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<document id="C7DE0C4B672EEBF76A708F71C35C9DBC" ID-CLB-Dataset="132960" ID-DOI="10.11646/phytotaxa.559.2.7" ID-GBIF-Dataset="8d205305-4c48-47dd-9d00-3d1b0b660aa1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7021601" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661410718254" checkinUser="plazi" docAuthor="Geethakumary, M. P., Deepu, S., Kissling, J. &amp; Pandurangan, A. G." docDate="2022" docId="0397E93AFF8AF16607EBFF32DA0FF0DA" docLanguage="en" docName="phytotaxa.559.2.7.pdf" docOrigin="Phytotaxa 559 (2)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Exacum laxiflorum Geethakumary, Deepu, Kissling &amp; Pandurangan A. Habit 2022, comb. et stat. nov." docType="treatment" docVersion="8" lastPageNumber="190" masterDocId="FFAE9142FF8FF1600763FFADD959F006" masterDocTitle="Revisiting the taxonomy of Exacum courtallense (Gentianaceae) and recognizing E. courtallense var. laxiflorum at the species rank" masterLastPageNumber="192" masterPageNumber="185" pageNumber="190" updateTime="1699373709092" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="67624593F5AF5B66AD15111636112F40" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="BE2C3EA7E632E2CE5EBC8E7FACAADBF9">Revisiting the taxonomy of Exacum courtallense (Gentianaceae) and recognizing E. courtallense var. laxiflorum at the species rank</mods:title>
</mods:titleInfo>
<mods:name id="5300FD66920965F3E8BA1163D42C66AB" type="personal">
<mods:role id="3DD36C909EDC4F27543173AC25304FF7">
<mods:roleTerm id="BC225DF34904D9035AA0871989C47243">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="97759D2ADA1F8A90DEEF9AF425812507">Geethakumary, M. P.</mods:namePart>
<mods:nameIdentifier id="414275562B6FA010A154541B1B6CEA51" type="ORCID">0000-0002-2136-5987</mods:nameIdentifier>
<mods:affiliation id="43A70EB24103CC5336C137B9907EE1A6">Plant Systematic and Evolutionary Science Division, Jawaharlal Nehru Tropical Botanic Garden and Research Institute, Karimancode P. O., Thiruvananthapuram, Kerala 695562, India &amp; geethatbgt @ gmail. com; https: // orcid. org / 0000 - 0002 - 2136 - 5987</mods:affiliation>
<mods:nameIdentifier id="76F220EC027C0BFE7200191D2AD3A63D" type="email">geethatbgt@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="AA5078491E9547754EE6706A99AF35E7" type="personal">
<mods:role id="3409B4FDF69EAB455A38DD1B46C31D31">
<mods:roleTerm id="2C227D4E079AF2498320BC982D6F6C7E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA551456A6C057E03D5FCA53F8EFE92B">Deepu, S.</mods:namePart>
<mods:affiliation id="BEB5E8890F7D8BF2DD9C3422F69B850A">Plant Systematic and Evolutionary Science Division, Jawaharlal Nehru Tropical Botanic Garden and Research Institute, Karimancode P. O., Thiruvananthapuram, Kerala 695562, India &amp; Forest Ecology Department, Kerala Forest Research Institute, Thrissur, Kerala 680653, India</mods:affiliation>
</mods:name>
<mods:name id="C2673647B13BCDA123646ADD1DC365B7" type="personal">
<mods:role id="0C4D4C2E638E44D3F6CF3C6F6ABEAC91">
<mods:roleTerm id="8B8B57EAA790CE0479E671D40F34BF69">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="483FE3B1657CAEFFB02440C585CA0929">Kissling, J.</mods:namePart>
<mods:nameIdentifier id="D0DD55C1A5E7167328BF5E0B6B9E0900" type="ORCID">0000-0002-1078-4000</mods:nameIdentifier>
<mods:affiliation id="28F9D46F9E15DB8F4A9A62AA1E530DDF">Institute of biology, University of Neuchâtel, Rue Emile-Argand 11, 2000 Neuchâtel, Switzerland &amp; jonathan. kissling @ unine. ch; https: // orcid. org / 0000 - 0002 - 1078 - 4000</mods:affiliation>
<mods:nameIdentifier id="A484EAAE0751ADFC1AF3836681972B48" type="email">jonathan.kissling@unine.ch</mods:nameIdentifier>
</mods:name>
<mods:name id="B8B32A28214AC9FB9A474F60FB4BCBCB" type="personal">
<mods:role id="F219D134EFCE4F7A1CFB47B7A9D7C4A4">
<mods:roleTerm id="EDBD2C563C7A8E5957B5A7741442F55B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="69210415A51CF53A3181A03718154D5A">Pandurangan, A. G.</mods:namePart>
<mods:affiliation id="3B7CE94B87FDD39AD02DA55CA52D88D3">Plant Systematic and Evolutionary Science Division, Jawaharlal Nehru Tropical Botanic Garden and Research Institute, Karimancode P. O., Thiruvananthapuram, Kerala 695562, India &amp; Centre for Innovation in Science and Social Action, Thiruvananthapuram, Kerala 695010, India</mods:affiliation>
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<mods:title id="5A3592C8B06277CF10FCC77A3BEDDB0C">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="A22083F26F308F33EF178D3A8C2CAB86">
<mods:date id="C8E65822F47C5E419291F4FF46D018C0">2022</mods:date>
<mods:detail id="D5066C36C6320DB9F2628AA2BFB58A5C" type="pubDate">
<mods:number id="0A2268FB214C2CB2CA7731C9D3EC75AB">2022-08-23</mods:number>
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<mods:detail id="A20D3666FBE55DF7F43F034E19248A76" type="volume">
<mods:number id="657627123592520C8103EB203ECEC2B8">559</mods:number>
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<mods:classification id="3D1FF55E5501BC2A6C6E16B2B4407F62">journal article</mods:classification>
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<mods:identifier id="2494CB266A352BA40D13D440E6A85534" type="DOI">10.11646/phytotaxa.559.2.7</mods:identifier>
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<treatment id="0397E93AFF8AF16607EBFF32DA0FF0DA" ID-DOI="http://doi.org/10.5281/zenodo.7021609" ID-GBIF-Taxon="199891832" ID-Zenodo-Dep="7021609" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA" httpUri="http://treatment.plazi.org/id/0397E93AFF8AF16607EBFF32DA0FF0DA" lastPageId="6" lastPageNumber="190" pageId="5" pageNumber="190">
<subSubSection id="C3240BA7FF8AF16507EBFF32DD80F0BF" box="[136,1241,158,185]" pageId="5" pageNumber="190" type="nomenclature">
<paragraph id="8B81582CFF8AF16507EBFF32DD80F0BF" blockId="5.[136,1241,158,185]" box="[136,1241,158,185]" pageId="5" pageNumber="190">
<heading id="D0C9EF40FF8AF16507EBFF32DD80F0BF" box="[136,1241,158,185]" fontSize="11" level="2" pageId="5" pageNumber="190" reason="3">
<taxonomicName id="4C3E23AFFF8AF16507EBFF32DD5BF0BE" ID-CoL="9VGDR" authority="(Gamble) Geethakumary, Deepu, Kissling &amp; Pandurangan" authorityName="Geethakumary, Deepu, Kissling &amp; Pandurangan A. Habit" authorityYear="2022" baseAuthorityName="Geethakumary, Deepu, Kissling &amp; Pandurangan A. Habit" baseAuthorityYear="2022" box="[136,1026,158,185]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="190" phylum="Tracheophyta" rank="species" species="laxiflorum" status="comb. et stat. nov.">
<emphasis id="B94A843EFF8AF16507EBFF32D832F0BF" bold="true" box="[136,363,158,185]" italics="true" pageId="5" pageNumber="190">Exacum laxiflorum</emphasis>
(Gamble) Geethakumary, Deepu, Kissling &amp; Pandurangan
</taxonomicName>
,
<taxonomicNameLabel id="A2793945FF8AF165036EFF33DD80F0BF" box="[1037,1241,158,185]" pageId="5" pageNumber="190">
<emphasis id="B94A843EFF8AF165036EFF33DD88F0BE" box="[1037,1233,158,184]" italics="true" pageId="5" pageNumber="190">comb. et stat. nov</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3240BA7FF8AF16507EBFF6ADBECF0DD" box="[136,693,198,220]" pageId="5" pageNumber="190" type="reference_group">
<paragraph id="8B81582CFF8AF16507EBFF6ADBECF0DD" blockId="5.[136,1452,198,364]" box="[136,693,198,220]" pageId="5" pageNumber="190">
<treatmentCitationGroup id="AB2E7F02FF8AF16507EBFF6ADBECF0DD" box="[136,693,198,220]" pageId="5" pageNumber="190">
<treatmentCitation id="0A9F7E3DFF8AF16507EBFF6ADBE9F0DD" author="Gamble, J. S." box="[136,688,198,220]" page=")" pageId="5" pageNumber="190" year="1923">
<taxonomicName id="4C3E23AFFF8AF16507EBFF6ADBE9F0DD" ID-CoL="9T97V" authority="Gamble (1923: 873)" authorityName="Gamble" authorityPageNumber="873" authorityYear="1923" box="[136,688,198,220]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="190" phylum="Tracheophyta" rank="variety" species="courtallense" variety="laxiflorum">
<emphasis id="B94A843EFF8AF16507EBFF6AD80BF0DD" box="[136,338,198,220]" italics="true" pageId="5" pageNumber="190">Exacum courtallense</emphasis>
var.
