treatments-xml/data/69/00/28/6900287EFFC1167C16CB18DFFE4D47D5.xml

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<mods:title id="A18BEC78C700F285E1F9AE02883E1A0E">Description of Coeliccia lephuocdieui sp. nov. from the Central Highlands of Vietnam (Odonata: Zygoptera: Platycnemididae) with notes on its congeners</mods:title>
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<mods:namePart id="24A9891ACC41351E07C3AA19582D10F9">Phan, Quoc Toan</mods:namePart>
<mods:affiliation id="EAA2D3DF50552B06515309485089A70D">Center for Entomology &amp; Parasitology Research, Institute of Research &amp; Training of Medicine, Biology &amp; Pharmacy, Duy Tan University, 3 Quang Trung, Da Nang city, Vietnam</mods:affiliation>
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<mods:namePart id="F5A006BC22311D67B8FE5B9B19543A43">Ngo, Quoc Phu</mods:namePart>
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<mods:namePart id="4F74D97D0CF5535ABA0CC817CF89E86E">Bui, Anh Phong</mods:namePart>
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<taxonomicName id="26A9E2EBFFC1167816CB18DFFE0B40FB" ID-CoL="9X2BQ" authority="Phan &amp; Ngo &amp; Bui, 2020" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[151,427,225,252]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="70" phylum="Arthropoda" rank="species" species="lephuocdieui" status="sp. nov.">
<emphasis id="D3DD457AFFC1167816CB18DFFE0B40FB" bold="true" box="[151,427,225,252]" italics="true" pageId="1" pageNumber="70">Coeliccia lephuocdieui</emphasis>
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<taxonomicNameLabel id="C8EEF801FFC1167817EF18DCFDB040FB" box="[435,528,226,252]" pageId="1" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="E1169968FFC1167816CB193BFDED4118" blockId="1.[151,589,225,288]" box="[151,589,261,288]" pageId="1" pageNumber="70">
(
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, 
<figureCitation id="799285EDFFC116781744193BFE954118" box="[280,309,261,287]" captionStart="FIGURE 2" captionStartId="2.[151,250,910,935]" captionTargetBox="[151,1436,181,884]" captionTargetId="figure@2.[151,1436,181,884]" captionTargetPageId="2" captionText="FIGURE 2. Head and thorax of male, lateral view. (2a) C. lephuocdieui sp. nov., holotype male; (2b) C. scutellum, Kon Ka Kinh National Park." figureDoi="http://doi.org/10.5281/zenodo.3864988" httpUri="https://zenodo.org/record/3864988/files/figure.png" pageId="1" pageNumber="70">2a</figureCitation>
, 
<figureCitation id="799285EDFFC11678171C193BFEED4118" box="[320,333,261,287]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="1" pageNumber="70">3</figureCitation>
a–b, 4a–b, 5, 6a–b, 8a)
</paragraph>
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<emphasis id="D3DD457AFFC1167816CB1970FEE8416F" bold="true" box="[151,328,333,360]" pageId="1" pageNumber="70">Type material.</emphasis>
<materialsCitation id="51C19335FFC11678170D1970FE2941A8" ID-GBIF-Occurrence="2628393302" collectingDate="2019-04-21" collectionCode="T" collectorName="Kon Ka Kinh NP &amp; Q. P. Ngo" elevation="1489" latitude="14.2586" location="Mang Yang District" longLatPrecision="7" longitude="108.3786" municipality="Ayun Commune" pageId="1" pageNumber="70" specimenCount="1" specimenCount-male="1" stateProvince="Gia Lai" typeStatus="holotype">
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<typeStatus id="3E1227CAFFC11678170D1970FE62416F" box="[337,450,334,360]" pageId="1" pageNumber="70" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="F7AF52E1FFC11678178D1970FE48416F" box="[465,488,334,360]" pageId="1" pageNumber="70" type="male">♂</specimenCount>
, a fully mature male, mountain top near the headquarters of the 
<collectorName id="4C5CFCBEFFC1167812881970FA3C416F" box="[1236,1436,333,360]" pageId="1" pageNumber="70">Kon Ka Kinh NP</collectorName>
(
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, 
<geoCoordinate id="849DFFAFFFC11678177C194FFE3E418C" box="[288,414,369,395]" degrees="108.3786" direction="east" orientation="longitude" pageId="1" pageNumber="70" precision="5" value="108.3786">108.3786E</geoCoordinate>
; altitude 
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<elevation id="6A847E5BFFC116781455194FFDC3418B" box="[521,611,369,396]" metricMagnitude="3" metricUnit="m" metricValue="1.489" pageId="1" pageNumber="70" unit="m" value="1489.0">1489 m</elevation>
</quantity>
), 
<collectingMunicipality id="01720312FFC116781424194CFC95418C" box="[632,821,369,396]" pageId="1" pageNumber="70">Ayun Commune</collectingMunicipality>
, 
<location id="E476CFB3FFC11678151E194CFB86418C" LSID="urn:lsid:plazi:treatment:6900287EFFC1167C16CB18DFFE4D47D5:E476CFB3FFC11678151E194CFB86418C" box="[834,1062,369,396]" latitude="14.2586" longLatPrecision="7" longitude="108.3786" municipality="Ayun Commune" name="Mang Yang District" pageId="1" pageNumber="70" stateProvince="Gia Lai">Mang Yang District</location>
, 
<collectingRegion id="236D578AFFC11678126F194FFB71418B" box="[1075,1233,369,396]" country="Vietnam" name="Gia Lai" pageId="1" pageNumber="70">Gia Lai Prov.</collectingRegion>
, 
<date id="9517BFA8FFC116781282194FFAFE418C" box="[1246,1374,369,395]" pageId="1" pageNumber="70" value="2019-04-21">
<collectingDate id="85534640FFC116781282194FFAFE418C" box="[1246,1374,369,395]" pageId="1" pageNumber="70" value="2019-04-21">21 iv 2019</collectingDate>
</date>
, Q. 
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. Phan &amp; 
<collectorName id="4C5CFCBEFFC1167816AF19ABFEF941B7" box="[243,345,405,432]" pageId="1" pageNumber="70">Q.P. Ngo</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="E1169968FFC11678169B1984FE0E41F0" blockId="1.[151,1437,333,792]" pageId="1" pageNumber="70">
<materialsCitation id="51C19335FFC11678169B1984FCFE41D4" ID-GBIF-Occurrence="2628393331" box="[199,862,441,468]" collectingDate="2019-04-21" collectionCode="T" collectorName="Kon Ka Kinh NP &amp; Q. P. Ngo" elevation="1489" latitude="14.2586" location="Mang Yang District" longLatPrecision="7" longitude="108.3786" municipality="Ayun Commune" pageId="1" pageNumber="70" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Gia Lai" typeStatus="paratype">
<emphasis id="D3DD457AFFC11678169B1984FEE841D3" bold="true" box="[199,328,442,468]" pageId="1" pageNumber="70">
<typeStatus id="3E1227CAFFC11678169B1984FEE341D3" box="[199,323,442,468]" pageId="1" pageNumber="70" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="F7AF52E1FFC1167817131987FE2F41D3" box="[335,399,441,468]" pageId="1" pageNumber="70" type="male">3 ♂♂</specimenCount>
, 
<specimenCount id="F7AF52E1FFC1167817C71987FE7A41D3" box="[411,474,441,468]" pageId="1" pageNumber="70" type="female">3 ♀♀</specimenCount>
, all mature, same data as holotype
</materialsCitation>
; 
<materialsCitation id="51C19335FFC1167815341987FE0E41F0" ID-GBIF-Occurrence="2628393314" collectingDate="2018-04-04" collectionCode="T" collectorName="To Van Quang" elevation="1489" latitude="14.2586" location="Mang Yang District" longLatPrecision="7" longitude="108.3786" municipality="Ayun Commune" pageId="1" pageNumber="70" specimenCount="1" specimenCount-male="1" stateProvince="Gia Lai" typeStatus="paratype">
<specimenCount id="F7AF52E1FFC1167815341987FC3441D3" box="[872,916,441,468]" pageId="1" pageNumber="70" type="male">1 ♂</specimenCount>
, fully mature, same location as holotype, 
<date id="9517BFA8FFC1167813381987FF7241F0" pageId="1" pageNumber="70" value="2018-04-04">
<collectingDate id="85534640FFC1167813381987FF7241F0" pageId="1" pageNumber="70" value="2018-04-04">04 iv 2018</collectingDate>
</date>
, 
<collectorName id="4C5CFCBEFFC11678168119E0FEDE41F0" box="[221,382,477,504]" pageId="1" pageNumber="70">To Van Quang</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="E1169968FFC11678169B1A3CFE524238" blockId="1.[151,1437,333,792]" pageId="1" pageNumber="70">
<emphasis id="D3DD457AFFC11678169B1A3CFE2B421B" bold="true" box="[199,395,513,540]" pageId="1" pageNumber="70">Type deposition.</emphasis>
<materialsCitation id="51C19335FFC1167817CD1A3FFE4E4238" ID-GBIF-Occurrence="2628393311" country="Vietnam" location="Zoological Collection of Duy Tan University" pageId="1" pageNumber="70" specimenCount="1" stateProvince="Da Nang">
All 
<typeStatus id="3E1227CAFFC1167817E01A3CFE4C421B" box="[444,492,514,540]" pageId="1" pageNumber="70">type</typeStatus>
specimens of the new species are deposited in the 
<location id="E476CFB3FFC11678127F1A3FFEAE4238" LSID="urn:lsid:plazi:treatment:6900287EFFC1167C16CB18DFFE4D47D5:E476CFB3FFC11678127F1A3FFEAE4238" country="Vietnam" name="Zoological Collection of Duy Tan University" pageId="1" pageNumber="70" stateProvince="Da Nang">Zoological Collection of Duy Tan University</location>
, 
<collectingRegion id="236D578AFFC1167817461A18FE204247" box="[282,384,550,576]" country="Vietnam" name="Da Nang" pageId="1" pageNumber="70">Da Nang</collectingRegion>
, 
<collectingCountry id="99BED9F8FFC1167817D71A1BFE4E4238" box="[395,494,549,575]" name="Vietnam" pageId="1" pageNumber="70">Vietnam</collectingCountry>
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.
