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<document id="1377F7ADE422EDC283E91B031EAA6F87" ID-CLB-Dataset="45448" ID-DOI="10.5281/zenodo.280057" ID-GBIF-Dataset="e6234795-c9d7-447e-aefd-b3469b6f57b5" ID-ISSN="1175-5326" ID-Zenodo-Dep="280057" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460438720851" checkinUser="plazi" docAuthor="Geissler, Peter, Hartmann, Timo &amp; Neang, Thy" docDate="2012" docId="03FC39414F01FF81FF55FD9911F37CD1" docLanguage="en" docName="zt03190p068.pdf" docOrigin="Zootaxa 3190" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lygosoma veunsaiensis Geissler, Hartmann &amp; Neang, 2012, sp. nov." docType="treatment" docVersion="9" lastPageNumber="63" masterDocId="FFC541394F03FF89FFC2FF77127D7B47" masterDocTitle="A new species of the genus Lygosoma Hardwicke &amp; Gray, 1827 (Squamata: Scincidae) from northeastern Cambodia, with an updated identification key to the genus Lygosoma in mainland Southeast Asia" masterLastPageNumber="68" masterPageNumber="56" pageNumber="58" updateTime="1698332598785" updateUser="ExternalLinkService">
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<mods:title id="0E959CB2110B5F9E1C5116638CBA4DD2">A new species of the genus Lygosoma Hardwicke &amp; Gray, 1827 (Squamata: Scincidae) from northeastern Cambodia, with an updated identification key to the genus Lygosoma in mainland Southeast Asia</mods:title>
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<mods:namePart id="D090DC3D316F7BB117E1D85F44F78C46">Geissler, Peter</mods:namePart>
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<mods:namePart id="6B44D28C5213233E9A6A160E6250B782">Hartmann, Timo</mods:namePart>
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<mods:namePart id="0C9A478D252D8BF5E2BA3117D78CEA10">Neang, Thy</mods:namePart>
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<treatment id="03FC39414F01FF81FF55FD9911F37CD1" ID-DOI="http://doi.org/10.5281/zenodo.6167562" ID-GBIF-Taxon="119507270" ID-Zenodo-Dep="6167562" LSID="urn:lsid:plazi:treatment:03FC39414F01FF81FF55FD9911F37CD1" httpUri="http://treatment.plazi.org/id/03FC39414F01FF81FF55FD9911F37CD1" lastPageId="8" lastPageNumber="63" pageId="2" pageNumber="58">
<subSubSection id="C34FDBDC4F01FF8BFF55FD99135E786E" pageId="2" pageNumber="58" type="nomenclature">
<paragraph id="8BEA88574F01FF8BFF55FD99106A784F" blockId="2.[151,535,750,809]" box="[151,535,750,776]" pageId="2" pageNumber="58">
<heading id="D0A23F3B4F01FF8BFF55FD99106A784F" bold="true" box="[151,535,750,776]" fontSize="11" level="1" pageId="2" pageNumber="58" reason="1">
<emphasis id="B92154454F01FF8BFF55FD99106A784F" bold="true" box="[151,535,750,776]" pageId="2" pageNumber="58">
<taxonomicName id="4C55F3D44F01FF8BFF55FD9913CF784F" box="[151,434,750,776]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="58" phylum="Chordata" rank="species" species="veunsaiensis" status="sp. nov.">
<emphasis id="B92154454F01FF8BFF55FD9913CF784F" bold="true" box="[151,434,750,776]" italics="true" pageId="2" pageNumber="58">Lygosoma veunsaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212E93E4F01FF8BFE7BFD99106A784F" box="[441,535,750,776]" pageId="2" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA88574F01FF8BFF55FC66135E786E" blockId="2.[151,535,750,809]" box="[151,291,785,809]" pageId="2" pageNumber="58">
<figureCitation id="136E94D24F01FF8BFF55FC66135D786E" box="[151,288,785,809]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="3.[151,250,1948,1971]" captionStartId-1="4.[151,250,851,874]" captionStartId-2="4.[151,250,1595,1618]" captionStartId-3="7.[151,250,1144,1167]" captionTargetBox-0="[209,1377,445,1907]" captionTargetBox-1="[151,1392,193,733]" captionTargetBox-2="[170,1436,985,1572]" captionTargetBox-3="[151,1436,193,1122]" captionTargetId-0="figure@3.[209,1378,445,1907]" captionTargetId-1="figure@4.[151,1436,193,830]" captionTargetId-2="figure@4.[151,1436,954,1574]" captionTargetId-3="figure@7.[151,1436,193,1123]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="4" captionTargetPageId-3="7" captionText-0="FIGURE 2. Head and anterior part of body of the holotype of Lygosoma veunsaiensis sp. nov. (CBC 00808): a) dorsal view and b) ventral view. Photo: Peter Geissler." captionText-1="FIGURE 3. Lateral view of head and anterior part of body of the holotype of Lygosoma veunsaiensis sp. nov. (CBC 00808). Photo: Peter Geissler." captionText-2="FIGURE 4. Holotype of Lygosoma veunsaiensis sp. nov. (CBC 00808): a) dorsolateral view and b) ventral view. Photo: Peter Geissler." captionText-3="FIGURE 5. Holotype of Lygosoma veunsaiensis sp. nov. (CBC 0808) in life. Photo: Gabor Csorba." httpUri-0="https://zenodo.org/record/280059/files/figure.png" httpUri-1="https://zenodo.org/record/280060/files/figure.png" httpUri-2="https://zenodo.org/record/280061/files/figure.png" httpUri-3="https://zenodo.org/record/280062/files/figure.png" pageId="2" pageNumber="58">Figures 25</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34FDBDC4F01FF8BFF55FC2D133378FE" pageId="2" pageNumber="58" type="materials_examined">
<paragraph id="8BEA88574F01FF8BFF55FC2D133378FE" blockId="2.[151,1437,857,2033]" pageId="2" pageNumber="58">
<emphasis id="B92154454F01FF8BFF55FC2D13767835" bold="true" box="[151,267,858,882]" pageId="2" pageNumber="58">
<typeStatus id="54EE36F54F01FF8BFF55FC2D137B7835" box="[151,262,858,882]" pageId="2" pageNumber="58" type="holotype">Holotype</typeStatus>
.
