treatments-xml/data/03/B2/C0/03B2C019B109FFF4F3CDFED5FD552A28.xml
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<document id="1A9A66729C177E69C03540442EF13AB5" ID-CLB-Dataset="127383" ID-DOI="10.1080/00222933.2022.2092429" ID-GBIF-Dataset="3cf6a07e-1580-4cfd-b8b6-74f51dbd72d7" ID-ISSN="1464-5262" ID-Zenodo-Dep="7012096" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660989733513" checkinUser="plazi" docAuthor="Carvalho, Edrielly Carolinne &amp; Vaz-de-Mello, Fernando Z." docDate="2022" docId="03B2C019B109FFF4F3CDFED5FD552A28" docLanguage="en" docName="JNATHIST.56.13-16.769-828.pdf" docOrigin="Journal of Natural History 56 (13 - 16)" docSource="http://dx.doi.org/10.1080/00222933.2022.2092429" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Bolbapium modestum" docType="treatment" docVersion="7" lastPageNumber="804" masterDocId="FF8BB861B12BFFD7F26FFFF8FFC32B1C" masterDocTitle="A taxonomic revision of Bolbapium Boucomont, 1910 (Coleoptera: Scarabaeoidea: Geotrupidae)" masterLastPageNumber="828" masterPageNumber="769" pageNumber="803" updateTime="1699370716173" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6EA873AD0A30B16307D30F4FB9C5571A">A taxonomic revision of Bolbapium Boucomont, 1910 (Coleoptera: Scarabaeoidea: Geotrupidae)</mods:title>
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<mods:namePart id="9151DF869E7E28FCF4B9D16811AF0F52">Carvalho, Edrielly Carolinne</mods:namePart>
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<mods:namePart id="15DC558A4ADF738DAC8EC81A8C9027B7">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="671D68376FBEE4770983BB550765B8A3">Journal of Natural History</mods:title>
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<mods:date id="8E0E670167F716ACB0C24815C64364F7">2022</mods:date>
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<treatment id="03B2C019B109FFF4F3CDFED5FD552A28" ID-DOI="http://doi.org/10.5281/zenodo.7017292" ID-GBIF-Taxon="199724772" ID-Zenodo-Dep="7017292" LSID="urn:lsid:plazi:treatment:03B2C019B109FFF4F3CDFED5FD552A28" httpUri="http://treatment.plazi.org/id/03B2C019B109FFF4F3CDFED5FD552A28" lastPageId="35" lastPageNumber="804" pageId="34" pageNumber="803">
<subSubSection id="C3012284B109FFF5F3CDFED5FC422A5B" box="[418,897,301,327]" pageId="34" pageNumber="803" type="nomenclature">
<paragraph id="8BA4710FB109FFF5F3CDFED5FC422A5B" blockId="34.[418,897,301,327]" box="[418,897,301,327]" pageId="34" pageNumber="803">
<heading id="D0ECC663B109FFF5F3CDFED5FC422A5B" box="[418,897,301,327]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="803" reason="5">
<taxonomicName id="4C1B0A8CB109FFF5F3CDFED5FC422A5B" authority="(Castelnau, 1840)" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[418,897,301,327]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="modestum">
<emphasis id="B96FAD1DB109FFF5F3CDFED5FD6C2A5B" bold="true" box="[418,687,301,327]" italics="true" pageId="34" pageNumber="803">Bolbapium modestum</emphasis>
(
<bibRefCitation id="EF8A0CFEB109FFF5F0D2FED5FCB82A5B" author="Castelnau" box="[701,891,301,327]" pageId="34" pageNumber="803" pagination="564" refId="ref38516" refString="Castelnau le compte de [= F. L. N. de Laporte]. 1840. Histoire Naturelle des Insectes Coleopteres. Vol. 2. Paris: P. Dumenil; p. 564." type="book chapter" year="1840">Castelnau, 1840</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3012284B109FFF5F3BFFE9EFC902A9C" box="[464,851,358,384]" pageId="34" pageNumber="803" type="description">
<paragraph id="8BA4710FB109FFF5F3BFFE9EFC902A9C" blockId="34.[464,851,358,384]" box="[464,851,358,384]" pageId="34" pageNumber="803">
(
