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<document id="E563536266FFE08B81ECD3E4F90C9CD4" ID-DOI="10.11646/zootaxa.4786.2.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3874789" ID-ZooBank="B2D53D68-E116-42E1-B2E8-3C78B0AD1473" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591189312045" checkinUser="plazi" docAuthor="Gupta, Devanshu, Chandra, Kailash, Ghosh, Joyjit &amp; Das, Priyanka" docDate="2020" docId="86417A763B45FF9FFF1FFF2CE176F08B" docLanguage="en" docName="zootaxa.4786.2.9.pdf" docOrigin="Zootaxa 4786 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bolboceras arunachalensis Chandra, Gupta &amp; Ghosh 2020, new species" docType="treatment" docVersion="3" lastPageNumber="279" masterDocId="7A78020E3B47FF9DFF88FFB5E438F668" masterDocTitle="A new species and new records of the genus Bolboceras Kirby, 1819 (Coleoptera Scarabaeoidea: Geotrupidae: Bolboceratinae) from India" masterLastPageNumber="282" masterPageNumber="277" pageNumber="279" updateTime="1700276380027" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F16435D1E12C2DBA982B3FFB4C2A412C">A new species and new records of the genus Bolboceras Kirby, 1819 (Coleoptera Scarabaeoidea: Geotrupidae: Bolboceratinae) from India</mods:title>
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<mods:name id="E272FAC1F1EDE58F59C22D3B62F6EB41" type="personal">
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<mods:namePart id="AE7EDE5269C7415200A602F4610B3627">Gupta, Devanshu</mods:namePart>
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<mods:namePart id="27C47FC2738C1EA11618EB67E66367CE">Chandra, Kailash</mods:namePart>
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<mods:namePart id="71B647D8F1C26090993007C54C45FFCA">Ghosh, Joyjit</mods:namePart>
</mods:name>
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<mods:namePart id="DA8606590F3C1F6E06A7517DEEF60703">Das, Priyanka</mods:namePart>
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<mods:title id="4163D399DFB05041D924F99E43E62817">Zootaxa</mods:title>
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<mods:part id="1CD5E555CF135CDC900B7A0E84868655">
<mods:date id="8DFF678CA5CA1E8EE4F6991AA8AC9217">2020</mods:date>
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<mods:number id="DAC91FFEC616351B082DAE7F0BA892CA">2020-06-02</mods:number>
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<mods:number id="A8DC3D23B9F27B0A7531A010CFD0094F">4786</mods:number>
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<mods:classification id="B638EC2C45F0BD456EB6DAECBC8AD987">journal article</mods:classification>
<mods:identifier id="28E5696694D636F7DB7CD28F170F16C8" type="DOI">10.11646/zootaxa.4786.2.9</mods:identifier>
<mods:identifier id="A12965D8B65FAD795062B14B7D249789" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E8394EAAB0BB46CBD1C53225A6557BDA" type="ZooBank">B2D53D68-E116-42E1-B2E8-3C78B0AD1473</mods:identifier>
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<treatment id="86417A763B45FF9FFF1FFF2CE176F08B" LSID="urn:lsid:plazi:treatment:86417A763B45FF9FFF1FFF2CE176F08B" httpUri="http://treatment.plazi.org/id/86417A763B45FF9FFF1FFF2CE176F08B" lastPageNumber="279" pageId="2" pageNumber="279">
<subSubSection id="46F298EB3B45FF9FFF1FFF2CE7F5F6DB" box="[151,973,153,180]" pageId="2" pageNumber="279" type="nomenclature">
<paragraph id="0E57CB603B45FF9FFF1FFF2CE7F5F6DB" blockId="2.[151,973,153,216]" box="[151,973,153,180]" pageId="2" pageNumber="279">