</paragraph>
</subSubSection>
<subSubSection id="A9B3CAE3FFC11678169B1A74FB6B428F" pageId="1" pageNumber="70" type="etymology">
<paragraph id="E1169968FFC11678169B1A74FB6B428F" blockId="1.[151,1437,333,792]" pageId="1" pageNumber="70">
<emphasis id="D3DD457AFFC11678169B1A74FEEE4263" bold="true" box="[199,334,586,612]" pageId="1" pageNumber="70">Etymology.</emphasis>
The species is named 
<taxonomicName id="26A9E2EBFFC1167814131A77FD404264" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[591,736,585,611]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="70" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC1167814131A77FD404264" box="[591,736,585,611]" italics="true" pageId="1" pageNumber="70">lephuocdieui</emphasis>
</taxonomicName>
in memory of our friend, Mr Le Phuoc Dieu (born 
<date id="9517BFA8FFC1167813751A77FF5C4280" pageId="1" pageNumber="70" value="1984-12-07">December 07, 1984</date>
, deceased 
<date id="9517BFA8FFC11678172F1A50FDB24280" box="[371,530,621,648]" pageId="1" pageNumber="70" value="2019-05-24">May 24, 2019</date>
) of 
<collectingRegion id="236D578AFFC1167814621A53FCBE4280" box="[574,798,621,647]" country="Vietnam" name="Quang Tri" pageId="1" pageNumber="70">Quang Tri Province</collectingRegion>
, 
<collectingCountry id="99BED9F8FFC1167815751A53FC2C4280" box="[809,908,621,647]" name="Vietnam" pageId="1" pageNumber="70">Vietnam</collectingCountry>
. A noun in the genitive case.
</paragraph>
</subSubSection>
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<emphasis id="D3DD457AFFC11678169B1AAFFE4542AB" bold="true" box="[199,485,657,684]" pageId="1" pageNumber="70">
Description of 
<typeStatus id="3E1227CAFFC11678172B1AACFE4042AB" box="[375,480,658,684]" pageId="1" pageNumber="70" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="D3DD457AFFC1167817B01AAFFD8942AC" box="[492,553,657,683]" italics="true" pageId="1" pageNumber="70">Head</emphasis>
(
<figureCitation id="799285EDFFC1167814641AAFFD2E42AC" box="[568,654,657,683]" captionStart="FIGURE 2" captionStartId="2.[151,250,910,935]" captionTargetBox="[151,1436,181,884]" captionTargetId="figure@2.[151,1436,181,884]" captionTargetPageId="2" captionText="FIGURE 2. Head and thorax of male, lateral view. (2a) C. lephuocdieui sp. nov., holotype male; (2b) C. scutellum, Kon Ka Kinh National Park." figureDoi="http://doi.org/10.5281/zenodo.3864988" httpUri="https://zenodo.org/record/3864988/files/figure.png" pageId="1" pageNumber="70">Figs 2a</figureCitation>
, 
<figureCitation id="799285EDFFC1167814C61AAFFD1742AB" box="[666,695,657,684]" captionStart="FIGURE 5" captionStartId="5.[151,250,1616,1641]" captionTargetBox="[151,1436,528,1592]" captionTargetId="figure@5.[151,1436,525,1592]" captionTargetPageId="5" captionText="FIGURE 5. Structure features of Coeliccia lephuocdieui sp. nov. (5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="1" pageNumber="70">5c</figureCitation>
). Labrum, anteclypeus, postclypeus, genae and mandibles shining black; front of head and antennae matte black except for top of first segment dark yellowish. Dorsum of head matte black excluding two narrow stripes adjacent to posterior ocelli and two oval yellowish postocular spots. Compound eyes bicoloured in life, black above and green below (
<figureCitation id="799285EDFFC1167814A91AC3FCE64310" box="[757,838,765,791]" captionStart="FIGURE 8" captionStartId="9.[151,250,1783,1808]" captionTargetBox="[151,1436,597,1757]" captionTargetId="figure@9.[151,1436,597,1757]" captionTargetPageId="9" captionText="FIGURE 8. Habitus of Coeliccia spp. in nature. (8a) C. lephuocdieui sp. nov. in the type location, 21 April 2019 (with a red parasitic mite on the dorsum synthorax of the male); (8b) C. scutellum, Son Tra Nature Reserve, 25 April 2018. Photographed by the first author." figureDoi="http://doi.org/10.5281/zenodo.3865001" httpUri="https://zenodo.org/record/3865001/files/figure.png" pageId="1" pageNumber="70">Fig. 8a</figureCitation>
).
</paragraph>
<caption id="B5D6C9E0FFC1167816CB1FF4FB5C47E5" ID-DOI="http://doi.org/10.5281/zenodo.3864986" ID-Zenodo-Dep="3864986" box="[151,1276,1994,2019]" httpUri="https://zenodo.org/record/3864986/files/figure.png" pageId="1" pageNumber="70" startId="1.[151,250,1994,2019]" targetBox="[184,1404,824,1957]" targetPageId="1">
<paragraph id="E1169968FFC1167816CB1FF4FB5C47E5" blockId="1.[151,1276,1994,2019]" box="[151,1276,1994,2019]" pageId="1" pageNumber="70">
<emphasis id="D3DD457AFFC1167816CB1FF4FEAD47E4" bold="true" box="[151,269,1994,2019]" pageId="1" pageNumber="70">FIGURE 1</emphasis>
. Habitus of 
<taxonomicName id="26A9E2EBFFC1167817D61FF5FD8B47E4" authority="Phan &amp; Ngo &amp; Bui, 2020" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[394,555,1995,2019]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="70" phylum="Arthropoda" rank="species" species="lephuocdieui" status="sp. nov.">
<emphasis id="D3DD457AFFC1167817D61FF5FD8B47E4" box="[394,555,1995,2019]" italics="true" pageId="1" pageNumber="70">C. lephuocdieui</emphasis>
</taxonomicName>
<emphasis id="D3DD457AFFC11678146D1FF5FD2147E4" bold="true" box="[561,641,1995,2019]" pageId="1" pageNumber="70">
<taxonomicNameLabel id="C8EEF801FFC11678146D1FF5FD2147E4" box="[561,641,1995,2019]" pageId="1" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(1a) holotype male &amp; (1b) paratype female. Scale bar is 1 cm.
</paragraph>
</caption>
<caption id="B5D6C9E0FFC2167B16CB1BB0FEC443CC" ID-DOI="http://doi.org/10.5281/zenodo.3864988" ID-Zenodo-Dep="3864988" httpUri="https://zenodo.org/record/3864988/files/figure.png" pageId="2" pageNumber="71" startId="2.[151,250,910,935]" targetBox="[151,1436,181,884]" targetPageId="2">
<paragraph id="E1169968FFC2167B16CB1BB0FEC443CC" blockId="2.[151,1437,910,971]" pageId="2" pageNumber="71">
<emphasis id="D3DD457AFFC2167B16CB1BB0FEAF43A0" bold="true" box="[151,271,910,935]" pageId="2" pageNumber="71">FIGURE 2</emphasis>
. Head and thorax of male, lateral view. (2a) 
<taxonomicName id="26A9E2EBFFC2167B14BC1BB1FC2243A1" authority="Phan &amp; Ngo &amp; Bui, 2020" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[736,898,910,934]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="71" phylum="Arthropoda" rank="species" species="lephuocdieui" status="sp. nov.">
<emphasis id="D3DD457AFFC2167B14BC1BB1FC2243A1" box="[736,898,910,934]" italics="true" pageId="2" pageNumber="71">C. lephuocdieui</emphasis>
</taxonomicName>
<emphasis id="D3DD457AFFC2167B15D61BB0FC7C43A1" bold="true" box="[906,988,910,934]" pageId="2" pageNumber="71">
<taxonomicNameLabel id="C8EEF801FFC2167B15D61BB0FC7C43A1" box="[906,988,910,934]" pageId="2" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male; (2b) 
<emphasis id="D3DD457AFFC2167B12E11BB1FAE443A1" box="[1213,1348,910,934]" italics="true" pageId="2" pageNumber="71">
<taxonomicName id="26A9E2EBFFC2167B12E11BB1FAE043A1" authorityName="Laidlaw" authorityYear="1932" box="[1213,1344,910,934]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="71" phylum="Arthropoda" rank="species" species="scutellum">C. scutellum</taxonomicName>
,
</emphasis>
Kon Ka Kinh National Park.
</paragraph>
</caption>
<paragraph id="E1169968FFC2167B169B1BC7FD1444A4" blockId="2.[151,1437,1017,2016]" pageId="2" pageNumber="71">
<emphasis id="D3DD457AFFC2167B169B1BC7FEB84414" box="[199,280,1017,1043]" italics="true" pageId="2" pageNumber="71">Thorax</emphasis>
(
<figureCitation id="799285EDFFC2167B177A1BC7FED74414" box="[294,375,1017,1043]" captionStart="FIGURE 2" captionStartId="2.[151,250,910,935]" captionTargetBox="[151,1436,181,884]" captionTargetId="figure@2.[151,1436,181,884]" captionTargetPageId="2" captionText="FIGURE 2. Head and thorax of male, lateral view. (2a) C. lephuocdieui sp. nov., holotype male; (2b) C. scutellum, Kon Ka Kinh National Park." figureDoi="http://doi.org/10.5281/zenodo.3864988" httpUri="https://zenodo.org/record/3864988/files/figure.png" pageId="2" pageNumber="71">Fig. 2a</figureCitation>
). Prothorax: black apart from dark yellow marking on middle pronotal lobe. Synthorax: mesepisternum and mesepimeron entirely black; metepisternum and metepimeron with two distinct large yellow markings as in 
<figureCitation id="799285EDFFC2167B168F1C7FFE9E445C" box="[211,318,1089,1115]" captionStart="FIGURE 2" captionStartId="2.[151,250,910,935]" captionTargetBox="[151,1436,181,884]" captionTargetId="figure@2.[151,1436,181,884]" captionTargetPageId="2" captionText="FIGURE 2. Head and thorax of male, lateral view. (2a) C. lephuocdieui sp. nov., holotype male; (2b) C. scutellum, Kon Ka Kinh National Park." figureDoi="http://doi.org/10.5281/zenodo.3864988" httpUri="https://zenodo.org/record/3864988/files/figure.png" pageId="2" pageNumber="71">Figure 2a</figureCitation>
(pattern greenish-yellow in living individual, 
<figureCitation id="799285EDFFC2167B151F1C7FFC34445C" box="[835,916,1089,1115]" captionStart="FIGURE 8" captionStartId="9.[151,250,1783,1808]" captionTargetBox="[151,1436,597,1757]" captionTargetId="figure@9.[151,1436,597,1757]" captionTargetPageId="9" captionText="FIGURE 8. Habitus of Coeliccia spp. in nature. (8a) C. lephuocdieui sp. nov. in the type location, 21 April 2019 (with a red parasitic mite on the dorsum synthorax of the male); (8b) C. scutellum, Son Tra Nature Reserve, 25 April 2018. Photographed by the first author." figureDoi="http://doi.org/10.5281/zenodo.3865001" httpUri="https://zenodo.org/record/3865001/files/figure.png" pageId="2" pageNumber="71">Fig. 8a</figureCitation>
). Mesinfraepisternum and metinfraepisternum entirely black. 