<emphasis id="B94A843EFF8AF16506E2FF6BD8BCF0DD" box="[385,485,198,219]" italics="true" pageId="5" pageNumber="190">laxiflorum</emphasis>
<bibRefCitation id="EFAF25DDFF8AF1650688FF6BDBE9F0DD" author="Gamble, J. S." box="[491,688,198,220]" pageId="5" pageNumber="190" refId="ref4321" refString="Gamble, J. S. (1923) The Flora of Presidency of Madras, Vol. 2. Adlard and Son Ltd., London, 875 pp." type="book" year="1923">Gamble (1923: 873)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3240BA7FF8AF16507EBFF47DB2FF122" pageId="5" pageNumber="190" type="materials_examined">
<paragraph id="8B81582CFF8AF16507EBFF47DB2FF122" blockId="5.[136,1452,198,364]" pageId="5" pageNumber="190">
<materialsCitation id="3B565271FF8AF16507EBFF47D9A9F122" ID-GBIF-Occurrence="3897114332" collectionCode="MH, K" country="India" county="Beddome" location="Type" municipality="Travancore" pageId="5" pageNumber="190" specimenCode="MH00002366, K000759944" specimenCount="1" typeStatus="lectotype">
<location id="8EE10EF7FF8AF16507EBFF47D9E3F0F9" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF16507EBFF47D9E3F0F9" box="[136,186,234,255]" country="India" county="Beddome" municipality="Travancore" name="Type" pageId="5" pageNumber="190">Type</location>
:—
<collectingCountry id="F32918BCFF8AF16507B4FF47D847F0F9" box="[215,286,234,255]" name="India" pageId="5" pageNumber="190">INDIA</collectingCountry>
.
<collectingMunicipality id="6BE5C256FF8AF165064BFF47D8CEF0F9" box="[296,407,234,255]" pageId="5" pageNumber="190">Travancore</collectingMunicipality>
, 1873,
<emphasis id="B94A843EFF8AF16506BCFF47DB3BF106" box="[479,610,234,256]" italics="true" pageId="5" pageNumber="190">
<collectingCounty id="62E020A0FF8AF16506BCFF47DB61F0F9" box="[479,568,234,255]" pageId="5" pageNumber="190">Beddome</collectingCounty>
s.n.
</emphasis>
(
<typeStatus id="5485E68EFF8AF165050CFF47DB90F106" box="[623,713,234,256]" pageId="5" pageNumber="190" type="lectotype">lectotype</typeStatus>
<collectionCode id="ED2FC0E9FF8AF16505B2FF47DBA1F0F9" box="[721,760,234,255]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="5" pageNumber="190">MH</collectionCode>
[barcode
<specimenCode id="DB98F057FF8AF1650439FF47DABFF106" box="[858,998,234,256]" collectionCode="MH" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="5" pageNumber="190">MH00002366</specimenCode>
], designated by
<bibRefCitation id="EFAF25DDFF8AF16503E6FF47D9B5F122" author="Henry, A. N. &amp; Swaminathan, M. S." pageId="5" pageNumber="190" pagination="456 - 459" refId="ref4585" refString="Henry, A. N. &amp; Swaminathan, M. S. (1983) Rare or new Exacum L. (Gentianaceae) from Southern India. Journal of Bombay Natural History Society 80: 456 - 459." type="journal article" year="1983">Henry &amp; Swaminathan (1983: 458)</bibRefCitation>
;
</materialsCitation>
<materialsCitation id="3B565271FF8AF1650795FEA3DB2BF122" ID-GBIF-Occurrence="3897114340" box="[246,626,270,292]" collectionCode="MH, K" country="India" county="Beddome" location="Type" municipality="Travancore" pageId="5" pageNumber="190" specimenCode="MH00002366, K000759944" specimenCount="1" typeStatus="isolectotype">
isolectotype
<collectionCode id="ED2FC0E9FF8AF1650613FEA3D8D8F125" box="[368,385,270,291]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="190" type="Herbarium">K</collectionCode>
[barcode
<specimenCode id="DB98F057FF8AF1650682FEA3DB3FF122" box="[481,614,270,292]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="190" type="Herbarium">K000759944</specimenCode>
])
</materialsCitation>
.
</paragraph>
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<subSubSection id="C3240BA7FF8AF16507EBFE9EDC5EF141" box="[136,1287,306,328]" pageId="5" pageNumber="190" type="reference_group">
<paragraph id="8B81582CFF8AF16507EBFE9EDC5EF141" blockId="5.[136,1452,198,364]" box="[136,1287,306,328]" pageId="5" pageNumber="190">
<taxonomicName id="4C3E23AFFF8AF16507EBFE9EDBCAF14E" ID-CoL="7LRST" authority="M. Mohanan" authorityName="M. Mohanan" authorityYear="1994" box="[136,659,306,328]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="190" phylum="Tracheophyta" rank="variety" species="courtallense" variety="boneccordense">
<emphasis id="B94A843EFF8AF16507EBFE9ED80BF141" box="[136,338,306,328]" italics="true" pageId="5" pageNumber="190">Exacum courtallense</emphasis>
var.
<emphasis id="B94A843EFF8AF16506E2FE9FDB49F141" box="[385,528,306,327]" italics="true" pageId="5" pageNumber="190">boneccordense</emphasis>
M. Mohanan
</taxonomicName>
<emphasis id="B94A843EFF8AF16505FBFE9FDBF2F141" box="[664,683,306,327]" italics="true" pageId="5" pageNumber="190">in</emphasis>
M. Mohanan &amp; A.N. Henry (1994: 305, as “
<emphasis id="B94A843EFF8AF1650339FE9FDDB7F141" box="[1114,1262,306,327]" italics="true" pageId="5" pageNumber="190">boneccordensis</emphasis>
”).
</paragraph>
</subSubSection>
<subSubSection id="C3240BA7FF8AF16507EBFEFBDC3BF16D" box="[136,1378,342,364]" pageId="5" pageNumber="190" type="materials_examined">
<paragraph id="8B81582CFF8AF16507EBFEFBDC3BF16D" blockId="5.[136,1452,198,364]" box="[136,1378,342,364]" pageId="5" pageNumber="190">
<materialsCitation id="3B565271FF8AF16507EBFEFBDC07F16D" ID-GBIF-Occurrence="3897114328" box="[136,1374,342,364]" collectionCode="CAL, MH" collectorName="M. Mohanan" country="India" location="Thiruvananthapuram district" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
Type:—
<collectingCountry id="F32918BCFF8AF16507B4FEFBD847F16D" box="[215,286,342,363]" name="India" pageId="5" pageNumber="190">INDIA</collectingCountry>
.
<collectingRegion id="49FA96CEFF8AF1650645FEFBD831F16A" box="[294,360,342,364]" country="India" name="Kerala" pageId="5" pageNumber="190">Kerala</collectingRegion>
,
<location id="8EE10EF7FF8AF1650611FEFBDBDFF16A" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF1650611FEFBDBDFF16A" box="[370,646,342,364]" country="India" name="Thiruvananthapuram district" pageId="5" pageNumber="190" stateProvince="Kerala">Thiruvananthapuram district</location>
, Bonacaud,
<emphasis id="B94A843EFF8AF165059FFEFADA7DF16D" box="[764,804,342,364]" italics="true" pageId="5" pageNumber="190">s. d.</emphasis>
,
<emphasis id="B94A843EFF8AF165044CFEFADAB4F16D" box="[815,1005,342,364]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF165044CFEFADAF2F16D" box="[815,939,342,364]" pageId="5" pageNumber="190">M. Mohanan</collectorName>
63225
</emphasis>
(
<typeStatus id="5485E68EFF8AF165049AFEFBDD16F16A" box="[1017,1103,342,364]" pageId="5" pageNumber="190" type="holotype">holotype</typeStatus>
<collectionCode id="ED2FC0E9FF8AF1650336FEFBDDDDF16D" box="[1109,1156,342,363]" country="India" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="5" pageNumber="190" type="Herbarium">CAL</collectionCode>
n.v.; isotype
<collectionCode id="ED2FC0E9FF8AF1650266FEFBDC75F16D" box="[1285,1324,342,363]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="5" pageNumber="190">MH</collectionCode>
n.v.)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3240BA7FF8AF16507EBFE37DD48F38E" pageId="5" pageNumber="190" type="description">
<paragraph id="8B81582CFF8AF16507EBFE37DD48F38E" blockId="5.[136,1452,410,2093]" pageId="5" pageNumber="190">
<emphasis id="B94A843EFF8AF16507EBFE37D995F1B2" box="[136,204,410,436]" italics="true" pageId="5" pageNumber="190">Herbs</emphasis>
,
<quantity id="4CC6F5C9FF8AF16507B9FE37D80EF1B3" box="[218,343,410,437]" metricMagnitude="-1" metricUnit="m" metricValue="8.35" metricValueMax="15.2" metricValueMin="1.5" pageId="5" pageNumber="190" unit="cm" value="83.5" valueMax="152.0" valueMin="15.0">15–152 cm</quantity>
tall. Stems much branched, terete at the base, sharply 4-angular above, young branches 4-lineolate, internodes
<quantity id="4CC6F5C9FF8AF1650665FE13D8DDF1DF" box="[262,388,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="9.5" metricValueMin="1.0" pageId="5" pageNumber="190" unit="mm" value="52.5" valueMax="95.0" valueMin="10.0">10–95 mm</quantity>
.