</emphasis>
CBC 0 0 808, adult male, collected by Gabor Csorba on
<date id="FFEBAE974F01FF8BFC69FC2E16067836" box="[939,1147,857,881]" pageId="2" pageNumber="58" value="2010-02-24">24 February 2010</date>
at O'Kasieb campsite at
<geoCoordinate id="EE61EE904F01FF8BFF55FC0A134678D2" box="[151,315,893,917]" direction="north" orientation="latitude" pageId="2" pageNumber="58" precision="1" value="14.016722">14°0100.2N</geoCoordinate>
,
<geoCoordinate id="EE61EE904F01FF8BFE88FC0A138478D2" box="[330,505,893,917]" direction="east" orientation="longitude" pageId="2" pageNumber="58" precision="1" value="106.7525">106°4509.0E</geoCoordinate>
,
<quantity id="4CAD25B24F01FF8BFDC5FC0A102D78D2" box="[519,592,893,917]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="2" pageNumber="58" unit="m" value="114.0">114 m</quantity>
in disturbed semi-deciduous forest in Seun Sai District, Ratanakiri Province,
<collectingCountry id="F342C8C74F01FF8BFF16FCD6133678FE" box="[212,331,929,953]" name="Cambodia" pageId="2" pageNumber="58">Cambodia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34FDBDC4F01FF8BFF0BFCB2138D7FF2" pageId="2" pageNumber="58" type="diagnosis">
<paragraph id="8BEA88574F01FF8BFF0BFCB2138D7FF2" blockId="2.[151,1437,857,2033]" pageId="2" pageNumber="58">
<emphasis id="B92154454F01FF8BFF0BFCB2133E7899" bold="true" box="[201,323,965,990]" pageId="2" pageNumber="58">Diagnosis.</emphasis>
<taxonomicName id="4C55F3D44F01FF8BFE88FCB1102C789A" box="[330,593,966,989]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="58" phylum="Chordata" rank="species" species="veunsaiensis" status="sp. nov.">
<emphasis id="B92154454F01FF8BFE88FCB1102C789A" box="[330,593,966,989]" italics="true" pageId="2" pageNumber="58">Lygosoma veunsaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212E93E4F01FF8BFD9AFCB210CF789A" box="[600,690,965,989]" pageId="2" pageNumber="58" rank="species">
<emphasis id="B92154454F01FF8BFD9AFCB210CF789A" bold="true" box="[600,690,965,989]" pageId="2" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
is differentiated from all other congeners by the following combination of characters: (1) body elongated, SVL approximately 5.4 times of HlL; (2) supranasals distinct, separated from each other by frontonasal; (3) frontoparietals paired; (4) supraciliaries nine; (5) supralabials five; (6) lower eyelid scaly; (7) infralabials five; (8) external ear openings absent; (9) midbody scale rows 22; (10) paravertebral scales 51; (11) ventrals in 49 transverse rows; (12) subdigital lamellae on fourth finger five; (13) subdigital lamellae on fourth toe nine; (14) a light stripe present on outer edge of the dorsum; and a dark dorsolateral stripe present, from behind the eye to the tail.
</paragraph>
</subSubSection>
<subSubSection id="C34FDBDC4F01FF8BFF0BFBB5164B7FBA" pageId="2" pageNumber="58" type="etymology">
<paragraph id="8BEA88574F01FF8BFF0BFBB5164B7FBA" blockId="2.[151,1437,857,2033]" pageId="2" pageNumber="58">
<emphasis id="B92154454F01FF8BFF0BFBB5132D7F9D" bold="true" box="[201,336,1218,1242]" pageId="2" pageNumber="58">Etymology.</emphasis>
We name this new species after the
<typeStatus id="54EE36F54F01FF8BFD36FBB511597F9D" box="[756,804,1218,1242]" pageId="2" pageNumber="58">type</typeStatus>
locality to underscore the importance of Veun Sai Proposed Protected Forest for the conservation of Cambodias threatened biodiversity.
</paragraph>
</subSubSection>
<subSubSection id="C34FDBDC4F01FF8DFF0BFA7E10B77CB4" lastPageId="4" lastPageNumber="60" pageId="2" pageNumber="58" type="description">
<paragraph id="8BEA88574F01FF8AFF0BFA7E16607AC0" blockId="2.[151,1437,857,2033]" lastBlockId="3.[151,1436,151,391]" lastPageId="3" lastPageNumber="59" pageId="2" pageNumber="58">
<emphasis id="B92154454F01FF8BFF0BFA7E13977E65" bold="true" box="[201,490,1289,1314]" pageId="2" pageNumber="58">
Description of
<typeStatus id="54EE36F54F01FF8BFEBEFA7D13987E65" box="[380,485,1290,1314]" pageId="2" pageNumber="58" type="holotype">holotype</typeStatus>
.
</emphasis>
Male specimen; SVL 33.6 mm, tail tip lost, for further measurements see Table 1. Head distinct from neck; snout rounded; rostral almost three times wider than long, forming a transverse narrow suture with frontonasal, separated from supranasals; frontonasal 1.3 times as wide as long; prefrontals small, widely separated from each other by frontal, touching loreal as well as anterior preoculars laterally; frontal large, elongate, diamond shaped, widened anteriorly and sharpened posterioly, longer than its distance to the tip of the snout, in contact with frontonasal, prefrontals, first supraciliary, first and second supraoculars; frontoparietals paired, in broad contact medially at anterior half and widely separated posteriorly, bordered by frontal anteriorly, second, third and fourth supraoculars laterally, interparietal, and parietals posteriorly; interparietal diamond shaped, widened anteriorly and sharpened posteriorly, bearing an eye spot; parietals subrectangular, in contact posteriorly, bordered by seven dorsal scales posteriorly and two small temporals laterally, enlarged nuchals absent; three temporals, anterior one smaller. Nostril within nasal; nasal in contact with rostral, supranasal, frontonasal, first supralabial and anterior loreal; two small loreals on the right side, similar in size, vertically elongated, bordered by first and second supralabials, nasal, frontonasal, prefrontals, and anterior preoculars; one large loreal on the left side, subtriangular in shape; two small subrectangular preoculars bordered by smaller presuboculars, forming a transition to the lower eyelid; four supraoculars; nine supraciliaries; lower eyelid scaly; two enlarged postoculars, the lower one bordering the fifth supralabial; five supralabials, the fifth largest; external ear openings absent, covered by scales at small depressed cavity behind mandible. Mental wider than long, in contact with the postmental and first infralabials; postmental single, bordering mental, first infralabials and first pair of chin shields; first pair of chin shields in broad contact medially; second pair of chin shields separated from each other by a small gular scale, in contact with first pair of chin shields anteriorly, second, third and fourth infralabials laterally and three gular scales posteriorly; five infralabials (Figs. 2,3); body elongated, the distance between axilla and groin almost five times the length of the forelimb; 22 midbody scale rows; 51 paravertebral scales; dorsal and ventral scales smooth, enlarged, similar in size; ventrals in 49 transverse rows; scales on flanks and throat smaller than those on body and venter; three enlarged precloacals, the medial largest, the outer scales overlap the inner; 63 undivided subcaudal scales, anterior four paired, followed by 59 undivided scales (incomplete as the tail tip is missing); forelimb and hind limb short, pentadactyl; fingers and toes widely separated when adpressed; the first finger and toe less than one-half the second; five smooth subdigital lamellae on fourth finger; nine smooth lamellae on fourth toe. Selected scalation traits across all Southeast Asian taxa are given in Table 1.
</paragraph>
<caption id="DF2AD8DF4F00FF8AFF55F8EB106A7C96" httpUri="https://zenodo.org/record/280059/files/figure.png" pageId="3" pageNumber="59" targetBox="[209,1377,445,1907]" targetPageId="3">
<paragraph id="8BEA88574F00FF8AFF55F8EB106A7C96" blockId="3.[151,1436,1948,2001]" pageId="3" pageNumber="59">
<emphasis id="B92154454F00FF8AFF55F8EB136F7CF5" bold="true" box="[151,274,1948,1971]" pageId="3" pageNumber="59">FIGURE 2.</emphasis>
Head and anterior part of body of the holotype of
<taxonomicName id="4C55F3D44F00FF8AFCC9F8E911857CF4" box="[779,1016,1950,1971]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="59" phylum="Chordata" rank="species" species="veunsaiensis" status="sp. nov.">
<emphasis id="B92154454F00FF8AFCC9F8E911857CF4" box="[779,1016,1950,1971]" italics="true" pageId="3" pageNumber="59">Lygosoma veunsaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212E93E4F00FF8AFC3FF8EA16307CF5" box="[1021,1101,1949,1970]" pageId="3" pageNumber="59" rank="species">
<emphasis id="B92154454F00FF8AFC3FF8EA16307CF5" bold="true" box="[1021,1101,1949,1970]" pageId="3" pageNumber="59">sp. nov.</emphasis>
</taxonomicNameLabel>
(CBC00808): a) dorsal view and b) ventral view. Photo: Peter Geissler.