<figureCitation id="13206D8AB109FFF5F3B7FE9EFD862A9C" box="[472,581,358,384]" captionStart="Figure 6" captionStartId="13.[160,225,1433,1456]" captionTargetBox="[166,1149,148,1396]" captionTargetId="figure-3@13.[163,1153,142,1401]" captionTargetPageId="13" captionText="Figure 6. (a). Pronotum with almost rectangular structure, Bolbapium modestum (Castelnau, 1840); (b). head with tubercle adjacent to each eye, Bolbapium borgmeieri Martínez, 1976; (c). head without tubercles, Bolbapium matheusi Carvalho and Vaz-de-Mello, sp. nov.; (d). pronotum with curved structure towards the head, without angulations, Bolbapium howdeni Martínez, 1976; (e). head with tubercles wider than long; pronotum with rectangular structure, Bolbapium sampaioorum Carvalho and Vaz-de-Mello, sp. nov.; (f). head with tubercle narrow adjacent to each eye; pronotum with curved structure, Bolbapium caesum (Klug, 1843)." figureDoi="http://doi.org/10.5281/zenodo.7012114" httpUri="https://zenodo.org/record/7012114/files/figure.png" pageId="34" pageNumber="803">Figures 6</figureCitation>
(a), 10(a), (b), (c), 15(b))
</paragraph>
</subSubSection>
<subSubSection id="C3012284B109FFF5F2CFFE63FD392900" pageId="34" pageNumber="803" type="reference_group">
<paragraph id="8BA4710FB109FFF5F2CFFE63FCA72AA8" blockId="34.[160,1156,411,540]" box="[160,868,411,436]" pageId="34" pageNumber="803">
<treatmentCitationGroup id="AB0B5621B109FFF5F2CFFE63FCA72AA8" box="[160,868,411,436]" pageId="34" pageNumber="803">
<treatmentCitation id="0ABA571EB109FFF5F2CFFE63FDB72AA8" author="Castelnau" box="[160,628,411,436]" page="105" pageId="34" pageNumber="803" year="1840">
<taxonomicName id="4C1B0A8CB109FFF5F2CFFE63FDB72AA8" authority="Castelnau, 1840: 105" authorityName="Castelnau" authorityPageNumber="105" authorityYear="1840" box="[160,628,411,436]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="modestus">
<emphasis id="B96FAD1DB109FFF5F2CFFE63FE462AA8" box="[160,389,411,436]" italics="true" pageId="34" pageNumber="803">Bolboceras modestus</emphasis>
<bibRefCitation id="EF8A0CFEB109FFF5F3E3FE63FDB72AA8" author="Castelnau" box="[396,628,411,436]" pageId="34" pageNumber="803" pagination="564" refId="ref38516" refString="Castelnau le compte de [= F. L. N. de Laporte]. 1840. Histoire Naturelle des Insectes Coleopteres. Vol. 2. Paris: P. Dumenil; p. 564." type="book chapter" year="1840">Castelnau, 1840: 105</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA4710FB109FFF5F2CFFE46FC6E2AE5" blockId="34.[160,1156,411,540]" pageId="34" pageNumber="803">
<treatmentCitationGroup id="AB0B5621B109FFF5F2CFFE46FC6E2AE5" pageId="34" pageNumber="803">
<treatmentCitation id="0ABA571EB109FFF5F2CFFE46FCE72ACB" author="Gemminger D &amp; Harold B" box="[160,804,446,471]" page="1079" pageId="34" pageNumber="803" year="1869">
<taxonomicName id="4C1B0A8CB109FFF5F2CFFE46FCE72ACB" authority="Gemminger &amp; Harold, 1869" authorityName="Gemminger &amp; Harold, 1869" authorityPageNumber="1079" authorityYear="1869" box="[160,804,446,471]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="modestum">
<emphasis id="B96FAD1DB109FFF5F2CFFE46FE4C2ACB" box="[160,399,446,471]" italics="true" pageId="34" pageNumber="803">Bolboceras modestum</emphasis>
:
<bibRefCitation id="EF8A0CFEB109FFF5F3F3FE46FCE72ACB" author="Gemminger D &amp; Harold B" box="[412,804,446,471]" pageId="34" pageNumber="803" refId="ref38607" refString="Gemminger D, and Harold B. 1869. Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Monachii: Gummi." type="book" year="1869">Gemminger and Harold 1869: 1079</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0ABA571EB109FFF5F1DBFE46FF2F2AE5" author="Luederwaldt H" page="430" pageId="34" pageNumber="803" year="1931">
<bibRefCitation id="EF8A0CFEB109FFF5F1DBFE46FF2F2AE5" author="Luederwaldt H" pageId="34" pageNumber="803" pagination="427 - 454" refId="ref39038" refString="Luederwaldt H. 1931. As especies Sul-Americanas de Bolboceras. Revista Do Museu Paulista. 17: 427 - 454." type="journal article" year="1931">Luederwaldt 1931: 430</bibRefCitation>
