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<document id="B16A2D7A2478CD52A6A929E03185397D" ID-CLB-Dataset="51174" ID-DOI="10.11646/zootaxa.4066.4.5" ID-GBIF-Dataset="24f49927-38fd-40c9-9d5d-102b8b883c7f" ID-ISSN="1175-5326" ID-Zenodo-Dep="258909" ID-ZooBank="15920248-D234-4FE9-8A30-3EE461B92198" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459689165899" checkinUser="plazi" docAuthor="Bian, Xun &amp; Shi, Fuming" docDate="2016" docId="6576FD57724BFFC0FF3FF9C99AD9FEC2" docLanguage="en" docName="zootaxa.4066.4.5.pdf" docOrigin="Zootaxa 4066 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eugryllacris" docType="key" docVersion="9" lastPageNumber="440" masterDocId="994F852F724AFFC2FFA8FF809F45FFE0" masterDocTitle="Review of the genus Eugryllacris Karny, 1937 (Orthoptera: Gryllacridinae) from China" masterLastPageNumber="450" masterPageNumber="438" pageNumber="439" updateTime="1698209159626" updateUser="ExternalLinkService">
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<mods:title id="FF853F6004ED26399FA3060ED04B74B3">Review of the genus Eugryllacris Karny, 1937 (Orthoptera: Gryllacridinae) from China</mods:title>
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<mods:namePart id="7037C5A95DE69054426715D043C68909">Bian, Xun</mods:namePart>
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<mods:name id="F1BD02B3873CF9CE00AA0F2BCD9B45FE" type="personal">
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<mods:namePart id="F1354601BA793481B1C826BB9527D196">Shi, Fuming</mods:namePart>
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<mods:title id="C5B9CFD02DC3CC6A731DD3987FE4ACFB">Zootaxa</mods:title>
</mods:titleInfo>
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<mods:date id="97F10E18E1ECAE446E75DE7D27156430">2016</mods:date>
<mods:detail id="3445C0BA7FD719103DA45AD89BFE11E4" type="volume">
<mods:number id="A1426639FA61F6CD04261A483ABBA644">4066</mods:number>
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<mods:detail id="D71C868A081ED9F211D69BDCEE793002" type="issue">
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<mods:classification id="156A59F79BD3557E5047CF77111D6948">journal article</mods:classification>
<mods:identifier id="1E26D7A5199BCDF6CA2F95E77355E7E2" type="CLB-Dataset">51174</mods:identifier>
<mods:identifier id="B1C61D0C080544456139D3D6E04ECA6D" type="DOI">10.11646/zootaxa.4066.4.5</mods:identifier>
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<mods:identifier id="E794977A14D499F96890D45B2D8F200F" type="ISSN">1175-5326</mods:identifier>
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<treatment id="6576FD57724BFFC0FF3FF9C99AD9FEC2" ID-DOI="http://doi.org/10.5281/zenodo.6058511" ID-GBIF-Taxon="119333379" ID-Zenodo-Dep="6058511" LSID="urn:lsid:plazi:treatment:6576FD57724BFFC0FF3FF9C99AD9FEC2" httpUri="http://treatment.plazi.org/id/6576FD57724BFFC0FF3FF9C99AD9FEC2" lastPageId="2" lastPageNumber="440" pageId="1" pageNumber="439">
<subSubSection id="A5C51FCA724BFFC3FF3FF9C99D9CF983" box="[151,729,1609,1635]" pageId="1" pageNumber="439" type="nomenclature">
<paragraph id="ED604C41724BFFC3FF3FF9C99D9CF983" blockId="1.[151,729,1609,1635]" box="[151,729,1609,1635]" pageId="1" pageNumber="439">
<heading id="B628FB2D724BFFC3FF3FF9C99D9CF983" bold="true" box="[151,729,1609,1635]" fontSize="11" level="1" pageId="1" pageNumber="439" reason="1">
<emphasis id="DFAB9053724BFFC3FF3FF9C99D9CF983" bold="true" box="[151,729,1609,1635]" pageId="1" pageNumber="439">