</paragraph>
</caption>
<caption id="DF2AD8DF4F07FF8DFF55FC24130E78CF" httpUri="https://zenodo.org/record/280060/files/figure.png" pageId="4" pageNumber="60" targetBox="[151,1392,193,733]" targetPageId="4">
<paragraph id="8BEA88574F07FF8DFF55FC24130E78CF" blockId="4.[151,1437,851,904]" pageId="4" pageNumber="60">
<emphasis id="B92154454F07FF8DFF55FC241369782F" bold="true" box="[151,276,851,874]" pageId="4" pageNumber="60">FIGURE 3.</emphasis>
Lateral view of head and anterior part of body of the holotype of
<taxonomicName id="4C55F3D44F07FF8DFC00FC2316CC782E" box="[962,1201,852,873]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="60" phylum="Chordata" rank="species" species="veunsaiensis" status="sp. nov.">
<emphasis id="B92154454F07FF8DFC00FC2316CC782E" box="[962,1201,852,873]" italics="true" pageId="4" pageNumber="60">Lygosoma veunsaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212E93E4F07FF8DFB7BFC231776782E" box="[1209,1291,852,873]" pageId="4" pageNumber="60" rank="species">
<emphasis id="B92154454F07FF8DFB7BFC231776782E" bold="true" box="[1209,1291,852,873]" pageId="4" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
(CBC00808). Photo: Peter Geissler.
</paragraph>
</caption>
<caption id="DF2AD8DF4F07FF8DFF55F94C128D7D37" httpUri="https://zenodo.org/record/280061/files/figure.png" pageId="4" pageNumber="60" targetBox="[170,1436,985,1572]" targetPageId="4">
<paragraph id="8BEA88574F07FF8DFF55F94C128D7D37" blockId="4.[151,1436,1595,1648]" pageId="4" pageNumber="60">
<emphasis id="B92154454F07FF8DFF55F94C136E7D17" bold="true" box="[151,275,1595,1618]" pageId="4" pageNumber="60">FIGURE 4.</emphasis>
Holotype of
<taxonomicName id="4C55F3D44F07FF8DFE5EF94B10F77D16" box="[412,650,1596,1617]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="60" phylum="Chordata" rank="species" species="veunsaiensis" status="sp. nov.">
<emphasis id="B92154454F07FF8DFE5EF94B10F77D16" box="[412,650,1596,1617]" italics="true" pageId="4" pageNumber="60">Lygosoma veunsaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212E93E4F07FF8DFD53F94B109F7D16" box="[657,738,1596,1617]" pageId="4" pageNumber="60" rank="species">
<emphasis id="B92154454F07FF8DFD53F94B109F7D16" bold="true" box="[657,738,1596,1617]" pageId="4" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
(CBC00808): a) dorsolateral view and b) ventral view. Photo: Peter Geissler.
</paragraph>
</caption>
<paragraph id="8BEA88574F07FF8DFF0BF9E017737CEC" blockId="4.[151,1436,1687,2035]" pageId="4" pageNumber="60">
<emphasis id="B92154454F07FF8DFF0BF9E0106F7DF7" bold="true" box="[201,530,1687,1712]" pageId="4" pageNumber="60">Coloration in preservative.</emphasis>
Upper surface of head and body brownish with dark brown mottling, darker brown and lighter irregular spotted pattern on tail; flanks lighter; supraocular darker; small dark eye spot present on posterior interparietal; two dorsolateral light beige stripes, bordered dorsally by a thin dark line, starting on neck and running down each side of the back, becoming less distinct behind the shoulder (
<figureCitation id="136E94D24F07FF8DFB42F87416BA7C5C" box="[1152,1223,1795,1819]" captionStart="FIGURE 2" captionStartId="3.[151,250,1948,1971]" captionTargetBox="[209,1377,445,1907]" captionTargetId="figure@3.[209,1378,445,1907]" captionTargetPageId="3" captionText="FIGURE 2. Head and anterior part of body of the holotype of Lygosoma veunsaiensis sp. nov. (CBC 00808): a) dorsal view and b) ventral view. Photo: Peter Geissler." httpUri="https://zenodo.org/record/280059/files/figure.png" pageId="4" pageNumber="60">Fig. 2</figureCitation>
b). Neck and back between these two stripes stained with dark brown. Lateral stripe (dark brown), bordering the light beige dorsolateral stripes ventrally, starting at the hind margin of nasal, running down on head and flanks reaching the tail (
<figureCitation id="136E94D24F07FF8DFAA1F83C12DA7CC0" captionStart="FIGURE 3" captionStartId="4.[151,250,851,874]" captionTargetBox="[151,1392,193,733]" captionTargetId="figure@4.[151,1436,193,830]" captionTargetPageId="4" captionText="FIGURE 3. Lateral view of head and anterior part of body of the holotype of Lygosoma veunsaiensis sp. nov. (CBC 00808). Photo: Peter Geissler." httpUri="https://zenodo.org/record/280060/files/figure.png" pageId="4" pageNumber="60">Figs. 3</figureCitation>
,
<figureCitation id="136E94D24F07FF8DFF71F81812B97CC0" box="[179,196,1903,1927]" captionStart="FIGURE 4" captionStartId="4.[151,250,1595,1618]" captionTargetBox="[170,1436,985,1572]" captionTargetId="figure@4.[151,1436,954,1574]" captionTargetPageId="4" captionText="FIGURE 4. Holotype of Lygosoma veunsaiensis sp. nov. (CBC 00808): a) dorsolateral view and b) ventral view. Photo: Peter Geissler." httpUri="https://zenodo.org/record/280061/files/figure.png" pageId="4" pageNumber="60">4</figureCitation>
). Beige coloration below the lateral stripe, with a pale series of brown spots forming a lateral line on the flank. Lateral side of tail beige, stained with tiny dark grey spots. Ventral surface whitish beige, immaculate.
</paragraph>
<paragraph id="8BEA88574F07FF8DFF0BF8C010B77CB4" blockId="4.[151,1436,1687,2035]" pageId="4" pageNumber="60">
<emphasis id="B92154454F07FF8DFF0BF8C013E17C97" bold="true" box="[201,412,1975,2000]" pageId="4" pageNumber="60">Coloration in life.</emphasis>
Reddish brown ground color on head and body, becoming greyish on the upper tail surface (
<figureCitation id="136E94D24F07FF8DFF5DF8AC129E7CB4" box="[159,227,2011,2035]" captionStart="FIGURE 5" captionStartId="7.[151,250,1144,1167]" captionTargetBox="[151,1436,193,1122]" captionTargetId="figure@7.[151,1436,193,1123]" captionTargetPageId="7" captionText="FIGURE 5. Holotype of Lygosoma veunsaiensis sp. nov. (CBC 0808) in life. Photo: Gabor Csorba." httpUri="https://zenodo.org/record/280062/files/figure.png" pageId="4" pageNumber="60">Fig. 5</figureCitation>
). Color pattern the same as in preservative.