<heading id="551F7C0C3B45FF9FFF1FFF2CE7F5F6DB" bold="true" box="[151,973,153,180]" fontSize="11" level="1" pageId="2" pageNumber="279" reason="1">
<emphasis id="3C9C17723B45FF9FFF1FFF2CE7F5F6DB" bold="true" box="[151,973,153,180]" pageId="2" pageNumber="279">
<taxonomicName id="C9E8B0E33B45FF9FFF1FFF2CE708F6DB" authority="Chandra, Gupta &amp; Ghosh" authorityName="Chandra, Gupta &amp; Ghosh" authorityYear="2020" box="[151,816,153,180]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="3C9C17723B45FF9FFF1FFF2CE5D8F6DC" bold="true" box="[151,480,153,180]" italics="true" pageId="2" pageNumber="279">Bolboceras arunachalensis</emphasis>
Chandra, Gupta &amp; Ghosh
</taxonomicName>
,
<taxonomicNameLabel id="27AFAA093B45FF9FFCB5FF2FE7F5F6DB" box="[829,973,153,180]" pageId="2" pageNumber="279" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="46F298EB3B45FF9FFF1FFF08E52AF6B0" box="[151,274,189,216]" pageId="2" pageNumber="279" type="description">
<paragraph id="0E57CB603B45FF9FFF1FFF08E52AF6B0" blockId="2.[151,973,153,216]" box="[151,274,189,216]" pageId="2" pageNumber="279">
(
<figureCitation id="96D3D7E53B45FF9FFF29FF08E532F6B0" box="[161,266,189,216]" captionStart="FIGURES 16" captionStartId="1.[151,264,1945,1970]" captionTargetBox="[225,1375,189,1909]" captionTargetId="figure@1.[209,1379,181,1920]" captionTargetPageId="1" captionText="FIGURES 16. Bolboceras arunachalensis Chandra, Gupta &amp; Ghosh, new species (holotype). 1, habitus, dorsal view; 2, parameres, dorsal (frontal) view; 3, head and clypeus; 4, pronotum; 5, scutellum; 6, elytra." figureDoi="http://doi.org/10.5281/zenodo.3874791" httpUri="https://zenodo.org/record/3874791/files/figure.png" pageId="2" pageNumber="279">
Figs. 1
<emphasis id="3C9C17723B45FF9FFF65FF0BE4C3F6BF" box="[237,251,190,215]" italics="true" pageId="2" pageNumber="279"></emphasis>
6
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="46F298EB3B45FF9FFF1FFEB3E7F6F70F" pageId="2" pageNumber="279" type="materials_examined">
<paragraph id="0E57CB603B45FF9FFF1FFEB3E6D2F748" blockId="2.[151,1437,261,1764]" box="[151,746,261,288]" pageId="2" pageNumber="279">
<materialsCitation id="BE80C13D3B45FF9FFF1FFEB3E6DFF748" box="[151,743,261,288]" country="India" location="Papumpare" pageId="2" pageNumber="279" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<emphasis id="3C9C17723B45FF9FFF1FFEB3E50EF748" bold="true" box="[151,310,261,288]" pageId="2" pageNumber="279">
<typeStatus id="D15375C23B45FF9FFF1FFEB3E4EAF748" box="[151,210,262,288]" pageId="2" pageNumber="279">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="76FF8BF03B45FF9FFEB4FEB0E540F777" box="[316,376,261,287]" name="India" pageId="2" pageNumber="279">India</collectingCountry>
,
<collectingRegion id="CC2C05823B45FF9FFE0AFEB0E662F777" box="[386,602,261,287]" country="India" name="Arunachal Pradesh" pageId="2" pageNumber="279">Arunachal Pradesh</collectingRegion>
,
<location id="0B379DBB3B45FF9FFDEDFEB3E6DFF748" LSID="urn:lsid:plazi:treatment:86417A763B45FF9FFF1FFF2CE176F08B:0B379DBB3B45FF9FFDEDFEB3E6DFF748" box="[613,743,262,288]" country="India" name="Papumpare" pageId="2" pageNumber="279" stateProvince="Arunachal Pradesh">Papumpare</location>
</materialsCitation>
.
</paragraph>
<paragraph id="0E57CB603B45FF9FFF4FFE9FE7F6F70F" blockId="2.[151,1437,261,1764]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFE9FE61CF72C" bold="true" box="[199,548,297,324]" pageId="2" pageNumber="279">
Type material (
<specimenCount id="18EE00E93B45FF9FFE0AFE9CE62FF72C" box="[386,535,297,324]" pageId="2" pageNumber="279" type="generic">2 specimens</specimenCount>
).