Key to known
<taxonomicName id="2ADF37C2724BFFC3FEE4F9C99EA7F983" box="[332,482,1609,1635]" class="Insecta" family="Haglidae" genus="Eugryllacris" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="DFAB9053724BFFC3FEE4F9C99EA7F983" bold="true" box="[332,482,1609,1635]" italics="true" pageId="1" pageNumber="439">Eugryllacris</emphasis>
</taxonomicName>
species from
<collectingCountry id="95C80CD1724BFFC3FD24F9C99D9CF983" box="[652,729,1609,1635]" name="China" pageId="1" pageNumber="439">China</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A5C51FCA724BFFC0FF3FF90F9AD9FEC2" lastPageId="2" lastPageNumber="440" pageId="1" pageNumber="439" type="key">
<keyStep id="562B54E4724BFFC3FF3FF90F9ADEF922" pageId="1" pageNumber="439">
<paragraph id="ED604C41724BFFC3FF3FF90F9AD9F945" blockId="1.[151,1436,1679,2024]" box="[151,1436,1679,1701]" pageId="1" pageNumber="439">
<keyLead id="562EEF74724BFFC3FF3FF90F9AD9F945" box="[151,1436,1679,1701]" pageId="1" pageNumber="439">
<specimenCount id="FBD987C8724BFFC3FF3FF90F9E6DF944" box="[151,296,1679,1700]" pageId="1" pageNumber="439" type="female">1 Female</specimenCount>
seventh abdominal sternite wider than long (
<figureCitation id="75E450C4724BFFC3FD70F90F9C4BF944" box="[728,782,1679,1700]" captionStart="FIGURE 1" captionStartId="2.[151,250,1116,1138]" captionTargetBox="[151,1436,774,1091]" captionTargetId="figure@2.[151,1436,774,1091]" captionTargetPageId="2" captionText="FIGURE 1. Eugryllacris cylindrigera (Karny, 1926) com. nov., female: A. process of seventh abdominal sternite and subgenital plate in ventral view (Karny, 1926: Fig. 11); B. holotype habitus in lateral view (Eades et al., 2015)." httpUri="https://zenodo.org/record/258910/files/figure.png" pageId="1" pageNumber="439">Fig. 1</figureCitation>
A)...........
<taxonomicName id="2ADF37C2724BFFC3FC1CF90F9A70F944" authority="Karny, 1926" authorityName="Karny" authorityYear="1926" box="[948,1333,1679,1701]" class="Insecta" family="Haglidae" genus="Eugryllacris" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="cylindrigera">
<emphasis id="DFAB9053724BFFC3FC1CF90F9BEDF945" box="[948,1192,1679,1701]" italics="true" pageId="1" pageNumber="439">Eugryllacris cylindrigera</emphasis>
(
<bibRefCitation id="894E31B0724BFFC3FB1DF90F9A6BF944" author="Karny" box="[1205,1326,1679,1700]" pageId="1" pageNumber="450" refString="Karny, H. H. (1926) Gryllacridae (China-Ausbeute von R. Mell). Mitteilungen aus dem Zoologischen Museum in Berlin, 12, 257 - 394." type="journal article" year="1926">Karny, 1926</bibRefCitation>
)
</taxonomicName>
<emphasis id="DFAB9053724BFFC3FA93F90F9AD9F945" bold="true" box="[1339,1436,1679,1701]" pageId="1" pageNumber="439">com. nov.</emphasis>
</keyLead>
</paragraph>
<paragraph id="ED604C41724BFFC3FF3FF92D9ADEF922" blockId="1.[151,1436,1679,2024]" box="[151,1435,1709,1730]" pageId="1" pageNumber="439">
<keyLead id="562EEF74724BFFC3FF3FF92D9ADEF922" box="[151,1435,1709,1730]" pageId="1" pageNumber="439">- Female seventh abdominal sternite longer than wide.......................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="562B54E4724BFFC3FF3FF94A9AD9F91C" pageId="1" pageNumber="439">
<paragraph id="ED604C41724BFFC3FF3FF94A9AD9F93F" blockId="1.[151,1436,1679,2024]" box="[151,1436,1738,1759]" pageId="1" pageNumber="439">
<keyLead id="562EEF74724BFFC3FF3FF94A9AD9F93F" box="[151,1436,1738,1759]" pageId="1" pageNumber="439">2 Process of female seventh abdominal sternite longer, reaching the middle area of subgenital plate...................... 3</keyLead>