</paragraph>
</subSubSection>
<caption id="DF2AD8DF4F04FF8EFF55FB0F16FB7FC9" box="[151,1158,1144,1167]" httpUri="https://zenodo.org/record/280062/files/figure.png" pageId="7" pageNumber="63" targetBox="[151,1436,193,1122]" targetPageId="7">
<paragraph id="8BEA88574F04FF8EFF55FB0F16FB7FC9" blockId="7.[151,1158,1144,1167]" box="[151,1158,1144,1167]" pageId="7" pageNumber="63">
<emphasis id="B92154454F04FF8EFF55FB0F136F7FC9" bold="true" box="[151,274,1144,1167]" pageId="7" pageNumber="63">FIGURE 5.</emphasis>
Holotype of
<taxonomicName id="4C55F3D44F04FF8EFE5BFB0D10FB7FC8" box="[409,646,1146,1167]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="63" phylum="Chordata" rank="species" species="veunsaiensis" status="sp. nov.">
<emphasis id="B92154454F04FF8EFE5BFB0D10FB7FC8" box="[409,646,1146,1167]" italics="true" pageId="7" pageNumber="63">Lygosoma veunsaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212E93E4F04FF8EFD4EFB0E10A07FC9" box="[652,733,1145,1166]" pageId="7" pageNumber="63" rank="species">
<emphasis id="B92154454F04FF8EFD4EFB0E10A07FC9" bold="true" box="[652,733,1145,1166]" pageId="7" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
(CBC0808) in life. Photo: Gabor Csorba.
</paragraph>
</caption>
<subSubSection id="C34FDBDC4F04FF81FF0BFBCC11F37CD1" lastPageId="8" lastPageNumber="64" pageId="7" pageNumber="63" type="distribution">
<paragraph id="8BEA88574F04FF8EFF0BFBCC177C7FBF" blockId="7.[151,1436,1211,2008]" pageId="7" pageNumber="63">
<emphasis id="B92154454F04FF8EFF0BFBCC131F7F93" bold="true" box="[201,354,1211,1236]" pageId="7" pageNumber="63">Distribution.</emphasis>
The scincid
<taxonomicName id="4C55F3D44F04FF8EFE32FBCB108B7F94" box="[496,758,1212,1235]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="63" phylum="Chordata" rank="species" species="veunsaiensis" status="sp. nov.">
<emphasis id="B92154454F04FF8EFE32FBCB108B7F94" box="[496,758,1212,1235]" italics="true" pageId="7" pageNumber="63">Lygosoma veunsaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212E93E4F04FF8EFD3FFBCB112D7F93" box="[765,848,1212,1236]" pageId="7" pageNumber="63" rank="species">
<emphasis id="B92154454F04FF8EFD3FFBCB112D7F93" bold="true" box="[765,848,1212,1236]" pageId="7" pageNumber="63">sp. nov</emphasis>
</taxonomicNameLabel>
is currently known only from the
<typeStatus id="54EE36F54F04FF8EFB0EFBCB16817F93" box="[1228,1276,1212,1236]" pageId="7" pageNumber="63">type</typeStatus>
locality in the Proposed Protected Forest in Veun Sai District, Ratanakiri Province, northeastern
<collectingCountry id="F342C8C74F04FF8EFBEEFB9716E27FBF" box="[1068,1183,1248,1272]" name="Cambodia" pageId="7" pageNumber="63">Cambodia</collectingCountry>
(
<figureCitation id="136E94D24F04FF8EFB6DFB97168D7FBF" box="[1199,1264,1248,1272]" captionStart="FIGURE 1" captionStartId="1.[151,250,1449,1472]" captionTargetBox="[155,1435,474,1425]" captionTargetId="figure@1.[151,1436,469,1428]" captionTargetPageId="1" captionText="FIGURE 1. Map of the Kingdom of Cambodia, showing the type locality of Lygosoma veunsaiensis sp. nov., the Veun Sai Proposed Protected Forest, Veun Sai District, Ratanakiri Province." httpUri="https://zenodo.org/record/280058/files/figure.png" pageId="7" pageNumber="63">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BEA88574F04FF8EFF0BFA7216BF7E06" blockId="7.[151,1436,1211,2008]" pageId="7" pageNumber="63">
<emphasis id="B92154454F04FF8EFF0BFA7213F17E5A" bold="true" box="[201,396,1284,1309]" pageId="7" pageNumber="63">Natural history.</emphasis>
The specimen was found on the ground, near a
<quantity id="4CAD25B24F04FF8EFC1BFA7216747E5A" box="[985,1033,1285,1309]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="63" unit="m" value="3.0">3 m</quantity>
wide slow moving section of the O'Kasieb stream during the day in the dry season in disturbed semi-deciduous evergreen forest.
</paragraph>
<paragraph id="8BEA88574F04FF81FF0BFA3911F37CD1" blockId="7.[151,1436,1211,2008]" lastBlockId="8.[151,1436,151,1942]" lastPageId="8" lastPageNumber="64" pageId="7" pageNumber="63">
<emphasis id="B92154454F04FF8EFF0BFA3913127E20" bold="true" box="[201,367,1358,1383]" pageId="7" pageNumber="63">Comparisons.</emphasis>
The new species is distinguishable from all other species of the genus
<taxonomicName id="4C55F3D44F04FF8EFB49FA27136A7ECC" authority="Hardwicke &amp; Gray, 1827" authorityName="Hardwicke &amp; Gray" authorityYear="1827" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="63" phylum="Chordata" rank="genus">
<emphasis id="B92154454F04FF8EFB49FA2716837E20" box="[1163,1278,1360,1383]" italics="true" pageId="7" pageNumber="63">Lygosoma</emphasis>
<bibRefCitation id="EFC4F5A64F04FF8EFAC6FA39136A7ECC" author="Hardwicke" pageId="7" pageNumber="63" refString="Hardwicke, T. &amp; Gray, J. E. (1827) A synopsis of the species of saurian reptiles, collected in India by Major-General Hardwicke. The Zoological Journal, London, 3, 214 - 229." type="journal article" year="1827">Hardwicke &amp; Gray, 1827</bibRefCitation>
</taxonomicName>
known to occur in mainland Southeast Asia (
<bibRefCitation id="EFC4F5A64F04FF8EFCF0FA0411C67ECC" author="Smith" box="[818,955,1395,1419]" pageId="7" pageNumber="63" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Volume II. Sauria. Taylor &amp; Francis, London, 440 pp." type="book" year="1935">Smith 1935</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F04FF8EFC09FA04161D7ECC" author="Smith" box="[971,1120,1395,1419]" pageId="7" pageNumber="63" refString="Smith, M. A. (1937 a) A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum, 39, 213 - 234." type="journal article" year="1937" yearSuffix="a">Smith 1937a</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F04FF8EFBADFA0416827ECC" author="Shreve" box="[1135,1279,1395,1419]" pageId="7" pageNumber="63" refString="Shreve, B. (1940) Reptiles and amphibians from Burma with descriptions of three new skinks. Proceeding of the New England Zoological Club, 18, 17 - 26." type="journal article" year="1940">Shreve 1940</bibRefCitation>
<bibRefCitation id="EFC4F5A64F04FF8EFAC8FA0417E57ECC" author="Taylor" box="[1290,1432,1395,1419]" pageId="7" pageNumber="63" refString="Taylor, E. H. (1963) The Lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F04FF8EFF55FAEE13537EF7" author="Bourret" box="[151,302,1432,1456]" pageId="7" pageNumber="63" refString="Bourret, R. (2009) Les Lezards de l'Indochine. Edition Chimaira, Frankfurt am Main, 624 pp. (In French)." type="book" year="2009">Bourret 2009</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F04FF8EFEFAFAEE106D7EF7" author="Nguyen" box="[312,528,1432,1457]" pageId="7" pageNumber="63" refString="Nguyen, V. S., Ho, T. C. &amp; Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main, 768 pp." type="book" year="2009">
Nguyen
<emphasis id="B92154454F04FF8EFE55FAEE13B27EF7" box="[407,463,1432,1456]" italics="true" pageId="7" pageNumber="63">et al.</emphasis>
2009
</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F04FF8EFDD9FAEE10F57EF7" author="Das" box="[539,648,1432,1456]" pageId="7" pageNumber="63" refString="Das, I. (2010) A field guide to the reptiles of South-East Asia. New Holland Publishers, London, 376 pp." type="book" year="2010">Das 2010</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F04FF8EFD50FAEF11117EF7" author="Geissler" box="[658,876,1432,1456]" pageId="7" pageNumber="63" refString="Geissler, P., Nguyen, Q. T., Phung, M. T., Van Devender, R. W., Hartmann, T., Farkas, B., Ziegler, T. &amp; Bohme, W. (2011) A review of Indochinese skinks of the genus Lygosoma Hardwicke &amp; Gray, 1827 (Squamata: Scincidae), with natural history notes and an identification key. Biologia, 66 / 6, 1159 - 1176." type="journal article" year="2011">
Geissler
<emphasis id="B92154454F04FF8EFD37FAEE11507EF7" box="[757,813,1432,1456]" italics="true" pageId="7" pageNumber="63">et al.</emphasis>
2011
</bibRefCitation>
) by a combination of scalation and coloration features.
