,1519,1545]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="14CBFD3AFFEBA90EFF11F9F9FF2AF9E6" box="[151,193,1555,1581]" pageId="37" pageNumber="38" type="male">2 ♂</specimenCount>
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(Ps.-
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& HBUARA#2021-41303)
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</materialsCitation>
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,
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<materialsCitation id="B2A53CEEFFEBA90EFC2BF9F9FC88F99A" ID-GBIF-Occurrence="3946861325" collectingDate="2021-09-28" collectionCode="MSWU-HBUARA" collectorName="Zegang Feng & Yanmeng Hou & Lu Zhang & Liu Fu" country="China" county="Zhenxiong County" elevation="110" latitude="27.360672" location="Qixin Village" longLatPrecision="1" longitude="104.82131" municipality="Zhongtun Town" pageId="37" pageNumber="38" specimenCode="MSWU-HBUARA#2021-41304" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
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<specimenCount id="14CBFD3AFFEBA90EFC2BF9F9FC33F9E6" box="[941,984,1555,1581]" pageId="37" pageNumber="38" type="female">2 ♀</specimenCount>
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(Ps.-
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|
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& HBUARA#2021-41305), all with the same data as the holotype
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||
</materialsCitation>
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.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="4AD76538FFEBA90EFF41F9B1FDC2F952" pageId="37" pageNumber="38" type="etymology">
|
||
<paragraph id="027236B3FFEBA90EFF41F9B1FDC2F952" blockId="37.[151,1437,1446,2013]" pageId="37" pageNumber="38">
|
||
<emphasis id="30B9EAA1FFEBA90EFF41F9B1FEA5F9BE" bold="true" box="[199,334,1627,1653]" pageId="37" pageNumber="38">Etymology.</emphasis>
|
||
The specific name is derived from the Latin word “
|
||
<taxonomicName id="C5CD4D30FFEBA90EFC17F9B1FBFAF9BE" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[913,1041,1627,1653]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="spinulentus">
|
||
<emphasis id="30B9EAA1FFEBA90EFC17F9B1FBFAF9BE" box="[913,1041,1627,1653]" italics="true" pageId="37" pageNumber="38">spinulentus</emphasis>
|
||
</taxonomicName>
|
||
”, meaning spinous, which refers to the characters of more coxal spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AD76538FFEBA90EFF41F948FD9DF8BA" pageId="37" pageNumber="38" type="diagnosis">
|
||
<paragraph id="027236B3FFEBA90EFF41F948FD9DF8BA" blockId="37.[151,1437,1446,2013]" pageId="37" pageNumber="38">
|
||
<emphasis id="30B9EAA1FFEBA90EFF41F948FE6DF90B" bold="true" box="[199,390,1698,1728]" pageId="37" pageNumber="38">
|
||
Diagnosis (
|
||
<specimenCount id="14CBFD3AFFEBA90EFECBF94EFE89F90B" box="[333,354,1700,1728]" pageId="37" pageNumber="38" type="male">♂</specimenCount>
|
||
<specimenCount id="14CBFD3AFFEBA90EFEE4F94EFE9CF90B" box="[354,375,1700,1728]" pageId="37" pageNumber="38" type="female">♀</specimenCount>
|
||
).
|
||
</emphasis>
|
||
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed and small, triangular; posterior margin of carapace with 2 setae; tergites I–III each with 2 setae. Pedipalps slender, femur 8.39–8.81 (
|
||
<specimenCount id="14CBFD3AFFEBA90EFA86F901FAFCF8CE" box="[1280,1303,1771,1797]" pageId="37" pageNumber="38" type="male">♂</specimenCount>
|
||
), 7.40–7.74 (
|
||
<specimenCount id="14CBFD3AFFEBA90EFF24F8E5FF5FF8E2" box="[162,180,1807,1833]" pageId="37" pageNumber="38" type="female">♀</specimenCount>
|
||
) times longer than broad; chela 8.00–8.28 (
|
||
<specimenCount id="14CBFD3AFFEBA90EFD23F8E5FD57F8E2" box="[677,700,1807,1833]" pageId="37" pageNumber="38" type="male">♂</specimenCount>
|
||
), 7.34–8.42 (
|
||
<specimenCount id="14CBFD3AFFEBA90EFCD3F8E5FC8CF8E2" box="[853,871,1807,1833]" pageId="37" pageNumber="38" type="female">♀</specimenCount>
|
||
) times longer than broad; only fixed chelal finger with intercalary teeth and a modified accessory tooth (
|
||
<emphasis id="30B9EAA1FFEBA90EFD7AF8D9FCF9F886" box="[764,786,1843,1869]" italics="true" pageId="37" pageNumber="38">td</emphasis>
|
||
) on dorso-antiaxial face; chemosensory setae (
|
||
<emphasis id="30B9EAA1FFEBA90EFAA5F8D9FAD1F887" box="[1315,1338,1843,1868]" italics="true" pageId="37" pageNumber="38">sc</emphasis>
|
||
) present on dorsum of chelal hand; sensilla present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AD76538FFEBA901FF41F890FD8FFE66" lastPageId="42" lastPageNumber="43" pageId="37" pageNumber="38" type="description">
|
||
<paragraph id="027236B3FFEBA90EFF41F890FBFAF85E" blockId="37.[151,1437,1446,2013]" box="[199,1041,1914,1941]" pageId="37" pageNumber="38">
|
||
<emphasis id="30B9EAA1FFEBA90EFF41F890FEB2F85F" bold="true" box="[199,345,1914,1940]" pageId="37" pageNumber="38">Description.</emphasis>
|
||
Males (
|
||
<typeStatus id="DD768811FFEBA90EFE35F891FDFCF85E" box="[435,535,1915,1941]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="DD768811FFEBA90EFDCBF891FD51F85E" box="[589,698,1915,1941]" pageId="37" pageNumber="38" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<figureCitation id="9AF62A36FFEBA90EFD56F891FCD1F85E" box="[720,826,1915,1941]" captionStart="FIGURE 27" captionStartId="38.[151,250,715,739]" captionTargetBox="[151,1437,181,691]" captionTargetId="figure-20@38.[151,1437,181,691]" captionTargetPageId="38" captionText="FIGURE 27. Yingshan Cave, type locality of Lagynochthonius spinulentus sp. nov., A. Entrance; B. Live male of L. spinulentus sp. nov. in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251687" httpUri="https://zenodo.org/record/7251687/files/figure.png" pageId="37" pageNumber="38">Figs 27B</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFEBA90EFCC0F891FC92F85E" box="[838,889,1915,1941]" captionStart="FIGURE 28" captionStartId="38.[151,250,1556,1580]" captionTargetBox="[193,1436,811,1518]" captionTargetId="figure-49@38.[151,1437,800,1533]" captionTargetPageId="38" captionText="FIGURE 28. Lagynochthonius spinulentus sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251691" httpUri="https://zenodo.org/record/7251691/files/figure.png" pageId="37" pageNumber="38">28A</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFEBA90EFC02F891FC32F85E" box="[900,985,1915,1941]" captionStart="FIGURE 29" captionStartId="40.[151,250,1848,1872]" captionTargetBox="[151,1435,181,1824]" captionTargetId="figure-20@40.[151,1436,181,1824]" captionTargetPageId="40" captionText="FIGURE 29. Lagynochthonius spinulentus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251695" httpUri="https://zenodo.org/record/7251695/files/figure.png" pageId="37" pageNumber="38">29A–D</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFEBA90EFC62F891FBEFF85E" box="[996,1028,1915,1941]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="37" pageNumber="38">30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="027236B3FFEBA90EFF41F875FEE0F816" blockId="37.[151,1437,1446,2013]" pageId="37" pageNumber="38">