</emphasis>
<materialsCitation id="BE80C13D3B45FF9FFDA5FE9CE5A8F70F" collectingDate="1996-10-23" collectionCode="NZSI" collectorName="S. K. Mondol" country="India" location="Papumpare" pageId="2" pageNumber="279" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<typeStatus id="D15375C23B45FF9FFDA5FE9CE6A0F72B" box="[557,664,297,323]" pageId="2" pageNumber="279" type="holotype">Holotype</typeStatus>
, male: “
<collectingCountry id="76FF8BF03B45FF9FFD7FFE9CE70EF72B" box="[759,822,297,323]" name="India" pageId="2" pageNumber="279">India</collectingCountry>
,
<collectingRegion id="CC2C05823B45FF9FFCCAFE9CE024F72B" box="[834,1052,297,323]" country="India" name="Arunachal Pradesh" pageId="2" pageNumber="279">Arunachal Pradesh</collectingRegion>
,
<location id="0B379DBB3B45FF9FFBA1FE9FE092F72C" LSID="urn:lsid:plazi:treatment:86417A763B45FF9FFF1FFF2CE176F08B:0B379DBB3B45FF9FFBA1FE9FE092F72C" box="[1065,1194,298,324]" country="India" name="Papumpare" pageId="2" pageNumber="279" stateProvince="Arunachal Pradesh">Papumpare</location>
,
<date id="7A56EDA03B45FF9FFB3FFE9CE112F72B" box="[1207,1322,297,323]" pageId="2" pageNumber="279" value="1996-10-23">
<collectingDate id="6A1214483B45FF9FFB3FFE9CE112F72B" box="[1207,1322,297,323]" pageId="2" pageNumber="279" value="1996-10-23">23.x.1996</collectingDate>
</date>
, collected
<collectorName id="A31DAEB63B45FF9FFF1FFEF8E514F70F" box="[151,300,333,359]" pageId="2" pageNumber="279">S.K. Mondol</collectorName>
” (
<collectionCode id="68F953A53B45FF9FFECCFEF8E5B0F70F" box="[324,392,333,359]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="279">NZSI</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="BE80C13D3B45FF9FFE13FEFBE7F2F70F" box="[411,970,333,360]" collectingDate="1996-10-23" collectionCode="NZSI" collectorName="S. K. Mondol" country="India" location="Papumpare" pageId="2" pageNumber="279" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<typeStatus id="D15375C23B45FF9FFE13FEFBE638F700" box="[411,512,334,360]" pageId="2" pageNumber="279" type="paratype">Paratype</typeStatus>
, male: same data as for holotype (
<collectionCode id="68F953A53B45FF9FFCF5FEF8E7F9F70F" box="[893,961,333,359]" country="India" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="2" pageNumber="279">NZSI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="46F298EB3B45FF9FFF4FFEC4E1B0F2CC" pageId="2" pageNumber="279" type="description">
<paragraph id="0E57CB603B45FF9FFF4FFEC4E05BF7E3" blockId="2.[151,1437,261,1764]" box="[199,1123,369,396]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFEC4E569F7E4" bold="true" box="[199,337,369,396]" pageId="2" pageNumber="279">Description</emphasis>
(
<typeStatus id="D15375C23B45FF9FFEE8FEC4E5FEF7E3" box="[352,454,369,395]" pageId="2" pageNumber="279" type="holotype">holotype</typeStatus>
, male). Shape subglobular. Dark brown, more or less shiny.
</paragraph>
<paragraph id="0E57CB603B45FF9FFF4FFE20E7C6F40B" blockId="2.[151,1437,261,1764]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFE20E53CF7C7" box="[199,260,405,431]" italics="true" pageId="2" pageNumber="279">Head</emphasis>
(
<figureCitation id="96D3D7E53B45FF9FFE9CFE20E560F7C7" box="[276,344,405,431]" captionStart="FIGURES 16" captionStartId="1.[151,264,1945,1970]" captionTargetBox="[225,1375,189,1909]" captionTargetId="figure@1.[209,1379,181,1920]" captionTargetPageId="1" captionText="FIGURES 16. Bolboceras arunachalensis Chandra, Gupta &amp; Ghosh, new species (holotype). 1, habitus, dorsal view; 2, parameres, dorsal (frontal) view; 3, head and clypeus; 4, pronotum; 5, scutellum; 6, elytra." figureDoi="http://doi.org/10.5281/zenodo.3874791" httpUri="https://zenodo.org/record/3874791/files/figure.png" pageId="2" pageNumber="279">Fig. 3</figureCitation>
). Labrum anterior margin concave-shaped, apical ridge bisinuated. Lateral ridges of clypeus convergent to a single anteromedian elevated point, with apex broadly rounded; perimarginal disc distinct and K-shaped. Clypeus closely punctured; frons smooth with few punctures; surface before interocular ridge nearly smooth; genal angle rounded. Anterior edge of eye canthus short, with a small indistinct anterolateral tubercle, thence arcuate at tip, surface rugo-punctate. Paraocular ridge fine, rounded, extended posteriorly beyond side of interocular elevation. Transverse interocular ridge long and straight, not reaching paraocular ridges.