<taxonomicName id="4C55F3D44F04FF8EFF1CFAC9139A7E92" box="[222,487,1470,1493]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="63" phylum="Chordata" rank="species" species="veunsaiensis" status="sp. nov.">
<emphasis id="B92154454F04FF8EFF1CFAC9139A7E92" box="[222,487,1470,1493]" italics="true" pageId="7" pageNumber="63">Lygosoma veunsaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212E93E4F04FF8EFE33FACA10307E92" box="[497,589,1469,1493]" pageId="7" pageNumber="63" rank="species">
<emphasis id="B92154454F04FF8EFE33FACA10307E92" bold="true" box="[497,589,1469,1493]" pageId="7" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
differs: from
<taxonomicName id="4C55F3D44F04FF8EFD30FAC9163C7E92" authority="Gray, 1846" authorityName="Gray" authorityYear="1846" box="[754,1089,1469,1493]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="63" phylum="Chordata" rank="species" species="albopunctata">
<emphasis id="B92154454F04FF8EFD30FAC911DB7E92" box="[754,934,1469,1493]" italics="true" pageId="7" pageNumber="63">L. albopunctata</emphasis>
(
<bibRefCitation id="EFC4F5A64F04FF8EFC75FACA16447E92" author="Gray" box="[951,1081,1469,1493]" pageId="7" pageNumber="63" refString="Gray, J. E. (1846) Descriptions of some new species of Indian Lizards. Annals and Magazine of Natural History, (1) 18, 429 - 430." type="journal article" year="1846">Gray, 1846</bibRefCitation>
)
</taxonomicName>
by having medially separated supranasals instead of supranasals being in contact medially, having five supralabials instead of seven, lacking external ear openings instead of having oval ear openings, having 22 midbody scale rows instead of 2628, and having 51 paravertebral scales instead of 6372 (
<bibRefCitation id="EFC4F5A64F04FF8EFD7AF95C110C7D04" author="Boulenger" box="[696,881,1579,1603]" pageId="7" pageNumber="63" refString="Boulenger, G. A. (1887) Catalogue of the lizards in the British Museum (Natural History). Volume III. British Museum (Natural History), London, xii + 575 pp." type="book" year="1887">Boulenger 1887</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F04FF8EFCB9F95C167C7D04" author="Smith" box="[891,1025,1579,1603]" pageId="7" pageNumber="63" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Volume II. Sauria. Taylor &amp; Francis, London, 440 pp." type="book" year="1935">Smith 1935</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F04FF8EFB93F95B172B7D04" authority="Smith, 1937" authorityName="Smith" authorityYear="1937" box="[1105,1366,1579,1603]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="63" phylum="Chordata" rank="species" species="angeli">
<emphasis id="B92154454F04FF8EFB93F95B16C97D04" box="[1105,1204,1579,1603]" italics="true" pageId="7" pageNumber="63">L. angeli</emphasis>
(Smith, 1937)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, lacking external ear openings instead of having small ear openings, having five supralabials instead of seven, having 22 midbody scale rows instead of 30, and having fewer paravertebral scales (51 vs. 107115) (
<bibRefCitation id="EFC4F5A64F04FF8EFBC5F9EE16E37DF6" author="Smith" box="[1031,1182,1689,1713]" pageId="7" pageNumber="63" refString="Smith, M. A. (1937 b) Un nouveau Lezard de Cochinchine. Bulletin du Museum National d'Histoire Naturelle, 9, 366. (In French)." type="journal article" year="1937" yearSuffix="b">Smith 1937b</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F04FF8EFB69F9EE17F27DF6" author="Geissler" box="[1195,1423,1689,1714]" pageId="7" pageNumber="63" refString="Geissler, P., Nguyen, Q. T., Phung, M. T., Van Devender, R. W., Hartmann, T., Farkas, B., Ziegler, T. &amp; Bohme, W. (2011) A review of Indochinese skinks of the genus Lygosoma Hardwicke &amp; Gray, 1827 (Squamata: Scincidae), with natural history notes and an identification key. Biologia, 66 / 6, 1159 - 1176." type="journal article" year="2011">
Geissler
<emphasis id="B92154454F04FF8EFAD2F9EC17367DF5" box="[1296,1355,1690,1714]" italics="true" pageId="7" pageNumber="63">et al.</emphasis>
2011
</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F04FF8EFF1AF9B710477D91" authority="Theobald, 1868" authorityName="Theobald" authorityYear="1868" box="[216,570,1726,1751]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="63" phylum="Chordata" rank="species" species="anguinum">
<emphasis id="B92154454F04FF8EFF1AF9B713177D91" box="[216,362,1727,1751]" italics="true" pageId="7" pageNumber="63">L. anguinum</emphasis>
(
<bibRefCitation id="EFC4F5A64F04FF8EFEBEF9C9104F7D91" author="Theobald" box="[380,562,1726,1750]" pageId="7" pageNumber="63" refString="Theobald, W. (1868) Catalogue of the reptiles of British Birma, embracing the provinces of Pegu, Martaban, and Tenasserim; with descriptions of new or little-known species. The Journal of the Linnean Society of London, 10, 4 - 67." type="journal article" year="1868">Theobald, 1868</bibRefCitation>
)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, lacking external ear openings instead of having small rounded ear openings, having five supralabials instead of seven, having scaly lower eyelids instead of eyelids bearing transparent discs, and by the presence of dark dorsolateral stripes instead of having a uniform olive-brown back (
<bibRefCitation id="EFC4F5A64F04FF8EFBAAF85A17647C02" author="Theobald" box="[1128,1305,1837,1861]" pageId="7" pageNumber="63" refString="Theobald, W. (1868) Catalogue of the reptiles of British Birma, embracing the provinces of Pegu, Martaban, and Tenasserim; with descriptions of new or little-known species. The Journal of the Linnean Society of London, 10, 4 - 67." type="journal article" year="1868">Theobald 1868</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F04FF8EFAE5F85A12AE7C2E" author="Boulenger" pageId="7" pageNumber="63" refString="Boulenger, G. A. (1887) Catalogue of the lizards in the British Museum (Natural History). Volume III. British Museum (Natural History), London, xii + 575 pp." type="book" year="1887">Boulenger 1887</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F04FF8EFEE4F824100A7C2E" authority="Bartlett, 1895" authorityName="Bartlett" authorityYear="1895" box="[294,631,1873,1898]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="63" phylum="Chordata" rank="species" species="bampfyldei">
<emphasis id="B92154454F04FF8EFEE4F82413BC7C2D" box="[294,449,1874,1898]" italics="true" pageId="7" pageNumber="63">L. bampfyldei</emphasis>
(
<bibRefCitation id="EFC4F5A64F04FF8EFE13F82610127C2E" author="Bartlett" box="[465,623,1873,1897]" pageId="7" pageNumber="63" refString="Bartlett, E. (1895) The crocodiles and lizards of Borneo in the Sarawak Museum, with descriptions of supposed new species, and the variation of colours in the several species during life. Journal of the Straits Branch of the Royal Asiatic Society, 28, 73 - 96." type="journal article" year="1895">Bartlett, 1895</bibRefCitation>
)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, lacking external ear openings instead of having small ear openings with three projecting lobules, lacking a deep groove from below the eye to nostril, having five supralabials instead of six, having 22 midbody scale rows instead of 3840, having nine subdigital lamellae on fourth toe instead of 14 (