|
||
Colour (
|
||
<figureCitation id="9AF62A36FFEBA90EFEA2F875FE64F872" box="[292,399,1951,1977]" captionStart="FIGURE 27" captionStartId="38.[151,250,715,739]" captionTargetBox="[151,1437,181,691]" captionTargetId="figure-20@38.[151,1437,181,691]" captionTargetPageId="38" captionText="FIGURE 27. Yingshan Cave, type locality of Lagynochthonius spinulentus sp. nov., A. Entrance; B. Live male of L. spinulentus sp. nov. in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251687" httpUri="https://zenodo.org/record/7251687/files/figure.png" pageId="37" pageNumber="38">Figs 27B</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFEBA90EFE1DF875FE25F872" box="[411,462,1951,1977]" captionStart="FIGURE 28" captionStartId="38.[151,250,1556,1580]" captionTargetBox="[193,1436,811,1518]" captionTargetId="figure-49@38.[151,1437,800,1533]" captionTargetPageId="38" captionText="FIGURE 28. Lagynochthonius spinulentus sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251691" httpUri="https://zenodo.org/record/7251691/files/figure.png" pageId="37" pageNumber="38">28A</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFEBA90EFE5CF875FDD9F872" box="[474,562,1951,1977]" captionStart="FIGURE 29" captionStartId="40.[151,250,1848,1872]" captionTargetBox="[151,1435,181,1824]" captionTargetId="figure-20@40.[151,1436,181,1824]" captionTargetPageId="40" captionText="FIGURE 29. Lagynochthonius spinulentus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251695" httpUri="https://zenodo.org/record/7251695/files/figure.png" pageId="37" pageNumber="38">29A–D</figureCitation>
|
||
): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
|
||
</paragraph>
|
||
<caption id="56B2663BFFE8A90DFF11FD21FE10FCCC" ID-DOI="http://doi.org/10.5281/zenodo.7251687" ID-Zenodo-Dep="7251687" httpUri="https://zenodo.org/record/7251687/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,250,715,739]" targetBox="[151,1437,181,691]" targetPageId="38">
|
||
<paragraph id="027236B3FFE8A90DFF11FD21FE10FCCC" blockId="38.[151,1436,715,776]" pageId="38" pageNumber="39">
|
||
<emphasis id="30B9EAA1FFE8A90DFF11FD21FEF4FD28" bold="true" box="[151,287,715,739]" pageId="38" pageNumber="39">FIGURE 27.</emphasis>
|
||
Yingshan Cave, type locality of
|
||
<taxonomicName id="C5CD4D30FFE8A90DFDE2FD21FC60FD28" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[612,907,715,739]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="spinulentus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFE8A90DFDE2FD21FC60FD28" box="[612,907,715,739]" italics="true" pageId="38" pageNumber="39">Lagynochthonius spinulentus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFE8A90DFC17FD26FC0BFD2F" bold="true" box="[913,992,716,740]" pageId="38" pageNumber="39">
|
||
<taxonomicNameLabel id="2B8A57DAFFE8A90DFC17FD26FC0BFD2F" box="[913,992,716,740]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A. Entrance; B. Live male of
|
||
<taxonomicName id="C5CD4D30FFE8A90DFA89FD26FA77FD28" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[1295,1436,715,739]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="spinulentus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFE8A90DFA89FD26FA77FD28" box="[1295,1436,715,739]" italics="true" pageId="38" pageNumber="39">L. spinulentus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFE8A90DFF11FD1AFF03FCC3" bold="true" box="[151,232,752,776]" pageId="38" pageNumber="39">
|
||
<taxonomicNameLabel id="2B8A57DAFFE8A90DFF11FD1AFF03FCC3" box="[151,232,752,776]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in its natural environment.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="56B2663BFFE8A90DFF11F9FEFECEF99B" ID-DOI="http://doi.org/10.5281/zenodo.7251691" ID-Zenodo-Dep="7251691" httpUri="https://zenodo.org/record/7251691/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,250,1556,1580]" targetBox="[193,1436,811,1518]" targetPageId="38">
|
||
<paragraph id="027236B3FFE8A90DFF11F9FEFECEF99B" blockId="38.[151,1436,1556,1616]" pageId="38" pageNumber="39">
|
||
<emphasis id="30B9EAA1FFE8A90DFF11F9FEFEC8F9E7" bold="true" box="[151,291,1556,1580]" pageId="38" pageNumber="39">FIGURE 28.</emphasis>
|
||
<taxonomicName id="C5CD4D30FFE8A90DFEAAF9FEFDBCF9E7" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[300,599,1556,1580]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="spinulentus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFE8A90DFEAAF9FEFDBCF9E7" box="[300,599,1556,1580]" italics="true" pageId="38" pageNumber="39">Lagynochthonius spinulentus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFE8A90DFDE7F9FEFD5EF9E7" bold="true" box="[609,693,1556,1580]" pageId="38" pageNumber="39">
|
||
<taxonomicNameLabel id="2B8A57DAFFE8A90DFDE7F9FEFD5EF9E7" box="[609,693,1556,1580]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="027236B3FFE8A90DFF41F994FA72F872" blockId="38.[151,1436,1662,2013]" pageId="38" pageNumber="39">
|
||
Cephalothorax (
|
||
<figureCitation id="9AF62A36FFE8A90DFEF9F994FE06F953" box="[383,493,1662,1688]" captionStart="FIGURE 29" captionStartId="40.[151,250,1848,1872]" captionTargetBox="[151,1435,181,1824]" captionTargetId="figure-20@40.[151,1436,181,1824]" captionTargetPageId="40" captionText="FIGURE 29. Lagynochthonius spinulentus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251695" httpUri="https://zenodo.org/record/7251695/files/figure.png" pageId="38" pageNumber="39">Figs 29B</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFE8A90DFE7AF994FDDAF953" box="[508,561,1662,1688]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="38" pageNumber="39">30A</figureCitation>
|
||
): carapace 0.96–1.03 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 13–14 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
|
||
<figureCitation id="9AF62A36FFE8A90DFD7BF874FC8FF873" box="[765,868,1950,1976]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="38" pageNumber="39">Fig. 30C</figureCitation>
|
||
); intercoxal tubercle absent; without sub-oral seta.
