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<mods:title id="F69181047F5D43DCE320448A80CD27BB">Taxonomic review of the lac insect genus Paratachardina Balachowsky (Hemiptera: Coccoidea: Kerriidae), with a revised key to genera of Kerriidae and description of two new species</mods:title>
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<mods:namePart id="C23E5758761391B3D780D1F53B4BD32F">Kondo, Takumasa</mods:namePart>
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<treatment id="0397AD19FF85FFAAC6CDFDE045CD6A76" ID-DOI="http://doi.org/10.5281/zenodo.6247479" ID-GBIF-Taxon="119375703" ID-Zenodo-Dep="6247479" LSID="urn:lsid:plazi:treatment:0397AD19FF85FFAAC6CDFDE045CD6A76" httpUri="http://treatment.plazi.org/id/0397AD19FF85FFAAC6CDFDE045CD6A76" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<subSubSection id="C3244F84FF85FFABC6CDFDE0431D6B45" box="[151,560,591,617]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B811C0FFF85FFABC6CDFDE0431D6B45" blockId="16.[151,560,591,617]" box="[151,560,591,617]" pageId="16" pageNumber="17">
<heading id="D0C9AB63FF85FFABC6CDFDE0431D6B45" bold="true" box="[151,560,591,617]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C3E678CFF85FFABC6CDFDE0431D6B45" ID-CoL="75QD4" authority="Varshney" authorityName="Varshney" box="[151,560,591,617]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithila">
<emphasis id="B94AC01DFF85FFABC6CDFDE0431D6B45" bold="true" box="[151,560,591,617]" pageId="16" pageNumber="17">
<emphasis id="B94AC01DFF85FFABC6CDFDE0409F6B45" bold="true" box="[151,434,591,617]" italics="true" pageId="16" pageNumber="17">Paratachardina mithila</emphasis>
Varshney
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3244F84FF85FFABC6CDFD3545656BFD" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8B811C0FFF85FFABC6CDFD3543CB6B9D" blockId="16.[151,1096,666,721]" box="[151,742,666,689]" pageId="16" pageNumber="17">
<taxonomicName id="4C3E678CFF85FFABC6CDFD35435B6B9D" ID-CoL="75QD4" authority="Varshney, 1968: 489" authorityName="Varshney" authorityPageNumber="489" authorityYear="1968" box="[151,630,666,689]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithila">
<emphasis id="B94AC01DFF85FFABC6CDFD3540BF6B9D" box="[151,402,666,689]" italics="true" pageId="16" pageNumber="17">Paratachardina mithila</emphasis>
Varshney, 1968: 489
</taxonomicName>
; 1977: 58.
</paragraph>
<paragraph id="8B811C0FFF85FFABC6CDFD1545656BFD" blockId="16.[151,1096,666,721]" box="[151,1096,698,721]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB2E3B21FF85FFABC6CDFD1545656BFD" box="[151,1096,698,721]" pageId="16" pageNumber="17">
<taxonomicName id="4C3E678CFF85FFABC6CDFD1543586BFD" ID-CoL="762D4" authority="Varshney, 1997: 30" authorityName="Varshney" authorityPageNumber="30" authorityYear="1997" box="[151,629,698,721]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithilae">
<emphasis id="B94AC01DFF85FFABC6CDFD1540B36BFD" box="[151,414,698,721]" italics="true" pageId="16" pageNumber="17">Paratachardina mithilae</emphasis>
<treatmentCitation id="0A9F3A1EFF85FFABC7FFFD1543586BFD" author="Varshney" box="[421,629,698,721]" page="30" pageId="16" pageNumber="17" year="1997">
<bibRefCitation id="EFAF61FEFF85FFABC7FFFD1543606BFD" author="Varshney" box="[421,589,698,721]" pageId="16" pageNumber="41" refString="Varshney, R. K. (1997) An up-to-date list of the lac insects of the world. Proceedings of the National Seminar: Lac Industry -- Challenges and Solutions, 27 - 31." type="proceedings paper" year="1997">Varshney, 1997</bibRefCitation>
: 30
</treatmentCitation>
</taxonomicName>
. Incorrect subsequent spelling [see 'Notes'].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3244F84FF85FFAAC6CDFCAD45CD6A76" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8B811C0FFF85FFABC6CDFCAD42936A40" blockId="16.[151,1437,770,2036]" pageId="16" pageNumber="17">
<emphasis id="B94AC01DFF85FFABC6CDFCAD40B76A30" bold="true" box="[151,410,770,796]" pageId="16" pageNumber="17">
<typeStatus id="5485A2ADFF85FFABC6CDFCAD41F96A30" box="[151,212,770,796]" pageId="16" pageNumber="17">Type</typeStatus>
data.