<emphasis id="B94A843EFF8AF16506F3FE12D887F1DE" box="[400,478,447,472]" italics="true" pageId="5" pageNumber="190">Leaves</emphasis>
opposite, subsessile, narrowly elliptic to lanceolate, 15–88 ×
<quantity id="4CC6F5C9FF8AF16503C0FE13DC4AF1DF" box="[1187,1299,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="5" pageNumber="190" unit="mm" value="12.5" valueMax="20.0" valueMin="5.0">5–20 mm</quantity>
, chartaceous, glabrous, green, base attenuate, petiolate, apex long acuminate, 3–veined (midvein with one secondary vein on each side), tips recurved, veins diverging from the base of the lamina reaching to the apex prominent. Inflorescence terminal or axillary dichasial lax cymes, rarely solitary from terminal axis; peduncles
<quantity id="4CC6F5C9FF8AF1650483FD87DD00F243" box="[992,1113,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.5" metricValueMin="1.5" pageId="5" pageNumber="190" unit="mm" value="35.0" valueMax="55.0" valueMin="15.0">15–55 mm</quantity>
long; pedicels
<quantity id="4CC6F5C9FF8AF1650261FD87DC35F243" box="[1282,1388,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="2.5" metricValueMin="0.6" pageId="5" pageNumber="190" unit="mm" value="15.5" valueMax="25.0" valueMin="6.0">6–25 mm</quantity>
long, enlarging in fruit (
<quantity id="4CC6F5C9FF8AF165063DFDE3D886F26E" box="[350,479,590,616]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="5" pageNumber="190" unit="mm" value="20.0" valueMax="30.0" valueMin="10.0">10–30 mm</quantity>
), bent towards one side.
<emphasis id="B94A843EFF8AF1650460FDE3DA1DF26E" box="[771,836,590,616]" italics="true" pageId="5" pageNumber="190">Calyx</emphasis>
5-lobed; tube campanulate,
<quantity id="4CC6F5C9FF8AF16503ECFDE3DDB6F26F" box="[1167,1263,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="190" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long; lobes
<quantity id="4CC6F5C9FF8AF16502E2FDE3D9EDF28B" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="5" pageNumber="190" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
long acuminate at apex, winged; wings
<quantity id="4CC6F5C9FF8AF165051BFDDFDB8CF28B" box="[632,725,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="190" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
broad, wings narrowing at the apex, abruptly narrowing to base, accrescent in fruit, with prominent veins.
<emphasis id="B94A843EFF8AF1650504FD3BDBE4F2B6" box="[615,701,662,688]" italics="true" pageId="5" pageNumber="190">Corolla</emphasis>
tube
<quantity id="4CC6F5C9FF8AF1650463FD3BDA03F2B7" box="[768,858,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="190" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
long, green; lobes 5, broadly obovate-oblong, 8–11 ×
<quantity id="4CC6F5C9FF8AF16507C3FD17D857F2D3" box="[160,270,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="5" pageNumber="190" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
, subacute at apex, long persistent in fruit with prominent nerves but eventually deciduous, blue towards purplish.
<emphasis id="B94A843EFF8AF1650791FD72D816F2FF" box="[242,335,735,761]" italics="true" pageId="5" pageNumber="190">Stamens</emphasis>
5,
<quantity id="4CC6F5C9FF8AF1650610FD73D8A3F2FF" box="[371,506,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.5" metricValueMin="6.5" pageId="5" pageNumber="190" unit="mm" value="8.0" valueMax="9.5" valueMin="6.5">6.5–9.5 mm</quantity>
long; filaments
<emphasis id="B94A843EFF8AF16505D2FD72DB92F2FE" box="[689,715,735,760]" italics="true" pageId="5" pageNumber="190">ca</emphasis>
.
<quantity id="4CC6F5C9FF8AF16505B9FD73DA42F2FF" box="[730,795,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="190" unit="mm" value="4.0">4 mm</quantity>
long, attached at distal portion of the corolla tube; anthers bottle-shaped with long narrow neck,
<quantity id="4CC6F5C9FF8AF1650548FCAFDBC5F31B" box="[555,668,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="5" pageNumber="190" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">5–5.5 mm</quantity>
long, slightly curved with the convex back face, base cordate, opening by apical pores that later widen to slits to the base, without papilla.
<emphasis id="B94A843EFF8AF1650408FC8ADAE8F347" box="[875,945,807,833]" italics="true" pageId="5" pageNumber="190">Ovary</emphasis>
ellipsoid, ca.
<quantity id="4CC6F5C9FF8AF1650329FC8BDDDAF346" box="[1098,1155,806,832]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="190" unit="mm" value="4.0">4mm</quantity>
long; style
<quantity id="4CC6F5C9FF8AF1650260FC8BDC34F347" box="[1283,1389,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="5" pageNumber="190" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
long, curved, persistent; stigma capitate.
<emphasis id="B94A843EFF8AF1650579FCE7DB2FF362" box="[538,630,842,868]" italics="true" pageId="5" pageNumber="190">Capsule</emphasis>
ovoid, 6–9 ×
<quantity id="4CC6F5C9FF8AF165047BFCE7DA20F363" box="[792,889,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="190" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, dehiscing septicidally, 2-valved; style persistent.
<emphasis id="B94A843EFF8AF16507EBFCC3D991F38E" box="[136,200,878,904]" italics="true" pageId="5" pageNumber="190">Seeds</emphasis>
many, irregularly rhomboid, angular, with shallowly sunken sides (
<figureCitation id="130544A9FF8AF16504DDFCC3DD5DF38E" box="[958,1028,878,904]" captionStart="FIGURE 2" captionStartId="4.[136,229,2028,2050]" captionTargetBox="[196,1389,206,1985]" captionTargetId="figure-29@4.[186,1401,187,2001]" captionTargetPageId="4" captionText="FIGURE 2. Exacum laxiflorum (Gamble) Geethakumary, Deepu, Kissling &amp; Pandurangan A. Habit; B. Flower; C. Calyx; D. Petal; E. Stamen; F. Pistil; G. Mature fruit split open; H. Seeds." figureDoi="http://doi.org/10.5281/zenodo.7021611" httpUri="https://zenodo.org/record/7021611/files/figure.png" pageId="5" pageNumber="190">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3240BA7FF8AF16507DEFC3FDA69F3F2" pageId="5" pageNumber="190" type="discussion">
<paragraph id="8B81582CFF8AF16507DEFC3FDA69F3F2" blockId="5.[136,1452,410,2093]" pageId="5" pageNumber="190">
<emphasis id="B94A843EFF8AF16507DEFC3FD9ACF3AA" bold="true" box="[189,245,914,940]" pageId="5" pageNumber="190">Note</emphasis>
:—We searched for the
<typeStatus id="5485E68EFF8AF1650567FC3EDB6DF3AB" box="[516,564,915,941]" pageId="5" pageNumber="190">type</typeStatus>
specimens of
<taxonomicName id="4C3E23AFFF8AF16505BAFC3EDDFEF3AA" authorityName="M. Mohanan" authorityYear="1994" box="[729,1191,914,941]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="190" phylum="Tracheophyta" rank="variety" species="courtallense" variety="boneccordense">
<emphasis id="B94A843EFF8AF16505BAFC3EDA9CF3AA" box="[729,965,914,940]" italics="true" pageId="5" pageNumber="190">Exacum courtallense</emphasis>
var.
<emphasis id="B94A843EFF8AF1650363FC3FDDFEF3AA" box="[1024,1191,914,940]" italics="true" pageId="5" pageNumber="190">boneccordense</emphasis>
</taxonomicName>
in both CAL and MH, and we were unable to locate them. However, for this study, specimens collected from the
<typeStatus id="5485E68EFF8AF16503E5FC1ADDEFF3D7" box="[1158,1206,951,977]" pageId="5" pageNumber="190">type</typeStatus>
locality matching the drawing in the protologue were used for logical conclusions.
</paragraph>
</subSubSection>
<subSubSection id="C3240BA7FF8AF16507DEFC53DA0EF43A" pageId="5" pageNumber="190" type="biology_ecology">
<paragraph id="8B81582CFF8AF16507DEFC53DD4CF41E" blockId="5.[136,1452,410,2093]" box="[189,1045,1022,1049]" pageId="5" pageNumber="190">
<emphasis id="B94A843EFF8AF16507DEFC53D860F41E" bold="true" box="[189,313,1022,1048]" pageId="5" pageNumber="190">Phenology</emphasis>
:—Flowering and fruiting were observed from July to December.