|
||
</paragraph>
|
||
<paragraph id="027236B3FFE8A90CFF41F828FD87FE42" blockId="38.[151,1436,1662,2013]" lastBlockId="39.[151,1437,151,1977]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
|
||
Chelicera (
|
||
<figureCitation id="9AF62A36FFE8A90DFEC4F828FE47F817" box="[322,428,1986,2012]" captionStart="FIGURE 29" captionStartId="40.[151,250,1848,1872]" captionTargetBox="[151,1435,181,1824]" captionTargetId="figure-20@40.[151,1436,181,1824]" captionTargetPageId="40" captionText="FIGURE 29. Lagynochthonius spinulentus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251695" httpUri="https://zenodo.org/record/7251695/files/figure.png" pageId="38" pageNumber="39">Figs 29C</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFE8A90DFE3FF828FE06F817" box="[441,493,1986,2012]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="38" pageNumber="39">30B</figureCitation>
|
||
): large, about as long as carapace, 2.44–2.47 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 21–23 teeth, distal one largest; movable finger with 14–15 retrorse contiguous small teeth; galea completely vestigial (
|
||
<figureCitation id="9AF62A36FFE9A90CFBF7FEE9FB33FED6" box="[1137,1240,259,285]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="39" pageNumber="40">Fig. 30B</figureCitation>
|
||
). Serrula exterior with 24–25 and serrula interior with 14–15 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
|
||
<figureCitation id="9AF62A36FFE9A90CFE70FE85FDB4FE42" box="[502,607,367,393]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="39" pageNumber="40">Fig. 30D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="027236B3FFE9A90CFF41FE79FD32FBFE" blockId="39.[151,1437,151,1977]" pageId="39" pageNumber="40">
|
||
Pedipalp (
|
||
<figureCitation id="9AF62A36FFE9A90CFEB1FE79FE49FE66" box="[311,418,403,429]" captionStart="FIGURE 29" captionStartId="40.[151,250,1848,1872]" captionTargetBox="[151,1435,181,1824]" captionTargetId="figure-20@40.[151,1436,181,1824]" captionTargetPageId="40" captionText="FIGURE 29. Lagynochthonius spinulentus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251695" httpUri="https://zenodo.org/record/7251695/files/figure.png" pageId="39" pageNumber="40">Figs 29A</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFE9A90CFE2BFE79FDEAFE66" box="[429,513,403,429]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="39" pageNumber="40">30E–G</figureCitation>
|
||
): long and slender, trochanter 1.70–1.94, femur 8.39–8.81, patella 2.57–2.84, chela 8.00–8.28, hand 3.08–3.19 times longer than broad; femur 2.61–2.80 times longer than patella; movable chelal finger 1.45–1.61 times longer than hand and 0.58–0.60 times longer than chela. Setae generally long and acuminate; 2 distal lyrifissures present on patella (
|
||
<figureCitation id="9AF62A36FFE9A90CFDE7FE15FD21FDD2" box="[609,714,511,537]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="39" pageNumber="40">Fig. 30E</figureCitation>
|
||
). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
|
||
<emphasis id="30B9EAA1FFE9A90CFAA8FDADFAAFFDAA" box="[1326,1348,583,609]" italics="true" pageId="39" pageNumber="40">ib</emphasis>
|
||
and
|
||
<emphasis id="30B9EAA1FFE9A90CFAFAFDADFA76FDAA" box="[1404,1437,583,609]" italics="true" pageId="39" pageNumber="40">isb</emphasis>
|
||
situated close together, submedially on dorsum of chelal hand;
|
||
<emphasis id="30B9EAA1FFE9A90CFCE9FD81FC62FD4E" box="[879,905,619,645]" italics="true" pageId="39" pageNumber="40">eb</emphasis>
|
||
,
|
||
<emphasis id="30B9EAA1FFE9A90CFC1CFD81FC54FD4E" box="[922,959,619,645]" italics="true" pageId="39" pageNumber="40">esb</emphasis>
|
||
and
|
||
<emphasis id="30B9EAA1FFE9A90CFC7DFD81FBFEFD4F" box="[1019,1045,619,644]" italics="true" pageId="39" pageNumber="40">ist</emphasis>
|
||
forming a straight oblique row at base of fixed chelal finger;
|
||
<emphasis id="30B9EAA1FFE9A90CFE4BFD65FE36FD63" box="[461,477,655,680]" italics="true" pageId="39" pageNumber="40">it</emphasis>
|
||
slightly distal to
|
||
<emphasis id="30B9EAA1FFE9A90CFD24FD65FD2AFD63" box="[674,705,655,680]" italics="true" pageId="39" pageNumber="40">est</emphasis>
|
||
, situated subdistally;
|
||
<emphasis id="30B9EAA1FFE9A90CFC31FD65FC20FD63" box="[951,971,655,680]" italics="true" pageId="39" pageNumber="40">et</emphasis>
|
||
slightly near to tip of fixed chelal finger, very close to chelal teeth;
|
||
<emphasis id="30B9EAA1FFE9A90CFE46FD59FE31FD06" box="[448,474,691,717]" italics="true" pageId="39" pageNumber="40">dx</emphasis>
|
||
situated distal to
|
||
<emphasis id="30B9EAA1FFE9A90CFD22FD59FD53FD07" box="[676,696,691,716]" italics="true" pageId="39" pageNumber="40">et</emphasis>
|
||
;
|
||
<emphasis id="30B9EAA1FFE9A90CFD4EFD59FD0AFD06" box="[712,737,691,717]" italics="true" pageId="39" pageNumber="40">sb</emphasis>
|
||
closer to
|
||