</treatmentCitation>
, 436, 437 (review, key),
<bibRefCitation id="EF8A0CFEB109FFF5F399FE18FD552AE5" author="Martinez A" box="[502,662,480,505]" pageId="34" pageNumber="803" pagination="531 - 551" refId="ref39092" refString="Martinez A. 1976. Contribution al conocimento de los Bolboceratini sudamericanos (Coleoptera, Scarabaeidae, Geotrupinae, Bolboceratini). Studia Entomologica. 19 (1 / 4): 531 - 551." type="journal article" year="1976">Martínez 1976</bibRefCitation>
(checklist,
<emphasis id="B96FAD1DB109FFF5F17CFE18FC662AE5" box="[787,933,480,505]" italics="true" pageId="34" pageNumber="803">incertae sedis</emphasis>
)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA4710FB109FFF5F2CFFDFBFD392900" blockId="34.[160,1156,411,540]" box="[160,762,515,540]" pageId="34" pageNumber="803">
<treatmentCitationGroup id="AB0B5621B109FFF5F2CFFDFBFD392900" box="[160,762,515,540]" pageId="34" pageNumber="803">
<treatmentCitation id="0ABA571EB109FFF5F2CFFDFBFD412900" author="Blackwelder RE" box="[160,642,515,540]" page="220" pageId="34" pageNumber="803" year="1944">
<taxonomicName id="4C1B0A8CB109FFF5F2CFFDFBFD412900" authority=": Blackwelder 1944: 220" authorityName="Blackwelder" authorityPageNumber="220" authorityYear="1944" box="[160,642,515,540]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B96FAD1DB109FFF5F2CFFDFBFEB82900" box="[160,379,515,540]" italics="true" pageId="34" pageNumber="803">Bolboceras modesta</emphasis>
:
<bibRefCitation id="EF8A0CFEB109FFF5F3E7FDFBFD412900" author="Blackwelder RE" box="[392,642,515,540]" pageId="34" pageNumber="803" pagination="1 - 1492" refId="ref38308" refString="Blackwelder RE. 1944. Checklist of the Coleopterous insects of Mexico, Central America The West Indies, and South America, Part 2. Smithsonian Inst US Nat Mus. 185: 1 - 1492." type="journal article" year="1944">Blackwelder 1944: 220</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(checklist)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3012284B109FFF4F2CFFDB3FD552A28" lastPageId="35" lastPageNumber="804" pageId="34" pageNumber="803" type="diagnosis">
<paragraph id="8BA4710FB109FFF5F2CFFDB3FED72979" blockId="34.[159,1156,587,717]" box="[160,276,587,613]" pageId="34" pageNumber="803">
<emphasis id="B96FAD1DB109FFF5F2CFFDB3FED72979" bold="true" box="[160,276,587,613]" italics="true" pageId="34" pageNumber="803">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BA4710FB109FFF5F2CFFD96FB4729D1" blockId="34.[159,1156,587,717]" pageId="34" pageNumber="803">
Frons with interocular tubercles between eyes and depression in the central region between the tubercles and the clypeus. Pronotum mesoanteriorly with almost rectangular structure in dorsal view;the structure measures less than or equal to half the interocular width (
<figureCitation id="13206D8AB109FFF5F668FD4CFB9829D1" box="[1031,1115,692,717]" captionStart="Figure 6" captionStartId="13.[160,225,1433,1456]" captionTargetBox="[166,1149,148,1396]" captionTargetId="figure-3@13.[163,1153,142,1401]" captionTargetPageId="13" captionText="Figure 6. (a). Pronotum with almost rectangular structure, Bolbapium modestum (Castelnau, 1840); (b). head with tubercle adjacent to each eye, Bolbapium borgmeieri Martínez, 1976; (c). head without tubercles, Bolbapium matheusi Carvalho and Vaz-de-Mello, sp. nov.; (d). pronotum with curved structure towards the head, without angulations, Bolbapium howdeni Martínez, 1976; (e). head with tubercles wider than long; pronotum with rectangular structure, Bolbapium sampaioorum Carvalho and Vaz-de-Mello, sp. nov.; (f). head with tubercle narrow adjacent to each eye; pronotum with curved structure, Bolbapium caesum (Klug, 1843)." figureDoi="http://doi.org/10.5281/zenodo.7012114" httpUri="https://zenodo.org/record/7012114/files/figure.png" pageId="34" pageNumber="803">Figure6</figureCitation>
(a)).