</paragraph>
<paragraph id="8B81582CFF8AF16507DEFB8FDA0EF43A" blockId="5.[136,1452,410,2093]" box="[189,855,1058,1084]" pageId="5" pageNumber="190">
<emphasis id="B94A843EFF8AF16507DEFB8FD89AF43A" bold="true" box="[189,451,1058,1084]" pageId="5" pageNumber="190">Chromosome number</emphasis>
:—2n=68 (
<bibRefCitation id="EFAF25DDFF8AF165055EFB8FDA17F43A" author="Mallikarjuna, M. B. &amp; Scheriff, A. &amp; Krishnappa, D. G." box="[573,846,1058,1084]" pageId="5" pageNumber="190" pagination="766 - 767" refId="ref4844" refString="Mallikarjuna, M. B., Scheriff, A. &amp; Krishnappa, D. G. (1987) Chromosome number reports XCVII (ed. A. Love). Taxon 36: 766 - 767. https: // doi. org / 10.1002 / j. 1996 - 8175.1987. tb 00894. x" type="journal article" year="1987">
Mallikarjuna
<emphasis id="B94A843EFF8AF16505B5FB8EDA56F43A" box="[726,783,1058,1084]" italics="true" pageId="5" pageNumber="190">et al.</emphasis>
1987
</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3240BA7FF8AF16507DEFBEBD80EF4AE" pageId="5" pageNumber="190" type="distribution">
<paragraph id="8B81582CFF8AF16507DEFBEBD80EF4AE" blockId="5.[136,1452,410,2093]" pageId="5" pageNumber="190">
<emphasis id="B94A843EFF8AF16507DEFBEBD8B1F466" bold="true" box="[189,488,1094,1120]" pageId="5" pageNumber="190">Distribution and Habitat</emphasis>
:—Occurs in the evergreen and semi-evergreen forests of the Southern Western Ghats in
<collectingRegion id="49FA96CEFF8AF16507CBFBC7D9AAF482" box="[168,243,1130,1156]" country="India" name="Kerala" pageId="5" pageNumber="190">Kerala</collectingRegion>
and
<collectingRegion id="49FA96CEFF8AF1650653FBC7D8E2F482" box="[304,443,1130,1156]" country="India" name="Tamil Nadu" pageId="5" pageNumber="190">Tamil Nadu</collectingRegion>
. This species has a wider distribution than
<taxonomicName id="4C3E23AFFF8AF16504DBFBC6DD3CF482" authorityName="Arnott. A. Habit" authorityYear="1839" box="[952,1125,1130,1156]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="190" phylum="Tracheophyta" rank="species" species="courtallense">
<emphasis id="B94A843EFF8AF16504DBFBC6DD3CF482" box="[952,1125,1130,1156]" italics="true" pageId="5" pageNumber="190">E. courtallense</emphasis>
</taxonomicName>
and is recorded from many districts in
<collectingRegion id="49FA96CEFF8AF1650666FB23D80AF4AE" box="[261,339,1166,1192]" country="India" name="Kerala" pageId="5" pageNumber="190">Kerala</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3240BA7FF8AF16507DEFB1FDBEFF5EA" pageId="5" pageNumber="190" type="conservation">
<paragraph id="8B81582CFF8AF16507DEFB1FDBEFF5EA" blockId="5.[136,1452,410,2093]" pageId="5" pageNumber="190">
<emphasis id="B94A843EFF8AF16507DEFB1FD8EFF4CA" bold="true" box="[189,438,1202,1228]" pageId="5" pageNumber="190">Conservation status:</emphasis>
—Based on the information from field surveys and herbarium data, the species is known from 13 subpopulations. Some of these subpopulations are under stress due to a decline in the quality of habitat. The species is annual, and it is observed to have undergone a reduction of over 50% over a period of 10 years due to a decline in area of occupancy (AOO) and quality of habitat, as evident by the fact that some subpopulations seen in the past are currently not present. Some subpopulations occur along the trek paths and wall cuttings along the roads, which are subjected to annual clearing, posing a threat to these subpopulations. Based on the available population information, the AOO of the species is not expected to be greater than
<quantity id="4CC6F5C9FF8AF16504D7FA27DD53F5A2" box="[948,1034,1418,1445]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="190" unit="km" value="500.0">500 km</quantity>
<superScript id="7C4BF564FF8AF1650369FA27DD4BF59E" attach="left" box="[1034,1042,1418,1432]" fontSize="6" pageId="5" pageNumber="190">2</superScript>
. Given the species has a low AOO, suffered population decline over the past, and the continuing threat due to the decline in habitat quality, the species is assessed as Endangered [EN A2ac; B2b(ii, iv, v)].
</paragraph>
</subSubSection>
<subSubSection id="C3240BA7FF8AF16607DEFA5BDA0FF0DA" lastPageId="6" lastPageNumber="191" pageId="5" pageNumber="190" type="materials_examined">
<paragraph id="8B81582CFF8AF16607DEFA5BDA0FF0DA" blockId="5.[136,1452,410,2093]" lastBlockId="6.[136,1451,158,221]" lastPageId="6" lastPageNumber="191" pageId="5" pageNumber="190">
<emphasis id="B94A843EFF8AF16507DEFA5BD8E8F616" bold="true" box="[189,433,1526,1552]" pageId="5" pageNumber="190">Specimens examined</emphasis>
:—
<materialsCitation id="3B565271FF8AF16506B0FA5BD86FF632" ID-GBIF-Occurrence="3897114317" collectingDate="2012-09-22" collectionCode="TBGT" collectorName="Geethakumary" country="India" location="Bhoothathankettu" municipality="Ernakulam District" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<collectingCountry id="F32918BCFF8AF16506B0FA5BDB71F616" box="[467,552,1526,1552]" name="India" pageId="5" pageNumber="190">INDIA</collectingCountry>
.
<collectingRegion id="49FA96CEFF8AF165054EFA5BDB23F616" box="[557,634,1526,1552]" country="India" name="Kerala" pageId="5" pageNumber="190">Kerala</collectingRegion>
,
<collectingMunicipality id="6BE5C256FF8AF16505E2FA5BDA0FF616" box="[641,854,1526,1552]" pageId="5" pageNumber="190">Ernakulam District</collectingMunicipality>
,
<location id="8EE10EF7FF8AF165043FFA5BDD7FF616" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF165043FFA5BDD7FF616" box="[860,1062,1526,1552]" country="India" municipality="Ernakulam District" name="Bhoothathankettu" pageId="5" pageNumber="190" stateProvince="Kerala">Bhoothathankettu</location>
,
<date id="FF807EECFF8AF165034EFA5BDC59F616" box="[1069,1280,1526,1552]" pageId="5" pageNumber="190" value="2012-09-22">
<collectingDate id="EFC48704FF8AF165034EFA5BDC59F616" box="[1069,1280,1526,1552]" pageId="5" pageNumber="190" value="2012-09-22">22September2012</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF1650264FA5BD997F632" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF1650264FA5BDCF5F616" box="[1287,1452,1526,1552]" pageId="5" pageNumber="190">Geethakumary</collectorName>
70936
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16507BDF9B7D874F632" box="[222,301,1562,1588]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B565271FF8AF1650621F9B7DD09F632" ID-GBIF-Occurrence="3897114309" box="[322,1104,1562,1589]" collectingDate="1984-07-07" collectionCode="MH" collectorName="C. N. Mohanan" country="India" location="Kulamavu" municipality="Idukki District" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<collectingMunicipality id="6BE5C256FF8AF1650621F9B7D8B3F632" box="[322,490,1562,1588]" pageId="5" pageNumber="190">Idukki District</collectingMunicipality>
,
<location id="8EE10EF7FF8AF1650694F9B7DB36F632" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF1650694F9B7DB36F632" box="[503,623,1562,1588]" country="India" municipality="Idukki District" name="Kulamavu" pageId="5" pageNumber="190" stateProvince="Kerala">Kulamavu</location>
,
<date id="FF807EECFF8AF1650518F9B6DA5AF632" box="[635,771,1562,1589]" pageId="5" pageNumber="190" value="1984-07-07">
<collectingDate id="EFC48704FF8AF1650518F9B6DA5AF632" box="[635,771,1562,1589]" pageId="5" pageNumber="190" value="1984-07-07">7 July 1984</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF1650473F9B6DD5CF632" box="[784,1029,1562,1589]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF1650473F9B6DAEFF632" box="[784,950,1562,1589]" pageId="5" pageNumber="190">C.N. Mohanan</collectorName>
82012
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650375F9B7DD10F632" box="[1046,1097,1562,1588]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="5" pageNumber="190">MH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF165033EF9B7DA16F65E" ID-GBIF-Occurrence="3897114316" collectingDate="1821-12-26" collectionCode="MH" collectorName="R. H. Beddome" country="India" location="Travancore" municipality="Thiruvananthapuram District" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<collectingMunicipality id="6BE5C256FF8AF165033EF9B7DCFEF632" box="[1117,1447,1562,1588]" pageId="5" pageNumber="190">Thiruvananthapuram District</collectingMunicipality>
,
<location id="8EE10EF7FF8AF16507EBF992D85EF65F" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF16507EBF992D85EF65F" box="[136,263,1599,1625]" country="India" municipality="Thiruvananthapuram District" name="Travancore" pageId="5" pageNumber="190" stateProvince="Kerala">Travancore</location>
hills,
<date id="FF807EECFF8AF165062FF993DB7BF65E" box="[332,546,1598,1624]" pageId="5" pageNumber="190" value="1821-12-26">
<collectingDate id="EFC48704FF8AF165062FF993DB7BF65E" box="[332,546,1598,1624]" pageId="5" pageNumber="190" value="1821-12-26">26 December 1821</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF165054DF992DA5CF65E" box="[558,773,1598,1624]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF165054DF992DB8EF65E" box="[558,727,1598,1624]" pageId="5" pageNumber="190">R. H. Beddome</collectorName>
s.n.