<emphasis id="30B9EAA1FFE9A90CFCD7FD59FC8FFD07" box="[849,868,691,716]" italics="true" pageId="39" pageNumber="40">st</emphasis>
|
||
than to
|
||
<emphasis id="30B9EAA1FFE9A90CFC47FD59FC24FD06" box="[961,975,691,717]" italics="true" pageId="39" pageNumber="40">b</emphasis>
|
||
;
|
||
<emphasis id="30B9EAA1FFE9A90CFC58FD59FC07FD06" box="[990,1004,691,717]" italics="true" pageId="39" pageNumber="40">b</emphasis>
|
||
and
|
||
<emphasis id="30B9EAA1FFE9A90CFBA3FD59FBC6FD07" box="[1061,1069,691,716]" italics="true" pageId="39" pageNumber="40">t</emphasis>
|
||
situated subdistally,
|
||
<emphasis id="30B9EAA1FFE9A90CFA9FFD59FACAFD07" box="[1305,1313,691,716]" italics="true" pageId="39" pageNumber="40">t</emphasis>
|
||
situated at same level as
|
||
<emphasis id="30B9EAA1FFE9A90CFEB4FD3DFEBAFD3B" box="[306,337,727,752]" italics="true" pageId="39" pageNumber="40">est</emphasis>
|
||
and distal to
|
||
<emphasis id="30B9EAA1FFE9A90CFE60FD3DFE1FFD3A" box="[486,500,727,753]" italics="true" pageId="39" pageNumber="40">b</emphasis>
|
||
(
|
||
<figureCitation id="9AF62A36FFE9A90CFD84FD3DFD8DFD3A" box="[514,614,727,753]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="39" pageNumber="40">Fig. 30F</figureCitation>
|
||
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
|
||
<emphasis id="30B9EAA1FFE9A90CFF7CFD11FEFFFCDF" box="[250,276,763,788]" italics="true" pageId="39" pageNumber="40">ist</emphasis>
|
||
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 30–32 macrodenticles, slightly retrorse and pointed, plus 12–
|
||
<specimenCount id="14CBFD3AFFE9A90CFBA5FCA9FB55FC96" box="[1059,1214,835,861]" pageId="39" pageNumber="40" type="intercalary">13 intercalary</specimenCount>
|
||
microdenticles and a modified accessory tooth on dorso-antiaxial face (
|
||
<emphasis id="30B9EAA1FFE9A90CFD65FC8DFD12FC4A" box="[739,761,871,897]" italics="true" pageId="39" pageNumber="40">td</emphasis>
|
||
, slightly distal to
|
||
<emphasis id="30B9EAA1FFE9A90CFC40FC8DFC0BFC4A" box="[966,992,871,897]" italics="true" pageId="39" pageNumber="40">dx</emphasis>
|
||
),
|
||
<quantity id="C5359B56FFE9A90CFC7FFC8DFBB6FC4A" box="[1017,1117,871,897]" metricMagnitude="0" metricUnit="m" metricValue="1.1303" metricValueMax="1.1684" metricValueMin="1.0922" pageId="39" pageNumber="40" unit="in" value="44.5" valueMax="46.0" valueMin="43.0">43–46 in</quantity>
|
||
total; movable chelal finger with 28–30 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, without intercalary teeth but with a tubercle between fifth and sixth teeth (
|
||
<figureCitation id="9AF62A36FFE9A90CFC05FC45FC0CFC02" box="[899,999,943,969]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="39" pageNumber="40">Fig. 30F</figureCitation>
|
||
). Fixed chelal finger with sensilla
|
||
<emphasis id="30B9EAA1FFE9A90CFAE8FC45FA6FFC02" box="[1390,1412,943,969]" italics="true" pageId="39" pageNumber="40">af</emphasis>
|
||
<subScript id="9E4934F6FFE9A90CFA02FC2BFA77FC04" attach="left" box="[1412,1436,961,975]" fontSize="6" pageId="39" pageNumber="40">1–2</subScript>
|
||
close together, near tip; movable chelal finger with 4 sensilla:
|
||
<emphasis id="30B9EAA1FFE9A90CFCD9FC39FC6AFC27" box="[863,897,979,1004]" italics="true" pageId="39" pageNumber="40">am</emphasis>
|
||
<subScript id="9E4934F6FFE9A90CFC04FC0FFC71FC38" attach="left" box="[898,922,997,1011]" fontSize="6" pageId="39" pageNumber="40">1–2</subScript>
|
||
near tip,
|
||
<emphasis id="30B9EAA1FFE9A90CFB8FFC39FBFCFC27" box="[1033,1047,979,1004]" italics="true" pageId="39" pageNumber="40">p</emphasis>
|
||
<subScript id="9E4934F6FFE9A90CFB91FC0FFBF4FC38" attach="left" box="[1047,1055,997,1011]" fontSize="6" pageId="39" pageNumber="40">1</subScript>
|
||
slightly proximad of
|
||
<emphasis id="30B9EAA1FFE9A90CFA9EFC39FADAFC26" box="[1304,1329,979,1005]" italics="true" pageId="39" pageNumber="40">sb</emphasis>
|
||
and very close to chelal teeth,
|
||
<emphasis id="30B9EAA1FFE9A90CFE0CFC1DFE73FBDB" box="[394,408,1015,1040]" italics="true" pageId="39" pageNumber="40">p</emphasis>
|
||
<subScript id="9E4934F6FFE9A90CFE1EFBE3FE4BFBDC" attach="left" box="[408,416,1033,1047]" fontSize="6" pageId="39" pageNumber="40">2</subScript>
|
||
distad of
|
||
<emphasis id="30B9EAA1FFE9A90CFD92FC1DFDC6FBDA" box="[532,557,1015,1041]" italics="true" pageId="39" pageNumber="40">sb</emphasis>
|
||
(
|
||
<figureCitation id="9AF62A36FFE9A90CFDB8FC1DFD4FFBDA" box="[574,676,1015,1041]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="39" pageNumber="40">Fig. 30F</figureCitation>
|
||
). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
|
||
<figureCitation id="9AF62A36FFE9A90CFDE5FBF1FD27FBFE" box="[611,716,1051,1077]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="39" pageNumber="40">Fig. 30G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="027236B3FFE9A90CFF41FBD5FADCFB0E" blockId="39.[151,1437,151,1977]" pageId="39" pageNumber="40">