<emphasis id="D3DD457AFFC2167B17631C5BFED44479" box="[319,372,1125,1150]" italics="true" pageId="2" pageNumber="71">Legs</emphasis>
(
<figureCitation id="799285EDFFC2167B17DF1C5BFE764478" box="[387,470,1125,1151]" captionStart="FIGURE 2" captionStartId="2.[151,250,910,935]" captionTargetBox="[151,1436,181,884]" captionTargetId="figure@2.[151,1436,181,884]" captionTargetPageId="2" captionText="FIGURE 2. Head and thorax of male, lateral view. (2a) C. lephuocdieui sp. nov., holotype male; (2b) C. scutellum, Kon Ka Kinh National Park." figureDoi="http://doi.org/10.5281/zenodo.3864988" httpUri="https://zenodo.org/record/3864988/files/figure.png" pageId="2" pageNumber="71">Fig. 2a</figureCitation>
). Coxae and trochanters black with dark yellowish marking. Femur and tibia pale yellowish with black lines. Spines and claws black.
</paragraph>
<paragraph id="E1169968FFC2167B169B1C93FBCA44C0" blockId="2.[151,1437,1017,2016]" box="[199,1130,1197,1223]" pageId="2" pageNumber="71">
<emphasis id="D3DD457AFFC2167B169B1C93FEAB44C1" box="[199,267,1197,1222]" italics="true" pageId="2" pageNumber="71">Wings</emphasis>
. Hyaline, 18 px in FW and 
<quantity id="2651348DFFC2167B141F1C93FDDC44C0" box="[579,636,1197,1223]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="2" pageNumber="71" unit="in" value="17.0">17 in</quantity>
HW. Pt brown, covering 2 underlying cells.
</paragraph>
<paragraph id="E1169968FFC2167B169B1CEFFDE54533" blockId="2.[151,1437,1017,2016]" pageId="2" pageNumber="71">
<emphasis id="D3DD457AFFC2167B169B1CEFFE9144EC" box="[199,305,1233,1259]" italics="true" pageId="2" pageNumber="71">Abdomen</emphasis>
(
<figureCitation id="799285EDFFC2167B17631CEFFE3244EC" box="[319,402,1233,1259]" captionStart="FIGURE 1" captionStartId="1.[151,250,1994,2019]" captionTargetBox="[184,1404,824,1957]" captionTargetId="figure@1.[169,1418,813,1969]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of C. lephuocdieui sp. nov. (1a) holotype male &amp; (1b) paratype female. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3864986" httpUri="https://zenodo.org/record/3864986/files/figure.png" pageId="2" pageNumber="71">Figs 1a</figureCitation>
, 
<figureCitation id="799285EDFFC2167B17C11CEFFE0A44EB" box="[413,426,1233,1260]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="2" pageNumber="71">3</figureCitation>
a–b). S1 largely whitish, black dorsally; S2–6 black dorsally, whitish ventrally with subapical white annulus from S3–6; S8–9 largely black, ventral margin with yellowish stripe; S10 black dorsally (
<figureCitation id="799285EDFFC2167B13621CCBFA2E4508" box="[1342,1422,1269,1295]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="2" pageNumber="71">Fig. 3a</figureCitation>
), lateral and ventral yellowish (
<figureCitation id="799285EDFFC2167B17B91D27FD984533" box="[485,568,1305,1332]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="2" pageNumber="71">Fig. 3b</figureCitation>
).
</paragraph>
<paragraph id="E1169968FFC2167B169B1D03FC7345C4" blockId="2.[151,1437,1017,2016]" pageId="2" pageNumber="71">
<emphasis id="D3DD457AFFC2167B169B1D03FE2B4550" box="[199,395,1341,1367]" italics="true" pageId="2" pageNumber="71">Anal appendages</emphasis>
(
<figureCitation id="799285EDFFC2167B17C01D03FE7E455F" box="[412,478,1341,1368]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="2" pageNumber="71">Fig. 3</figureCitation>
a–b) yellowish apart from black ventral margin of cerci and tip of paraprocts. Cerci as long as S10, ventral side with blackish carina, forming a point. Paraprocts slightly longer than cerci, gently rising with tip curved medially and ending in a black tooth. In dorsal view (
<figureCitation id="799285EDFFC2167B15891DBBFB874598" box="[981,1063,1413,1439]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="2" pageNumber="71">Fig. 3a</figureCitation>
), cerci yellowish, apex smoothly rounded but in lateral view (
<figureCitation id="799285EDFFC2167B17881D97FD8645C3" box="[468,550,1449,1476]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="2" pageNumber="71">Fig. 3b</figureCitation>
), cerci expanding near base, tip acute.
</paragraph>
<paragraph id="E1169968FFC2167B169B1DF3FDDF460C" blockId="2.[151,1437,1017,2016]" pageId="2" pageNumber="71">
<emphasis id="D3DD457AFFC2167B169B1DF3FEC345E0" box="[199,355,1485,1511]" italics="true" pageId="2" pageNumber="71">Genital ligula</emphasis>
(
<figureCitation id="799285EDFFC2167B172F1DF3FE1545EF" box="[371,437,1485,1512]" captionStart="FIGURE 4" captionStartId="4.[151,250,1074,1099]" captionTargetBox="[159,1428,187,1039]" captionTargetId="figure@4.[151,1436,181,1049]" captionTargetPageId="4" captionText="FIGURE 4. Genital ligula of Coeliccia spp. in dorsal and lateral view. (4a, b) C. lephuocdieui sp. nov., holotype male; (4c, d) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864992" httpUri="https://zenodo.org/record/3864992/files/figure.png" pageId="2" pageNumber="71">Fig. 4</figureCitation>
a–b) bifurcate distally with a pair of long flagella. Terminal segment with a large triangular-shaped flap distally and thin apical fold.
</paragraph>
<paragraph id="E1169968FFC2167B169B1E2BFC664628" blockId="2.[151,1437,1017,2016]" box="[199,966,1557,1583]" pageId="2" pageNumber="71">Measurements (in mm). HW 30; abdomen including appendages 46.</paragraph>
<paragraph id="E1169968FFC2167B169B1E06FC6846B8" blockId="2.[151,1437,1017,2016]" pageId="2" pageNumber="71">
<emphasis id="D3DD457AFFC2167B169B1E06FD844654" bold="true" box="[199,548,1592,1619]" pageId="2" pageNumber="71">
Variation in 
<typeStatus id="3E1227CAFFC2167B173F1E07FE6E4654" box="[355,462,1593,1619]" pageId="2" pageNumber="71" type="paratype">paratype</typeStatus>
males.
</emphasis>
One male smaller: HW 
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, abdomen including appendages 
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, but higher number of px, 
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FW and 
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HW. In another 
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male, prothorax entirely black, not dark yellowish as in 
<typeStatus id="3E1227CAFFC2167B168E1EBFFE94469C" box="[210,308,1665,1691]" pageId="2" pageNumber="71" type="holotype">holotype</typeStatus>
and HW 
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, abdomen including appendages 
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. The other 
<typeStatus id="3E1227CAFFC2167B12681EBFFB34469C" box="[1076,1172,1665,1691]" pageId="2" pageNumber="71" type="paratype">paratype</typeStatus>
male with HW 
<quantity id="2651348DFFC2167B131B1EBFFA38469C" box="[1351,1432,1665,1691]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="71" unit="mm" value="30.0">30 mm</quantity>
, abdomen including appendages 
<quantity id="2651348DFFC2167B17A31E9BFDF046B8" box="[511,592,1701,1727]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="2" pageNumber="71" unit="mm" value="48.0">48 mm</quantity>
, the number of px as in 
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.
</paragraph>
<paragraph id="E1169968FFC2167B169B1EF6FF674790" blockId="2.[151,1437,1017,2016]" pageId="2" pageNumber="71">
<emphasis id="D3DD457AFFC2167B169B1EF6FE6E46E4" bold="true" box="[199,462,1736,1763]" pageId="2" pageNumber="71">Description of female.</emphasis>
<emphasis id="D3DD457AFFC2167B17891EF7FDB246E4" box="[469,530,1737,1763]" italics="true" pageId="2" pageNumber="71">Head</emphasis>
(
<figureCitation id="799285EDFFC2167B147D1EF7FDC246E4" box="[545,610,1736,1763]" captionStart="FIGURE 5" captionStartId="5.[151,250,1616,1641]" captionTargetBox="[151,1436,528,1592]" captionTargetId="figure@5.[151,1436,525,1592]" captionTargetPageId="5" captionText="FIGURE 5. Structure features of Coeliccia lephuocdieui sp. nov. (5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="2" pageNumber="71">Fig. 5</figureCitation>
a–b). Labrum and postclypeus entirely shining black. Anteclypeus yellow with two black spots left and right on lower margin. Mandibles and genae yellow, this pattern connecting above with postclypeus. Head dorsally matte black with two long yellow stripes each extending from eye margin to posterior ocellus, two oval yellow spots adjacent the anterior ocellus and two oblong yellowish postocular spots. Antennae black, top of first segment yellow. Compound eyes in life as in the male (
<figureCitation id="799285EDFFC2167B15881F67FB864774" box="[980,1062,1881,1907]" captionStart="FIGURE 8" captionStartId="9.[151,250,1783,1808]" captionTargetBox="[151,1436,597,1757]" captionTargetId="figure@9.[151,1436,597,1757]" captionTargetPageId="9" captionText="FIGURE 8. Habitus of Coeliccia spp. in nature. (8a) C. lephuocdieui sp. nov. in the type location, 21 April 2019 (with a red parasitic mite on the dorsum synthorax of the male); (8b) C. scutellum, Son Tra Nature Reserve, 25 April 2018. Photographed by the first author." figureDoi="http://doi.org/10.5281/zenodo.3865001" httpUri="https://zenodo.org/record/3865001/files/figure.png" pageId="2" pageNumber="71">Fig. 8a</figureCitation>
), black dorsally and greenish below.