<typeStatus id="5485A2ADFF85FFABC77AFCAD40B96A30" box="[288,404,770,796]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
,
</emphasis>
adult female.
<emphasis id="B94AC01DFF85FFABC465FCAD422C6A30" bold="true" box="[575,769,770,796]" pageId="16" pageNumber="17">
<collectingCountry id="F3295C9FFF85FFABC465FCAD43B76A30" box="[575,666,770,796]" name="India" pageId="16" pageNumber="17">INDIA</collectingCountry>
: Assam,
</emphasis>
Shillong, in the gardens of Ward Lake, coll. R. K.
<bibRefCitation id="EFAF61FEFF85FFABC315FCAD40306A68" author="Varshney" pageId="16" pageNumber="41" refString="Varshney, R. K. (1967) A note on the several species of Indian coccids (Homopt., Coccoidea), proposed by E. E. Green as nomina nuda. Entomologist's Monthly Magazine, 102 (1223 - 1225), 77 - 79." type="journal article" year="1967">
Varshney,
<date id="FF803ACFFF85FFABC68AFC8540306A68" box="[208,285,810,836]" pageId="16" pageNumber="17" value="1967-01">i.1967</date>
</bibRefCitation>
, on
<taxonomicName id="4C3E678CFF85FFABC70BFC85431B6A6F" box="[337,566,810,835]" class="Magnoliopsida" family="Rosaceae" genus="Photinia" kingdom="Plantae" order="Rosales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="notoniana">
<emphasis id="B94AC01DFF85FFABC70BFC85431B6A6F" box="[337,566,810,835]" italics="true" pageId="16" pageNumber="17">Photinia notoniana</emphasis>
</taxonomicName>
<taxonomicName id="4C3E678CFF85FFABC465FC85422A6A6F" box="[575,775,810,836]" pageId="16" pageNumber="17" rank="variety" variety="macrophylla">
var.
<emphasis id="B94AC01DFF85FFABC42EFC85422A6A6F" box="[628,775,810,835]" italics="true" pageId="16" pageNumber="17">macrophylla</emphasis>
</taxonomicName>
.
<emphasis id="B94AC01DFF85FFABC54DFC8542B36A68" bold="true" box="[791,926,810,836]" pageId="16" pageNumber="17">
<typeStatus id="5485A2ADFF85FFABC54DFC8542B46A68" box="[791,921,810,836]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
:
</emphasis>
same data as
<typeStatus id="5485A2ADFF85FFABC213FC8545826A68" box="[1097,1199,810,836]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
except some specimens coll.
<date id="FF803ACFFF85FFABC74CFCFD40426A40" box="[278,367,850,876]" pageId="16" pageNumber="17" value="1967-06">vi.1967</date>
or
<date id="FF803ACFFF85FFABC7CCFCFD40D26A40" box="[406,511,850,876]" pageId="16" pageNumber="17" value="1970-08">viii.1970</date>
(
<collectionCode id="ED2F84CAFF85FFABC455FCFD437B6A40" box="[527,598,850,876]" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="16" pageNumber="17">NZSI</collectionCode>
). [
<typeStatus id="5485A2ADFF85FFABC42AFCFD43946A40" box="[624,697,850,876]" pageId="16" pageNumber="17">Types</typeStatus>
not seen; see 'Notes'.]
</paragraph>
<paragraph id="8B811C0FFF85FFABC69CFCD5404D6AB8" blockId="16.[151,1437,770,2036]" box="[198,352,890,916]" pageId="16" pageNumber="17">Adult female</paragraph>
<paragraph id="8B811C0FFF85FFABC69DFC0D44016D18" blockId="16.[151,1437,770,2036]" pageId="16" pageNumber="17">
The following descriptions of unmounted and mounted material are adapted from
<bibRefCitation id="EFAF61FEFF85FFABC296FC0D44B56A90" author="Varshney" box="[1228,1432,930,956]" pageId="16" pageNumber="41" refString="Varshney, R. K. (1977) Taxonomic studies on lac insects of India (Homoptera: Tachardiidae). Oriental Insects, Supplement 5 (1976), 1 - 97." type="journal article" year="1977">Varshney (1977)</bibRefCitation>
.