|
||
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 2: 4: 4: 4: 5: 5: 5: 4: T2T: 0, tergites VII–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 13–15: 11–12: 8–9: 8: 9: 9: 9: 7: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 11–13 marginal setae on each side,
|
||
<quantity id="C5359B56FFE9A90CFB91FB40FB91FB0E" box="[1047,1146,1194,1221]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" metricValueMax="8.89" metricValueMin="8.382" pageId="39" pageNumber="40" unit="in" value="34.0" valueMax="35.0" valueMin="33.0">33–35 in</quantity>
|
||
total (
|
||
<figureCitation id="9AF62A36FFE9A90CFB44FB41FAC0FB0E" box="[1218,1323,1195,1221]" captionStart="FIGURE 29" captionStartId="40.[151,250,1848,1872]" captionTargetBox="[151,1435,181,1824]" captionTargetId="figure-20@40.[151,1436,181,1824]" captionTargetPageId="40" captionText="FIGURE 29. Lagynochthonius spinulentus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251695" httpUri="https://zenodo.org/record/7251695/files/figure.png" pageId="39" pageNumber="40">Fig. 29D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="027236B3FFE9A90CFF41FB25FADCFA0A" blockId="39.[151,1437,151,1977]" pageId="39" pageNumber="40">
|
||
Legs (
|
||
<figureCitation id="9AF62A36FFE9A90CFE8CFB25FE68FB22" box="[266,387,1231,1257]" captionStart="FIGURE 30" captionStartId="41.[151,250,1868,1892]" captionTargetBox="[182,1405,181,1844]" captionTargetId="figure-19@41.[182,1405,181,1844]" captionTargetPageId="41" captionText="FIGURE 30. Lagynochthonius spinulentus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="39" pageNumber="40">Fig.30H, I</figureCitation>
|
||
): generally typical, long and slender.Fine granulation present on anterodorsal faces of trochanter IV, femur I, IV and patella I, IV. Femur of leg I 1.90–1.98 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.18–2.21 times longer than tibia. Femoropatella of leg IV 4.11–4.17 times longer than deep; tibia 6.92–8.10 times longer than deep; with basal tactile setae on both tarsal segments: basitarsus 3.89–4.13 times longer than deep (TS= 0.27–0.29), telotarsus 15.17–16.33 times longer than deep and 2.76–2.80 times longer than basitarsus (TS= 0.25–0.29). Setae of leg I (trochanter to tibia) 2–3: 12–14: 10–11: 14–17, setae of leg IV (trochanter to basitarsus) 2–3: 3: 4–5: 10–13: 7–8. Arolium slightly shorter than the claws, not divided; claws simple.
|
||
</paragraph>
|
||
<paragraph id="027236B3FFE9A90CFF41FA21FB62F9BE" blockId="39.[151,1437,151,1977]" pageId="39" pageNumber="40">
|
||
<materialsCitation id="B2A53CEEFFE9A90CFF41FA21FB6DF9BE" ID-GBIF-Occurrence="3946861302" collectionCode="IV" pageId="39" pageNumber="40" specimenCode="III 5-6, IV 5" specimenCount="1" typeStatus="paratype">
|
||
Adult females (
|
||
<typeStatus id="DD768811FFE9A90CFEF2FA21FE08FA2E" box="[372,483,1483,1509]" pageId="39" pageNumber="40" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<figureCitation id="9AF62A36FFE9A90CFE7FFA21FD8FFA2E" box="[505,612,1483,1509]" captionStart="FIGURE 28" captionStartId="38.[151,250,1556,1580]" captionTargetBox="[193,1436,811,1518]" captionTargetId="figure-49@38.[151,1437,800,1533]" captionTargetPageId="38" captionText="FIGURE 28. Lagynochthonius spinulentus sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251691" httpUri="https://zenodo.org/record/7251691/files/figure.png" pageId="39" pageNumber="40">Figs 28B</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFE9A90CFDE9FA21FD4AFA2E" box="[623,673,1483,1509]" captionStart="FIGURE 29" captionStartId="40.[151,250,1848,1872]" captionTargetBox="[151,1435,181,1824]" captionTargetId="figure-20@40.[151,1436,181,1824]" captionTargetPageId="40" captionText="FIGURE 29. Lagynochthonius spinulentus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251695" httpUri="https://zenodo.org/record/7251695/files/figure.png" pageId="39" pageNumber="40">29E</figureCitation>
|
||
). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4,
|
||
<specimenCode id="526B9EC8FFE9A90CFF7FFA05FEACF9C2" box="[249,327,1518,1545]" collectionCode="III" pageId="39" pageNumber="40">III 5–6</specimenCode>
|
||
,
|
||
<specimenCode id="526B9EC8FFE9A90CFED2FA05FE63F9C2" box="[340,392,1518,1545]" collectionCode="IV" pageId="39" pageNumber="40">IV 5</specimenCode>
|
||
; with same tergal chaetotaxy as males; sternal chaetotaxy
|
||
<collectionCode id="64DCAE76FFE9A90CFB91FA05FBDEF9C2" box="[1047,1077,1519,1545]" pageId="39" pageNumber="40">IV</collectionCode>
|
||
–XII: 10–12: 8: 8–9: 8–10: 9: 9: 7: 0: 2; anterior genital operculum with 10 setae, posterior margin with 10–13 marginal setae,
|
||
<quantity id="C5359B56FFE9A90CFB2BF9F9FAFBF9E6" box="[1197,1296,1555,1581]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.842" metricValueMin="5.08" pageId="39" pageNumber="40" unit="in" value="21.5" valueMax="23.0" valueMin="20.0">20–23 in</quantity>