</paragraph>
<paragraph id="E1169968FFC2167A169B1F9FFCAD4118" blockId="2.[151,1437,1017,2016]" lastBlockId="3.[151,1436,152,323]" lastPageId="3" lastPageNumber="72" pageId="2" pageNumber="71">
<emphasis id="D3DD457AFFC2167B169B1F9FFEB847BC" box="[199,280,1953,1979]" italics="true" pageId="2" pageNumber="71">Thorax</emphasis>
(
<figureCitation id="799285EDFFC2167B17741F9FFE0A47BC" box="[296,426,1952,1979]" captionStart="FIGURE 5" captionStartId="5.[151,250,1616,1641]" captionTargetBox="[151,1436,528,1592]" captionTargetId="figure@5.[151,1436,525,1592]" captionTargetPageId="5" captionText="FIGURE 5. Structure features of Coeliccia lephuocdieui sp. nov. (5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="2" pageNumber="71">Figs 5a, 5d</figureCitation>
, 
<figureCitation id="799285EDFFC2167B17E41F9FFE6647BB" box="[440,454,1953,1980]" captionStart="FIGURE 6" captionStartId="6.[151,250,1269,1294]" captionTargetBox="[167,1425,189,1233]" captionTargetId="figure@6.[151,1436,181,1244]" captionTargetPageId="6" captionText="FIGURE 6. Posterior pronotal lobe of prothorax of Coeliccia spp., all female. (6a, b) C. lephuocdieui sp. nov.; (6c, d) C. scutellum, Kon Ka Kinh National Park; (6e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864997" httpUri="https://zenodo.org/record/3864997/files/figure.png" pageId="2" pageNumber="71">6</figureCitation>
a–b). Prothorax: Anterior, posterior and dorsal of middle pronotal lobes black, remaining area yellowish (
<figureCitation id="799285EDFFC2167B17281FFBFE4147E7" box="[372,481,1988,2016]" captionStart="FIGURE 5" captionStartId="5.[151,250,1616,1641]" captionTargetBox="[151,1436,528,1592]" captionTargetId="figure@5.[151,1436,525,1592]" captionTargetPageId="5" captionText="FIGURE 5. Structure features of Coeliccia lephuocdieui sp. nov. (5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="2" pageNumber="71">Fig. 5a, d</figureCitation>
). Posterior margin of posterior pronotal lobe slightly convex in lateral view (
<figureCitation id="799285EDFFC2167B13611FFBFA2E47E7" box="[1341,1422,1989,2016]" captionStart="FIGURE 6" captionStartId="6.[151,250,1269,1294]" captionTargetBox="[167,1425,189,1233]" captionTargetId="figure@6.[151,1436,181,1244]" captionTargetPageId="6" captionText="FIGURE 6. Posterior pronotal lobe of prothorax of Coeliccia spp., all female. (6a, b) C. lephuocdieui sp. nov.; (6c, d) C. scutellum, Kon Ka Kinh National Park; (6e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864997" httpUri="https://zenodo.org/record/3864997/files/figure.png" pageId="2" pageNumber="71">Fig. 6a</figureCitation>
), triangular-shaped in dorsal view (
<figureCitation id="799285EDFFC3167A144E18A7FDC440B3" box="[530,612,153,180]" captionStart="FIGURE 6" captionStartId="6.[151,250,1269,1294]" captionTargetBox="[167,1425,189,1233]" captionTargetId="figure@6.[151,1436,181,1244]" captionTargetPageId="6" captionText="FIGURE 6. Posterior pronotal lobe of prothorax of Coeliccia spp., all female. (6a, b) C. lephuocdieui sp. nov.; (6c, d) C. scutellum, Kon Ka Kinh National Park; (6e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864997" httpUri="https://zenodo.org/record/3864997/files/figure.png" pageId="3" pageNumber="72">Fig. 6b</figureCitation>
) and with two distinct lapels. Synthorax (
<figureCitation id="799285EDFFC3167A126118A7FB3040B4" box="[1085,1168,152,179]" captionStart="FIGURE 5" captionStartId="5.[151,250,1616,1641]" captionTargetBox="[151,1436,528,1592]" captionTargetId="figure@5.[151,1436,525,1592]" captionTargetPageId="5" captionText="FIGURE 5. Structure features of Coeliccia lephuocdieui sp. nov. (5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="3" pageNumber="72">Fig. 5a</figureCitation>
): mesepisternum with a long, narrow yellowish antehumeral stripe; mesepimeron entirely black; metepisternum and metepimeron mostly yellow except for narrow metepleural black stripe along lower margin of metepisternum. Mesinfraepisternum and metinfraepisternum entirely yellow. 
<emphasis id="D3DD457AFFC3167A14721938FDC34118" box="[558,611,262,287]" italics="true" pageId="3" pageNumber="72">Legs</emphasis>
as in the male.
</paragraph>
<paragraph id="E1169968FFC3167A169B1914FBDF4144" blockId="3.[151,1436,152,323]" box="[199,1151,296,323]" pageId="3" pageNumber="72">
<emphasis id="D3DD457AFFC3167A169B1914FEAB4144" box="[199,267,298,323]" italics="true" pageId="3" pageNumber="72">Wings</emphasis>
. Hyaline, 19 px in FW and 
<quantity id="2651348DFFC3167A141F1917FDDC4144" box="[579,636,297,323]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="3" pageNumber="72" unit="in" value="18.0">18 in</quantity>
HW. Pt brown, covering 1.5 underlying cells.
</paragraph>
<caption id="B5D6C9E0FFC3167A16CB1F1CFCBD4758" ID-DOI="http://doi.org/10.5281/zenodo.3864990" ID-Zenodo-Dep="3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="3" pageNumber="72" startId="3.[151,250,1826,1851]" targetBox="[151,1436,347,1800]" targetPageId="3">
<paragraph id="E1169968FFC3167A16CB1F1CFCBD4758" blockId="3.[151,1437,1826,1887]" pageId="3" pageNumber="72">
<emphasis id="D3DD457AFFC3167A16CB1F1CFEAE473C" bold="true" box="[151,270,1826,1851]" pageId="3" pageNumber="72">FIGURE 3</emphasis>
. Appendages 
<taxonomicName id="26A9E2EBFFC3167A17C21F1CFE5F473D" authorityName="Kirby" authorityYear="1890" box="[414,511,1826,1850]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="D3DD457AFFC3167A17C21F1CFE5F473D" box="[414,511,1826,1850]" italics="true" pageId="3" pageNumber="72">Coeliccia</emphasis>
</taxonomicName>
spp. in dorsal and lateral view. (3a, b) 
<taxonomicName id="26A9E2EBFFC3167A15CD1F1DFB92473D" authority="Phan &amp; Ngo &amp; Bui, 2020" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[913,1074,1826,1850]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="72" phylum="Arthropoda" rank="species" species="lephuocdieui" status="sp. nov.">
<emphasis id="D3DD457AFFC3167A15CD1F1DFB92473D" box="[913,1074,1826,1850]" italics="true" pageId="3" pageNumber="72">C. lephuocdieui</emphasis>
</taxonomicName>
<emphasis id="D3DD457AFFC3167A12651F1CFB2B473D" bold="true" box="[1081,1163,1826,1850]" pageId="3" pageNumber="72">
<taxonomicNameLabel id="C8EEF801FFC3167A12651F1CFB2B473D" box="[1081,1163,1826,1850]" pageId="3" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; (3c, d) 
<emphasis id="D3DD457AFFC3167A12B81F1DFACA473D" box="[1252,1386,1826,1850]" italics="true" pageId="3" pageNumber="72">
<taxonomicName id="26A9E2EBFFC3167A12B81F1DFAC6473D" authorityName="Laidlaw" authorityYear="1932" box="[1252,1382,1826,1850]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="72" phylum="Arthropoda" rank="species" species="scutellum">C. scutellum</taxonomicName>
,
</emphasis>
Kon Ka Kinh National Park; (3e, f) 
<taxonomicName id="26A9E2EBFFC3167A178E1F79FDF64759" authorityName="Asahina" authorityYear="1984" box="[466,598,1862,1886]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="72" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC3167A178E1F79FDF64759" box="[466,598,1862,1886]" italics="true" pageId="3" pageNumber="72">C. yamasakii</emphasis>
</taxonomicName>
, Phu Quoc Islands.