<bibRefCitation id="EFC4F5A64F04FF8EFAC2F8B717E57C9F" author="Bartlett" box="[1280,1432,1984,2008]" pageId="7" pageNumber="63" refString="Bartlett, E. (1895) The crocodiles and lizards of Borneo in the Sarawak Museum, with descriptions of supposed new species, and the variation of colours in the several species during life. Journal of the Straits Branch of the Royal Asiatic Society, 28, 73 - 96." type="journal article" year="1895">Bartlett 1895</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FF55FFE013497BE8" author="Grismer" box="[151,308,151,175]" pageId="8" pageNumber="64" refString="Neang, T., Holden, J., Eastoe, T., Seng, R., Ith, S. &amp; Grismer, L. L. (2011) A new species of Dibamus (Squamata: Dibamidae) from Phnom Samkos Wildlife Sanctuary, southwestern Cardamom Mountain, Cambodia. Zootaxa, 2828, 58 - 68." type="journal article" year="2011">Grismer 2011</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FE47FFEE165F7BE8" authority="Ziegler, Schmitz, Heidrich, Vu &amp; Nguyen, 2007" authorityName="Ziegler, Schmitz, Heidrich, Vu &amp; Nguyen" authorityYear="2007" box="[389,1058,151,176]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="boehmei">
<emphasis id="B92154454F0BFF81FE47FFEE107F7BF7" box="[389,514,152,176]" italics="true" pageId="8" pageNumber="64">L. boehmei</emphasis>
<bibRefCitation id="EFC4F5A64F0BFF81FDCBFFE0165F7BE8" author="Ziegler" box="[521,1058,151,176]" pageId="8" pageNumber="64" refString="Ziegler, T., Schmitz, A., Heidrich, A., Vu, N. T. &amp; Nguyen, Q. T. (2007) A new species of Lygosoma (Squamata: Sauria: Scincidae) from the central Truong Son, Vietnam, with notes on its molecular phylogenetic position. Revue Suisse de Zoologie, 114, 397 - 415." type="journal article" year="2007">Ziegler, Schmitz, Heidrich, Vu &amp; Nguyen, 2007</bibRefCitation>
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, lacking external ear openings instead of having small ovoid ear openings, having five supralabials instead of seven, having 22 midbody scale rows instead of 32, having 51 paravertebral scales instead of 66, and having nine subdigital lamellae on fourth toe instead of 14 (
<bibRefCitation id="EFC4F5A64F0BFF81FF5CFE5C130E7A04" author="Ziegler" box="[158,371,299,323]" pageId="8" pageNumber="64" refString="Ziegler, T., Schmitz, A., Heidrich, A., Vu, N. T. &amp; Nguyen, Q. T. (2007) A new species of Lygosoma (Squamata: Sauria: Scincidae) from the central Truong Son, Vietnam, with notes on its molecular phylogenetic position. Revue Suisse de Zoologie, 114, 397 - 415." type="journal article" year="2007">
Ziegler
<emphasis id="B92154454F0BFF81FF3AFE5B13527A04" box="[248,303,299,323]" italics="true" pageId="8" pageNumber="64">et al.</emphasis>
2007
</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FE06FE5B11757A04" authority="Gunther, 1864" authorityName="Gunther" authorityYear="1864" box="[452,776,299,323]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="bowringii">
<emphasis id="B92154454F0BFF81FE06FE5B10337A04" box="[452,590,299,323]" italics="true" pageId="8" pageNumber="64">L. bowringii</emphasis>
(
<bibRefCitation id="EFC4F5A64F0BFF81FD9FFE5C117D7A04" author="Gunther" box="[605,768,299,323]" pageId="8" pageNumber="64" refString="Gunther, A. C. L. G. (1864) The Reptiles of British India. Royal Society, London, xxvii + 452 pp." type="book" year="1864">Günther, 1864</bibRefCitation>
)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, lacking external ear openings instead of having small rounded ear openings, having five supralabials instead of seven supralabials, having 22 midbody scale rows instead of 28, having nine subdigital lamellae on fourth toe instead of 12 or 13, and lacking black and white spotted flanks (
<bibRefCitation id="EFC4F5A64F0BFF81FAFCFEEE12AF7A91" author="Gunther" pageId="8" pageNumber="64" refString="Gunther, A. C. L. G. (1864) The Reptiles of British India. Royal Society, London, xxvii + 452 pp." type="book" year="1864">Günther 1864</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FF1EFEC9131C7A91" author="Smith" box="[220,353,446,470]" pageId="8" pageNumber="64" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Volume II. Sauria. Taylor &amp; Francis, London, 440 pp." type="book" year="1935">Smith 1935</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FEA9FEC9138B7A91" author="Taylor" box="[363,502,446,470]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1963) The Lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FDC2FEC910A17A91" author="Geissler" box="[512,732,446,470]" pageId="8" pageNumber="64" refString="Geissler, P., Nguyen, Q. T., Phung, M. T., Van Devender, R. W., Hartmann, T., Farkas, B., Ziegler, T. &amp; Bohme, W. (2011) A review of Indochinese skinks of the genus Lygosoma Hardwicke &amp; Gray, 1827 (Squamata: Scincidae), with natural history notes and an identification key. Biologia, 66 / 6, 1159 - 1176." type="journal article" year="2011">
Geissler
<emphasis id="B92154454F0BFF81FDA1FEC810E77A91" box="[611,666,446,470]" italics="true" pageId="8" pageNumber="64">et al.</emphasis>
2011
</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FCEFFEC816047A91" authority="Smith, 1921" authorityName="Smith" authorityYear="1921" box="[813,1145,446,470]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="corpulentum">
<emphasis id="B92154454F0BFF81FCEFFEC811A57A91" box="[813,984,446,470]" italics="true" pageId="8" pageNumber="64">L. corpulentum</emphasis>
(
<bibRefCitation id="EFC4F5A64F0BFF81FC24FEC9160C7A91" author="Smith" box="[998,1137,446,470]" pageId="8" pageNumber="64" refString="Smith, M. A. (1921) New or little known reptiles and batrachians from southern Annam (Indo-China). Proceedings of the Zoological Society of London, 29, 423 - 440." type="journal article" year="1921">Smith, 1921</bibRefCitation>
)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, lacking external ear openings instead of having subcircular ear openings, having five supralabials instead of seven, having 22 midbody scale rows instead of 3640, having 51 paravertebral scales instead of 7886, and bearing dorsolateral stripes instead of having a uniformly brown back (
<bibRefCitation id="EFC4F5A64F0BFF81FE42FD26107A792E" author="Smith" box="[384,519,593,617]" pageId="8" pageNumber="64" refString="Smith, M. A. (1921) New or little known reptiles and batrachians from southern Annam (Indo-China). Proceedings of the Zoological Society of London, 29, 423 - 440." type="journal article" year="1921">Smith 1921</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FDD6FD261088792E" author="Geissler" box="[532,757,593,617]" pageId="8" pageNumber="64" refString="Geissler, P., Nguyen, Q. T., Phung, M. T., Van Devender, R. W., Hartmann, T., Farkas, B., Ziegler, T. &amp; Bohme, W. (2011) A review of Indochinese skinks of the genus Lygosoma Hardwicke &amp; Gray, 1827 (Squamata: Scincidae), with natural history notes and an identification key. Biologia, 66 / 6, 1159 - 1176." type="journal article" year="2011">
Geissler
<emphasis id="B92154454F0BFF81FDBAFD2510CF792E" box="[632,690,593,617]" italics="true" pageId="8" pageNumber="64">et al.</emphasis>
2011