|
||
total; leg
|
||
<collectionCode id="64DCAE76FFE9A90CFAF9F9F9FA77F9E6" box="[1407,1436,1555,1581]" pageId="39" pageNumber="40">IV</collectionCode>
|
||
with a long tactile seta on both tarsal segments: basitarsus 3.40–3.89 times longer than deep (TS= 0.26), telotarsus 13.43 times longer than deep and 2.69–2.76 times longer than basitarsus (TS= 0.28–0.33)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="027236B3FFE9A90CFF41F995FE45F872" blockId="39.[151,1437,151,1977]" pageId="39" pageNumber="40">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.98–2.05. Pedipalps: trochanter 0.33–0.34/0.17–0.20 (1.70–1.94), femur 1.41–1.51/0.16–0.18 (8.39–8.81), patella 0.54/0.19–0.21 (2.57–2.84), chela 2.07–2.16/0.25–0.27 (8.00–8.28), hand 0.77–0.86/0.25–0.27 (3.08– 3.19), movable chelal finger length 1.24–1.25. Chelicera 0.79–0.83/0.32–0.34 (2.44–2.47), movable finger length 0.42–0.43. Carapace 0.65–0.70/0.68 (0.96–1.03). Leg I: trochanter 0.21–0.25/0.16–0.17 (1.31–1.47), femur 0.80– 0.83/0.09–0.10 (8.30–8.89), patella 0.42/0.08–0.09 (4.67–5.25), tibia 0.38–0.39/0.07 (5.43–5.57), tarsus 0.83– 0.86/0.06–0.07 (12.29–13.83). Leg IV: trochanter 0.25–0.27/0.18 (1.39–1.50), femoropatella 1.15–1.21/0.28–0.29 (4.11–4.17), tibia 0.81–0.83/0.10–0.12 (6.92–8.10), basitarsus 0.33–0.35/0.08–0.09 (3.89–4.13), telotarsus 0.91– 0.98/0.06 (15.17–16.33).</paragraph>
|
||
<caption id="56B2663BFFE6A903FF11F8D2FF33F853" ID-DOI="http://doi.org/10.5281/zenodo.7251695" ID-Zenodo-Dep="7251695" httpUri="https://zenodo.org/record/7251695/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,250,1848,1872]" targetBox="[151,1435,181,1824]" targetPageId="40">
|
||
<paragraph id="027236B3FFE6A903FF11F8D2FF33F853" blockId="40.[151,1436,1848,1944]" pageId="40" pageNumber="41">
|
||
<emphasis id="30B9EAA1FFE6A903FF11F8D2FECAF89B" bold="true" box="[151,289,1848,1872]" pageId="40" pageNumber="41">FIGURE 29.</emphasis>
|
||
<taxonomicName id="C5CD4D30FFE6A903FEAFF8D2FDB9F89B" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[297,594,1848,1872]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="spinulentus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFE6A903FEAFF8D2FDB9F89B" box="[297,594,1848,1872]" italics="true" pageId="40" pageNumber="41">Lagynochthonius spinulentus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFE6A903FDDCF8D2FD47F89B" bold="true" box="[602,684,1848,1872]" pageId="40" pageNumber="41">
|
||
<taxonomicNameLabel id="2B8A57DAFFE6A903FDDCF8D2FD47F89B" box="[602,684,1848,1872]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="56B2663BFFE7A902FF11F8A6FA68F83E" ID-DOI="http://doi.org/10.5281/zenodo.7251699" ID-Zenodo-Dep="7251699" httpUri="https://zenodo.org/record/7251699/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,1868,1892]" targetBox="[182,1405,181,1844]" targetPageId="41">
|
||
<paragraph id="027236B3FFE7A902FF11F8A6FA68F83E" blockId="41.[151,1436,1868,2037]" pageId="41" pageNumber="42">
|
||
<emphasis id="30B9EAA1FFE7A902FF11F8A6FEF4F8AF" bold="true" box="[151,287,1868,1892]" pageId="41" pageNumber="42">FIGURE 30.</emphasis>
|
||
<taxonomicName id="C5CD4D30FFE7A902FEA3F8A6FDA7F8AF" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[293,588,1868,1892]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="spinulentus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFE7A902FEA3F8A6FDA7F8AF" box="[293,588,1868,1892]" italics="true" pageId="41" pageNumber="42">Lagynochthonius spinulentus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFE7A902FDD4F8A7FD49F8AE" bold="true" box="[594,674,1869,1893]" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="2B8A57DAFFE7A902FDD4F8A7FD49F8AE" box="[594,674,1869,1893]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="027236B3FFE4A901FF41FF7DFD8FFE66" blockId="42.[151,1437,150,1077]" pageId="42" pageNumber="43">Females: body length 2.33–2.75. Pedipalps: trochanter 0.39–0.40/0.20–0.21 (1.90–1.95), femur 1.47– 1.48/0.19–0.20 (7.40–7.74), patella 0.55–0.56/0.21–0.23 (2.39–2.67), chela 2.06–2.19/0.26–0.28 (7.34–8.42), hand 0.83–0.88/0.26–0.28 (2.96–3.38), movable chelal finger length 1.20–1.26. Chelicera 0.87–0.88/0.35–0.36 (2.42–2.51), movable finger length 0.46–0.48. Carapace 0.71–0.72/0.73–0.74 (0.96–0.99). Leg I: trochanter 0.22– 0.23/0.14–0.15 (1.47–1.64), femur 0.72–0.77/0.09–0.10 (7.70–8.00), patella 0.41–0.42/0.08 (5.13–5.25), tibia 0.38–0.40/0.08 (4.75–5.00), tarsus 0.83–0.89/0.06 (13.83–14.83). Leg IV: trochanter 0.31–0.32/0.19–0.20 (1.60– 1.63), femoropatella 1.16–1.20/0.27 (4.30–4.44), tibia 0.79/0.10–0.11 (7.18–7.90), basitarsus 0.34–0.35/0.09–0.10 (3.40–3.89), telotarsus 0.94/0.07 (13.43).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AD76538FFE4A901FF41FE5DFB4DFCF2" pageId="42" pageNumber="43" type="discussion">
|
||