</paragraph>
</caption>
<caption id="B5D6C9E0FFC4167D16CB1C0CFE424468" ID-DOI="http://doi.org/10.5281/zenodo.3864992" ID-Zenodo-Dep="3864992" httpUri="https://zenodo.org/record/3864992/files/figure.png" pageId="4" pageNumber="73" startId="4.[151,250,1074,1099]" targetBox="[159,1428,187,1039]" targetPageId="4">
<paragraph id="E1169968FFC4167D16CB1C0CFE424468" blockId="4.[151,1437,1074,1135]" pageId="4" pageNumber="73">
<emphasis id="D3DD457AFFC4167D16CB1C0CFEAD444C" bold="true" box="[151,269,1074,1099]" pageId="4" pageNumber="73">FIGURE 4</emphasis>
. Genital ligula of 
<taxonomicName id="26A9E2EBFFC4167D17941C0DFD89444C" authorityName="Kirby" authorityYear="1890" box="[456,553,1075,1099]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="D3DD457AFFC4167D17941C0DFD89444C" box="[456,553,1075,1099]" italics="true" pageId="4" pageNumber="73">Coeliccia</emphasis>
</taxonomicName>
spp. in dorsal and lateral view. (4a, b) 
<taxonomicName id="26A9E2EBFFC4167D15EA1C0DFBF7444C" authority="Phan &amp; Ngo &amp; Bui, 2020" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[950,1111,1075,1099]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="lephuocdieui" status="sp. nov.">
<emphasis id="D3DD457AFFC4167D15EA1C0DFBF7444C" box="[950,1111,1075,1099]" italics="true" pageId="4" pageNumber="73">C. lephuocdieui</emphasis>
</taxonomicName>
<emphasis id="D3DD457AFFC4167D12011C0DFB0E444C" bold="true" box="[1117,1198,1075,1099]" pageId="4" pageNumber="73">
<taxonomicNameLabel id="C8EEF801FFC4167D12011C0DFB0E444C" box="[1117,1198,1075,1099]" pageId="4" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male; (4c, d) 
<emphasis id="D3DD457AFFC4167D16CB1C69FE824468" box="[151,290,1111,1135]" italics="true" pageId="4" pageNumber="73">
<taxonomicName id="26A9E2EBFFC4167D16CB1C69FEBE4468" authorityName="Asahina" authorityYear="1984" box="[151,286,1111,1135]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="yamasakii">C. yamasakii</taxonomicName>
,
</emphasis>
Phu Quoc Islands.
</paragraph>
</caption>
<paragraph id="E1169968FFC4167D169B1CA3FDA34524" blockId="4.[151,1437,1180,2036]" pageId="4" pageNumber="73">
<emphasis id="D3DD457AFFC4167D169B1CA3FE9144B0" box="[199,305,1181,1207]" italics="true" pageId="4" pageNumber="73">Abdomen</emphasis>
(
<figureCitation id="799285EDFFC4167D171E1CA3FE3C44B0" box="[322,412,1181,1207]" captionStart="FIGURE 2" captionStartId="2.[151,250,910,935]" captionTargetBox="[151,1436,181,884]" captionTargetId="figure@2.[151,1436,181,884]" captionTargetPageId="2" captionText="FIGURE 2. Head and thorax of male, lateral view. (2a) C. lephuocdieui sp. nov., holotype male; (2b) C. scutellum, Kon Ka Kinh National Park." figureDoi="http://doi.org/10.5281/zenodo.3864988" httpUri="https://zenodo.org/record/3864988/files/figure.png" pageId="4" pageNumber="73">Figs 2b</figureCitation>
, 
<figureCitation id="799285EDFFC4167D17F61CA2FE6444B0" box="[426,452,1180,1207]" captionStart="FIGURE 5" captionStartId="5.[151,250,1616,1641]" captionTargetBox="[151,1436,528,1592]" captionTargetId="figure@5.[151,1436,525,1592]" captionTargetPageId="5" captionText="FIGURE 5. Structure features of Coeliccia lephuocdieui sp. nov. (5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="4" pageNumber="73">5e</figureCitation>
). S1 largely yellow with posterior margin and dorsum narrowly black; dorsal half S2 black, yellow ventrally; S3–7 dark brownish dorsally, ventral margin yellowish with subapical yellow annulus; S8 black with an oval yellowish spot laterally, lower margin with yellow stripe; S9–10 mostly yellowish, ventral and posterior margin black (
<figureCitation id="799285EDFFC4167D17FF1D37FE564524" box="[419,502,1288,1315]" captionStart="FIGURE 5" captionStartId="5.[151,250,1616,1641]" captionTargetBox="[151,1436,528,1592]" captionTargetId="figure@5.[151,1436,525,1592]" captionTargetPageId="5" captionText="FIGURE 5. Structure features of Coeliccia lephuocdieui sp. nov. (5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="4" pageNumber="73">Fig. 5e</figureCitation>
).
</paragraph>
<paragraph id="E1169968FFC4167D169B1D13FF74456B" blockId="4.[151,1437,1180,2036]" pageId="4" pageNumber="73">
<emphasis id="D3DD457AFFC4167D169B1D13FE2B4540" box="[199,395,1325,1351]" italics="true" pageId="4" pageNumber="73">Anal appendages</emphasis>
(
<figureCitation id="799285EDFFC4167D17C11D13FE4E4540" box="[413,494,1324,1351]" captionStart="FIGURE 5" captionStartId="5.[151,250,1616,1641]" captionTargetBox="[151,1436,528,1592]" captionTargetId="figure@5.[151,1436,525,1592]" captionTargetPageId="5" captionText="FIGURE 5. Structure features of Coeliccia lephuocdieui sp. nov. (5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="4" pageNumber="73">Fig. 5e</figureCitation>
). Cerci black, half S10; ovipositor black with yellow anterior marking dorsally at apex.
</paragraph>
<paragraph id="E1169968FFC4167D169B1D48FBAF4588" blockId="4.[151,1437,1180,2036]" box="[199,1039,1397,1424]" pageId="4" pageNumber="73">Measurements (in mm). Hind wing 30; abdomen including appendages 43.</paragraph>
<paragraph id="E1169968FFC4167D169B1DA6FD3345FB" blockId="4.[151,1437,1180,2036]" pageId="4" pageNumber="73">
<emphasis id="D3DD457AFFC4167D169B1DA6FD9245B4" bold="true" box="[199,562,1432,1459]" pageId="4" pageNumber="73">
Variation in 
<typeStatus id="3E1227CAFFC4167D17021DA7FE6945B4" box="[350,457,1433,1459]" pageId="4" pageNumber="73" type="paratype">paratype</typeStatus>
females.
</emphasis>
<typeStatus id="3E1227CAFFC4167D14651DA4FD3B45B3" box="[569,667,1434,1460]" pageId="4" pageNumber="73" type="paratype">Paratype</typeStatus>
females show no significant morphological differences from female used for the description apart from size, ranging from 
<quantity id="2651348DFFC4167D155E1D83FCDD45DF" box="[770,893,1469,1496]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="4" pageNumber="73" unit="mm" value="30.5" valueMax="31.0" valueMin="30.0">30–31 mm</quantity>
(HW), 
<quantity id="2651348DFFC4167D15891D83FBF045DF" box="[981,1104,1469,1496]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="4" pageNumber="73" unit="mm" value="42.5" valueMax="44.0" valueMin="41.0">41–44 mm</quantity>
(abdomen including appendages) and px 
<quantity id="2651348DFFC4167D17701DDFFE2F45FC" box="[300,399,1505,1531]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" metricValueMax="5.334" metricValueMin="4.826" pageId="4" pageNumber="73" unit="in" value="20.0" valueMax="21.0" valueMin="19.0">19–21 in</quantity>
FW and 
<quantity id="2651348DFFC4167D17A91DDFFDF845FC" box="[501,600,1505,1531]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="4" pageNumber="73" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">18–20 in</quantity>
HW.
</paragraph>
<paragraph id="E1169968FFC4167D169B1E3AFEC24643" blockId="4.[151,1437,1180,2036]" pageId="4" pageNumber="73">
<emphasis id="D3DD457AFFC4167D169B1E3AFE1C4618" bold="true" box="[199,444,1540,1567]" pageId="4" pageNumber="73">Habitat and ecology.</emphasis>
Adults were collected at a dry narrow, densely vegetated stream within a humid, pristine forest (
<figureCitation id="799285EDFFC4167D16BA1E17FE894643" box="[230,297,1577,1604]" captionStart="FIGURE 9" captionStartId="10.[151,250,1117,1142]" captionTargetBox="[152,1436,183,1092]" captionTargetId="figure@10.[151,1436,181,1092]" captionTargetPageId="10" captionText="FIGURE 9. Habitat of Coeliccia lephuocdieui sp. nov. (9a) a section of the type location; (9b) the second author was collecting aquatic insects in the type location. Photographed by the first author." figureDoi="http://doi.org/10.5281/zenodo.3865003" httpUri="https://zenodo.org/record/3865003/files/figure.png" pageId="4" pageNumber="73">Fig. 9</figureCitation>
a–b).