</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FC8BFD2516C7792E" authority="Taylor, 1962" authorityName="Taylor" authorityYear="1962" box="[841,1210,593,617]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="frontoparietale">
<emphasis id="B92154454F0BFF81FC8BFD25166D792E" box="[841,1040,593,617]" italics="true" pageId="8" pageNumber="64">L. frontoparietale</emphasis>
(
<bibRefCitation id="EFC4F5A64F0BFF81FBE2FD2616CF792E" author="Taylor" box="[1056,1202,593,617]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1962) New Oriental reptiles. The University of Kansas Science Bulletin, 43, 209 - 263." type="journal article" year="1962">Taylor, 1962</bibRefCitation>
)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, lacking external ear openings instead of having small ear openings, having five supra- and infralabials instead of seven supralabials and six infralabials, having 22 midbody scale rows instead of 2830, and by having nine subdigital lamellae on fourth toe instead of 13 (
<bibRefCitation id="EFC4F5A64F0BFF81FEFDFD9313B379BB" author="Taylor" box="[319,462,740,764]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1962) New Oriental reptiles. The University of Kansas Science Bulletin, 43, 209 - 263." type="journal article" year="1962">Taylor 1962</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FE1FFD93106479BB" author="Taylor" box="[477,537,740,764]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1963) The Lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">1963</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FDB2FD9111A979BB" authority="Taylor, 1962" authorityName="Taylor" authorityYear="1962" box="[624,980,740,765]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="haroldyoungi">
<emphasis id="B92154454F0BFF81FDB2FD91115A79BA" box="[624,807,741,765]" italics="true" pageId="8" pageNumber="64">L. haroldyoungi</emphasis>
(
<bibRefCitation id="EFC4F5A64F0BFF81FCFAFD9311B179BB" author="Taylor" box="[824,972,740,764]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1962) New Oriental reptiles. The University of Kansas Science Bulletin, 43, 209 - 263." type="journal article" year="1962">Taylor, 1962</bibRefCitation>
)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, lacking external ear openings instead of having ear openings, having five supra- and infralabials instead of nine supra- and infralabials, having 22 midbody scale rows instead of 3842, having 51 paravertebral scales instead of 141145, having nine subdigital lamellae on fourth toe instead of six or seven, and by bearing dorsolateral stripes instead of 2232 transverse dark bands (
<bibRefCitation id="EFC4F5A64F0BFF81FACBFC0017E578C8" author="Taylor" box="[1289,1432,887,911]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1962) New Oriental reptiles. The University of Kansas Science Bulletin, 43, 209 - 263." type="journal article" year="1962">Taylor 1962</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FF55FCEB12AE78F3" author="Taylor" box="[151,211,924,948]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1963) The Lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">1963</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FEE5FCE9103B78F3" authority="Smith, 1916" authorityName="Smith" authorityYear="1916" box="[295,582,924,949]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="herberti">
<emphasis id="B92154454F0BFF81FEE5FCE913DD78F2" box="[295,416,925,949]" italics="true" pageId="8" pageNumber="64">L. herberti</emphasis>
(
<bibRefCitation id="EFC4F5A64F0BFF81FE72FCEB104378F3" author="Smith" box="[432,574,924,948]" pageId="8" pageNumber="64" refString="Smith, M. A. (1916) Description of three new lizards and a new snake from Siam. The Journal of the Natural History Society of Siam, 2, 44 - 47." type="journal article" year="1916">Smith, 1916</bibRefCitation>
)
</taxonomicName>
it differes by having medially separated supranasals instead of supranasals being in contact medially, by having five supra- and infralabials instead of seven supralabials and six infralabials, by having 22 midbody scale rows instead of 2630, having smooth instead of carinated body scales, and having nine subdigital lamellae on fourth toe instead of 15 (
<bibRefCitation id="EFC4F5A64F0BFF81FD18FB7C111A7F64" author="Taylor" box="[730,871,1035,1059]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1963) The Lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FC75FB7B17687F64" authority="Gunther, 1864" authorityName="Gunther" authorityYear="1864" box="[951,1301,1035,1059]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="isodactylum">
<emphasis id="B92154454F0BFF81FC75FB7B16217F64" box="[951,1116,1035,1059]" italics="true" pageId="8" pageNumber="64">L. isodactylum</emphasis>
(
<bibRefCitation id="EFC4F5A64F0BFF81FBA8FB7C17707F64" author="Gunther" box="[1130,1293,1035,1059]" pageId="8" pageNumber="64" refString="Gunther, A. C. L. G. (1864) The Reptiles of British India. Royal Society, London, xxvii + 452 pp." type="book" year="1864">Günther, 1864</bibRefCitation>
)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, nasals not being fused with supranasals instead of being fused anteriorly, having five supralabials instead of seven, having 22 midbody scale rows instead of 3034, having 51 paravertebral scales instead of 8898, and lacking a transverse dark band between the ears (
<bibRefCitation id="EFC4F5A64F0BFF81FEA2FBE913827FF1" author="Gunther" box="[352,511,1182,1206]" pageId="8" pageNumber="64" refString="Gunther, A. C. L. G. (1864) The Reptiles of British India. Royal Society, London, xxvii + 452 pp." type="book" year="1864">Günther 1864</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FDC8FBE910EB7FF1" author="Taylor" box="[522,662,1182,1206]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1963) The Lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FD25FBE816697FF1" authority="Smith, 1917" authorityName="Smith" authorityYear="1917" box="[743,1044,1182,1206]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="koratense">
<emphasis id="B92154454F0BFF81FD25FBE8110F7FF1" box="[743,882,1182,1206]" italics="true" pageId="8" pageNumber="64">L. koratense</emphasis>
(
<bibRefCitation id="EFC4F5A64F0BFF81FC43FBE916717FF1" author="Smith" box="[897,1036,1182,1206]" pageId="8" pageNumber="64" refString="Smith, M. A. (1917) Descriptions of new reptiles and a new batrachian from Siam. The Journal of the Natural History Society of Siam, 2, 221 - 225." type="journal article" year="1917">Smith, 1917</bibRefCitation>
)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, lacking external ear openings instead of having ear openings, five supralabials instead of eight, having 22 midbody scale rows instead of 3234, having nine subdigital lamellae on fourth toe instead of 13 or 14, and the absence of a dark blotch on the base of each dorsal and lateral body scale (
<bibRefCitation id="EFC4F5A64F0BFF81FEAAFA4613937E0E" author="Smith" box="[360,494,1329,1353]" pageId="8" pageNumber="64" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Volume II. Sauria. Taylor &amp; Francis, London, 440 pp." type="book" year="1935">Smith 1935</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FE3BFA4610F97E0E" author="Taylor" box="[505,644,1329,1353]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1963) The Lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FD16FA45165E7E0E" authority="Stolicka, 1870" authorityName="Stolicka" authorityYear="1870" box="[724,1059,1329,1353]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="lineolatum">