</paragraph>
<paragraph id="0E57CB603B45FF9FFF4FFDDBE133F570" blockId="2.[151,1437,261,1764]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFDDBE50EF4EF" box="[199,310,622,647]" italics="true" pageId="2" pageNumber="279">Pronotum</emphasis>
(
<figureCitation id="96D3D7E53B45FF9FFEC0FDD8E5B6F4EF" box="[328,398,621,647]" captionStart="FIGURES 16" captionStartId="1.[151,264,1945,1970]" captionTargetBox="[225,1375,189,1909]" captionTargetId="figure@1.[209,1379,181,1920]" captionTargetPageId="1" captionText="FIGURES 16. Bolboceras arunachalensis Chandra, Gupta &amp; Ghosh, new species (holotype). 1, habitus, dorsal view; 2, parameres, dorsal (frontal) view; 3, head and clypeus; 4, pronotum; 5, scutellum; 6, elytra." figureDoi="http://doi.org/10.5281/zenodo.3874791" httpUri="https://zenodo.org/record/3874791/files/figure.png" pageId="2" pageNumber="279">Fig. 4</figureCitation>
). Anteromedially very slightly depressed (laterally), with large punctures; discoparamedian and lateral tubercles absent; pronotum elevated medially forming an inverted W-shaped carina; discomedian surface smooth, with punctured median groove, discolateral protrusion absent; basal margin of pronotum punctured; anterolateral point blunt; pronotum punctures vary from large to small, with large punctures arranged at anterior and lateral pronotal fovea and disco lateral protrusion, few punctures between lateral fovea and basomedian part.
</paragraph>
<paragraph id="0E57CB603B45FF9FFF4FFC94E6EFF553" blockId="2.[151,1437,261,1764]" box="[199,727,801,828]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFC94E50DF553" box="[199,309,801,827]" italics="true" pageId="2" pageNumber="279">Scutellum</emphasis>
(
<figureCitation id="96D3D7E53B45FF9FFECEFC94E5B1F554" box="[326,393,801,828]" captionStart="FIGURES 16" captionStartId="1.[151,264,1945,1970]" captionTargetBox="[225,1375,189,1909]" captionTargetId="figure@1.[209,1379,181,1920]" captionTargetPageId="1" captionText="FIGURES 16. Bolboceras arunachalensis Chandra, Gupta &amp; Ghosh, new species (holotype). 1, habitus, dorsal view; 2, parameres, dorsal (frontal) view; 3, head and clypeus; 4, pronotum; 5, scutellum; 6, elytra." figureDoi="http://doi.org/10.5281/zenodo.3874791" httpUri="https://zenodo.org/record/3874791/files/figure.png" pageId="2" pageNumber="279">Fig. 5</figureCitation>
). Finely punctured anteriorly.
</paragraph>
<paragraph id="0E57CB603B45FF9FFF4FFCF0E786F5EB" blockId="2.[151,1437,261,1764]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFCF0E535F537" box="[199,269,837,863]" italics="true" pageId="2" pageNumber="279">Elytra</emphasis>
(
<figureCitation id="96D3D7E53B45FF9FFE95FCF0E558F508" box="[285,352,837,864]" captionStart="FIGURES 16" captionStartId="1.[151,264,1945,1970]" captionTargetBox="[225,1375,189,1909]" captionTargetId="figure@1.[209,1379,181,1920]" captionTargetPageId="1" captionText="FIGURES 16. Bolboceras arunachalensis Chandra, Gupta &amp; Ghosh, new species (holotype). 1, habitus, dorsal view; 2, parameres, dorsal (frontal) view; 3, head and clypeus; 4, pronotum; 5, scutellum; 6, elytra." figureDoi="http://doi.org/10.5281/zenodo.3874791" httpUri="https://zenodo.org/record/3874791/files/figure.png" pageId="2" pageNumber="279">Fig. 6</figureCitation>
). Discal striae deeply impressed, striae finely punctured; stria 2 incomplete before reaching up to base; intervals convex and smooth; lateral part of elytron finely smooth.
</paragraph>
<paragraph id="0E57CB603B45FF9FFF4FFC3BE741F5A3" blockId="2.[151,1437,261,1764]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFC3BE4C4F5CF" box="[199,252,910,935]" italics="true" pageId="2" pageNumber="279">Legs</emphasis>
. Protibia hexadentate, denticles reduce in size from apex to end; terminal spur sharp and slender; apex of mesotibiae and metatibiae bilobed, their crest fringed with spines.
</paragraph>
<paragraph id="0E57CB603B45FF9FFF4FFC60E4ECF250" blockId="2.[151,1437,261,1764]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFC60E50CF587" box="[199,308,981,1007]" italics="true" pageId="2" pageNumber="279">Aedeagus</emphasis>
(
<figureCitation id="96D3D7E53B45FF9FFECFFC60E5B5F587" box="[327,397,981,1007]" captionStart="FIGURES 16" captionStartId="1.[151,264,1945,1970]" captionTargetBox="[225,1375,189,1909]" captionTargetId="figure@1.[209,1379,181,1920]" captionTargetPageId="1" captionText="FIGURES 16. Bolboceras arunachalensis Chandra, Gupta &amp; Ghosh, new species (holotype). 1, habitus, dorsal view; 2, parameres, dorsal (frontal) view; 3, head and clypeus; 4, pronotum; 5, scutellum; 6, elytra." figureDoi="http://doi.org/10.5281/zenodo.3874791" httpUri="https://zenodo.org/record/3874791/files/figure.png" pageId="2" pageNumber="279">Fig. 2</figureCitation>
). Parameres acuminate, slightly curved outwardly, tips pointed, sclerotized aedeagal stalks expanded downward, tips of stalks blunt, paramedian aedeagal struts thick, median aedeagal apparatus rounded at apex.