</paragraph>
<paragraph id="8BA4710FB109FFF5F2CFFD03FE842809" blockId="34.[160,1156,763,1517]" box="[160,327,763,789]" pageId="34" pageNumber="803">
<emphasis id="B96FAD1DB109FFF5F2CFFD03FE842809" bold="true" box="[160,327,763,789]" italics="true" pageId="34" pageNumber="803">Redescripition</emphasis>
</paragraph>
<paragraph id="8BA4710FB109FFF5F2CFFCE7FEDD2EF1" blockId="34.[160,1156,763,1517]" pageId="34" pageNumber="803">
<emphasis id="B96FAD1DB109FFF5F2CFFCE7FED72824" bold="true" box="[160,276,799,824]" pageId="34" pageNumber="803">Body size</emphasis>
: Length:
<quantity id="4CE3DCEAB109FFF5F3EBFCE7FDE42824" box="[388,551,799,824]" metricMagnitude="-3" metricUnit="m" metricValue="5.58" metricValueMax="5.64" metricValueMin="5.52" pageId="34" pageNumber="803" unit="mm" value="5.58" valueMax="5.64" valueMin="5.52">5.525.64 mm</quantity>
. Width:
<quantity id="4CE3DCEAB109FFF5F0E7FCE7FCE82824" box="[648,811,799,824]" metricMagnitude="-3" metricUnit="m" metricValue="4.46" metricValueMax="4.57" metricValueMin="4.35" pageId="34" pageNumber="803" unit="mm" value="4.46" valueMax="4.57" valueMin="4.35">4.354.57 mm</quantity>
.
<emphasis id="B96FAD1DB109FFF5F157FCE7FCB52824" bold="true" box="[824,886,799,824]" pageId="34" pageNumber="803">Head</emphasis>
: Frons with interocular tubercles between eyes, not connected by tranverse carina; region between the tubercles depressed and punctured.
<emphasis id="B96FAD1DB109FFF5F3A2FC9CFD862861" bold="true" box="[461,581,868,893]" pageId="34" pageNumber="803">Pronotum</emphasis>
: Pronotum mesoanteriorly with almost rectangular structure in dorsal view, the structure measures less than or equal to half the interocular width. Disc with sparse points, unevenly distributed.