</paragraph>
<paragraph id="ED604C41724BFFC3FF3FF9679AD9F91C" blockId="1.[151,1436,1679,2024]" box="[151,1436,1767,1788]" pageId="1" pageNumber="439">
<keyLead id="562EEF74724BFFC3FF3FF9679AD9F91C" box="[151,1436,1767,1788]" pageId="1" pageNumber="439">- Process of female seventh abdominal sternite shorter, only reaching the basal area of subgenital plate................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="562B54E4724BFFC3FF3FF8859AD9F892" pageId="1" pageNumber="439">
<paragraph id="ED604C41724BFFC3FF3FF8859AD9F8D8" blockId="1.[151,1436,1679,2024]" pageId="1" pageNumber="439">
<keyLead id="562EEF74724BFFC3FF3FF8859AD9F8D8" pageId="1" pageNumber="439">
3 Apical half of process of female abdominal sternite nearly straight, its apex with 1 shallow concavity in the middle (
<figureCitation id="75E450C4724BFFC3FA92F8859A37F8FA" box="[1338,1394,1797,1818]" captionStart="FIGURE 2" captionStartId="3.[151,250,1598,1620]" captionTargetBox="[151,1436,185,1579]" captionTargetId="figure@3.[151,1436,185,1580]" captionTargetPageId="3" captionText="FIGURE 2. Eugryllacris elongata Bian &amp; Shi sp. nov. A, H. head in frontal view; B – C, I – J. head and pronotum: B, I. dorsal view, C, J. lateral view; D – E, J – K. apex of abdomen: D, K. lateral view, E. ventral and slightly apical view, G. dorso-apical view; F, L. subgenital plate in ventral view; A – G. male; H – L. female." httpUri="https://zenodo.org/record/258911/files/figure.png" pageId="1" pageNumber="439">Fig. 2</figureCitation>
L)....................................................................
<taxonomicName id="2ADF37C2724BFFC3FBADF8A19A0CF8D7" authority="Bian &amp; Shi" authorityName="Bian &amp; Shi" box="[1029,1353,1825,1847]" class="Insecta" family="Haglidae" genus="Eugryllacris" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="elongata" status="sp. nov.">
<emphasis id="DFAB9053724BFFC3FBADF8A19B92F8D7" box="[1029,1239,1825,1847]" italics="true" pageId="1" pageNumber="439">Eugryllacris elongata</emphasis>
Bian &amp; Shi
</taxonomicName>
<taxonomicNameLabel id="C4982D28724BFFC3FAE7F8A29AD9F8D8" box="[1359,1436,1826,1848]" pageId="1" pageNumber="439" rank="species">
<emphasis id="DFAB9053724BFFC3FAE7F8A29AD9F8D8" bold="true" box="[1359,1436,1826,1848]" pageId="1" pageNumber="439">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="ED604C41724BFFC3FF3FF8BF9AD9F892" blockId="1.[151,1436,1679,2024]" pageId="1" pageNumber="439">
<keyLead id="562EEF74724BFFC3FF3FF8BF9AD9F892" pageId="1" pageNumber="439">
- Apical half of process of female abdominal sternite slightly curved dorsad, its apex faintly concave in the middle (
<figureCitation id="75E450C4724BFFC3FA80F8BF9A27F8B4" box="[1320,1378,1855,1876]" captionStart="FIGURE 6" captionStartId="9.[151,250,1055,1077]" captionTargetBox="[151,1435,195,1030]" captionTargetId="figure@9.[151,1436,195,1031]" captionTargetPageId="9" captionText="FIGURE 6. Eugryllacris xiei Bian &amp; Shi sp. nov. A, G. head in frontal view; B – C, H – I. head and pronotum: B, H. dorsal view, C, I. lateral view; D, F, K. apex of abdomen: D. K. lateral view, F. ventral and slightly apical view; K. subgenital plate in lateral view: E, J. subgenital plate in ventral view; A – F. male; G – K. female." httpUri="https://zenodo.org/record/258915/files/figure.png" pageId="1" pageNumber="439">Fig. 6</figureCitation>
J)..........................................................................