</paragraph>
</subSubSection>
<subSubSection id="A9B3CAE3FFC4167C169B1E72FC0D4761" lastPageId="5" lastPageNumber="74" pageId="4" pageNumber="73" type="diagnosis">
<paragraph id="E1169968FFC4167C169B1E72FE9B41F0" blockId="4.[151,1437,1180,2036]" lastBlockId="5.[151,1437,153,503]" lastPageId="5" lastPageNumber="74" pageId="4" pageNumber="73">
<emphasis id="D3DD457AFFC4167D169B1E72FE754660" bold="true" box="[199,469,1612,1639]" pageId="4" pageNumber="73">Differential Diagnosis.</emphasis>
Within 
<taxonomicName id="26A9E2EBFFC4167D146A1E73FD014660" authorityName="Kirby" authorityYear="1890" box="[566,673,1613,1639]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="D3DD457AFFC4167D146A1E73FD014660" box="[566,673,1613,1639]" italics="true" pageId="4" pageNumber="73">Coeliccia</emphasis>
</taxonomicName>
, the new species and 
<emphasis id="D3DD457AFFC4167D15F91E70FB9B4660" box="[933,1083,1613,1640]" italics="true" pageId="4" pageNumber="73">
<taxonomicName id="26A9E2EBFFC4167D15F91E70FB974660" authorityName="Laidlaw" authorityYear="1932" box="[933,1079,1613,1640]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="scutellum">C. scutellum</taxonomicName>
,
</emphasis>
<taxonomicName id="26A9E2EBFFC4167D12191E70FA354660" authority="Laidlaw, 1932" authorityName="Laidlaw" authorityYear="1932" box="[1093,1429,1613,1640]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="hainanense">
<emphasis id="D3DD457AFFC4167D12191E70FB494660" box="[1093,1257,1613,1640]" italics="true" pageId="4" pageNumber="73">C. hainanense</emphasis>
Laidlaw, 1932
</taxonomicName>
, 
<taxonomicName id="26A9E2EBFFC4167D16CB1E4CFE14468B" authority="(Selys, 1886)" baseAuthorityName="Selys" baseAuthorityYear="1886" box="[151,436,1649,1676]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="renifera">
<emphasis id="D3DD457AFFC4167D16CB1E4CFEB2468C" box="[151,274,1649,1676]" italics="true" pageId="4" pageNumber="73">C. renifera</emphasis>
(Selys, 1886)
</taxonomicName>
, 
<taxonomicName id="26A9E2EBFFC4167D179D1E4CFD42468C" authority="Asahina, 1970" authorityName="Asahina" authorityYear="1970" box="[449,738,1649,1676]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="svihleri">
<emphasis id="D3DD457AFFC4167D179D1E4CFD96468C" box="[449,566,1649,1676]" italics="true" pageId="4" pageNumber="73">C. svihleri</emphasis>
<bibRefCitation id="8538E499FFC4167D14631E4FFD42468C" author="Asahina, S." box="[575,738,1649,1675]" pageId="4" pageNumber="73" pagination="98 - 126" refId="ref5115" refString="Asahina, S. (1970) Burmese Odonata collected by Dr. Arthur Svihla with supplementary notes on Asiatic Ceriagrion species. Japanese Journal of Zoology, 16, 98 - 126." type="journal article" year="1970">Asahina, 1970</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="26A9E2EBFFC4167D15471E4CFC0F468C" authorityName="Asahina" authorityYear="1984" box="[795,943,1649,1676]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC4167D15471E4CFC0F468C" box="[795,943,1649,1676]" italics="true" pageId="4" pageNumber="73">C. yamasakii</emphasis>
</taxonomicName>
can be defined as a group characterized by the combination of following male characters: synthorax with three separated large yellowish (sometimes greenish) markings (apart from 
<taxonomicName id="26A9E2EBFFC4167D17941E84FDDC46D4" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[456,636,1721,1748]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC4167D17941E84FDDC46D4" box="[456,636,1721,1748]" italics="true" pageId="4" pageNumber="73">C. lephuocdieui</emphasis>
</taxonomicName>
), genital ligula with two thin apico-lateral flagella and abdominal tip (excluding 
<taxonomicName id="26A9E2EBFFC4167D174A1EE0FE6846F0" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[278,456,1757,1784]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC4167D174A1EE0FE6846F0" box="[278,456,1757,1784]" italics="true" pageId="4" pageNumber="73">C. lephuocdieui</emphasis>
</taxonomicName>
and 
<taxonomicName id="26A9E2EBFFC4167D17A21EE0FDD746F0" baseAuthorityName="Selys" baseAuthorityYear="1886" box="[510,631,1757,1784]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="renifera">
<emphasis id="D3DD457AFFC4167D17A21EE0FDD746F0" box="[510,631,1757,1784]" italics="true" pageId="4" pageNumber="73">C. renifera</emphasis>
</taxonomicName>
) including appendages yellowish (
<bibRefCitation id="8538E499FFC4167D15A11EE3FB3D46F0" author="Asahina, S." box="[1021,1181,1757,1783]" pageId="4" pageNumber="73" pagination="98 - 126" refId="ref5115" refString="Asahina, S. (1970) Burmese Odonata collected by Dr. Arthur Svihla with supplementary notes on Asiatic Ceriagrion species. Japanese Journal of Zoology, 16, 98 - 126." type="journal article" year="1970">Asahina 1970</bibRefCitation>
, 
<bibRefCitation id="8538E499FFC4167D12F41EE3FB4346F0" author="Asahina, S." box="[1192,1251,1757,1783]" pageId="4" pageNumber="73" pagination="2 - 20" refId="ref5149" refString="Asahina, S. (1984) A list of the Odonata recorded from Thailand, 6: Platycemididae, genus Coeliccia. Tombo, 27, 2 - 20." type="journal article" year="1984">1984</bibRefCitation>
, 
<bibRefCitation id="8538E499FFC4167D12B21EE2FA8946F0" author="Asahina, S." box="[1262,1321,1756,1783]" pageId="4" pageNumber="73" pagination="2 - 13" refId="ref5180" refString="Asahina, S. (1985) Further contributions to the taxonomy of Southasiatic Coeliccia species (Odonata, Platycnemididae). Cho cho, 8 (2), 2 - 13." type="journal article" year="1985">1985</bibRefCitation>
; 
<bibRefCitation id="8538E499FFC4167D13691EE3FE86471C" author="Steinhoff, P. O. M. &amp; Uhl, G." pageId="4" pageNumber="73" pagination="257 - 276" refId="ref5561" refString="Steinhoff, P. O. M. &amp; Uhl, G. (2015) Taxonomy and nomenclature of some mainland SE-Asian Coeliccia species (Odonata, Platycnemididae) using micro-CT analysis. Zootaxa, 4059 (2), 257 - 276. https: // doi. org / 10.11646 / zootaxa. 4059.2.2" type="journal article" year="2015">Steinhoff &amp; Uhl 2015</bibRefCitation>
). 
<taxonomicName id="26A9E2EBFFC4167D17671F3FFDE0471C" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[315,576,1793,1819]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC4167D17671F3FFDE0471C" box="[315,576,1793,1819]" italics="true" pageId="4" pageNumber="73">Coeliccia lephuocdieui</emphasis>
</taxonomicName>
differs from all of the above species by lacking the dorsal yellow marking on its synthorax (
<figureCitation id="799285EDFFC4167D173C1F1BFE104738" box="[352,432,1829,1855]" captionStart="FIGURE 1" captionStartId="1.[151,250,1994,2019]" captionTargetBox="[184,1404,824,1957]" captionTargetId="figure@1.[169,1418,813,1969]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of C. lephuocdieui sp. nov. (1a) holotype male &amp; (1b) paratype female. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3864986" httpUri="https://zenodo.org/record/3864986/files/figure.png" pageId="4" pageNumber="73">Fig. 1a</figureCitation>
) and combination of the morphological differences in the male appendages and female prothoracic structures. The following comparisons are based on the new species and two Vietnamese species 
<taxonomicName id="26A9E2EBFFC4167D13DF1F74FEA24780" authorityName="Laidlaw" authorityYear="1932" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="scutellum">
<emphasis id="D3DD457AFFC4167D13DF1F74FEA24780" italics="true" pageId="4" pageNumber="73">C. scutellum</emphasis>
</taxonomicName>
and 
<taxonomicName id="26A9E2EBFFC4167D17671F50FE6F4780" authorityName="Asahina" authorityYear="1984" box="[315,463,1901,1928]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC4167D17671F50FE6F4780" box="[315,463,1901,1928]" italics="true" pageId="4" pageNumber="73">C. yamasakii</emphasis>
</taxonomicName>
apart from 
<taxonomicName id="26A9E2EBFFC4167D14091F50FD574780" authorityName="Laidlaw" authorityYear="1932" box="[597,759,1901,1928]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="hainanense">
<emphasis id="D3DD457AFFC4167D14091F50FD574780" box="[597,759,1901,1928]" italics="true" pageId="4" pageNumber="73">C. hainanense</emphasis>
</taxonomicName>
which we were not examined. Male of 
<taxonomicName id="26A9E2EBFFC4167D12E71F50FACE4780" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[1211,1390,1901,1928]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC4167D12E71F50FACE4780" box="[1211,1390,1901,1928]" italics="true" pageId="4" pageNumber="73">C. lephuocdieui</emphasis>
</taxonomicName>
can be easily separated from both 
<taxonomicName id="26A9E2EBFFC4167D17BB1FACFDD347AC" authorityName="Laidlaw" authorityYear="1932" box="[487,627,1937,1964]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="scutellum">
<emphasis id="D3DD457AFFC4167D17BB1FACFDD347AC" box="[487,627,1937,1964]" italics="true" pageId="4" pageNumber="73">C. scutellum</emphasis>
</taxonomicName>
and 
<taxonomicName id="26A9E2EBFFC4167D14F51FACFC9B47AC" authorityName="Asahina" authorityYear="1984" box="[681,827,1937,1964]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC4167D14F51FACFC9B47AC" box="[681,827,1937,1964]" italics="true" pageId="4" pageNumber="73">C. yamasakii</emphasis>
</taxonomicName>
by following characters: lacking the large oval yellow antehumeral stripe (
<figureCitation id="799285EDFFC4167D17261F8BFE6B47C8" box="[378,459,1973,1999]" captionStart="FIGURE 1" captionStartId="1.[151,250,1994,2019]" captionTargetBox="[184,1404,824,1957]" captionTargetId="figure@1.[169,1418,813,1969]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of C. lephuocdieui sp. nov. (1a) holotype male &amp; (1b) paratype female. Scale bar is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3864986" httpUri="https://zenodo.org/record/3864986/files/figure.png" pageId="4" pageNumber="73">Fig. 1a</figureCitation>
); dorsal S10 of 
<taxonomicName id="26A9E2EBFFC4167D14DA1F88FC9947C8" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[646,825,1973,2000]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC4167D14DA1F88FC9947C8" box="[646,825,1973,2000]" italics="true" pageId="4" pageNumber="73">C. lephuocdieui</emphasis>
</taxonomicName>
black, yellowish latero-ventrally (
<figureCitation id="799285EDFFC4167D129E1F8BFAA447D7" box="[1218,1284,1973,2000]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="4" pageNumber="73">Fig. 3</figureCitation>
a–b) whereas dorsal (in 
<taxonomicName id="26A9E2EBFFC4167D17551FE4FE3C47F4" authorityName="Asahina" authorityYear="1984" box="[265,412,2009,2036]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC4167D17551FE4FE3C47F4" box="[265,412,2009,2036]" italics="true" pageId="4" pageNumber="73">C. yamasakii</emphasis>
</taxonomicName>
) or all (in 
<taxonomicName id="26A9E2EBFFC4167D14481FE4FD0047F4" authorityName="Laidlaw" authorityYear="1932" box="[532,672,2009,2036]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="scutellum">
<emphasis id="D3DD457AFFC4167D14481FE4FD0047F4" box="[532,672,2009,2036]" italics="true" pageId="4" pageNumber="73">C. scutellum</emphasis>
</taxonomicName>
) S10 yellowish (