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650476F993DA11F65E" box="[789,840,1598,1624]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="5" pageNumber="190">MH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF1650439F992DC14F65E" ID-GBIF-Occurrence="3897114333" box="[858,1357,1598,1625]" collectionCode="MH" collectorName="R. H. Beddome" country="India" location="Travancore" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF1650439F992DA82F65F" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF1650439F992DA82F65F" box="[858,987,1599,1625]" country="India" name="Travancore" pageId="5" pageNumber="190" stateProvince="Kerala">Travancore</location>
, 1873,
<emphasis id="B94A843EFF8AF165034EF992DC5AF65E" box="[1069,1283,1598,1624]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF165034EF992DD8FF65E" box="[1069,1238,1598,1624]" pageId="5" pageNumber="190">R. H. Beddome</collectorName>
s.n.
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650270F993DC1FF65E" box="[1299,1350,1598,1624]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="5" pageNumber="190">MH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF165023AF992D8B0F67A" ID-GBIF-Occurrence="3897114338" collectionCode="K, UCT" country="India" location="Neyyar Dam" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF165023AF992D998F67B" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF165023AF992D998F67B" country="India" name="Neyyar Dam" pageId="5" pageNumber="190" stateProvince="Kerala">Neyyar Dam</location>
,
<emphasis id="B94A843EFF8AF16507ADF9CED8CBF67A" box="[206,402,1635,1660]" italics="true" pageId="5" pageNumber="190">
<collectionCode id="ED2FC0E9FF8AF16507ADF9CED9BDF67A" box="[206,228,1635,1660]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="190" type="Herbarium">K</collectionCode>
. Rajappan, s.n.
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16506C0F9CFD8B8F67A" box="[419,481,1634,1660]" pageId="5" pageNumber="190">UCT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF1650696F9CEDC47F67A" ID-GBIF-Occurrence="3897114324" box="[501,1310,1634,1661]" collectingDate="1973-10-01" collectionCode="MH" collectorName="J. Joseph" country="India" elevation="500" location="Bonaccord" municipality="Forest" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<collectingMunicipality id="6BE5C256FF8AF1650696F9CEDB62F67B" box="[501,571,1635,1661]" pageId="5" pageNumber="190">Forest</collectingMunicipality>
near
<location id="8EE10EF7FF8AF165051FF9CFDBA1F67A" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF165051FF9CFDBA1F67A" box="[636,760,1634,1660]" country="India" municipality="Forest" name="Bonaccord" pageId="5" pageNumber="190" stateProvince="Kerala">Bonaccord</location>
,
<quantity id="4CC6F5C9FF8AF1650466F9CFDA16F67B" box="[773,847,1634,1661]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="190" unit="m" value="500.0">
<elevation id="0013BF1FFF8AF1650466F9CFDA16F67B" box="[773,847,1634,1661]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="190" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="FF807EECFF8AF165043FF9CFDD56F67A" box="[860,1039,1634,1660]" pageId="5" pageNumber="190" value="1973-10-01">
<collectingDate id="EFC48704FF8AF165043FF9CFDD56F67A" box="[860,1039,1634,1660]" pageId="5" pageNumber="190" value="1973-10-01">1 October 1973</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF1650378F9CEDD8AF67A" box="[1051,1235,1634,1660]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF1650378F9CEDDDCF67A" box="[1051,1157,1634,1660]" pageId="5" pageNumber="190">J. Joseph</collectorName>
44484
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650387F9CFDC4EF67A" box="[1252,1303,1634,1660]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="5" pageNumber="190">MH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF1650248F9CFDB9EF6A6" ID-GBIF-Occurrence="3897114307" collectingDate="1978-03-22" collectionCode="MH" collectorName="M. Mohanan" country="India" elevation="600" location="Bonaccord" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF1650248F9CFDCFEF67A" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF1650248F9CFDCFEF67A" box="[1323,1447,1634,1660]" country="India" name="Bonaccord" pageId="5" pageNumber="190" stateProvince="Kerala">Bonaccord</location>
,
<quantity id="4CC6F5C9FF8AF16507EBF92BD98BF6A7" box="[136,210,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="190" unit="m" value="600.0">
<elevation id="0013BF1FFF8AF16507EBF92BD98BF6A7" box="[136,210,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="190" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="FF807EECFF8AF16507BCF92BD8C9F6A6" box="[223,400,1670,1696]" pageId="5" pageNumber="190" value="1978-03-22">
<collectingDate id="EFC48704FF8AF16507BCF92BD8C9F6A6" box="[223,400,1670,1696]" pageId="5" pageNumber="190" value="1978-03-22">22 March 1978</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF16506FFF92ADB25F6A6" box="[412,636,1670,1696]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF16506FFF92ADB74F6A6" box="[412,557,1670,1696]" pageId="5" pageNumber="190">M. Mohanan</collectorName>
54736
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16505EFF92BDBE6F6A6" box="[652,703,1670,1696]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="5" pageNumber="190">MH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF16505B7F92BDC3CF6A6" ID-GBIF-Occurrence="3897114331" box="[724,1381,1670,1697]" collectingDate="1978-08-03" collectionCode="MH" collectorName="M. Mohanan" country="India" elevation="575" location="Bonaccord" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<emphasis id="B94A843EFF8AF16505B7F92BDA59F6A6" box="[724,768,1670,1696]" italics="true" pageId="5" pageNumber="190">ibid</emphasis>
., +
<quantity id="4CC6F5C9FF8AF165044DF92BDA21F6A7" box="[814,888,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="5.75" pageId="5" pageNumber="190" unit="m" value="575.0">
<elevation id="0013BF1FFF8AF165044DF92BDA21F6A7" box="[814,888,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="5.75" pageId="5" pageNumber="190" unit="m" value="575.0">575 m</elevation>
</quantity>
,
<date id="FF807EECFF8AF16504E6F92BDD77F6A6" box="[901,1070,1670,1696]" pageId="5" pageNumber="190" value="1978-08-03">
<collectingDate id="EFC48704FF8AF16504E6F92BDD77F6A6" box="[901,1070,1670,1696]" pageId="5" pageNumber="190" value="1978-08-03">3 August 1978</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF1650359F92ADC43F6A6" box="[1082,1306,1670,1696]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF1650359F92ADD92F6A6" box="[1082,1227,1670,1696]" pageId="5" pageNumber="190">M. Mohanan</collectorName>
58523
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650249F92BDC04F6A6" box="[1322,1373,1670,1696]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="5" pageNumber="190">MH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF1650211F92BDBE5F6C2" ID-GBIF-Occurrence="3897114325" collectingDate="1991-05-18" collectionCode="TBGT" collectorName="N. Mohanan" country="India" elevation="700" location="Bonaccord" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<emphasis id="B94A843EFF8AF1650211F92BDCC7F6A6" box="[1394,1438,1670,1696]" italics="true" pageId="5" pageNumber="190">ibid</emphasis>
.,
<quantity id="4CC6F5C9FF8AF16507EBF907D988F6C3" box="[136,209,1706,1733]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="190" unit="m" value="700.0">
<elevation id="0013BF1FFF8AF16507EBF907D988F6C3" box="[136,209,1706,1733]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="190" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="FF807EECFF8AF16507BFF907D82BF6C2" box="[220,370,1706,1733]" pageId="5" pageNumber="190" value="1991-05-18">
<collectingDate id="EFC48704FF8AF16507BFF907D82BF6C2" box="[220,370,1706,1733]" pageId="5" pageNumber="190" value="1991-05-18">18 May 1991</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF165061DF906DB0FF6C2" box="[382,598,1706,1732]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF165061DF906DB50F6C2" box="[382,521,1706,1732]" pageId="5" pageNumber="190">N. Mohanan</collectorName>
10818
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650507F907DBEAF6C2" box="[612,691,1706,1732]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF16505A5F907D9BEF6EE" ID-GBIF-Occurrence="3897114336" collectingDate="1992-10-30" collectionCode="TBGT" collectorName="E S Santhoshkumar &amp; M. A. Jabbar" country="India" elevation="700" location="Bonaccord" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<emphasis id="B94A843EFF8AF16505A5F907DBABF6C2" box="[710,754,1706,1732]" italics="true" pageId="5" pageNumber="190">ibid</emphasis>
.,
<date id="FF807EECFF8AF1650464F907DA9CF6C2" box="[775,965,1706,1732]" pageId="5" pageNumber="190" value="1992-10-30">
<collectingDate id="EFC48704FF8AF1650464F907DA9CF6C2" box="[775,965,1706,1732]" pageId="5" pageNumber="190" value="1992-10-30">30 October 1992</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF16504ACF906DCF2F6C2" box="[975,1451,1706,1733]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF16504ACF906DDF0F6C2" box="[975,1193,1706,1733]" pageId="5" pageNumber="190">E S Santhoshkumar</collectorName>