<emphasis id="B96FAD1DB109FFF5F164FC51FC0828DE" bold="true" box="[779,971,937,962]" pageId="34" pageNumber="803">Male terminalia</emphasis>
: In dorsal view, greatly separated parameres and posterior margin with projection beyond the apex of the parameres (broken right paramere) (
<figureCitation id="13206D8AB109FFF5F02FFC17FD6F2F14" box="[576,684,1007,1032]" captionStart="Figure 10" captionStartId="43.[160,225,1353,1376]" captionTargetBox="[166,1137,159,1318]" captionTargetId="figure-55@43.[163,1153,142,1322]" captionTargetPageId="43" captionText="Figure 10. Aedeagus. Bolbapium modestum (Castelnau, 1840) (a). dorsal view; (b). ventral view; (c). lateral view. Bolbapium parcepunctatum (Arrow, 1913) (d). dorsal view; (e). ventral view; (f). lateral view. Bolbapium santanaorum Carvalho and Vaz-de-Mello, sp. nov. (g). dorsal view; (h). ventral view; (i). lateral view. Bolbapium borgmeieri Martínez, 1976 (j). dorsal view; (k). ventral view; (l). lateral view." figureDoi="http://doi.org/10.5281/zenodo.7012122" httpUri="https://zenodo.org/record/7012122/files/figure.png" pageId="34" pageNumber="803">Figure 10</figureCitation>
(a)); In lateral view, parameres presenting curvature close to 45°, with the base released and the apex thin (
<figureCitation id="13206D8AB109FFF5F1F9FBEAFBC02F37" box="[918,1027,1042,1067]" captionStart="Figure 10" captionStartId="43.[160,225,1353,1376]" captionTargetBox="[166,1137,159,1318]" captionTargetId="figure-55@43.[163,1153,142,1322]" captionTargetPageId="43" captionText="Figure 10. Aedeagus. Bolbapium modestum (Castelnau, 1840) (a). dorsal view; (b). ventral view; (c). lateral view. Bolbapium parcepunctatum (Arrow, 1913) (d). dorsal view; (e). ventral view; (f). lateral view. Bolbapium santanaorum Carvalho and Vaz-de-Mello, sp. nov. (g). dorsal view; (h). ventral view; (i). lateral view. Bolbapium borgmeieri Martínez, 1976 (j). dorsal view; (k). ventral view; (l). lateral view." figureDoi="http://doi.org/10.5281/zenodo.7012122" httpUri="https://zenodo.org/record/7012122/files/figure.png" pageId="34" pageNumber="803">Figure 10</figureCitation>
(c)). Genital capsule with rounded lateral margins, with setae in both the lateral and central lobes.
<emphasis id="B96FAD1DB109FFF5F2CFFBAFFE572F6C" bold="true" box="[160,404,1111,1136]" pageId="34" pageNumber="803">Sexual dimorphism</emphasis>
: This species does not feature sexual dimorphism.
<emphasis id="B96FAD1DB109FFF5F67CFBAFFB472F6C" bold="true" box="[1043,1156,1111,1136]" pageId="34" pageNumber="803">Remarks:</emphasis>
<bibRefCitation id="EF8A0CFEB109FFF5F2CFFB81FE9F2F8E" author="Castelnau" box="[160,348,1145,1170]" pageId="34" pageNumber="803" pagination="564" refId="ref38516" refString="Castelnau le compte de [= F. L. N. de Laporte]. 1840. Histoire Naturelle des Insectes Coleopteres. Vol. 2. Paris: P. Dumenil; p. 564." type="book chapter" year="1840">Castelnau (1840)</bibRefCitation>
described the species apparently based on a single specimen collected in
<collectingCountry id="F30C319FB109FFF5F2AFFB64FEED2FA9" box="[192,302,1180,1205]" name="Colombia" pageId="34" pageNumber="803">Colombia</collectingCountry>
; the original description is simple and does not mention the number of specimens examined. The author mentions that the specimen belongs to the collection of M. Gory; however, the
<typeStatus id="54A0CFADB109FFF5F3D8FB19FE292FE6" box="[439,490,1249,1274]" pageId="34" pageNumber="803">type</typeStatus>
was never found. It is possible the
<typeStatus id="54A0CFADB109FFF5F1CAFB19FC1B2FE6" box="[933,984,1249,1274]" pageId="34" pageNumber="803">type</typeStatus>
is among the miscellany in MNHN; however, no one has been able to analyse it. After the review by