</paragraph>
<paragraph id="0E57CB603B45FF9FFF4FFBF7E1B0F2CC" blockId="2.[151,1437,261,1764]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFBF7E550F233" box="[199,360,1090,1115]" italics="true" pageId="2" pageNumber="279">Measurements</emphasis>
: Body length:
<quantity id="C91066853B45FF9FFD84FBF4E645F234" box="[524,637,1089,1116]" metricMagnitude="-2" metricUnit="m" metricValue="1.152" pageId="2" pageNumber="279" unit="mm" value="11.52">11.52 mm</quantity>
, width: 7.0 mm. Head, length in dorsal view excluding labrum and mandibles:
<quantity id="C91066853B45FF9FFF6FFBD0E575F2E8" box="[231,333,1125,1152]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" pageId="2" pageNumber="279" unit="mm" value="2.33">2.33 mm</quantity>
; interocular ridge:
<quantity id="C91066853B45FF9FFD94FBD0E6BAF2E8" box="[540,642,1125,1152]" metricMagnitude="-3" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="279" unit="mm" value="1.46">1.46 mm</quantity>
; ocular distance:
<quantity id="C91066853B45FF9FFCC8FBD0E79EF2E8" box="[832,934,1125,1152]" metricMagnitude="-3" metricUnit="m" metricValue="2.16" pageId="2" pageNumber="279" unit="mm" value="2.16">2.16 mm</quantity>
. Pronotum length:
<quantity id="C91066853B45FF9FFBFEFBD0E0E4F2E8" box="[1142,1244,1125,1152]" metricMagnitude="-3" metricUnit="m" metricValue="3.67" pageId="2" pageNumber="279" unit="mm" value="3.67">3.67 mm</quantity>
; width:
<quantity id="C91066853B45FF9FFABBFBD0E1A1F2E8" box="[1331,1433,1125,1152]" metricMagnitude="-3" metricUnit="m" metricValue="6.78" pageId="2" pageNumber="279" unit="mm" value="6.78">6.78 mm</quantity>
. Scutellar shield length:
<quantity id="C91066853B45FF9FFE15FB3CE63AF2CC" box="[413,514,1161,1188]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="2" pageNumber="279" unit="mm" value="1.08">1.08 mm</quantity>
, width:
<quantity id="C91066853B45FF9FFDD3FB3CE6F9F2CC" box="[603,705,1161,1188]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" pageId="2" pageNumber="279" unit="mm" value="1.07">1.07 mm</quantity>
. Elytra sutural length:
<quantity id="C91066853B45FF9FFC33FB3CE01AF2CC" box="[955,1058,1161,1188]" metricMagnitude="-3" metricUnit="m" metricValue="4.18" pageId="2" pageNumber="279" unit="mm" value="4.18">4.18 mm</quantity>
. Genital capsule width:
<quantity id="C91066853B45FF9FFAA3FB3CE1BCF2CC" box="[1323,1412,1161,1188]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="279" unit="mm" value="0.9">0.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="46F298EB3B45FF9FFF4FFB1BE62AF283" pageId="2" pageNumber="279" type="etymology">
<paragraph id="0E57CB603B45FF9FFF4FFB1BE62AF283" blockId="2.[151,1437,261,1764]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFB1BE576F2A0" bold="true" box="[199,334,1198,1224]" pageId="2" pageNumber="279">Etymology.</emphasis>
The specific name
<taxonomicName id="C9E8B0E33B45FF9FFDADFB18E6E8F2AF" authorityName="Chandra, Gupta &amp; Ghosh" authorityYear="2020" box="[549,720,1197,1223]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="arunachalensis">
<emphasis id="3C9C17723B45FF9FFDADFB18E6E8F2AF" box="[549,720,1197,1223]" italics="true" pageId="2" pageNumber="279">arunachalensis</emphasis>
</taxonomicName>
refers to the place where the new species has been collected, the state of
<collectingRegion id="CC2C05823B45FF9FFF66FB64E5FEF283" box="[238,454,1233,1259]" country="India" name="Arunachal Pradesh" pageId="2" pageNumber="279">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="76FF8BF03B45FF9FFE59FB64E636F283" box="[465,526,1233,1259]" name="India" pageId="2" pageNumber="279">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="46F298EB3B45FF9FFF4FFB40E0C5F367" box="[199,1277,1269,1296]" pageId="2" pageNumber="279" type="distribution">