&amp;
<collectorName id="26CB3DFAFF8AF16503AFF906DC07F6C2" box="[1228,1374,1706,1733]" pageId="5" pageNumber="190">M. A. Jabbar</collectorName>
14727
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16507F3F963D986F6EE" box="[144,223,1742,1768]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF1650795F963DA55F6EE" ID-GBIF-Occurrence="3897114303" box="[246,780,1742,1769]" collectingDate="1992-10-30" collectionCode="TBGT" collectorName="Geethakumary" country="India" elevation="600" location="Bonaccord" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<emphasis id="B94A843EFF8AF1650795F963D87BF6EE" box="[246,290,1742,1768]" italics="true" pageId="5" pageNumber="190">ibid</emphasis>
., ±
<quantity id="4CC6F5C9FF8AF1650630F963D8C6F6EF" box="[339,415,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="190" unit="m" value="600.0">
<elevation id="0013BF1FFF8AF1650630F963D8C6F6EF" box="[339,415,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="190" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<emphasis id="B94A843EFF8AF16506CDF963DBFBF6EE" box="[430,674,1742,1768]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF16506CDF963DB0AF6EE" box="[430,595,1742,1768]" pageId="5" pageNumber="190">Geethakumary</collectorName>
53681
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16505D7F963DA5AF6EE" box="[692,771,1742,1768]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF1650479F963DCFEF6EE" ID-GBIF-Occurrence="3897114334" box="[794,1447,1742,1769]" collectingDate="2014-12-11" collectionCode="TBGT" collectorName="Geethakumary" country="India" elevation="600" location="Bonaccord" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<emphasis id="B94A843EFF8AF1650479F963DA1FF6EE" box="[794,838,1742,1768]" italics="true" pageId="5" pageNumber="190">ibid</emphasis>
.,
<date id="FF807EECFF8AF165043DF963DD62F6EE" box="[862,1083,1742,1768]" pageId="5" pageNumber="190" value="2014-12-11">
<collectingDate id="EFC48704FF8AF165043DF963DD62F6EE" box="[862,1083,1742,1768]" pageId="5" pageNumber="190" value="2014-12-11">11 December 2014</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF165032AF963DC67F6EE" box="[1097,1342,1742,1768]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF165032AF963DDB7F6EE" box="[1097,1262,1742,1768]" pageId="5" pageNumber="190">Geethakumary</collectorName>
80885
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650233F963DCC6F6EE" box="[1360,1439,1742,1768]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF16507EBF95FDAC9F70A" ID-GBIF-Occurrence="3897114337" box="[136,912,1778,1805]" collectingDate="2014-12-11" collectionCode="TBGT" collectorName="Geethakumary" country="India" elevation="700" location="Bonaccord" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<emphasis id="B94A843EFF8AF16507EBF95FD9EDF70A" box="[136,180,1778,1804]" italics="true" pageId="5" pageNumber="190">ibid</emphasis>
., +
<quantity id="4CC6F5C9FF8AF1650789F95FD875F70A" box="[234,300,1778,1804]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="190" unit="m" value="700.0">
<elevation id="0013BF1FFF8AF1650789F95FD875F70A" box="[234,300,1778,1804]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="190" unit="m" value="700.0">700m</elevation>
</quantity>
,
<date id="FF807EECFF8AF165065EF95FDB47F70A" box="[317,542,1778,1804]" pageId="5" pageNumber="190" value="2014-12-11">
<collectingDate id="EFC48704FF8AF165065EF95FDB47F70A" box="[317,542,1778,1804]" pageId="5" pageNumber="190" value="2014-12-11">11 December 2014</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF165054DF95FDA7CF70A" box="[558,805,1778,1804]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF165054DF95FDB8AF70A" box="[558,723,1778,1804]" pageId="5" pageNumber="190">Geethakumary</collectorName>
80885
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF165045AF95FDAD1F70A" box="[825,904,1778,1804]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF16504C2F95FD9BEF736" ID-GBIF-Occurrence="3897114305" collectingDate="2011-08-12" collectionCode="TBGT" collectorName="Geethakumary" country="India" elevation="700" location="Bonaccord" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<emphasis id="B94A843EFF8AF16504C2F95FDA94F70A" box="[929,973,1778,1804]" italics="true" pageId="5" pageNumber="190">ibid</emphasis>
.,
<date id="FF807EECFF8AF1650484F95FDDFDF70A" box="[999,1188,1778,1804]" pageId="5" pageNumber="190" value="2011-08-12">
<collectingDate id="EFC48704FF8AF1650484F95FDDFDF70A" box="[999,1188,1778,1804]" pageId="5" pageNumber="190" value="2011-08-12">12 August 2011</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF16503D7F95FDCF2F70A" box="[1204,1451,1778,1804]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF16503D7F95FDC00F70A" box="[1204,1369,1778,1804]" pageId="5" pageNumber="190">Geethakumary</collectorName>
67929
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16507F3F8BBD986F736" box="[144,223,1814,1840]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF1650796F8BBDAD8F736" ID-GBIF-Occurrence="3897114314" box="[245,897,1814,1841]" collectingDate="2011-11-16" collectionCode="TBGT" collectorName="Geethakumary" country="India" elevation="700" location="Bonaccord" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<emphasis id="B94A843EFF8AF1650796F8BBD878F736" box="[245,289,1814,1840]" italics="true" pageId="5" pageNumber="190">ibid</emphasis>
.,
<date id="FF807EECFF8AF165065BF8BBDB4EF736" box="[312,535,1814,1840]" pageId="5" pageNumber="190" value="2011-11-16">
<collectingDate id="EFC48704FF8AF165065BF8BBDB4EF736" box="[312,535,1814,1840]" pageId="5" pageNumber="190" value="2011-11-16">16 November 2011</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF1650547F8BBDA40F736" box="[548,793,1814,1840]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF1650547F8BBDB90F736" box="[548,713,1814,1840]" pageId="5" pageNumber="190">Geethakumary</collectorName>
69883
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650449F8BBDA20F736" box="[810,889,1814,1840]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF16504EEF8BBD86CF752" ID-GBIF-Occurrence="3897114329" collectingDate="2014-12-11" collectionCode="TBGT" collectorName="Geethakumary" country="India" location="Athirumala" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF16504EEF8BBDD49F736" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF16504EEF8BBDD49F736" box="[909,1040,1814,1840]" country="India" name="Athirumala" pageId="5" pageNumber="190" stateProvince="Kerala">Athirumala</location>
,
<date id="FF807EECFF8AF165037EF8BBDDA0F736" box="[1053,1273,1814,1840]" pageId="5" pageNumber="190" value="2014-12-11">
<collectingDate id="EFC48704FF8AF165037EF8BBDDA0F736" box="[1053,1273,1814,1840]" pageId="5" pageNumber="190" value="2014-12-11">11 December 2014</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF1650264F8BBD997F752" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF1650264F8BBDCF5F736" box="[1287,1452,1814,1840]" pageId="5" pageNumber="190">Geethakumary</collectorName>
80887
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16507BEF897D875F752" box="[221,300,1850,1876]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF1650623F897DD68F752" ID-GBIF-Occurrence="3897114341" box="[320,1073,1850,1877]" collectingDate="1988-10-12" collectionCode="CALI, TBGT" collectorName="N. Mohanan" country="India" elevation="1100" location="Athirumala" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<emphasis id="B94A843EFF8AF1650623F897D835F752" box="[320,364,1850,1876]" italics="true" pageId="5" pageNumber="190">ibid</emphasis>
.,
<quantity id="4CC6F5C9FF8AF16506E2F897D881F753" box="[385,472,1850,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="190" unit="m" value="1100.0">
<elevation id="0013BF1FFF8AF16506E2F897D881F753" box="[385,472,1850,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="190" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date id="FF807EECFF8AF1650687F897DBFDF752" box="[484,676,1850,1876]" pageId="5" pageNumber="190" value="1988-10-12">
<collectingDate id="EFC48704FF8AF1650687F897DBFDF752" box="[484,676,1850,1876]" pageId="5" pageNumber="190" value="1988-10-12">12 October 1988</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF16505CCF896DA22F752" box="[687,891,1850,1876]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF16505CCF896DA62F752" box="[687,827,1850,1876]" pageId="5" pageNumber="190">N. Mohanan</collectorName>
4215
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16504E9F897DA89F752" box="[906,976,1850,1876]" country="India" lsid="urn:lsid:biocol.org:col:14488" name="University of Calicut" pageId="5" pageNumber="190" type="Herbarium">CALI</collectionCode>
,