<bibRefCitation id="EF8A0CFEB109FFF5F2CFFADFFE832E5C" author="Martinez A" box="[160,320,1319,1344]" pageId="34" pageNumber="803" pagination="531 - 551" refId="ref39092" refString="Martinez A. 1976. Contribution al conocimento de los Bolboceratini sudamericanos (Coleoptera, Scarabaeidae, Geotrupinae, Bolboceratini). Studia Entomologica. 19 (1 / 4): 531 - 551." type="journal article" year="1976">Martínez 1976</bibRefCitation>
, the species was considered
<emphasis id="B96FAD1DB109FFF5F0EAFADFFCD42E5C" box="[645,791,1319,1344]" italics="true" pageId="34" pageNumber="803">incertae sedis</emphasis>
and later works do not mention it. Earlier, however,
<bibRefCitation id="EF8A0CFEB109FFF5F315FAB1FD872E7E" author="Luederwaldt H" box="[378,580,1353,1378]" pageId="34" pageNumber="803" pagination="427 - 454" refId="ref39038" refString="Luederwaldt H. 1931. As especies Sul-Americanas de Bolboceras. Revista Do Museu Paulista. 17: 427 - 454." type="journal article" year="1931">Luederwaldt 1931</bibRefCitation>
mentioned the species among those that currently belong to the genus
<taxonomicName id="4C1B0A8CB109FFF5F3F4FA94FDD32E99" authorityName=": Howden" authorityYear="1954" box="[411,528,1388,1413]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="genus">
<emphasis id="B96FAD1DB109FFF5F3F4FA94FDD32E99" box="[411,528,1388,1413]" italics="true" pageId="34" pageNumber="803">Bolbapium</emphasis>
</taxonomicName>
. Considering this, and carefully analysing the original description and species distribution, the transfer of the species to the genus
<taxonomicName id="4C1B0A8CB109FFF5F199FA77FBA82EB4" authorityName=": Howden" authorityYear="1954" box="[1014,1131,1423,1448]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="genus">
<emphasis id="B96FAD1DB109FFF5F199FA77FBA82EB4" box="[1014,1131,1423,1448]" italics="true" pageId="34" pageNumber="803">Bolbapium</emphasis>
</taxonomicName>
is here proposed. Thus,
<taxonomicName id="4C1B0A8CB109FFF5F3CBFA49FD5E2ED6" authorityName=": Gemminger and Harold 1869:" authorityYear="1079" box="[420,669,1457,1482]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="modestum">
<emphasis id="B96FAD1DB109FFF5F3CBFA49FD5E2ED6" box="[420,669,1457,1482]" italics="true" pageId="34" pageNumber="803">Bolboceras modestum</emphasis>
</taxonomicName>
is now considered
<taxonomicName id="4C1B0A8CB109FFF5F1E3FA49FB402ED6" authorityName="Carvalho &amp; Vaz-de-Mello" authorityYear="2022" baseAuthorityName=": Gemminger and Harold 1869:" baseAuthorityYear="1840" box="[908,1155,1457,1482]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="modestum" status="comb. nov.">
<emphasis id="B96FAD1DB109FFF5F1E3FA49FB402ED6" box="[908,1155,1457,1482]" italics="true" pageId="34" pageNumber="803">Bolbapium modestum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C1066B109FFF5F2CFFA2CFEDD2EF1" box="[160,286,1492,1517]" pageId="34" pageNumber="803" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BA4710FB109FFF5F2CFF9E4FE6C2DDC" blockId="34.[160,1156,1564,1728]" pageId="34" pageNumber="803">
<emphasis id="B96FAD1DB109FFF5F2CFF9E4FE602D2A" bold="true" box="[160,419,1564,1590]" italics="true" pageId="34" pageNumber="803">Differential diagnosis.</emphasis>
<taxonomicName id="4C1B0A8CB109FFF5F3DCF9E5FD602D2A" authorityName="Carvalho &amp; Vaz-de-Mello" authorityYear="2022" baseAuthorityName=": Gemminger and Harold 1869:" baseAuthorityYear="1840" box="[435,675,1565,1590]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="modestum">
<emphasis id="B96FAD1DB109FFF5F3DCF9E5FD602D2A" box="[435,675,1565,1590]" italics="true" pageId="34" pageNumber="803">Bolbapium modestum</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName id="4C1B0A8CB109FFF5F181F9E5FBBD2D2A" authorityName="Martinez" authorityYear="1976" box="[1006,1150,1565,1590]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="borgmeieri">