<taxonomicName id="2ADF37C2724BFFC3FB9FF8DC9A0CF892" authority="Bian &amp; Shi" authorityName="Bian &amp; Shi" box="[1079,1353,1884,1906]" class="Insecta" family="Haglidae" genus="Eugryllacris" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="xiei" status="sp. nov.">
<emphasis id="DFAB9053724BFFC3FB9FF8DC9B92F892" box="[1079,1239,1884,1906]" italics="true" pageId="1" pageNumber="439">Eugryllacris xiei</emphasis>
Bian &amp; Shi
</taxonomicName>
<taxonomicNameLabel id="C4982D28724BFFC3FAE7F8DC9AD9F892" box="[1359,1436,1884,1906]" pageId="1" pageNumber="439" rank="species">
<emphasis id="DFAB9053724BFFC3FAE7F8DC9AD9F892" bold="true" box="[1359,1436,1884,1906]" pageId="1" pageNumber="439">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="562B54E4724BFFC3FF3FF8FA9ADEF84C" pageId="1" pageNumber="439">
<paragraph id="ED604C41724BFFC3FF3FF8FA9AD9F870" blockId="1.[151,1436,1679,2024]" box="[151,1436,1913,1936]" pageId="1" pageNumber="439">
<keyLead id="562EEF74724BFFC3FF3FF8FA9AD9F870" box="[151,1436,1913,1936]" pageId="1" pageNumber="439">
4 The lateral lobes of male subgenital plate shorter than styli (
<figureCitation id="75E450C4724BFFC3FC88F8FA9C1CF86F" box="[800,857,1914,1935]" captionStart="FIGURE 7" captionStartId="10.[151,250,1162,1184]" captionTargetBox="[183,1398,297,1136]" captionTargetId="figure@10.[183,1399,297,1136]" captionTargetPageId="10" captionText="FIGURE 7. Eugryllacris lobulis Bian &amp; Shi sp. nov. male: A. head in frontal view; B – C. head and pronotum: B. dorsal view, C. lateral view; D, E. apex of abdomen: D. apical and ventral view, E. ventral view." httpUri="https://zenodo.org/record/258916/files/figure.png" pageId="1" pageNumber="439">Fig. 7</figureCitation>
E).............
<taxonomicName id="2ADF37C2724BFFC3FBB0F8F99A0CF86F" authority="Bian &amp; Shi" authorityName="Bian &amp; Shi" box="[1048,1353,1913,1935]" class="Insecta" family="Haglidae" genus="Eugryllacris" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="lobulis" status="sp. nov.">
<emphasis id="DFAB9053724BFFC3FBB0F8F99B92F86F" box="[1048,1239,1913,1935]" italics="true" pageId="1" pageNumber="439">Eugryllacris lobulis</emphasis>
Bian &amp; Shi
</taxonomicName>
<taxonomicNameLabel id="C4982D28724BFFC3FAE7F8FA9AD9F870" box="[1359,1436,1914,1936]" pageId="1" pageNumber="439" rank="species">
<emphasis id="DFAB9053724BFFC3FAE7F8FA9AD9F870" bold="true" box="[1359,1436,1914,1936]" pageId="1" pageNumber="439">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="ED604C41724BFFC3FF3FF8179ADEF84C" blockId="1.[151,1436,1679,2024]" box="[151,1435,1943,1964]" pageId="1" pageNumber="439">
<keyLead id="562EEF74724BFFC3FF3FF8179ADEF84C" box="[151,1435,1943,1964]" pageId="1" pageNumber="439">- The lateral lobes of male subgenital plate as long as styli...................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="562B54E4724BFFC0FF3FF8359ADEFF4C" lastPageId="2" lastPageNumber="440" pageId="1" pageNumber="439">
<paragraph id="ED604C41724BFFC3FF3FF8359AD9F808" blockId="1.[151,1436,1679,2024]" pageId="1" pageNumber="439">
<keyLead id="562EEF74724BFFC3FF3FF8359AD9F808" pageId="1" pageNumber="439">
5 The lateral lobes of male subgenital plate on posterior margin obviously curved ventrad (
<figureCitation id="75E450C4724BFFC3FB9DF8359B28F82A" box="[1077,1133,1973,1994]" captionStart="FIGURE 8" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[177,1414,1264,1917]" captionTargetId="figure@10.[177,1414,1264,1918]" captionTargetPageId="10" captionText="FIGURE 8. Eugryllacris fanjingshanensis Bian &amp; Shi sp. nov. male: A. head in frontal view; B – C. head and pronotum: B. dorsal view, C. lateral view; D, E. apex of abdomen: D. lateral view, E. apical and slightly ventral view; F. subgenital plate in ventral view." httpUri="https://zenodo.org/record/258917/files/figure.png" pageId="1" pageNumber="439">Fig. 8</figureCitation>
E)...................................................................................
<taxonomicName id="2ADF37C2724BFFC3FC1FF8529A0CF807" authority="Bian &amp; Shi" authorityName="Bian &amp; Shi" box="[951,1353,2002,2024]" class="Insecta" family="Haglidae" genus="Eugryllacris" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="fanjingshanensis" status="sp. nov.">
<emphasis id="DFAB9053724BFFC3FC1FF8529B92F808" box="[951,1239,2002,2024]" italics="true" pageId="1" pageNumber="439">Eugryllacris fanjingshanensis</emphasis>
Bian &amp; Shi
</taxonomicName>
<taxonomicNameLabel id="C4982D28724BFFC3FAE7F8529AD9F808" box="[1359,1436,2002,2024]" pageId="1" pageNumber="439" rank="species">
<emphasis id="DFAB9053724BFFC3FAE7F8529AD9F808" bold="true" box="[1359,1436,2002,2024]" pageId="1" pageNumber="439">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="ED604C417248FFC0FF3FFF179ADEFF4C" blockId="2.[151,1436,151,290]" box="[151,1435,151,172]" pageId="2" pageNumber="440">
<keyLead id="562EEF747248FFC0FF3FFF179ADEFF4C" box="[151,1435,151,172]" pageId="2" pageNumber="440">- The lateral lobes of male subgenital plate not curved ventrad................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="562B54E47248FFC0FF3FFF359AD9FEC2" pageId="2" pageNumber="440">
<paragraph id="ED604C417248FFC0FF3FFF359AD9FF08" blockId="2.[151,1436,151,290]" pageId="2" pageNumber="440">
<keyLead id="562EEF747248FFC0FF3FFF359AD9FF08" pageId="2" pageNumber="440">
6 Posterior margin of male subgenital plate deeply concave, the lateral lobes nearly narrowly rounded (
<figureCitation id="75E450C47248FFC0FB02FF359BA6FF2A" box="[1194,1251,181,202]" captionStart="FIGURE 4" captionStartId="6.[151,250,1810,1832]" captionTargetBox="[151,1436,195,1785]" captionTargetId="figure@6.[151,1436,195,1786]" captionTargetPageId="6" captionText="FIGURE 4. Eugryllacris longifissa Bian &amp; Shi sp. nov. A, G. head in frontal view; B – C, H – I. head and pronotum: B, H. dorsal view, C, I. lateral view; D – E, J. apex of abdomen: D. ventro-lateral view, E. apical and slightly ventral view, J. lateral view; F, K. subgenital plate in ventral view; A – F. male; G – K. female." httpUri="https://zenodo.org/record/258913/files/figure.png" pageId="2" pageNumber="440">Fig. 4</figureCitation>
F)...............................................................................
<taxonomicName id="2ADF37C27248FFC0FC54FF519A0CFF07" authority="Bian &amp; Shi" authorityName="Bian &amp; Shi" box="[1020,1353,209,231]" class="Insecta" family="Haglidae" genus="Eugryllacris" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="440" phylum="Arthropoda" rank="species" species="longifissa" status="sp. nov.">
<emphasis id="DFAB90537248FFC0FC54FF519B92FF07" box="[1020,1239,209,231]" italics="true" pageId="2" pageNumber="440">Eugryllacris longifissa</emphasis>
Bian &amp; Shi
</taxonomicName>
<taxonomicNameLabel id="C4982D287248FFC0FAE7FF529AD9FF08" box="[1359,1436,210,232]" pageId="2" pageNumber="440" rank="species">
<emphasis id="DFAB90537248FFC0FAE7FF529AD9FF08" bold="true" box="[1359,1436,210,232]" pageId="2" pageNumber="440">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="ED604C417248FFC0FF3FFF6F9AD9FEC2" blockId="2.[151,1436,151,290]" pageId="2" pageNumber="440">
<keyLead id="562EEF747248FFC0FF3FFF6F9AD9FEC2" pageId="2" pageNumber="440">
- Posterior margin of male subgenital plate with 1 triangular concavity, the lateral lobes nearly triangular (
<figureCitation id="75E450C47248FFC0FB61FF6F9A47FEE4" box="[1225,1282,239,260]" captionStart="FIGURE 5" captionStartId="7.[151,250,1938,1960]" captionTargetBox="[151,1436,1022,1914]" captionTargetId="figure@7.[151,1436,1022,1914]" captionTargetPageId="7" captionText="FIGURE 5. Eugryllacris bifoliata Bian &amp; Shi sp. nov. A, F. head in frontal view; B – C, G – H. head and pronotum: B, G. dorsal view, C, H. lateral view; D, J. apex of abdomen in lateral view; K. subgenital plate in lateral view: E, L. subgenital plate in ventral view; A – E. male; F – L. female." httpUri="https://zenodo.org/record/258914/files/figure.png" pageId="2" pageNumber="440">Fig. 5</figureCitation>
E)..............................................................................
<taxonomicName id="2ADF37C27248FFC0FBA0FE8C9A0CFEC2" authority="Bian &amp; Shi" authorityName="Bian &amp; Shi" box="[1032,1353,268,290]" class="Insecta" family="Haglidae" genus="Eugryllacris" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="440" phylum="Arthropoda" rank="species" species="bifoliata" status="sp. nov.">
<emphasis id="DFAB90537248FFC0FBA0FE8C9B92FEC2" box="[1032,1239,268,290]" italics="true" pageId="2" pageNumber="440">Eugryllacris bifoliata</emphasis>
Bian &amp; Shi
</taxonomicName>
<taxonomicNameLabel id="C4982D287248FFC0FAE7FE8C9AD9FEC2" box="[1359,1436,268,290]" pageId="2" pageNumber="440" rank="species">
<emphasis id="DFAB90537248FFC0FAE7FE8C9AD9FEC2" bold="true" box="[1359,1436,268,290]" pageId="2" pageNumber="440">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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