<emphasis id="B94AC01DFF85FFABC6CDFC6540886AC8" bold="true" box="[151,421,970,996]" pageId="16" pageNumber="17">Unmounted material.</emphasis>
Lac test of adult female almost round, brownish black, with three small openings on top for brachial and anal orifices; with 16 conspicuous longitudinal ridges that divide the test into sectors; a circular spot on the middle of each ridge, probably corresponding to marginal duct cluster openings.
</paragraph>
<paragraph id="8B811C0FFF85FFABC69DFBED43016CC0" blockId="16.[151,1437,770,2036]" pageId="16" pageNumber="17">
<emphasis id="B94AC01DFF85FFABC69DFBED40986D70" bold="true" box="[199,437,1090,1116]" pageId="16" pageNumber="17">Mounted material.</emphasis>
Body trilobed, 2.53.0 mm long, 2.83.0 mm wide. Brachia short, 103 µm long. Each brachial plate oval, distal half slightly larger, each 68120 µm long, 5170 µm wide; pseudospines totalling 4450, occupying about two-thirds area of brachial plate center, with gaps on upper large portion. Anterior spiracles each 137 µm long, 86 µm wide, situated far away from brachial plates, spiracular pores with 5- loculi. Dorsal spine small, conical, 6870 µm long, with a hollow, not pointed tip; membranous pedicel of dorsal spine well developed, 70103 um long and 70103 µm wide. Anal tubercle well developed, 86170 µm long, 120140 µm wide; supra-anal plate subequal or slightly longer than its maximum width. Anal ring not divided in sectors; supra-anal plate forming a cup-shaped cavity. Anal fringe of few acute lobes, with narrow and deep clefts. Anal ring setae just reach, or slightly protrude past anal fringe. Antennae minute and obscure. Marginal duct clusters in 8 pairs, each roughly round, poorly demarcated, with ducts arranged irregularly. Ventral duct clusters present.
</paragraph>
<paragraph id="8B811C0FFF85FFABC69DFA55429C6E00" blockId="16.[151,1437,770,2036]" pageId="16" pageNumber="17">
<emphasis id="B94AC01DFF85FFABC69DFA5540396F38" bold="true" box="[199,276,1530,1556]" pageId="16" pageNumber="17">Notes.</emphasis>
Subsequent to his original description,
<bibRefCitation id="EFAF61FEFF85FFABC559FA5542E66F38" author="Varshney" box="[771,971,1530,1556]" pageId="16" pageNumber="41" refString="Varshney, R. K. (1997) An up-to-date list of the lac insects of the world. Proceedings of the National Seminar: Lac Industry -- Challenges and Solutions, 27 - 31." type="proceedings paper" year="1997">Varshney (1997)</bibRefCitation>
listed the species name as &quot;
<taxonomicName id="4C3E678CFF85FFABC377FA5544B96F38" box="[1325,1428,1530,1556]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithilae">mithilae</taxonomicName>
&quot; rather than &quot;
<taxonomicName id="4C3E678CFF85FFABC770F98D40AB6F10" box="[298,390,1570,1596]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithila">mithila</taxonomicName>
&quot;, without giving an explanation for his action. Varshney's (1968, 1977) descriptions do not specify the etymology of the name &quot;
<taxonomicName id="4C3E678CFF85FFABC437F9E543E46F48" box="[621,713,1610,1636]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithila">mithila</taxonomicName>
&quot;, and do not indicate whether it should be regarded as a noun or an adjective. According to the Article 31.2.2 of the
<emphasis id="B94AC01DFF85FFABC54FF9DC446B6FA0" box="[789,1350,1651,1676]" italics="true" pageId="16" pageNumber="17">International Code of Zoological Nomenclature</emphasis>
(
<collectionCode id="ED2F84CAFF85FFABC30CF9DD44B16FA0" box="[1366,1436,1650,1676]" pageId="16" pageNumber="17">ICZN</collectionCode>