<paragraph id="0E57CB603B45FF9FFF4FFB40E0C5F367" blockId="2.[151,1437,261,1764]" box="[199,1277,1269,1296]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFB40E561F378" bold="true" box="[199,345,1269,1296]" pageId="2" pageNumber="279">Distribution</emphasis>
(
<figureCitation id="96D3D7E53B45FF9FFEE2FB40E595F378" box="[362,429,1269,1296]" captionStart="FIGURE 7" captionStartId="3.[151,250,1624,1649]" captionTargetBox="[161,1428,357,1591]" captionTargetId="figure@3.[151,1436,345,1599]" captionTargetPageId="3" captionText="FIGURE 7. Map of India and Arunachal Pradesh showing type locality of Bolboceras arunachalensis Chandra, Gupta &amp; Ghosh, new species." figureDoi="http://doi.org/10.5281/zenodo.3874793" httpUri="https://zenodo.org/record/3874793/files/figure.png" pageId="2" pageNumber="279">Fig. 7</figureCitation>
). So far known only from the
<typeStatus id="D15375C23B45FF9FFC88FB43E708F378" box="[768,816,1270,1296]" pageId="2" pageNumber="279">type</typeStatus>
locality, Papumpare,
<collectingRegion id="CC2C05823B45FF9FFBA9FB40E0C1F367" box="[1057,1273,1269,1295]" country="India" name="Arunachal Pradesh" pageId="2" pageNumber="279">Arunachal Pradesh</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="46F298EB3B45FF9FFF4FFAACE176F08B" pageId="2" pageNumber="279" type="diagnosis">
<paragraph id="0E57CB603B45FF9FFF4FFAACE61AF38F" blockId="2.[151,1437,261,1764]" pageId="2" pageNumber="279">
<emphasis id="3C9C17723B45FF9FFF4FFAACE5E8F35C" bold="true" box="[199,464,1305,1332]" pageId="2" pageNumber="279">Differential diagnosis.</emphasis>
<taxonomicName id="C9E8B0E33B45FF9FFE51FAACE735F35B" authority="Gupta &amp; Chandra &amp; Ghosh &amp; Das, 2020" authorityName="Gupta &amp; Chandra &amp; Ghosh &amp; Das" authorityYear="2020" box="[473,781,1305,1331]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="3C9C17723B45FF9FFE51FAACE735F35B" box="[473,781,1305,1331]" italics="true" pageId="2" pageNumber="279">Bolboceras arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27AFAA093B45FF9FFC9CFAAFE7A5F35B" box="[788,925,1305,1332]" pageId="2" pageNumber="279" rank="species">new species</taxonomicNameLabel>
is delineated mainly by the unique structure of the aedeagus. The newly described species belongs to
<taxonomicName id="C9E8B0E33B45FF9FFC96FA8BE790F33F" authorityName="Westwood" authorityYear="1848" box="[798,936,1342,1367]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="3C9C17723B45FF9FFC96FA8BE790F33F" box="[798,936,1342,1367]" italics="true" pageId="2" pageNumber="279">B. nigricans</emphasis>
</taxonomicName>
group, because of lateral ridges of the clypeus convergent to a single anteromedian elevated point (
<figureCitation id="96D3D7E53B45FF9FFC99FAD4E76DF313" box="[785,853,1377,1403]" captionStart="FIGURES 16" captionStartId="1.[151,264,1945,1970]" captionTargetBox="[225,1375,189,1909]" captionTargetId="figure@1.[209,1379,181,1920]" captionTargetPageId="1" captionText="FIGURES 16. Bolboceras arunachalensis Chandra, Gupta &amp; Ghosh, new species (holotype). 1, habitus, dorsal view; 2, parameres, dorsal (frontal) view; 3, head and clypeus; 4, pronotum; 5, scutellum; 6, elytra." figureDoi="http://doi.org/10.5281/zenodo.3874791" httpUri="https://zenodo.org/record/3874791/files/figure.png" pageId="2" pageNumber="279">Fig. 3</figureCitation>
). Within the