<figureCitation id="799285EDFFC4167D153C1FE7FC0247F3" box="[864,930,2009,2036]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="4" pageNumber="73">Fig. 3</figureCitation>
c–f); cerci of 
<taxonomicName id="26A9E2EBFFC4167D12671FE4FB4D47F4" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[1083,1261,2009,2036]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC4167D12671FE4FB4D47F4" box="[1083,1261,2009,2036]" italics="true" pageId="4" pageNumber="73">C. lephuocdieui</emphasis>
</taxonomicName>
as long as S10, black ventrally, expanding basally, tip acute in lateral view (
<figureCitation id="799285EDFFC5167C156B18A7FC2840B3" box="[823,904,153,180]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="5" pageNumber="74">Fig. 3b</figureCitation>
) while cerci of 
<taxonomicName id="26A9E2EBFFC5167C126418A7FB6340B4" authorityName="Laidlaw" authorityYear="1932" box="[1080,1219,153,179]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="scutellum">
<emphasis id="D3DD457AFFC5167C126418A7FB6340B4" box="[1080,1219,153,179]" italics="true" pageId="5" pageNumber="74">C. scutellum</emphasis>
</taxonomicName>
clearly longer than S10, entirely yellowish with tip smoothly rounded (
<figureCitation id="799285EDFFC5167C14931883FCAF40DF" box="[719,783,189,216]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="5" pageNumber="74">Fig. 3</figureCitation>
c–d). The new species differs from 
<taxonomicName id="26A9E2EBFFC5167C12C91883FA8640D0" authorityName="Asahina" authorityYear="1984" box="[1173,1318,189,215]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC5167C12C91883FA8640D0" box="[1173,1318,189,215]" italics="true" pageId="5" pageNumber="74">C. yamasakii</emphasis>
</taxonomicName>
by several differential characters: genae and the pattern between eye margin and antennae entirely black (
<figureCitation id="799285EDFFC5167C129F18DFFAB340FC" box="[1219,1299,224,251]" captionStart="FIGURE 5" captionStartId="5.[151,250,1616,1641]" captionTargetBox="[151,1436,528,1592]" captionTargetId="figure@5.[151,1436,525,1592]" captionTargetPageId="5" captionText="FIGURE 5. Structure features of Coeliccia lephuocdieui sp. nov. (5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="5" pageNumber="74">Fig. 5c</figureCitation>
) but blue in 
<taxonomicName id="26A9E2EBFFC5167C16CB193BFE8C4118" authorityName="Asahina" authorityYear="1984" box="[151,300,261,287]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC5167C16CB193BFE8C4118" box="[151,300,261,287]" italics="true" pageId="5" pageNumber="74">C. yamasakii</emphasis>
</taxonomicName>
(
<figureCitation id="799285EDFFC5167C1760193BFE324127" box="[316,402,261,288]" captionStart="FIGURE 7" captionStartId="8.[151,250,1303,1328]" captionTargetBox="[151,1436,181,1276]" captionTargetId="figure@8.[151,1436,181,1278]" captionTargetPageId="8" captionText="FIGURE 7. Structure features of Coeliccia yamasakii, Phu Quoc Islands. (7a) head &amp; thorax of male, lateral view; (7b) head of male, frontal view; (7c) head of female, frontal view; (7d) head &amp; thorax of female, lateral view; (7e) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864999" httpUri="https://zenodo.org/record/3864999/files/figure.png" pageId="5" pageNumber="74">Fig. 7b</figureCitation>
); male synthorax lacking oval spots and the yellow pattern covering most of metepimeron (
<figureCitation id="799285EDFFC5167C16FC1917FF4F4144" box="[160,239,297,323]" captionStart="FIGURE 2" captionStartId="2.[151,250,910,935]" captionTargetBox="[151,1436,181,884]" captionTargetId="figure@2.[151,1436,181,884]" captionTargetPageId="2" captionText="FIGURE 2. Head and thorax of male, lateral view. (2a) C. lephuocdieui sp. nov., holotype male; (2b) C. scutellum, Kon Ka Kinh National Park." figureDoi="http://doi.org/10.5281/zenodo.3864988" httpUri="https://zenodo.org/record/3864988/files/figure.png" pageId="5" pageNumber="74">Fig. 2a</figureCitation>
) but these spots present in 
<taxonomicName id="26A9E2EBFFC5167C14431917FD124144" authorityName="Asahina" authorityYear="1984" box="[543,690,297,323]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC5167C14431917FD124144" box="[543,690,297,323]" italics="true" pageId="5" pageNumber="74">C. yamasakii</emphasis>
</taxonomicName>
and yellow pattern covering about posterior 1/3rd of metepimeron (
<figureCitation id="799285EDFFC5167C16FC1973FF524160" box="[160,242,333,359]" captionStart="FIGURE 7" captionStartId="8.[151,250,1303,1328]" captionTargetBox="[151,1436,181,1276]" captionTargetId="figure@8.[151,1436,181,1278]" captionTargetPageId="8" captionText="FIGURE 7. Structure features of Coeliccia yamasakii, Phu Quoc Islands. (7a) head &amp; thorax of male, lateral view; (7b) head of male, frontal view; (7c) head of female, frontal view; (7d) head &amp; thorax of female, lateral view; (7e) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864999" httpUri="https://zenodo.org/record/3864999/files/figure.png" pageId="5" pageNumber="74">Fig. 7a</figureCitation>
); genital ligula with the widely apical flap (
<figureCitation id="799285EDFFC5167C14831973FC82416F" box="[735,802,333,360]" captionStart="FIGURE 4" captionStartId="4.[151,250,1074,1099]" captionTargetBox="[159,1428,187,1039]" captionTargetId="figure@4.[151,1436,181,1049]" captionTargetPageId="4" captionText="FIGURE 4. Genital ligula of Coeliccia spp. in dorsal and lateral view. (4a, b) C. lephuocdieui sp. nov., holotype male; (4c, d) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864992" httpUri="https://zenodo.org/record/3864992/files/figure.png" pageId="5" pageNumber="74">Fig. 4</figureCitation>
a–b) whereas in 
<taxonomicName id="26A9E2EBFFC5167C15801973FBCF4160" authorityName="Asahina" authorityYear="1984" box="[988,1135,333,359]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC5167C15801973FBCF4160" box="[988,1135,333,359]" italics="true" pageId="5" pageNumber="74">C. yamasakii</emphasis>
</taxonomicName>
, apical portion of terminal segment strongly prominent (
<figureCitation id="799285EDFFC5167C1782194FFD81418B" box="[478,545,369,396]" captionStart="FIGURE 4" captionStartId="4.[151,250,1074,1099]" captionTargetBox="[159,1428,187,1039]" captionTargetId="figure@4.[151,1436,181,1049]" captionTargetPageId="4" captionText="FIGURE 4. Genital ligula of Coeliccia spp. in dorsal and lateral view. (4a, b) C. lephuocdieui sp. nov., holotype male; (4c, d) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864992" httpUri="https://zenodo.org/record/3864992/files/figure.png" pageId="5" pageNumber="74">Fig. 4</figureCitation>
c–d); paraprocts yellowish (
<figureCitation id="799285EDFFC5167C150A194FFC06418B" box="[854,934,369,396]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="5" pageNumber="74">Fig. 3b</figureCitation>
) but dark brownish in 
<taxonomicName id="26A9E2EBFFC5167C12FD194FFA93418C" authorityName="Asahina" authorityYear="1984" box="[1185,1331,369,395]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC5167C12FD194FFA93418C" box="[1185,1331,369,395]" italics="true" pageId="5" pageNumber="74">C. yamasakii</emphasis>
</taxonomicName>
(
<figureCitation id="799285EDFFC5167C131D194FFA2F418B" box="[1345,1423,369,396]" captionStart="FIGURE 3" captionStartId="3.[151,250,1826,1851]" captionTargetBox="[151,1436,347,1800]" captionTargetId="figure@3.[151,1436,345,1801]" captionTargetPageId="3" captionText="FIGURE 3. Appendages Coeliccia spp. in dorsal and lateral view. (3a, b) C. lephuocdieui sp. nov.; (3c, d) C. scutellum, Kon Ka Kinh National Park; (3e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864990" httpUri="https://zenodo.org/record/3864990/files/figure.png" pageId="5" pageNumber="74">Fig. 3f</figureCitation>
). In the female, posterior pronotal lobe of prothorax of 
<taxonomicName id="26A9E2EBFFC5167C14AD19ABFC0341A8" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[753,931,405,431]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC5167C14AD19ABFC0341A8" box="[753,931,405,431]" italics="true" pageId="5" pageNumber="74">C. lephuocdieui</emphasis>
</taxonomicName>
well developed (
<figureCitation id="799285EDFFC5167C123819ABFB0641B7" box="[1124,1190,405,432]" captionStart="FIGURE 6" captionStartId="6.[151,250,1269,1294]" captionTargetBox="[167,1425,189,1233]" captionTargetId="figure@6.[151,1436,181,1244]" captionTargetPageId="6" captionText="FIGURE 6. Posterior pronotal lobe of prothorax of Coeliccia spp., all female. (6a, b) C. lephuocdieui sp. nov.; (6c, d) C. scutellum, Kon Ka Kinh National Park; (6e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864997" httpUri="https://zenodo.org/record/3864997/files/figure.png" pageId="5" pageNumber="74">Fig. 6</figureCitation>
a–b) but this structure much smaller in 
<taxonomicName id="26A9E2EBFFC5167C17091987FE4941D4" authorityName="Asahina" authorityYear="1984" box="[341,489,441,467]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC5167C17091987FE4941D4" box="[341,489,441,467]" italics="true" pageId="5" pageNumber="74">C. yamasakii</emphasis>
</taxonomicName>
(
<figureCitation id="799285EDFFC5167C17A41987FD9B41D3" box="[504,571,441,468]" captionStart="FIGURE 6" captionStartId="6.[151,250,1269,1294]" captionTargetBox="[167,1425,189,1233]" captionTargetId="figure@6.[151,1436,181,1244]" captionTargetPageId="6" captionText="FIGURE 6. Posterior pronotal lobe of prothorax of Coeliccia spp., all female. (6a, b) C. lephuocdieui sp. nov.; (6c, d) C. scutellum, Kon Ka Kinh National Park; (6e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864997" httpUri="https://zenodo.org/record/3864997/files/figure.png" pageId="5" pageNumber="74">Fig. 6</figureCitation>
e–f); S10 largely yellowish (
<figureCitation id="799285EDFFC5167C15221987FC6D41D4" box="[894,973,440,467]" captionStart="FIGURE 5" captionStartId="5.[151,250,1616,1641]" captionTargetBox="[151,1436,528,1592]" captionTargetId="figure@5.[151,1436,525,1592]" captionTargetPageId="5" captionText="FIGURE 5. Structure features of Coeliccia lephuocdieui sp. nov. (5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="5" pageNumber="74">Fig. 5e</figureCitation>
) whereas S10 entirely black in 
<taxonomicName id="26A9E2EBFFC5167C13691987FF6C41F0" authorityName="Asahina" authorityYear="1984" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="D3DD457AFFC5167C13691987FF6C41F0" italics="true" pageId="5" pageNumber="74">C. yamasakii</emphasis>
</taxonomicName>
(
<figureCitation id="799285EDFFC5167C168719E3FE8D41F0" box="[219,301,477,503]" captionStart="FIGURE 7" captionStartId="8.[151,250,1303,1328]" captionTargetBox="[151,1436,181,1276]" captionTargetId="figure@8.[151,1436,181,1278]" captionTargetPageId="8" captionText="FIGURE 7. Structure features of Coeliccia yamasakii, Phu Quoc Islands. (7a) head &amp; thorax of male, lateral view; (7b) head of male, frontal view; (7c) head of female, frontal view; (7d) head &amp; thorax of female, lateral view; (7e) abdominal tip of female, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3864999" httpUri="https://zenodo.org/record/3864999/files/figure.png" pageId="5" pageNumber="74">Fig. 7e</figureCitation>
).
</paragraph>
<caption id="B5D6C9E0FFC5167C16CB1E6EFF6E46B6" ID-DOI="http://doi.org/10.5281/zenodo.3864995" ID-Zenodo-Dep="3864995" httpUri="https://zenodo.org/record/3864995/files/figure.png" pageId="5" pageNumber="74" startId="5.[151,250,1616,1641]" targetBox="[151,1436,528,1592]" targetPageId="5">
<paragraph id="E1169968FFC5167C16CB1E6EFF6E46B6" blockId="5.[151,1436,1616,1713]" pageId="5" pageNumber="74">
<emphasis id="D3DD457AFFC5167C16CB1E6EFEAE466E" bold="true" box="[151,270,1616,1641]" pageId="5" pageNumber="74">FIGURE 5</emphasis>
. Structure features of 
<taxonomicName id="26A9E2EBFFC5167C17AE1E6FFD7E466E" authority="Phan &amp; Ngo &amp; Bui, 2020" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[498,734,1617,1641]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lephuocdieui" status="sp. nov.">
<emphasis id="D3DD457AFFC5167C17AE1E6FFD7E466E" box="[498,734,1617,1641]" italics="true" pageId="5" pageNumber="74">Coeliccia lephuocdieui</emphasis>
</taxonomicName>
<emphasis id="D3DD457AFFC5167C14B91E6FFC96466E" bold="true" box="[741,822,1617,1641]" pageId="5" pageNumber="74">
<taxonomicNameLabel id="C8EEF801FFC5167C14B91E6FFC96466E" box="[741,822,1617,1641]" pageId="5" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(5a) head &amp; thorax of female, lateral view; (5b) head of female, frontal view; (5c) head of male, frontal view; (5d) prothorax of female, dorsal view; (5f) abdominal tip of female, lateral view.
</paragraph>
</caption>
<paragraph id="E1169968FFC5167C169B1EE1FC0D4761" blockId="5.[151,1437,1759,2002]" pageId="5" pageNumber="74">
General body colour of female of 
<taxonomicName id="26A9E2EBFFC5167C14181EDEFD5646FD" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[580,758,1760,1786]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC5167C14181EDEFD5646FD" box="[580,758,1760,1786]" italics="true" pageId="5" pageNumber="74">C. lephuocdieui</emphasis>
</taxonomicName>
same as in 
<taxonomicName id="26A9E2EBFFC5167C15241EDEFBA446FD" authorityName="Laidlaw" authorityYear="1932" box="[888,1028,1760,1786]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="scutellum">
<emphasis id="D3DD457AFFC5167C15241EDEFBA446FD" box="[888,1028,1760,1786]" italics="true" pageId="5" pageNumber="74">C. scutellum</emphasis>
</taxonomicName>
, but differs by structural differences of the posterior pronotal lobe of prothorax as follows: in lateral view, this structure is not strongly erect and elongat- ed as 
<taxonomicName id="26A9E2EBFFC5167C168A1F16FEC24745" authorityName="Laidlaw" authorityYear="1932" box="[214,354,1832,1858]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="scutellum">
<emphasis id="D3DD457AFFC5167C168A1F16FEC24745" box="[214,354,1832,1858]" italics="true" pageId="5" pageNumber="74">C. scutellum</emphasis>
</taxonomicName>
(
<figureCitation id="799285EDFFC5167C172D1F19FE7D4745" box="[369,477,1831,1858]" captionStart="FIGURE 6" captionStartId="6.[151,250,1269,1294]" captionTargetBox="[167,1425,189,1233]" captionTargetId="figure@6.[151,1436,181,1244]" captionTargetPageId="6" captionText="FIGURE 6. Posterior pronotal lobe of prothorax of Coeliccia spp., all female. (6a, b) C. lephuocdieui sp. nov.; (6c, d) C. scutellum, Kon Ka Kinh National Park; (6e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864997" httpUri="https://zenodo.org/record/3864997/files/figure.png" pageId="5" pageNumber="74">Fig. 6a, c</figureCitation>
); in dorsal view, posterior pronotal lobe of 
<taxonomicName id="26A9E2EBFFC5167C159C1F16FBD14745" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[960,1137,1832,1858]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC5167C159C1F16FBD14745" box="[960,1137,1832,1858]" italics="true" pageId="5" pageNumber="74">C. lephuocdieui</emphasis>
</taxonomicName>
broadly, triangular-shaped (
<figureCitation id="799285EDFFC5167C16FC1F75FF514761" box="[160,241,1867,1894]" captionStart="FIGURE 6" captionStartId="6.[151,250,1269,1294]" captionTargetBox="[167,1425,189,1233]" captionTargetId="figure@6.[151,1436,181,1244]" captionTargetPageId="6" captionText="FIGURE 6. Posterior pronotal lobe of prothorax of Coeliccia spp., all female. (6a, b) C. lephuocdieui sp. nov.; (6c, d) C. scutellum, Kon Ka Kinh National Park; (6e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864997" httpUri="https://zenodo.org/record/3864997/files/figure.png" pageId="5" pageNumber="74">Fig. 6b</figureCitation>
) but it is narrower and parallel sided in 
<taxonomicName id="26A9E2EBFFC5167C14EC1F72FC9C4761" authorityName="Laidlaw" authorityYear="1932" box="[688,828,1868,1894]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="scutellum">
<emphasis id="D3DD457AFFC5167C14EC1F72FC9C4761" box="[688,828,1868,1894]" italics="true" pageId="5" pageNumber="74">C. scutellum</emphasis>
</taxonomicName>
(
<figureCitation id="799285EDFFC5167C15101F75FC3F4761" box="[844,927,1867,1894]" captionStart="FIGURE 6" captionStartId="6.[151,250,1269,1294]" captionTargetBox="[167,1425,189,1233]" captionTargetId="figure@6.[151,1436,181,1244]" captionTargetPageId="6" captionText="FIGURE 6. Posterior pronotal lobe of prothorax of Coeliccia spp., all female. (6a, b) C. lephuocdieui sp. nov.; (6c, d) C. scutellum, Kon Ka Kinh National Park; (6e, f) C. yamasakii, Phu Quoc Islands." figureDoi="http://doi.org/10.5281/zenodo.3864997" httpUri="https://zenodo.org/record/3864997/files/figure.png" pageId="5" pageNumber="74">Fig. 6d</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A9B3CAE3FFC5167C169B1F4EFE4D47D5" pageId="5" pageNumber="74" type="discussion">
<paragraph id="E1169968FFC5167C169B1F4EFE4D47D5" blockId="5.[151,1437,1759,2002]" pageId="5" pageNumber="74">
<emphasis id="D3DD457AFFC5167C169B1F4EFE9B478D" bold="true" box="[199,315,1904,1930]" pageId="5" pageNumber="74">Remarks.</emphasis>
<taxonomicName id="26A9E2EBFFC5167C171E1F4EFDE5478D" authorityName="Phan &amp; Ngo &amp; Bui" authorityYear="2020" box="[322,581,1904,1930]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lephuocdieui">
<emphasis id="D3DD457AFFC5167C171E1F4EFDE5478D" box="[322,581,1904,1930]" italics="true" pageId="5" pageNumber="74">Coeliccia lephuocdieui</emphasis>
</taxonomicName>
and 
<taxonomicName id="26A9E2EBFFC5167C14271F4EFCA7478D" authorityName="Laidlaw" authorityYear="1932" box="[635,775,1904,1930]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="scutellum">
<emphasis id="D3DD457AFFC5167C14271F4EFCA7478D" box="[635,775,1904,1930]" italics="true" pageId="5" pageNumber="74">C. scutellum</emphasis>
</taxonomicName>
occur at different elevations in the mountain areas of Ayun Commune of 
<collectingRegion id="236D578AFFC5167C176F1FADFE5847AA" box="[307,504,1939,1965]" country="Vietnam" name="Gia Lai" pageId="5" pageNumber="74">Gia Lai Province</collectingRegion>
. The new species occurs at an approximate altitude of 
<quantity id="2651348DFFC5167C12391FADFB6247A9" box="[1125,1218,1939,1966]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="74" unit="m" value="1500.0">1,500 m</quantity>
compared to about 
<quantity id="2651348DFFC5167C16CB1F86FE8B47D5" box="[151,299,1976,2002]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="5" pageNumber="74" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">800–1,000 m</quantity>
for 
<taxonomicName id="26A9E2EBFFC5167C17061F86FE4547D5" authorityName="Laidlaw" authorityYear="1932" box="[346,485,1976,2002]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="scutellum">
<emphasis id="D3DD457AFFC5167C17061F86FE4547D5" box="[346,485,1976,2002]" italics="true" pageId="5" pageNumber="74">C. scutellum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>