1999), the name “
<taxonomicName id="4C3E678CFF85FFABC734F93540EA6F98" box="[366,455,1690,1716]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithila">mithila</taxonomicName>
” becomes a noun in apposition and should be retained as &quot;
<taxonomicName id="4C3E678CFF85FFABC2CBF93545C36F98" box="[1169,1262,1690,1716]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithila">mithila</taxonomicName>
&quot;. Even though Varshney (personal communication) emended &quot;
<taxonomicName id="4C3E678CFF85FFABC49DF96D420C6FF0" box="[711,801,1730,1756]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithila">mithila</taxonomicName>
&quot; to &quot;
<taxonomicName id="4C3E678CFF85FFABC501F96D42EF6FF0" box="[859,962,1730,1756]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithilae">mithilae</taxonomicName>
&quot; because the species was named after a woman, articles 31, 32 and 33 of the
<collectionCode id="ED2F84CAFF85FFABC417F94543BF6E28" box="[589,658,1770,1796]" pageId="16" pageNumber="17">ICZN</collectionCode>
(1999) make it clear that such an alteration to the species name is an incorrect subsequent spelling, as recognised by
<bibRefCitation id="EFAF61FEFF85FFABC4B2F8BD42836E00" author="Ben-Dov" box="[744,942,1810,1836]" pageId="16" pageNumber="39" refString="Ben-Dov, Y. (2006) A systematic catalogue of eight scale insect families (Hemiptera: Coccoidea) of the world: Aclerdidae, Asterolecaniidae, Beesoniidae, Carayonemidae, Conchaspididae, Dactylopiidae, Kerriidae and Lecanodiaspididae. Elsevier, Amsterdam, 368 pp." type="book" year="2006">Ben-Dov (2006)</bibRefCitation>
.
</paragraph>
<paragraph id="8B811C0FFF85FFABC69DF89544B46ED8" blockId="16.[151,1437,770,2036]" pageId="16" pageNumber="17">
According to
<bibRefCitation id="EFAF61FEFF85FFABC732F89543006E78" author="Varshney" box="[360,557,1850,1876]" pageId="16" pageNumber="41" refString="Varshney, R. K. (1977) Taxonomic studies on lac insects of India (Homoptera: Tachardiidae). Oriental Insects, Supplement 5 (1976), 1 - 97." type="journal article" year="1977">Varshney (1977)</bibRefCitation>
, this species is similar to
<emphasis id="B94AC01DFF85FFABC50DF89442826E78" box="[855,943,1851,1876]" italics="true" pageId="16" pageNumber="17">P. t h e a e</emphasis>
, from which it can be separated due to its larger adult female size, anal tubercle subequal in length and width, and pedicel of the dorsal spine not much longer than the length of the spine itself.
<typeStatus id="5485A2ADFF85FFABC421F82543986E88" box="[635,693,1930,1956]" pageId="16" pageNumber="17">Type</typeStatus>
material of
<taxonomicName id="4C3E678CFF85FFABC51EF82442986E88" box="[836,949,1931,1956]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithila">
<emphasis id="B94AC01DFF85FFABC51EF82442986E88" box="[836,949,1931,1956]" italics="true" pageId="16" pageNumber="17">P. mithila</emphasis>
</taxonomicName>
was not available in the present study, as we did not receive a reply to our request for a loan from the
<collectionCode id="ED2F84CAFF85FFABC50DF81D42B66EE0" box="[855,923,1970,1996]" httpUri="http://grbio.org/cool/7mhb-r8q3" name="Zoological Survey of India, National Zoological Collection" pageId="16" pageNumber="17">NZSI</collectionCode>
, and no
<typeStatus id="5485A2ADFF85FFABC5A7F81D451D6EE0" box="[1021,1072,1970,1996]" pageId="16" pageNumber="17">type</typeStatus>
material or non-type topotypic specimens could be located in any other museum.