<taxonomicName id="C9E8B0E33B45FF9FFC60FAD7E049F313" authorityName="Westwood" authorityYear="1848" box="[1000,1137,1378,1403]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="3C9C17723B45FF9FFC60FAD7E049F313" box="[1000,1137,1378,1403]" italics="true" pageId="2" pageNumber="279">B. nigricans</emphasis>
</taxonomicName>
group, new species shares K-shaped perimarginal ridges of male clypeus with
<taxonomicName id="C9E8B0E33B45FF9FFD72FA33E7BFF3F7" authorityName="Westwood" authorityYear="1848" box="[762,903,1414,1439]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="3C9C17723B45FF9FFD72FA33E7BFF3F7" box="[762,903,1414,1439]" italics="true" pageId="2" pageNumber="279">B. nigricans</emphasis>
</taxonomicName>
,
<taxonomicName id="C9E8B0E33B45FF9FFC1DFA33E075F3F7" authorityName="Krikken" authorityYear="2013" box="[917,1101,1413,1439]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="malabaricum">
<emphasis id="3C9C17723B45FF9FFC1DFA33E075F3F7" box="[917,1101,1413,1439]" italics="true" pageId="2" pageNumber="279">B. malabaricum</emphasis>
</taxonomicName>
,
<taxonomicName id="C9E8B0E33B45FF9FFBD4FA33E0D2F3F7" authorityName="Krikken" authorityYear="2013" box="[1116,1258,1414,1439]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="mimicans">
<emphasis id="3C9C17723B45FF9FFBD4FA33E0D2F3F7" box="[1116,1258,1414,1439]" italics="true" pageId="2" pageNumber="279">B. mimicans</emphasis>
</taxonomicName>
,
<taxonomicName id="C9E8B0E33B45FF9FFB70FA33E14EF3F7" authorityName="Krikken" authorityYear="2013" box="[1272,1398,1413,1439]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="schulzei">
<emphasis id="3C9C17723B45FF9FFB70FA33E14EF3F7" box="[1272,1398,1413,1439]" italics="true" pageId="2" pageNumber="279">B. schulzei</emphasis>
</taxonomicName>
,
<taxonomicName id="C9E8B0E33B45FF9FFA0CFA33E517F3AB" authorityName="Gupta &amp; Chandra" authorityYear="2016" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="bilaspuricans">
<emphasis id="3C9C17723B45FF9FFA0CFA33E517F3AB" italics="true" pageId="2" pageNumber="279">B. bilaspuricans</emphasis>
</taxonomicName>
,
<taxonomicName id="C9E8B0E33B45FF9FFEB6FA1FE5D0F3AB" authorityName="Gupta &amp; Chandra" authorityYear="2016" box="[318,488,1449,1475]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="darjeelicans">
<emphasis id="3C9C17723B45FF9FFEB6FA1FE5D0F3AB" box="[318,488,1449,1475]" italics="true" pageId="2" pageNumber="279">B. darjeelicans</emphasis>
</taxonomicName>
, and
<emphasis id="3C9C17723B45FF9FFDA0FA1FE6D0F3AB" box="[552,744,1449,1475]" italics="true" pageId="2" pageNumber="279">
<taxonomicName id="C9E8B0E33B45FF9FFDA0FA1FE6DDF3AB" authorityName="Kalawate &amp; Hillert" authorityYear="2018" box="[552,741,1449,1475]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="sahyadriensis">B. sahyadriensis</taxonomicName>
.
</emphasis>
While remaining two species,
<taxonomicName id="C9E8B0E33B45FF9FFBC2FA1FE0D8F3AB" authorityName="Krikken" authorityYear="2013" box="[1098,1248,1450,1475]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="extraneum">
<emphasis id="3C9C17723B45FF9FFBC2FA1FE0D8F3AB" box="[1098,1248,1450,1475]" italics="true" pageId="2" pageNumber="279">B. extraneum</emphasis>
</taxonomicName>
and
<taxonomicName id="C9E8B0E33B45FF9FFA92FA1FE1ADF3AB" authorityName="Krikken" authorityYear="2013" box="[1306,1429,1449,1475]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="insulare">
<emphasis id="3C9C17723B45FF9FFA92FA1FE1ADF3AB" box="[1306,1429,1449,1475]" italics="true" pageId="2" pageNumber="279">B. insulare</emphasis>
</taxonomicName>
, bear X-shaped perimarginal ridges.
</paragraph>
<paragraph id="0E57CB603B45FF9FFF4FFA44E176F08B" blockId="2.[151,1437,261,1764]" pageId="2" pageNumber="279">
Morphologically,
<taxonomicName id="C9E8B0E33B45FF9FFE18FA47E662F063" authority="Gupta &amp; Chandra &amp; Ghosh &amp; Das, 2020" authorityName="Gupta &amp; Chandra &amp; Ghosh &amp; Das" authorityYear="2020" box="[400,602,1521,1547]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="3C9C17723B45FF9FFE18FA47E662F063" box="[400,602,1521,1547]" italics="true" pageId="2" pageNumber="279">B. arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27AFAA093B45FF9FFDE9FA47E6D0F063" box="[609,744,1521,1548]" pageId="2" pageNumber="279" rank="species">new species</taxonomicNameLabel>
is similar to
<taxonomicName id="C9E8B0E33B45FF9FFCF2FA47E017F063" authorityName="Krikken" authorityYear="2013" box="[890,1071,1521,1547]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="malabaricum">
<emphasis id="3C9C17723B45FF9FFCF2FA47E017F063" box="[890,1071,1521,1547]" italics="true" pageId="2" pageNumber="279">B. malabaricum</emphasis>
</taxonomicName>
,
<taxonomicName id="C9E8B0E33B45FF9FFBB5FA47E0CCF063" authorityName="Gupta &amp; Chandra" authorityYear="2016" box="[1085,1268,1521,1547]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="bilaspuricans">
<emphasis id="3C9C17723B45FF9FFBB5FA47E0CCF063" box="[1085,1268,1521,1547]" italics="true" pageId="2" pageNumber="279">B. bilaspuricans</emphasis>
</taxonomicName>
, and
<taxonomicName id="C9E8B0E33B45FF9FFABAFA47E4C8F047" authorityName="Kalawate &amp; Hillert" authorityYear="2018" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="sahyadriensis">
<emphasis id="3C9C17723B45FF9FFABAFA47E4C8F047" italics="true" pageId="2" pageNumber="279">B. sahyadriensis</emphasis>
</taxonomicName>
, and closely related to
<taxonomicName id="C9E8B0E33B45FF9FFE7BF9A3E69FF047" authorityName="Krikken" authorityYear="2013" box="[499,679,1557,1583]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="malabaricum">
<emphasis id="3C9C17723B45FF9FFE7BF9A3E69FF047" box="[499,679,1557,1583]" italics="true" pageId="2" pageNumber="279">B. malabaricum</emphasis>
</taxonomicName>
. In
<taxonomicName id="C9E8B0E33B45FF9FFD5CF9A3E7A6F047" authority="Gupta &amp; Chandra &amp; Ghosh &amp; Das, 2020" authorityName="Gupta &amp; Chandra &amp; Ghosh &amp; Das" authorityYear="2020" box="[724,926,1557,1583]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="3C9C17723B45FF9FFD5CF9A3E7A6F047" box="[724,926,1557,1583]" italics="true" pageId="2" pageNumber="279">B. arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27AFAA093B45FF9FFC2DF9A3E016F047" box="[933,1070,1557,1584]" pageId="2" pageNumber="279" rank="species">new species</taxonomicNameLabel>
, the aedeagal stalks are long and expanded, pronotum with an inverted W-shaped carina medially, and the interocular ridge is long; whereas in
<taxonomicName id="C9E8B0E33B45FF9FFA0CF98FE514F01F" authorityName="Krikken" authorityYear="2013" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="malabaricum">
<emphasis id="3C9C17723B45FF9FFA0CF98FE514F01F" italics="true" pageId="2" pageNumber="279">B. malabaricum</emphasis>
</taxonomicName>
, the aedeagal stalks are small and narrow, pronotum with slightly developed discoparamedian and lateral tubercles, and the interocular ridge is small. In
<taxonomicName id="C9E8B0E33B45FF9FFD44F937E7ADF0F3" authority="Gupta &amp; Chandra &amp; Ghosh &amp; Das, 2020" authorityName="Gupta &amp; Chandra &amp; Ghosh &amp; Das" authorityYear="2020" box="[716,917,1665,1691]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="arunachalensis" status="sp. nov.">
<emphasis id="3C9C17723B45FF9FFD44F937E7ADF0F3" box="[716,917,1665,1691]" italics="true" pageId="2" pageNumber="279">B. arunachalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27AFAA093B45FF9FFC13F937E01BF0F3" box="[923,1059,1665,1692]" pageId="2" pageNumber="279" rank="species">new species</taxonomicNameLabel>
, the parameral tips are acuminate, but subangular in
<taxonomicName id="C9E8B0E33B45FF9FFEE8F913E62EF0D7" authorityName="Gupta &amp; Chandra" authorityYear="2016" box="[352,534,1701,1727]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="bilaspuricans">
<emphasis id="3C9C17723B45FF9FFEE8F913E62EF0D7" box="[352,534,1701,1727]" italics="true" pageId="2" pageNumber="279">B. bilaspuricans</emphasis>
</taxonomicName>
.
<taxonomicName id="C9E8B0E33B45FF9FFDADF910E77DF0D7" authorityName="Kalawate &amp; Hillert" authorityYear="2018" box="[549,837,1701,1727]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="sahyadriensis">
<emphasis id="3C9C17723B45FF9FFDADF910E77DF0D7" box="[549,837,1701,1727]" italics="true" pageId="2" pageNumber="279">Bolboceras sahyadriensis</emphasis>
</taxonomicName>
differs from the new species in having comparatively small and narrow aedeagal stalks and pronotum with well-developed discoparamedian and lateral tubercles.
</paragraph>
</subSubSection>
</treatment>
</document>