<paragraph id="027236B3FFE4A901FF41FE5DFB14FDAA" blockId="42.[151,1437,150,1077]" pageId="42" pageNumber="43">
|
||
<emphasis id="30B9EAA1FFE4A901FF41FE5DFED0FE1A" bold="true" box="[199,315,439,465]" pageId="42" pageNumber="43">Remarks.</emphasis>
|
||
<taxonomicName id="C5CD4D30FFE4A901FEC7FE5DFD63FE1A" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[321,648,439,465]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="spinulentus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFE4A901FEC7FE5DFD63FE1A" box="[321,648,439,465]" italics="true" pageId="42" pageNumber="43">Lagynochthonius spinulentus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFE4A901FD0BFE5DFD0DFE1A" bold="true" box="[653,742,439,465]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="2B8A57DAFFE4A901FD0BFE5DFD0DFE1A" box="[653,742,439,465]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="C5CD4D30FFE4A901FCF5FE5DFC1EFE1B" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[883,1013,439,464]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFE4A901FCF5FE5DFC1EFE1B" box="[883,1013,439,464]" italics="true" pageId="42" pageNumber="43">L. retrorsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFE4A901FC7CFE5DFBB8FE1A" bold="true" box="[1018,1107,439,465]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="2B8A57DAFFE4A901FC7CFE5DFBB8FE1A" box="[1018,1107,439,465]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having intercalary teeth on the fixed chelal finger and a pointed, triangular epistome. It differs by the longer chela (e.g. chela length 2.07–2.16
|
||
<emphasis id="30B9EAA1FFE4A901FF11FE15FF5DFDD3" box="[151,182,511,536]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
|
||
<quantity id="C5359B56FFE4A901FF39FE14FE8DFDD2" box="[191,358,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.59" metricValueMax="1.61" metricValueMin="1.57" pageId="42" pageNumber="43" unit="mm" value="1.59" valueMax="1.61" valueMin="1.57">1.57–1.61 mm</quantity>
|
||
, movable chelal finger length 1.24–1.25
|
||
<emphasis id="30B9EAA1FFE4A901FCB9FE15FCB6FDD3" box="[831,861,511,536]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
|
||
<quantity id="C5359B56FFE4A901FCE1FE15FBE4FDD2" box="[871,1039,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="9.45" metricValueMax="9.6" metricValueMin="9.3" pageId="42" pageNumber="43" unit="mm" value="0.945" valueMax="0.96" valueMin="0.93">0.93–0.96 mm</quantity>
|
||
; all in males), the dentation of the chelal fingers (the intercalary teeth small and pointed
|
||
<emphasis id="30B9EAA1FFE4A901FC8BFDC9FCC0FDF7" box="[781,811,547,572]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
|
||
round and obtuse, the teeth on movable chelal finger pointed and well-spaced
|
||
<emphasis id="30B9EAA1FFE4A901FE28FDADFE26FDAB" box="[430,461,583,608]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
|
||
retrorse and contiguous) and the number of setae on tergites IV (4
|
||
<emphasis id="30B9EAA1FFE4A901FB3AFDADFB31FDAB" box="[1212,1242,583,608]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
|
||
2).
|
||
</paragraph>
|
||
<paragraph id="027236B3FFE4A901FF41FD81FB54FD06" blockId="42.[151,1437,150,1077]" pageId="42" pageNumber="43">
|
||
<taxonomicName id="C5CD4D30FFE4A901FF41FD81FDFAFD4E" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[199,529,619,645]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="spinulentus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFE4A901FF41FD81FDFAFD4E" box="[199,529,619,645]" italics="true" pageId="42" pageNumber="43">Lagynochthonius spinulentus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFE4A901FD9CFD81FD9DFD4E" bold="true" box="[538,630,619,645]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="2B8A57DAFFE4A901FD9CFD81FD9DFD4E" box="[538,630,619,645]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="C5CD4D30FFE4A901FC97FD81FC0AFD4E" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[785,993,619,645]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="bailongtanensis">
|
||
<emphasis id="30B9EAA1FFE4A901FC97FD81FC0AFD4E" box="[785,993,619,645]" italics="true" pageId="42" pageNumber="43">L. bailongtanensis</emphasis>
|
||
</taxonomicName>
|
||
in the shape of epistome (pointed and triangular
|
||
<emphasis id="30B9EAA1FFE4A901FE8BFD65FECFFD63" box="[269,292,655,680]" italics="true" pageId="42" pageNumber="43">vs</emphasis>
|
||
. rounded and obtuse), the pattern of dentation on the chelal fingers (with intercalary teeth on fixed chelal finger
|
||
<emphasis id="30B9EAA1FFE4A901FEADFD59FEA9FD07" box="[299,322,691,716]" italics="true" pageId="42" pageNumber="43">vs</emphasis>
|
||
. without intercalary teeth), and the number of setae on tergites I–III (2
|
||
<emphasis id="30B9EAA1FFE4A901FBD9FD59FB95FD07" box="[1119,1150,691,716]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
|
||
3–4).