<emphasis id="B96FAD1DB109FFF5F181F9E5FBBD2D2A" box="[1006,1150,1565,1590]" italics="true" pageId="34" pageNumber="803">B. borgmeieri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1B0A8CB109FFF5F2CFF9C7FF3A2D44" authority="Carvalho &amp; Vaz-de-Mello, 2022" authorityName="Carvalho &amp; Vaz-de-Mello" authorityYear="2022" box="[160,249,1599,1624]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="boillyi" status="sp. nov.">
<emphasis id="B96FAD1DB109FFF5F2CFF9C7FF3A2D44" box="[160,249,1599,1624]" italics="true" pageId="34" pageNumber="803">B. boillyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C1066B109FFF5F290F9C7FE952D44" box="[255,342,1599,1624]" pageId="34" pageNumber="803" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="B96FAD1DB109FFF5F330F9C7FC1C2D44" box="[351,991,1599,1624]" italics="true" pageId="34" pageNumber="803">
<taxonomicName id="4C1B0A8CB109FFF5F330F9C7FE0F2D44" authorityName=": Martinez" authorityYear="1976" box="[351,460,1599,1624]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="caesum">B. caesum</taxonomicName>
,
<taxonomicName id="4C1B0A8CB109FFF5F3B9F9C7FD8D2D44" authorityName="Martinez" authorityYear="1976" box="[470,590,1599,1624]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="howdeni">B. howdeni</taxonomicName>
,
<taxonomicName id="4C1B0A8CB109FFF5F038F9C7FCE62D44" baseAuthorityName="Arrow" baseAuthorityYear="1913" box="[599,805,1599,1624]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="parcepunctatum">B. parcepunctatum</taxonomicName>
,
<taxonomicName id="4C1B0A8CB109FFF5F140F9C7FC1C2D44" authority="Carvalho &amp; Vaz-de-Mello, 2022" authorityName="Carvalho &amp; Vaz-de-Mello" authorityYear="2022" box="[815,991,1599,1624]" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="sampaioorum" status="sp. nov.">B. sampaioorum</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A25C1066B109FFF5F18AF9C7FBF82D44" box="[997,1083,1599,1624]" pageId="34" pageNumber="803" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1B0A8CB109FFF5F61FF9C7FEF02D67" authority="Carvalho &amp; Vaz-de-Mello, 2022" authorityName="Carvalho &amp; Vaz-de-Mello" authorityYear="2022" class="Insecta" family="Bolboceratidae" genus="Bolbapium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="803" phylum="Arthropoda" rank="species" species="santanaorum" status="sp. nov.">
<emphasis id="B96FAD1DB109FFF5F61FF9C7FEF02D67" italics="true" pageId="34" pageNumber="803">B. santanaorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C1066B109FFF5F353F99AFE562D67" box="[316,405,1634,1659]" pageId="34" pageNumber="803" rank="species">sp. nov.</taxonomicNameLabel>
in not presenting the transverse carina forehead interconnecting the interocular tubercles. However, it is easily differentiated by the almost rectangular structure in dorsal view.
</paragraph>
<paragraph id="8BA4710FB108FFF4F2CFFF76FD792BB4" blockId="35.[160,698,142,168]" box="[160,698,142,168]" pageId="35" pageNumber="804">
<emphasis id="B96FAD1DB108FFF4F2CFFF76FE152BB4" bold="true" box="[160,470,142,168]" italics="true" pageId="35" pageNumber="804">Geographical distribution.</emphasis>
<collectingCountry id="F30C319FB108FFF4F389FF77FD912BB4" box="[486,594,143,168]" name="Colombia" pageId="35" pageNumber="804">Colômbia</collectingCountry>
(
<collectingRegion id="49DFBFEDB108FFF4F030FF77FD732BB4" box="[607,688,143,168]" country="Colombia" name="Tolima" pageId="35" pageNumber="804">Tolima</collectingRegion>
).
</paragraph>
<paragraph id="8BA4710FB108FFF4F2CFFF2DFDE22BF3" blockId="35.[160,1155,213,308]" box="[160,545,213,239]" pageId="35" pageNumber="804">
<emphasis id="B96FAD1DB108FFF4F2CFFF2DFEBC2BF3" bold="true" box="[160,383,213,239]" italics="true" pageId="35" pageNumber="804">Non-type material.</emphasis>
<specimenCount id="9D1DBA86B108FFF4F3E0FF2EFDDE2BF3" box="[399,541,214,239]" pageId="35" pageNumber="804" type="generic">2 specimens</specimenCount>
.