<collectionCode id="ED2FC0E9FF8AF16504B8F897DD70F752" box="[987,1065,1850,1876]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF165035EF896DB4BF77E" ID-GBIF-Occurrence="3897114310" collectingDate="1990-11-06" collectionCode="TBGT" collectorName="N. Mohanan" country="India" elevation="500" location="Meenmutty" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF165035EF896DDE6F753" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF165035EF896DDE6F753" box="[1085,1215,1851,1877]" country="India" name="Meenmutty" pageId="5" pageNumber="190" stateProvince="Kerala">Meenmutty</location>
,
<quantity id="4CC6F5C9FF8AF16503A9F897DC4DF753" box="[1226,1300,1850,1877]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="190" unit="m" value="500.0">
<elevation id="0013BF1FFF8AF16503A9F897DC4DF753" box="[1226,1300,1850,1877]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="190" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="FF807EECFF8AF1650243F897D99AF77E" pageId="5" pageNumber="190" value="1990-11-06">
<collectingDate id="EFC48704FF8AF1650243F897D99AF77E" pageId="5" pageNumber="190" value="1990-11-06">6 November 1990</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF16507ACF8F2D8F2F77E" box="[207,427,1886,1912]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF16507ACF8F2D805F77E" box="[207,348,1886,1912]" pageId="5" pageNumber="190">N. Mohanan</collectorName>
10147
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16506D8F8F3DB53F77E" box="[443,522,1886,1912]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF165057CF8F3DC5BF77E" ID-GBIF-Occurrence="3897114339" box="[543,1282,1886,1913]" collectingDate="1991-02-05" collectionCode="TBGT" collectorName="N. Mohanan" country="India" elevation="900" location="Chemunji" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF165057CF8F3DBC9F77E" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF165057CF8F3DBC9F77E" box="[543,656,1886,1912]" country="India" name="Chemunji" pageId="5" pageNumber="190" stateProvince="Kerala">Chemunji</location>
,
<quantity id="4CC6F5C9FF8AF16505FEF8F3DBBEF77F" box="[669,743,1886,1913]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="190" unit="m" value="900.0">
<elevation id="0013BF1FFF8AF16505FEF8F3DBBEF77F" box="[669,743,1886,1913]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="190" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<date id="FF807EECFF8AF1650597F8F3DAEBF77E" box="[756,946,1886,1913]" pageId="5" pageNumber="190" value="1991-02-05">
<collectingDate id="EFC48704FF8AF1650597F8F3DAEBF77E" box="[756,946,1886,1913]" pageId="5" pageNumber="190" value="1991-02-05">5 February 1991</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF16504DCF8F2DDC2F77E" box="[959,1179,1886,1912]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF16504DCF8F2DD15F77E" box="[959,1100,1886,1912]" pageId="5" pageNumber="190">N. Mohanan</collectorName>
10325
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16503C8F8F3DDA3F77E" box="[1195,1274,1886,1912]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF165026CF8F2DB93F79A" ID-GBIF-Occurrence="3897114313" collectingDate="1991-07-30" collectionCode="TBGT" collectorName="N. Mohanan" country="India" elevation="700" location="Karamanayar" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF165026CF8F2DCF1F77F" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF165026CF8F2DCF1F77F" box="[1295,1448,1887,1913]" country="India" name="Karamanayar" pageId="5" pageNumber="190" stateProvince="Kerala">Karamanayar</location>
,
<quantity id="4CC6F5C9FF8AF16507EBF82FD98AF79B" box="[136,211,1922,1949]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="190" unit="m" value="700.0">
<elevation id="0013BF1FFF8AF16507EBF82FD98AF79B" box="[136,211,1922,1949]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="190" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="FF807EECFF8AF1650782F82FD82EF79A" box="[225,375,1922,1948]" pageId="5" pageNumber="190" value="1991-07-30">
<collectingDate id="EFC48704FF8AF1650782F82FD82EF79A" box="[225,375,1922,1948]" pageId="5" pageNumber="190" value="1991-07-30">30 July 1991</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF16506E6F82EDB3BF79A" box="[389,610,1922,1948]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF16506E6F82EDB4AF79A" box="[389,531,1922,1948]" pageId="5" pageNumber="190">N. Mohanan</collectorName>
10704
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650510F82FDB9BF79A" box="[627,706,1922,1948]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF16505BBF82ED82EF7C6" ID-GBIF-Occurrence="3897114323" collectingDate="1995-09-26" collectionCode="K, TBGT" country="India" elevation="660" location="Near Karamana River" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF16505BBF82EDA8AF79A" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF16505BBF82EDA8AF79A" box="[728,979,1922,1949]" country="India" name="Near Karamana River" pageId="5" pageNumber="190" stateProvince="Kerala">Near Karamana River</location>
,
<quantity id="4CC6F5C9FF8AF1650483F82FDD72F79B" box="[992,1067,1922,1949]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="5" pageNumber="190" unit="m" value="660.0">
<elevation id="0013BF1FFF8AF1650483F82FDD72F79B" box="[992,1067,1922,1949]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="5" pageNumber="190" unit="m" value="660.0">660 m</elevation>
</quantity>
,
<date id="FF807EECFF8AF165035AF82FDC40F79B" box="[1081,1305,1922,1949]" pageId="5" pageNumber="190" value="1995-09-26">
<collectingDate id="EFC48704FF8AF165035AF82FDC40F79B" box="[1081,1305,1922,1949]" pageId="5" pageNumber="190" value="1995-09-26">26 September 1995</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF1650245F82ED856F7C6" italics="true" pageId="5" pageNumber="190">
C.S.
<collectionCode id="ED2FC0E9FF8AF1650230F82EDC32F79B" box="[1363,1387,1923,1949]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="190" type="Herbarium">K</collectionCode>
. &amp; S. A.
<collectionCode id="ED2FC0E9FF8AF16507CBF80AD9E7F7C6" box="[168,190,1959,1984]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="190" type="Herbarium">K</collectionCode>
. 24359
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF165067CF80BD837F7C6" box="[287,366,1958,1984]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF16506E0F80ADAB6F7C6" ID-GBIF-Occurrence="3897114308" box="[387,1007,1958,1985]" collectionCode="TBGT" collectorName="Geethakumary" country="India" location="Karamanayar" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
near
<location id="8EE10EF7FF8AF16506D8F80ADB0DF7C7" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF16506D8F80ADB0DF7C7" box="[443,596,1959,1985]" country="India" name="Karamanayar" pageId="5" pageNumber="190" stateProvince="Kerala">Karamanayar</location>
,
<emphasis id="B94A843EFF8AF165053CF80BDBDFF7C6" box="[607,646,1958,1984]" italics="true" pageId="5" pageNumber="190">s.d.</emphasis>
,
<emphasis id="B94A843EFF8AF16505F6F80BDAD1F7C6" box="[661,904,1958,1984]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF16505F6F80BDA63F7C6" box="[661,826,1958,1984]" pageId="5" pageNumber="190">Geethakumary</collectorName>
63693
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16504F4F80BDABFF7C6" box="[919,998,1958,1984]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF165049AF80BDB1BF7E2" ID-GBIF-Occurrence="3897114326" collectingDate="1994-04-27" collectionCode="TBGT" collectorName="N. Mohanan &amp; T. Shaju" country="India" location="Agasthiyamala" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF165049AF80BDDF8F7C6" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF165049AF80BDDF8F7C6" box="[1017,1185,1958,1984]" country="India" name="Agasthiyamala" pageId="5" pageNumber="190" stateProvince="Kerala">Agasthiyamala</location>
MPCA,
<date id="FF807EECFF8AF1650264F80BDCFEF7C6" box="[1287,1447,1958,1984]" pageId="5" pageNumber="190" value="1994-04-27">
<collectingDate id="EFC48704FF8AF1650264F80BDCFEF7C6" box="[1287,1447,1958,1984]" pageId="5" pageNumber="190" value="1994-04-27">27 April 1994</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF16507EBF866D883F7E2" box="[136,474,1994,2021]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF16507EBF866D84CF7E2" box="[136,277,1994,2020]" pageId="5" pageNumber="190">N. Mohanan</collectorName>
&amp;
<collectorName id="26CB3DFAFF8AF165065EF866D8C3F7E2" box="[317,410,1994,2020]" pageId="5" pageNumber="190">T. Shaju</collectorName>
2055
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650689F867DB60F7E2" box="[490,569,1994,2020]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF165052CF867DC17F7E2" ID-GBIF-Occurrence="3897114304" box="[591,1358,1994,2021]" collectingDate="2001-08-14" collectionCode="TBGT" country="India" elevation="800" location="Cement" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF165052CF867DBFFF7E2" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF165052CF867DBFFF7E2" box="[591,678,1994,2020]" country="India" name="Cement" pageId="5" pageNumber="190" stateProvince="Kerala">Cement</location>
cana,
<quantity id="4CC6F5C9FF8AF1650592F867DA65F7E3" box="[753,828,1994,2021]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="190" unit="m" value="800.0">