<emphasis id="B92154454F0BFF81FD16FA4511177E0E" box="[724,874,1329,1353]" italics="true" pageId="8" pageNumber="64">L. lineolatum</emphasis>
(
<bibRefCitation id="EFC4F5A64F0BFF81FCBAFA4616667E0E" author="Stolicka" box="[888,1051,1329,1353]" pageId="8" pageNumber="64" refString="Stoliczka, F. (1870) Observations of some Indian and Malayan Amphibia and Reptilia. Annals and Magazine of Natural History, (4) 6, 105 - 109." type="journal article" year="1870">Stolicka, 1870</bibRefCitation>
)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, having two frontoparietals instead of one, having five supralabials instead of seven, lacking external ear openings instead of having moderately rounded ear openings, having five subdigital lamellae on fourth finger instead of 12, and lacking light longitudinal lines along the back and flanks (
<bibRefCitation id="EFC4F5A64F0BFF81FE42FAB310747E9B" author="Smith" box="[384,521,1476,1500]" pageId="8" pageNumber="64" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Volume II. Sauria. Taylor &amp; Francis, London, 440 pp." type="book" year="1935">Smith 1935</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FD99FAB111277E9B" authority="Shreve, 1940" authorityName="Shreve" authorityYear="1940" box="[603,858,1476,1501]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="popae">
<emphasis id="B92154454F0BFF81FD99FAB110C37E9A" box="[603,702,1478,1501]" italics="true" pageId="8" pageNumber="64">L. popae</emphasis>
<bibRefCitation id="EFC4F5A64F0BFF81FD07FAB311277E9B" author="Shreve" box="[709,858,1476,1500]" pageId="8" pageNumber="64" refString="Shreve, B. (1940) Reptiles and amphibians from Burma with descriptions of three new skinks. Proceeding of the New England Zoological Club, 18, 17 - 26." type="journal article" year="1940">Shreve, 1940</bibRefCitation>
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, lacking external ear openings instead of having minute subcircular ear openings, by having 22 instead of 24 scale rows around midbody, having 51 instead of 9295 paravertebral scales, and having a scaly lower eyelid instead of an eyelid bearing a transparent disc (
<bibRefCitation id="EFC4F5A64F0BFF81FB99F94416937D0C" author="Shreve" box="[1115,1262,1587,1611]" pageId="8" pageNumber="64" refString="Shreve, B. (1940) Reptiles and amphibians from Burma with descriptions of three new skinks. Proceeding of the New England Zoological Club, 18, 17 - 26." type="journal article" year="1940">Shreve 1940</bibRefCitation>
); from
<taxonomicName id="4C55F3D44F0BFF81FAFDF94313F17D28" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="punctata">
<emphasis id="B92154454F0BFF81FAFDF94312BE7D37" italics="true" pageId="8" pageNumber="64">L. punctata</emphasis>
(
<bibRefCitation id="EFC4F5A64F0BFF81FF11F92013F97D28" author="Linnaeus" box="[211,388,1623,1647]" pageId="8" pageNumber="64" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. 10 th Edition, Laurenti Salvi, Stockholm, 824 pp (In Latin)" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
it differs by having medially separated supranasals instead of supranasals being in contact medially, the absence of an undivided transparent disc in the lower eyelid, lacking external ear openings instead of having small oval ear openings, having 22 midbody scale rows instead of 2428, having nine subdigital lamellae on fourth toe instead of 1114 (
<bibRefCitation id="EFC4F5A64F0BFF81FDC3F9B110F77D99" author="Smith" box="[513,650,1734,1758]" pageId="8" pageNumber="64" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Volume II. Sauria. Taylor &amp; Francis, London, 440 pp." type="book" year="1935">Smith 1935</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FD54F9B0114F7D99" author="Bourret" box="[662,818,1734,1758]" pageId="8" pageNumber="64" refString="Bourret, R. (2009) Les Lezards de l'Indochine. Edition Chimaira, Frankfurt am Main, 624 pp. (In French)." type="book" year="2009">Bourret 2009</bibRefCitation>
); and from
<taxonomicName id="4C55F3D44F0BFF81FC75F9B0176A7D99" authority="Linnaeus, 1766" authorityName="Linnaeus" authorityYear="1766" box="[951,1303,1734,1758]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="quadrupes">
<emphasis id="B92154454F0BFF81FC75F9B016317D99" box="[951,1100,1734,1758]" italics="true" pageId="8" pageNumber="64">L. quadrupes</emphasis>
(
<bibRefCitation id="EFC4F5A64F0BFF81FB9FF9B117727D99" author="Linnaeus" box="[1117,1295,1734,1758]" pageId="8" pageNumber="64" refString="Linnaeus, C. (1766) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio duodecima, reformata. Tomus I, Pars I. Laurenti Salvi, Stockholm, 532 pp. (In Latin)." type="book" year="1766">Linnaeus, 1766</bibRefCitation>
)
</taxonomicName>
it difers by having distinct nasals and supranasals instead of nasals being fused with supranasals, having two frontoparietals instead of a single one, having five supra- and infralabials instead of six or seven, lacking external ear openings instead of having small punctiform ear openings, having 22 midbody scale rows instead of 2428, having 51 paravertebral scales instead of 109116, and having two broad dark dorsolateral stripes along flanks and tail (
<bibRefCitation id="EFC4F5A64F0BFF81FAF1F82E12AF7CD1" author="Linnaeus" pageId="8" pageNumber="64" refString="Linnaeus, C. (1766) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio duodecima, reformata. Tomus I, Pars I. Laurenti Salvi, Stockholm, 532 pp. (In Latin)." type="book" year="1766">Linnaeus 1766</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FF1FF809131F7CD1" author="Smith" box="[221,354,1918,1942]" pageId="8" pageNumber="64" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Volume II. Sauria. Taylor &amp; Francis, London, 440 pp." type="book" year="1935">Smith 1935</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FEAFF809138B7CD1" author="Taylor" box="[365,502,1918,1942]" pageId="8" pageNumber="64" refString="Taylor, E. H. (1963) The Lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FDC0F80810E77CD1" author="Bourret" box="[514,666,1918,1942]" pageId="8" pageNumber="64" refString="Bourret, R. (2009) Les Lezards de l'Indochine. Edition Chimaira, Frankfurt am Main, 624 pp. (In French)." type="book" year="2009">Bourret 2009</bibRefCitation>
,
<bibRefCitation id="EFC4F5A64F0BFF81FD67F80911FF7CD1" author="Geissler" box="[677,898,1918,1942]" pageId="8" pageNumber="64" refString="Geissler, P., Nguyen, Q. T., Phung, M. T., Van Devender, R. W., Hartmann, T., Farkas, B., Ziegler, T. &amp; Bohme, W. (2011) A review of Indochinese skinks of the genus Lygosoma Hardwicke &amp; Gray, 1827 (Squamata: Scincidae), with natural history notes and an identification key. Biologia, 66 / 6, 1159 - 1176." type="journal article" year="2011">
Geissler
<emphasis id="B92154454F0BFF81FCCAF808113D7CD1" box="[776,832,1918,1942]" italics="true" pageId="8" pageNumber="64">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>