</paragraph>
<paragraph id="8BA4710FB108FFF4F2AFFF01FD552A28" blockId="35.[160,1155,213,308]" pageId="35" pageNumber="804">
<materialsCitation id="3B737B52B108FFF4F2AFFF01FE1F2A0E" ID-GBIF-Occurrence="3892766683" box="[192,476,249,274]" collectorName="Chaparral" country="Colombia" location="COLOMBIA" pageId="35" pageNumber="804" specimenCount="1">
<collectingCountry id="F30C319FB108FFF4F2AFFF01FEFD2A0E" box="[192,318,249,274]" name="Colombia" pageId="35" pageNumber="804">COLOMBIA</collectingCountry>
,
<collectorName id="26EE14D9B108FFF4F329FF01FE772A0E" box="[326,436,249,274]" pageId="35" pageNumber="804">Chaparral</collectorName>
,
<date id="FFA557CFB108FFF4F3D2FF01FE1F2A0E" box="[445,476,249,274]" pageId="35" pageNumber="804" value="1998-08">viii</date>
</materialsCitation>
.1998, J
<materialsCitation id="3B737B52B108FFF4F05FFF01FD782A0E" ID-GBIF-Occurrence="3892766860" box="[560,699,249,274]" collectorName="Noriega" country="Colombia" isTailEnumeration="true" location="COLOMBIA" pageId="35" pageNumber="804" specimenCount="1">
.
<collectorName id="26EE14D9B108FFF4F057FF01FD532A0E" box="[568,656,249,274]" pageId="35" pageNumber="804">Noriega</collectorName>
leg
</materialsCitation>
. (1 CEMT).
<materialsCitation id="3B737B52B108FFF4F15FFF01FE532A28" ID-GBIF-Occurrence="3892767195" country="Colombia" elevation="300" latitude="4.0925" location="Meuse" longLatPrecision="21" longitude="-75.15445" pageId="35" pageNumber="804" specimenCount="1" stateProvince="Tolima">
<collectingRegion id="49DFBFEDB108FFF4F15FFF01FCBD2A0E" box="[816,894,249,274]" country="Colombia" name="Tolima" pageId="35" pageNumber="804">Tolima</collectingRegion>
,
<location id="8EC427D4B108FFF4F1E9FF01FC112A0E" LSID="urn:lsid:plazi:treatment:03B2C019B109FFF4F3CDFED5FD552A28:8EC427D4B108FFF4F1E9FF01FC112A0E" box="[902,978,249,274]" country="Colombia" latitude="4.0925" longLatPrecision="21" longitude="-75.15445" name="Meuse" pageId="35" pageNumber="804" stateProvince="Tolima">Meuse</location>
, BLT
<quantity id="4CE3DCEAB108FFF4F674FF01FB9E2A0E" box="[1051,1117,249,274]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="804" unit="m" value="300.0">
<elevation id="0036963CB108FFF4F674FF01FB9E2A0E" box="[1051,1117,249,274]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="804" unit="m" value="300.0">300m</elevation>
</quantity>
, [
<geoCoordinate id="EE2F17C8B108FFF4F604FF01FF2F2A28" degrees="4" direction="north" minutes="5" orientation="latitude" pageId="35" pageNumber="804" precision="15" seconds="33" value="4.0925">
4° 5
<emphasis id="B96FAD1DB108FFF4F2C1FEE3FF772A2F" bold="true" box="[174,180,283,307]" pageId="35" pageNumber="804">ʹ</emphasis>
33”N
</geoCoordinate>
,
<geoCoordinate id="EE2F17C8B108FFF4F298FEE3FEAD2A28" box="[247,366,283,308]" degrees="75" direction="west" minutes="9" orientation="longitude" pageId="35" pageNumber="804" precision="15" seconds="16" value="-75.15445">
75°9
<emphasis id="B96FAD1DB108FFF4F347FEE3FEED2A2F" bold="true" box="[296,302,283,307]" pageId="35" pageNumber="804">ʹ</emphasis>
16”W
</geoCoordinate>
],
<date id="FFA557CFB108FFF4F311FEE3FE532A28" box="[382,400,283,308]" pageId="35" pageNumber="804" value="1995-11">xi</date>
</materialsCitation>
.1995, EE leg. (1 CEMT).
</paragraph>
</subSubSection>
</treatment>
</document>