<elevation id="0013BF1FFF8AF1650592F867DA65F7E3" box="[753,828,1994,2021]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="190" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="FF807EECFF8AF165042AF867DD59F7E2" box="[841,1024,1994,2020]" pageId="5" pageNumber="190" value="2001-08-14">
<collectingDate id="EFC48704FF8AF165042AF867DD59F7E2" box="[841,1024,1994,2020]" pageId="5" pageNumber="190" value="2001-08-14">14 August 2001</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF165036EF866DDBFF7E2" box="[1037,1254,1994,2020]" italics="true" pageId="5" pageNumber="190">NWFP team 46630</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650394F867DC1FF7E2" box="[1271,1350,1994,2020]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF1650239F866DADBF80E" ID-GBIF-Occurrence="3897114311" collectingDate="2011-11-16" collectionCode="TBGT" collectorName="Geethakumary" country="India" location="Koviltherimala" municipality="Way" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<collectingMunicipality id="6BE5C256FF8AF1650239F866DCD4F7E3" box="[1370,1421,1995,2021]" pageId="5" pageNumber="190">Way</collectingMunicipality>
to
<location id="8EE10EF7FF8AF16507EBF843D86DF80E" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF16507EBF843D86DF80E" box="[136,308,2030,2056]" country="India" municipality="Way" name="Koviltherimala" pageId="5" pageNumber="190" stateProvince="Kerala">Koviltherimala</location>
,
<date id="FF807EECFF8AF1650623F843DB45F80E" box="[320,540,2030,2056]" pageId="5" pageNumber="190" value="2011-11-16">
<collectingDate id="EFC48704FF8AF1650623F843DB45F80E" box="[320,540,2030,2056]" pageId="5" pageNumber="190" value="2011-11-16">16 November 2011</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF165054BF843DA42F80E" box="[552,795,2030,2056]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF165054BF843DB94F80E" box="[552,717,2030,2056]" pageId="5" pageNumber="190">Geethakumary</collectorName>
69884
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF1650448F843DA23F80E" box="[811,890,2030,2056]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF16504EDF843DC73F80E" ID-GBIF-Occurrence="3897114330" box="[910,1322,2030,2057]" collectingDate="2011-11-16" collectionCode="TBGT" collectorName="Geethakumary" country="India" location="Koviltherimala" municipality="Way" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<emphasis id="B94A843EFF8AF16504EDF843DAE3F80E" box="[910,954,2030,2056]" italics="true" pageId="5" pageNumber="190">ibid</emphasis>
.,
<emphasis id="B94A843EFF8AF16504B3F843DD9AF80E" box="[976,1219,2030,2056]" italics="true" pageId="5" pageNumber="190">Geethakumary 69886</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF16503B1F843DC78F80E" box="[1234,1313,2030,2056]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF8AF1650255F843DB9FF82A" ID-GBIF-Occurrence="3897114312" collectingDate="2011-11-16" collectionCode="TBGT" collectorName="Geethakumary" country="India" location="Chemunji" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Kerala">
<location id="8EE10EF7FF8AF1650255F843DCFEF80E" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF1650255F843DCFEF80E" box="[1334,1447,2030,2056]" country="India" name="Chemunji" pageId="5" pageNumber="190" stateProvince="Kerala">Chemunji</location>
,
<date id="FF807EECFF8AF16507EBF7BFD83AF82A" box="[136,355,2066,2092]" pageId="5" pageNumber="190" value="2011-11-16">
<collectingDate id="EFC48704FF8AF16507EBF7BFD83AF82A" box="[136,355,2066,2092]" pageId="5" pageNumber="190" value="2011-11-16">16 November 2011</collectingDate>
</date>
,
<emphasis id="B94A843EFF8AF165060DF7BFDB39F82A" box="[366,608,2066,2092]" italics="true" pageId="5" pageNumber="190">
<collectorName id="26CB3DFAFF8AF165060DF7BFDB4AF82A" box="[366,531,2066,2092]" pageId="5" pageNumber="190">Geethakumary</collectorName>
69888
</emphasis>
(
<collectionCode id="ED2FC0E9FF8AF165050CF7BFDBE7F82A" box="[623,702,2066,2092]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="5" pageNumber="190" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B565271FF8AF16605B3F7BFDBB7F0BE" ID-GBIF-Occurrence="3897114320" collectingDate="1976-08-31" collectionCode="MH" collectorName="A. N. Henry" country="India" county="Kanyakumari District" elevation="1000" lastPageId="6" lastPageNumber="191" location="Muthukuzhivayal" municipality="Way" pageId="5" pageNumber="190" specimenCount="1" stateProvince="Tamil Nadu">
<collectingRegion id="49FA96CEFF8AF16505B3F7BFDA25F82A" box="[720,892,2066,2092]" country="India" name="Tamil Nadu" pageId="5" pageNumber="190">TAMILNADU</collectingRegion>
:
<collectingCounty id="62E020A0FF8AF16504E4F7BFDD24F82A" box="[903,1149,2066,2092]" pageId="5" pageNumber="190">Kanyakumari District</collectingCounty>
,
<collectingMunicipality id="6BE5C256FF8AF16503EBF7BEDDE2F82B" box="[1160,1211,2067,2093]" pageId="5" pageNumber="190">Way</collectingMunicipality>
to
<location id="8EE10EF7FF8AF16503BCF7BFDCFEF82A" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF8AF16503BCF7BFDCFEF82A" box="[1247,1447,2066,2092]" country="India" county="Kanyakumari District" municipality="Way" name="Muthukuzhivayal" pageId="5" pageNumber="190" stateProvince="Tamil Nadu">Muthukuzhivayal</location>
, ±
<quantity id="4CC6F5C9FF89F16607C2FF33D9A2F0BF" box="[161,251,158,185]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="191" unit="m" value="1000.0">
<elevation id="0013BF1FFF89F16607C2FF33D9A2F0BF" box="[161,251,158,185]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="191" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FF807EECFF89F166066AFF33D89AF0BE" box="[265,451,158,184]" pageId="6" pageNumber="191" value="1976-08-31">
<collectingDate id="EFC48704FF89F166066AFF33D89AF0BE" box="[265,451,158,184]" pageId="6" pageNumber="191" value="1976-08-31">31 August 1976</collectingDate>
</date>
,
<emphasis id="B94A843EFF89F16606B2FF32DBFBF0BE" box="[465,674,158,185]" italics="true" pageId="6" pageNumber="191">
<collectorName id="26CB3DFAFF89F16606B2FF32DB0BF0BE" box="[465,594,159,185]" pageId="6" pageNumber="191">A.N. Henry</collectorName>
48174
</emphasis>
(
<collectionCode id="ED2FC0E9FF89F16605D7FF33DBBEF0BE" box="[692,743,158,184]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="6" pageNumber="191">MH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF89F166059FFF33DC7BF0BE" ID-GBIF-Occurrence="3897114315" box="[764,1314,158,184]" collectionCode="TBGT" collectorName="Narayana Iyer" country="India" location="Thovazha" pageId="6" pageNumber="191" specimenCount="1" stateProvince="TAMILNADU">
<location id="8EE10EF7FF89F166059FFF33DA33F0BE" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF89F166059FFF33DA33F0BE" box="[764,874,158,184]" country="India" name="Thovazha" pageId="6" pageNumber="191" stateProvince="TAMILNADU">Thovazha</location>
hill,
<emphasis id="B94A843EFF89F16604C8FF33DDE0F0BE" box="[939,1209,158,184]" italics="true" pageId="6" pageNumber="191">
s.d.,
<collectorName id="26CB3DFAFF89F1660481FF32DDD0F0BE" box="[994,1161,159,184]" pageId="6" pageNumber="191">Narayana Iyer</collectorName>
s.n.
</emphasis>
(
<collectionCode id="ED2FC0E9FF89F16603A8FF33DC43F0BE" box="[1227,1306,158,184]" country="India" lsid="urn:lsid:biocol.org:col:14234" name="Tropical Botanic Garden and Research Institute" pageId="6" pageNumber="191" type="Herbarium">TBGT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B565271FF89F1660253FF33DA0BF0DA" ID-GBIF-Occurrence="3897114318" collectingDate="1916-09-18" collectionCode="MH" country="India" location="Mahendragiri" municipality="Tirunelveli District" pageId="6" pageNumber="191" specimenCount="1" stateProvince="TAMILNADU">
<collectingMunicipality id="6BE5C256FF89F1660253FF33D986F0DA" pageId="6" pageNumber="191">Tirunelveli District</collectingMunicipality>
,
<location id="8EE10EF7FF89F1660789FF6FD8DCF0DA" LSID="urn:lsid:plazi:treatment:0397E93AFF8AF16607EBFF32DA0FF0DA:8EE10EF7FF89F1660789FF6FD8DCF0DA" box="[234,389,194,220]" country="India" municipality="Tirunelveli District" name="Mahendragiri" pageId="6" pageNumber="191" stateProvince="TAMILNADU">Mahendragiri</location>
,
<date id="FF807EECFF89F16606F3FF6FDB34F0DA" box="[400,621,194,220]" pageId="6" pageNumber="191" value="1916-09-18">
<collectingDate id="EFC48704FF89F16606F3FF6FDB34F0DA" box="[400,621,194,220]" pageId="6" pageNumber="191" value="1916-09-18">18 September 1916</collectingDate>
</date>
,
<emphasis id="B94A843EFF89F166051BFF6FDBEDF0DA" box="[632,692,194,220]" italics="true" pageId="6" pageNumber="191">s.coll</emphasis>
.
<emphasis id="B94A843EFF89F16605A1FF6FDA51F0DA" box="[706,776,194,220]" italics="true" pageId="6" pageNumber="191">13249</emphasis>
(
<collectionCode id="ED2FC0E9FF89F1660474FF6FDA13F0DA" box="[791,842,194,220]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="6" pageNumber="191">MH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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