|
||
</paragraph>
|
||
<paragraph id="027236B3FFE4A901FF41FD3DFB4DFCF2" blockId="42.[151,1437,150,1077]" pageId="42" pageNumber="43">
|
||
<taxonomicName id="C5CD4D30FFE4A901FF41FD3DFDFAFD3A" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[199,529,727,753]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="spinulentus" status="sp. nov.">
|
||
<emphasis id="30B9EAA1FFE4A901FF41FD3DFDFAFD3A" box="[199,529,727,753]" italics="true" pageId="42" pageNumber="43">Lagynochthonius spinulentus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="30B9EAA1FFE4A901FD9CFD3DFD9DFD3A" bold="true" box="[538,630,727,753]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="2B8A57DAFFE4A901FD9CFD3DFD9DFD3A" box="[538,630,727,753]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="C5CD4D30FFE4A901FC96FD3DFC30FD3A" authorityName="Hou & Gao & Zhang" authorityYear="2022" box="[784,987,727,753]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis id="30B9EAA1FFE4A901FC96FD3DFC30FD3A" box="[784,987,727,753]" italics="true" pageId="42" pageNumber="43">L. laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
by the presence of intercalary teeth on both chelal fingers, the number of setae on tergites III (2
|
||
<emphasis id="30B9EAA1FFE4A901FCA4FD11FCABFCDF" box="[802,832,763,788]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
|
||
4), and a larger body (e.g. body length 1.98–2.05
|
||
<emphasis id="30B9EAA1FFE4A901FAF8FD11FA77FCDF" box="[1406,1436,763,788]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
|
||
<quantity id="C5359B56FFE4A901FF11FCF5FF16FCF2" box="[151,253,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="42" pageNumber="43" unit="mm" value="1.78">1.78 mm</quantity>
|
||
, chela length 2.07–2.16
|
||
<emphasis id="30B9EAA1FFE4A901FD89FCF5FDC6FCF3" box="[527,557,799,824]" italics="true" pageId="42" pageNumber="43">vs.</emphasis>
|
||
<quantity id="C5359B56FFE4A901FDB2FCF5FD70FCF2" box="[564,667,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="42" pageNumber="43" unit="mm" value="1.62">1.62 mm</quantity>
|
||
; all in males) (
|
||
<bibRefCitation id="665C4B42FFE4A901FCC6FCF5FC30FCF2" author="Li, Y. C. & Liu, H. & Shi, A. M." box="[832,987,799,825]" pageId="42" pageNumber="43" pagination="28 - 34" refId="ref39131" refString="Li, Y. C., Liu, H. & Shi, A. M. (2019) A new cave-dwelling species of Lagynochthonius (Arachnida: Pseudoscorpiones: Chthoniidae) from Yunnan Province, China. Zootaxa, 4571 (1), 28 - 34. https: // doi. org / 10.11646 / zootaxa. 4571.1.2" type="journal article" year="2019">
|
||
Li
|
||
<emphasis id="30B9EAA1FFE4A901FCE7FCF5FC79FCF2" box="[865,914,799,825]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="665C4B42FFE4A901FC61FCF5FB71FCF2" author="Hou, Y. M. & Gao, Z. Z. & Zhang, F." box="[999,1178,799,825]" pageId="42" pageNumber="43" pagination="65 - 83" refId="ref38684" refString="Hou, Y. M., Gao, Z. Z. & Zhang, F. (2022) Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China. ZooKeys, 1097, 65 - 83. https: // doi. org / 10.3897 / zookeys. 1097.82527" type="journal article" year="2022">
|
||
Hou
|
||
<emphasis id="30B9EAA1FFE4A901FB98FCF5FBA4FCF2" box="[1054,1103,799,825]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AD76538FFE4A901FF41FCA8FD8AFBFE" pageId="42" pageNumber="43" type="distribution">
|
||
<paragraph id="027236B3FFE4A901FF41FCA8FD8AFBFE" blockId="42.[151,1437,150,1077]" pageId="42" pageNumber="43">
|
||
<emphasis id="30B9EAA1FFE4A901FF41FCA8FE18FC97" bold="true" box="[199,499,834,861]" pageId="42" pageNumber="43">Distribution and habitat.</emphasis>
|
||
This species is only known from the
|
||
<typeStatus id="DD768811FFE4A901FC26FCA9FC3BFC96" box="[928,976,835,861]" pageId="42" pageNumber="43">type</typeStatus>
|
||
locality, Yingshan Cave (
|
||
<figureCitation id="9AF62A36FFE4A901FB72FCA9FAA5FC96" box="[1268,1358,835,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1542,1566]" captionTargetBox="[152,1433,567,1514]" captionTargetId="figure-169@3.[151,1436,561,1519]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of the type locality of troglomorphic Lagynochthonius species in Yunnan (China). A. L. retrorsus sp. nov. (Qinglong Cave); B. L. fengi sp. nov. (Unnamed Cave 2); C. L. xiaolinensis sp. nov. (Dayan Cave); D. L. xibaiensis sp. nov. (Hama Cave); E. L. spinulentus sp. nov. (Yingshan Cave); F. L. xinjiaoensis sp. nov. (Yueliang Cave); G. L. magnidentatus sp. nov. (Xiao Cave); H. L. bailongtanensis sp. nov. (Qianfo Cave); I. L. laoxueyanensis sp. nov. (Laoxueyan Cave); J. L. serratus sp. nov. (Unnamed Cave 3); K. L. yaowangguensis sp. nov. (Unnamed Cave 4); L. L. crassus sp. nov. (Unnamed Cave 1); M–N. L. minimus sp. nov. (M: Yanfen Cave, N: Cizhu Cave)." figureDoi="http://doi.org/10.5281/zenodo.7251604" httpUri="https://zenodo.org/record/7251604/files/figure.png" pageId="42" pageNumber="43">Figs 1E</figureCitation>
|
||
,
|
||
<figureCitation id="9AF62A36FFE4A901FADAFCA9FA64FC96" box="[1372,1423,835,861]" captionStart="FIGURE 27" captionStartId="38.[151,250,715,739]" captionTargetBox="[151,1437,181,691]" captionTargetId="figure-20@38.[151,1437,181,691]" captionTargetPageId="38" captionText="FIGURE 27. Yingshan Cave, type locality of Lagynochthonius spinulentus sp. nov., A. Entrance; B. Live male of L. spinulentus sp. nov. in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251687" httpUri="https://zenodo.org/record/7251687/files/figure.png" pageId="42" pageNumber="43">27A</figureCitation>
|
||
), which is located about
|
||
<quantity id="C5359B56FFE4A901FE1BFC8DFE3DFC4A" box="[413,470,871,897]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="km" value="2.0">2 km</quantity>
|
||
northwest of Qixin Village (Zhenxiong County) and near the top of a mountain. This limestone cave has one small and square entrance (about
|
||
<quantity id="C5359B56FFE4A901FC97FC61FCD7FC6E" box="[785,828,907,933]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="m" value="1.0">1 m</quantity>
|
||
high and
|
||
<quantity id="C5359B56FFE4A901FC2EFC61FC38FC6E" box="[936,979,907,933]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="m" value="1.0">1 m</quantity>
|
||
wide), but a huge and downward sloping interior space. The ground is covered with silt. About
|
||
<quantity id="C5359B56FFE4A901FD7CFC45FCAAFC02" box="[762,833,943,969]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="42" pageNumber="43" unit="m" value="300.0">300 m</quantity>
|
||
of the cave have beenexplored but the total length of the cave is unknown. All of the specimens were collected under stones within
|
||
<quantity id="C5359B56FFE4A901FB80FC39FB65FC26" box="[1030,1166,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="1.1" metricValueMax="2.0" metricValueMin="0.2" pageId="42" pageNumber="43" unit="m" value="110.0" valueMax="200.0" valueMin="20.0">20 to 200 m</quantity>
|
||
from the cave entrance. The cool, high-humidity and dark interior environment (temperature around 14°C on average, humidity over 90%) is suitable for the survival of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |