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<document id="5B6FD3D032C68DD0C576837E401865F5" ID-CLB-Dataset="5800" ID-DOI="10.2478/aemnp-2018-0032" ID-GBIF-Dataset="7a4ac9ad-4d2d-4808-9067-c02edb77fc23" ID-ISSN="1804-6487" ID-Zenodo-Dep="4504894" ID-ZooBank="8187447B-B951-4D73-973B-0AB6BB038A81" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612474315018" checkinUser="felipe" docAuthor="Bui, Van Bac &amp; Bonkowski, Michael" docDate="2018" docId="795D87BAFFBE6B1B85F7F95DABC8F79B" docLanguage="en" docName="ActaEntMusNatPra.58.2.407-418.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2)" docSource="http://dx.doi.org/10.2478/aemnp-2018-0032" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Synapsis Bates 1868" docType="key" docVersion="6" lastPageNumber="417" masterDocId="8564FFC2FFB76B118670FF9AA93FFF80" masterDocTitle="Synapsis puluongensis sp. nov. and redescription of S. horaki (Coleoptera: Scarabaeidae), with a key to Vietnamese species" masterLastPageNumber="418" masterPageNumber="407" pageNumber="416" updateTime="1698927446357" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="42BEE2E8BFA4C487CFF30F240BFE2E1F">Synapsis puluongensis sp. nov. and redescription of S. horaki (Coleoptera: Scarabaeidae), with a key to Vietnamese species</mods:title>
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<mods:namePart id="5BA42AA50EBECCF51BE6DC78738E48E3">Bui, Van Bac</mods:namePart>
<mods:affiliation id="964DD2664AB3BF626B448C7E1152E454">Institute of Zoology, Terrestrial Ecology, University of Cologne, Zülpicher Strasse 47 b, D- 50674 Cologne, Germany; e-mail: m. bonkowski @ unikoeln. de &amp; Department of Plant Protection, Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam; e-mails: buibac 80 @ gmail. com, vbui</mods:affiliation>
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<mods:namePart id="D42E45C0A8DAD8658210731F90FA989A">Bonkowski, Michael</mods:namePart>
<mods:affiliation id="47915411676C4C37AC22317859F74187">Institute of Zoology, Terrestrial Ecology, University of Cologne, Zülpicher Strasse 47 b, D- 50674 Cologne, Germany; e-mail: m. bonkowski @ unikoeln. de</mods:affiliation>
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Key to species of
<taxonomicName id="36F44D2FFFBE6B18821CF95CADE9F960" ID-CoL="7R23" authorityName="Bates" authorityYear="1868" box="[1132,1238,1734,1760]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="C380EABEFFBE6B18821CF95CADE9F960" bold="true" box="[1132,1238,1734,1760]" italics="true" pageId="9" pageNumber="416">Synapsis</emphasis>
</taxonomicName>
recorded from
<collectingCountry id="89E3763CFFBE6B18822BF970ADFAF884" box="[1115,1221,1770,1796]" name="Vietnam" pageId="9" pageNumber="416">Vietnam</collectingCountry>
</emphasis>
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<subSubSection id="B9EE6527FFBE6B18852AF880AC0DF85A" pageId="9" pageNumber="416" type="discussion">
<paragraph id="F14B36ACFFBE6B18852AF880ADFCF831" blockId="9.[828,1437,1818,2074]" pageId="9" pageNumber="416">
The key is based on the specimens examined by us in NMPC and VNUF as well as on literature data (
<bibRefCitation id="95654B5DFFBE6B188320F8A0ACA3F8D0" author="ARROW G. J." box="[1360,1436,1850,1873]" pageId="9" pageNumber="416" refId="ref7051" refString="ARROW G. J. 1931: The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor and Francis, London, 428 pp." type="book" year="1931">ARROW</bibRefCitation>
1931,
<bibRefCitation id="95654B5DFFBE6B18850EF8C0AD10F8F1" author="BALTHASAR V." box="[894,1071,1882,1905]" pageId="9" pageNumber="416" refId="ref7087" refString="BALTHASAR V. 1963: Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 1. Allgemeiner Teil, Systematischer Teil: 1. Scarabaeinae, 2. Coprinae (Pinotini, Coprini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prag, 391 pp." type="book" year="1963">BALTHASAR 1963</bibRefCitation>
,
<bibRefCitation id="95654B5DFFBE6B188249F8C0AC27F8F1" author="KRAL D. &amp; REJSEK J." box="[1081,1304,1882,1905]" pageId="9" pageNumber="416" pagination="267 - 270" refId="ref7420" refString="KRAL D. &amp; REJSEK J. 2000: Synapsis naxiorum sp. n. from Yunnan (Coleoptera: Scarabaeidae). Acta Societatis Zoologicae Bohemicae 64: 267 - 270." type="journal article" year="2000">KRÁL &amp; REJSEK 2000</bibRefCitation>
,
<bibRefCitation id="95654B5DFFBE6B188352F8C0ACA7F8F1" author="KRAL D." box="[1314,1432,1882,1905]" pageId="9" pageNumber="416" pagination="279 - 289" refId="ref7379" refString="KRAL D. 2002: Distribution and taxonomy of some Synapsis species, with description of S. strnadi sp. n. from Vietnam (Coleoptera: Scarabaeidae). Acta Societatis Zoologicae Bohemicae 66: 279 - 289." type="journal article" year="2002">KRÁL 2002</bibRefCitation>
,
<bibRefCitation id="95654B5DFFBE6B18854CF8E0ADA3F811" author="HANBOONSONG Y. &amp; MASUMOTO K." box="[828,1180,1914,1937]" pageId="9" pageNumber="416" pagination="453 - 462" refId="ref7308" refString="HANBOONSONG Y. &amp; MASUMOTO K. 1999: Dung beetles of Thailand, Part 1. Genus Synapsis. Elytra 27: 453 - 462." type="journal article" year="1999">HANBOONSONG &amp; MASUMOTO 1999</bibRefCitation>
,
<bibRefCitation id="95654B5DFFBE6B1882D5F8E0AC5FF811" author="OCHI T. &amp; KON M." box="[1189,1376,1914,1937]" pageId="9" pageNumber="416" pagination="91 - 95" refId="ref7545" refString="OCHI T. &amp; KON M. 2007: A new species of the genus Synapsis (Coleoptera, Scarabaeidae) from Laos. Elytra 35: 91 - 95." type="journal article" year="2007">OCHI &amp; KON 2007</bibRefCitation>
,
<bibRefCitation id="95654B5DFFBE6B188319F8E0ACA3F810" author="OCHI T. &amp; KON M. &amp; KAWAHARA M." box="[1385,1436,1914,1937]" pageId="9" pageNumber="416" pagination="191 - 199" refId="ref7576" refString="OCHI T., KON M. &amp; KAWAHARA M. 2008: Notes on the coprophagous scarabaeid beetles (Coleoptera: Scarabaeidae) from Southeast Asia (XIX). Three new taxa of Synapsis from Southeast Asia. Entomological Review of Japan 63: 191 - 199." type="journal article" year="2008">OCHI</bibRefCitation>
et al. 2008,
<bibRefCitation id="95654B5DFFBE6B1885C9F800AD88F831" author="ZIDEK J. &amp; POKORNY S." box="[953,1207,1946,1969]" pageId="9" pageNumber="416" pagination="1 - 21" refId="ref7657" refString="ZIDEK J. &amp; POKORNY S. 2010: Review of Synapsis Bates (Scarabaeidae: Scarabaeinae: Coprini), with description of a new species. Insecta Mundi 142: 1 - 21." type="journal article" year="2010">ZÍDEK &amp; POKORNÝ 2010</bibRefCitation>
).
</paragraph>
<paragraph id="F14B36ACFFBE6B18854CF85BAC0DF85A" blockId="9.[828,1437,1818,2074]" box="[828,1330,1985,2010]" pageId="9" pageNumber="416">
1(6) Hypomeral cavities present (
<figureCitation id="69CF2A29FFBE6B1882C6F85BADC9F85A" box="[1206,1270,1985,2010]" captionStart="Fig" captionStartId="3.[151,182,1474,1493]" captionTargetBox="[158,1431,469,1465]" captionTargetId="figure-137@3.[157,1431,469,1465]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of Synapsis species.AB elytral surface:A S. puluongensis sp. nov. with elytral striae strongly and densely punctate and interval 2 not swollen; B S. horaki Zídek &amp; Pokorný, 2010, with elytral striae impunctate and interval 2 swollen. CD metafemora: C S. puluongensis sp. nov.; D S. yama Gillet, 1911. EF setation of hypomeral cavities: E S. puluongensis sp. nov.; F S. birmanica Gillet, 1907. G elytron of S. naxiorum Král &amp; Rejsek, 2000 with weakly and sparsely punctate elytral striae and intervals weakly punctate. HI eye coloration in specimens of S. horaki Zídek &amp; Pokorný, 2010: H female; I male." figureDoi="http://doi.org/10.5281/zenodo.4504898" httpUri="https://zenodo.org/record/4504898/files/figure.png" pageId="9" pageNumber="416">Figs 2</figureCitation>
EF).
</paragraph>
</subSubSection>
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<paragraph id="F14B36ACFFBE6B18854CF878ACA2F79A" blockId="9.[828,1437,1818,2074]" pageId="9" pageNumber="416">
2(3) Elytral interval 2 near base not swollen (
<figureCitation id="69CF2A29FFBE6B18834CF87BACAFF87A" box="[1340,1424,2017,2042]" captionStart="Fig" captionStartId="3.[151,182,1474,1493]" captionTargetBox="[158,1431,469,1465]" captionTargetId="figure-137@3.[157,1431,469,1465]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of Synapsis species.AB elytral surface:A S. puluongensis sp. nov. with elytral striae strongly and densely punctate and interval 2 not swollen; B S. horaki Zídek &amp; Pokorný, 2010, with elytral striae impunctate and interval 2 swollen. CD metafemora: C S. puluongensis sp. nov.; D S. yama Gillet, 1911. EF setation of hypomeral cavities: E S. puluongensis sp. nov.; F S. birmanica Gillet, 1907. G elytron of S. naxiorum Král &amp; Rejsek, 2000 with weakly and sparsely punctate elytral striae and intervals weakly punctate. HI eye coloration in specimens of S. horaki Zídek &amp; Pokorný, 2010: H female; I male." figureDoi="http://doi.org/10.5281/zenodo.4504898" httpUri="https://zenodo.org/record/4504898/files/figure.png" pageId="9" pageNumber="416">Fig. 2A</figureCitation>
). ..........................................
<emphasis id="C380EABEFFBE6B1882EBF79BACA2F79A" bold="true" box="[1179,1437,2049,2074]" pageId="9" pageNumber="416">
<taxonomicName id="36F44D2FFFBE6B1882EBF79BAC7CF799" ID-CoL="7B7Y9" authority="Bui &amp; Bonkowski, 2018" authorityName="Bui &amp; Bonkowski" authorityYear="2018" box="[1179,1347,2049,2073]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="416" phylum="Arthropoda" rank="species" species="puluongensis" status="sp. nov.">
<emphasis id="C380EABEFFBE6B1882EBF79BAC7CF799" bold="true" box="[1179,1347,2049,2073]" italics="true" pageId="9" pageNumber="416">S. puluongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D8B357C5FFBE6B188339F798ACA2F79A" box="[1353,1437,2050,2074]" pageId="9" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<paragraph id="F14B36ACFFBD6B1B86E7FB1EACA1FB17" blockId="10.[151,1438,1156,1175]" box="[151,1438,1156,1175]" pageId="10" pageNumber="417">
Fig. 6. Dorsal habitus, ventral surface of femora and mesepisternal cavities.A, C, D
<taxonomicName id="36F44D2FFFBD6B1B851EFB1EADB0FB17" ID-CoL="7B7XX" authority="Boucomont, 1920" authorityName="Boucomont" authorityYear="1920" box="[878,1167,1156,1175]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="417" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="C380EABEFFBD6B1B851EFB1EAACFFB17" box="[878,1008,1156,1175]" italics="true" pageId="10" pageNumber="417">Synapsis ovalis</emphasis>
Boucomont, 1920
</taxonomicName>
; B, E, F
<taxonomicName id="36F44D2FFFBD6B1B8296FB1EACA6FB17" ID-CoL="7B7Y5" authority="Kral, 2002" authorityName="Kral" authorityYear="2002" box="[1254,1433,1156,1175]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="417" phylum="Arthropoda" rank="species" species="strnadi">
<emphasis id="C380EABEFFBD6B1B8296FB1EAC08FB17" box="[1254,1335,1156,1175]" italics="true" pageId="10" pageNumber="417">S. strnadi</emphasis>
<bibRefCitation id="95654B5DFFBD6B1B834CFB1EACA6FB17" author="KRAL D." box="[1340,1433,1156,1175]" pageId="10" pageNumber="417" pagination="279 - 289" refId="ref7379" refString="KRAL D. 2002: Distribution and taxonomy of some Synapsis species, with description of S. strnadi sp. n. from Vietnam (Coleoptera: Scarabaeidae). Acta Societatis Zoologicae Bohemicae 66: 279 - 289." type="journal article" year="2002">Král, 2002</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="F14B36ACFFBD6B1B86E7FB79AB80FB7B" blockId="10.[151,760,1251,2075]" box="[151,703,1251,1275]" pageId="10" pageNumber="417">
3(2) Elytral interval 2 near base swollen (
<figureCitation id="69CF2A29FFBD6B1B8413FB79AB8CFB7B" box="[611,691,1251,1275]" captionStart="Fig" captionStartId="3.[151,182,1474,1493]" captionTargetBox="[158,1431,469,1465]" captionTargetId="figure-137@3.[157,1431,469,1465]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of Synapsis species.AB elytral surface:A S. puluongensis sp. nov. with elytral striae strongly and densely punctate and interval 2 not swollen; B S. horaki Zídek &amp; Pokorný, 2010, with elytral striae impunctate and interval 2 swollen. CD metafemora: C S. puluongensis sp. nov.; D S. yama Gillet, 1911. EF setation of hypomeral cavities: E S. puluongensis sp. nov.; F S. birmanica Gillet, 1907. G elytron of S. naxiorum Král &amp; Rejsek, 2000 with weakly and sparsely punctate elytral striae and intervals weakly punctate. HI eye coloration in specimens of S. horaki Zídek &amp; Pokorný, 2010: H female; I male." figureDoi="http://doi.org/10.5281/zenodo.4504898" httpUri="https://zenodo.org/record/4504898/files/figure.png" pageId="10" pageNumber="417">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="F14B36ACFFBD6B1B86E7FA99ABC8FADB" blockId="10.[151,760,1251,2075]" pageId="10" pageNumber="417">
4(5) Metafemora in ventral view densely punctate, body length 17.524.0 mm. ...................................... ............................
<taxonomicName id="36F44D2FFFBD6B1B87EBFAD9ABC8FADB" ID-CoL="7B7XL" authority="Zidek &amp; Pokorny, 2010" authorityName="Zidek &amp; Pokorny" authorityYear="2010" box="[411,759,1347,1371]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="417" phylum="Arthropoda" rank="species" species="horaki">
<emphasis id="C380EABEFFBD6B1B87EBFAD9A8C2FADB" bold="true" box="[411,509,1347,1371]" italics="true" pageId="10" pageNumber="417">S. horaki</emphasis>
<bibRefCitation id="95654B5DFFBD6B1B8473FAD9ABC8FADB" author="ZIDEK J. &amp; POKORNY S." box="[515,759,1347,1371]" pageId="10" pageNumber="417" pagination="1 - 21" refId="ref7657" refString="ZIDEK J. &amp; POKORNY S. 2010: Review of Synapsis Bates (Scarabaeidae: Scarabaeinae: Coprini), with description of a new species. Insecta Mundi 142: 1 - 21." type="journal article" year="2010">Zídek &amp; Pokorný, 2010</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="F14B36ACFFBD6B1B86E7FAF9ABC8FA3B" blockId="10.[151,760,1251,2075]" pageId="10" pageNumber="417">
5(4) Metafemora in ventral view not punctate, body length 27.029.0 mm (
<figureCitation id="69CF2A29FFBD6B1B87A2FA19AB19FA1B" box="[466,550,1411,1435]" captionStart="Fig" captionStartId="3.[151,182,1474,1493]" captionTargetBox="[158,1431,469,1465]" captionTargetId="figure-137@3.[157,1431,469,1465]" captionTargetPageId="3" captionText="Fig. 2. Morphological details of Synapsis species.AB elytral surface:A S. puluongensis sp. nov. with elytral striae strongly and densely punctate and interval 2 not swollen; B S. horaki Zídek &amp; Pokorný, 2010, with elytral striae impunctate and interval 2 swollen. CD metafemora: C S. puluongensis sp. nov.; D S. yama Gillet, 1911. EF setation of hypomeral cavities: E S. puluongensis sp. nov.; F S. birmanica Gillet, 1907. G elytron of S. naxiorum Král &amp; Rejsek, 2000 with weakly and sparsely punctate elytral striae and intervals weakly punctate. HI eye coloration in specimens of S. horaki Zídek &amp; Pokorný, 2010: H female; I male." figureDoi="http://doi.org/10.5281/zenodo.4504898" httpUri="https://zenodo.org/record/4504898/files/figure.png" pageId="10" pageNumber="417">Fig. 2D</figureCitation>
). ............................... ..................................................
<taxonomicName id="36F44D2FFFBD6B1B8450FA39ABC8FA3B" ID-CoL="7B7Y7" authority="Gillet, 1911" authorityName="Gillet" authorityYear="1911" box="[544,759,1443,1467]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="417" phylum="Arthropoda" rank="species" species="yama">
<emphasis id="C380EABEFFBD6B1B8450FA39AB48FA3A" bold="true" box="[544,631,1443,1467]" italics="true" pageId="10" pageNumber="417">S. yama</emphasis>
<bibRefCitation id="95654B5DFFBD6B1B840DFA39ABC8FA3B" author="GILLET J. J. E." box="[637,759,1443,1467]" pageId="10" pageNumber="417" pagination="313 - 314" refId="ref7254" refString="GILLET J. J. E. 1911: Coprides nouveaux de la region orientale et remarques synonymiques. Annales de la Societe Entomologique de Belgique 55: 313 - 314." type="journal article" year="1911">Gillet, 1911</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="F14B36ACFFBD6B1B86E7FA59AB3DFA5B" blockId="10.[151,760,1251,2075]" box="[151,514,1475,1499]" pageId="10" pageNumber="417">6(7) Hypomeral cavities absent.</paragraph>
<paragraph id="F14B36ACFFBD6B1B86E7FA79A8DAF99B" blockId="10.[151,760,1251,2075]" pageId="10" pageNumber="417">7(10) Frons with minor horn or medial tubercle; mesepisternal cavities absent.</paragraph>
<paragraph id="F14B36ACFFBD6B1B86E7F9B9ABC8F9FB" blockId="10.[151,760,1251,2075]" pageId="10" pageNumber="417">
8(9) Genae not expanded, anterolateral angle of pronotum not dentate (
<figureCitation id="69CF2A29FFBD6B1B87EBF9D9A8D2F9DB" box="[411,493,1603,1627]" captionStart="Fig 5" captionStartId="9.[151,179,1097,1116]" captionTargetBox="[208,1397,202,1080]" captionTargetId="figure-17@9.[190,1397,185,1090]" captionTargetPageId="9" captionText="Fig 5. Dorsal habitus. A Synapsis simplex Sharp, 1875. B S. tridens Sharp, 1881" figureDoi="http://doi.org/10.5281/zenodo.4504904" httpUri="https://zenodo.org/record/4504904/files/figure.png" pageId="10" pageNumber="417">Fig. 5A</figureCitation>
). ........................................ ..............................................
<taxonomicName id="36F44D2FFFBD6B1B8478F9F9ABC8F9FB" ID-CoL="7B7XT" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[520,759,1635,1659]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="417" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="C380EABEFFBD6B1B8478F9F9AB4AF9FB" bold="true" box="[520,629,1635,1659]" italics="true" pageId="10" pageNumber="417">S. simplex</emphasis>
Sharp, 1875
</taxonomicName>
</paragraph>
<paragraph id="F14B36ACFFBD6B1B86E7F919ABC8F93B" blockId="10.[151,760,1251,2075]" pageId="10" pageNumber="417">
9(8) Genae expanded; anterolateral angle of pronotum tridentate (
<figureCitation id="69CF2A29FFBD6B1B8728F939A897F93B" box="[344,424,1699,1723]" captionStart="Fig 5" captionStartId="9.[151,179,1097,1116]" captionTargetBox="[208,1397,202,1080]" captionTargetId="figure-17@9.[190,1397,185,1090]" captionTargetPageId="9" captionText="Fig 5. Dorsal habitus. A Synapsis simplex Sharp, 1875. B S. tridens Sharp, 1881" figureDoi="http://doi.org/10.5281/zenodo.4504904" httpUri="https://zenodo.org/record/4504904/files/figure.png" pageId="10" pageNumber="417">Fig. 5B</figureCitation>
). ............
<taxonomicName id="36F44D2FFFBD6B1B847EF939ABC8F93B" ID-CoL="7B89W" authority="Sharp, 1881" authorityName="Sharp" authorityYear="1881" box="[526,759,1699,1723]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="417" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="C380EABEFFBD6B1B847EF939AB4BF93B" bold="true" box="[526,628,1699,1723]" italics="true" pageId="10" pageNumber="417">S. tridens</emphasis>
<bibRefCitation id="95654B5DFFBD6B1B840BF939ABC8F93B" author="SHARP D." box="[635,759,1699,1723]" pageId="10" pageNumber="417" refId="ref7626" refString="SHARP D. 1881: Note sur l'Ateuchus tmolus Fisch. Avec description d'une espece nouvelle du genre Synapsis. Annales de la Societe Entomologique de Belgique 25: xci-xcii." type="book" year="1881">Sharp, 1881</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="F14B36ACFFBD6B1B86E7F959AB20F89B" blockId="10.[151,760,1251,2075]" pageId="10" pageNumber="417">
10(11) Frons without minor horn or only with medial tubercle; mesepisternal cavities present and covered with red setae (
<figureCitation id="69CF2A29FFBD6B1B87F3F899A8FCF89B" box="[387,451,1795,1819]" captionStart="Fig" captionStartId="10.[151,182,1156,1175]" captionTargetBox="[196,1374,206,1137]" captionTargetId="figure-391@10.[187,1400,189,1157]" captionTargetPageId="10" captionText="Fig. 6. Dorsal habitus, ventral surface of femora and mesepisternal cavities.A, C, D Synapsis ovalis Boucomont, 1920; B, E, F S. strnadi Král, 2002." figureDoi="http://doi.org/10.5281/zenodo.4504906" httpUri="https://zenodo.org/record/4504906/files/figure.png" pageId="10" pageNumber="417">Figs 6</figureCitation>
AC, F).
</paragraph>
<paragraph id="F14B36ACFFBD6B1B86E7F8B9ABC8F81B" blockId="10.[151,760,1251,2075]" pageId="10" pageNumber="417">
11(12) Anterolateral angles of pronotum nearly rectangular (90°). Lateral angles of genae obtuse, rounded. Metafemora in ventral view sparsely punctate (
<figureCitation id="69CF2A29FFBD6B1B869AF819A85FF81B" box="[234,352,1923,1947]" captionStart="Fig" captionStartId="10.[151,182,1156,1175]" captionTargetBox="[196,1374,206,1137]" captionTargetId="figure-391@10.[187,1400,189,1157]" captionTargetPageId="10" captionText="Fig. 6. Dorsal habitus, ventral surface of femora and mesepisternal cavities.A, C, D Synapsis ovalis Boucomont, 1920; B, E, F S. strnadi Král, 2002." figureDoi="http://doi.org/10.5281/zenodo.4504906" httpUri="https://zenodo.org/record/4504906/files/figure.png" pageId="10" pageNumber="417">Figs 6A, D</figureCitation>
). ................
<taxonomicName id="36F44D2FFFBD6B1B87ABF819ABC8F81B" ID-CoL="7B7XX" authority="Boucomont, 1920" authorityName="Boucomont" authorityYear="1920" box="[475,759,1923,1947]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="417" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="C380EABEFFBD6B1B87ABF819AB0AF81B" bold="true" box="[475,565,1923,1947]" italics="true" pageId="10" pageNumber="417">S. ovalis</emphasis>
Boucomont, 1920
</taxonomicName>
</paragraph>
<paragraph id="F14B36ACFFBD6B1B86E7F839ABC8F79B" blockId="10.[151,760,1251,2075]" pageId="10" pageNumber="417">
12(11) Anterolateral angles of pronotum about 135°. Lateral angles of genae rather sharp, metafemora in ventral view densely punctate (
<figureCitation id="69CF2A29FFBD6B1B845AF879ABA6F87B" box="[554,665,2019,2043]" captionStart="Fig" captionStartId="10.[151,182,1156,1175]" captionTargetBox="[196,1374,206,1137]" captionTargetId="figure-391@10.[187,1400,189,1157]" captionTargetPageId="10" captionText="Fig. 6. Dorsal habitus, ventral surface of femora and mesepisternal cavities.A, C, D Synapsis ovalis Boucomont, 1920; B, E, F S. strnadi Král, 2002." figureDoi="http://doi.org/10.5281/zenodo.4504906" httpUri="https://zenodo.org/record/4504906/files/figure.png" pageId="10" pageNumber="417">Figs 6B, E</figureCitation>
). ............ .................................................
<taxonomicName id="36F44D2FFFBD6B1B846BF799ABC8F79B" ID-CoL="7B7Y5" authority="Kral, 2002" authorityName="Kral" authorityYear="2002" box="[539,759,2051,2075]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="417" phylum="Arthropoda" rank="species" species="strnadi">
<emphasis id="C380EABEFFBD6B1B846BF799ABBBF79B" bold="true" box="[539,644,2051,2075]" italics="true" pageId="10" pageNumber="417">S. strnadi</emphasis>
<bibRefCitation id="95654B5DFFBD6B1B84FAF799ABC8F79B" author="KRAL D." box="[650,759,2051,2075]" pageId="10" pageNumber="417" pagination="279 - 289" refId="ref7379" refString="KRAL D. 2002: Distribution and taxonomy of some Synapsis species, with description of S. strnadi sp. n. from Vietnam (Coleoptera: Scarabaeidae). Acta Societatis Zoologicae Bohemicae 66: 279 - 289." type="journal article" year="2002">Král, 2002</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>