<bibRefCitation id="EFAF61FEFF85FFABC4AFF87542976ED8" author="Varshney" box="[757,954,2010,2036]" pageId="16" pageNumber="41" refString="Varshney, R. K. (1977) Taxonomic studies on lac insects of India (Homoptera: Tachardiidae). Oriental Insects, Supplement 5 (1976), 1 - 97." type="journal article" year="1977">Varshney (1977)</bibRefCitation>
gave a key to separate
<taxonomicName id="4C3E678CFF85FFABC28DF87444646ED8" box="[1239,1353,2011,2036]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mithila">
<emphasis id="B94AC01DFF85FFABC28DF87444646ED8" box="[1239,1353,2011,2036]" italics="true" pageId="16" pageNumber="17">P. mithila</emphasis>
</taxonomicName>
and
<emphasis id="B94AC01DFF85FFABC3DCF87444B46ED8" box="[1414,1433,2011,2036]" italics="true" pageId="16" pageNumber="17">P.</emphasis>
</paragraph>
<paragraph id="8B811C0FFF84FFAAC6CDFF37424D699E" blockId="17.[151,864,152,178]" box="[151,864,152,178]" pageId="17" pageNumber="18">
<taxonomicName id="4C3E678CFF84FFAAC6CDFF3741FA699D" box="[151,215,152,177]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="theae">
<emphasis id="B94AC01DFF84FFAAC6CDFF3741FA699D" box="[151,215,152,177]" italics="true" pageId="17" pageNumber="18">theae</emphasis>
</taxonomicName>
as follows (
<bibRefCitation id="EFAF61FEFF84FFAAC73EFF37427F699E" author="Varshney" box="[356,850,152,178]" pageId="17" pageNumber="41" refString="Varshney, R. K. (1977) Taxonomic studies on lac insects of India (Homoptera: Tachardiidae). Oriental Insects, Supplement 5 (1976), 1 - 97." type="journal article" year="1977">Varshney 1977: 56, key couplet number 4</bibRefCitation>
):
</paragraph>
<paragraph id="8B811C0FFF84FFAAC6CDFF4744B76855" blockId="17.[151,1436,232,378]" pageId="17" pageNumber="18">
Anal tubercle slightly longer than its maximum width; pedicel of dorsal spine as long as spine itself ........ ............................................................................................................................................................
<taxonomicName id="4C3E678CFF84FFAAC31DFEBF44B76805" box="[1351,1434,272,297]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="mithila">
<emphasis id="B94AC01DFF84FFAAC31DFEBF44B76805" box="[1351,1434,272,297]" italics="true" pageId="17" pageNumber="18">mithila</emphasis>
</taxonomicName>
Anal tubercle distinctly broader than its maximum length; pedicel of dorsal spine much longer than the spine
<emphasis id="B94AC01DFF84FFAAC75DFECF44B76855" box="[263,1434,352,377]" italics="true" pageId="17" pageNumber="18">
.......................................................................................................................................................
<taxonomicName id="4C3E678CFF84FFAAC304FECF44B76855" box="[1374,1434,352,377]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="theae">theae</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8B811C0FFF84FFAAC69CFE1F45CD6A76" blockId="17.[151,1436,432,858]" pageId="17" pageNumber="18">
However, Varshneys (1977) description of
<taxonomicName id="4C3E678CFF84FFAAC49FFE1F421968E5" box="[709,820,432,457]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="mithila">
<emphasis id="B94AC01DFF84FFAAC49FFE1F421968E5" box="[709,820,432,457]" italics="true" pageId="17" pageNumber="18">P. mithila</emphasis>
</taxonomicName>
overlaps with his description of
<emphasis id="B94AC01DFF84FFAAC2EEFE1F442168E5" box="[1204,1292,432,457]" italics="true" pageId="17" pageNumber="18">P. t h e a e</emphasis>
in the character states used to separate them in the key. The minimum length and width of the anal tubercle of
<taxonomicName id="4C3E678CFF84FFAAC376FE7744B168DD" box="[1324,1436,472,497]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="mithila">
<emphasis id="B94AC01DFF84FFAAC376FE7744B168DD" box="[1324,1436,472,497]" italics="true" pageId="17" pageNumber="18">P. mithila</emphasis>
</taxonomicName>
given by Varshneys (1977) description is 86 µm and 120 µm, respectively, in which case, there must be specimens for which the anal tubercle is distinctly broader than its maximum length. On the other hand, the anal tubercle in the
<typeStatus id="5485A2ADFF84FFAAC711FDFF409F6B46" box="[331,434,592,618]" pageId="17" pageNumber="18" type="syntype">syntypes</typeStatus>
of
<emphasis id="B94AC01DFF84FFAAC786FDFF431B6B45" box="[476,566,592,617]" italics="true" pageId="17" pageNumber="18">P. t h e a e</emphasis>
herein studied are approximately as long as wide, with some specimens being slightly longer than wide, and others being slightly wider than long. Furthermore, the length of the pedicel of the dorsal spine also varies in
<emphasis id="B94AC01DFF84FFAAC42DFD0F43FC6B95" box="[631,721,672,697]" italics="true" pageId="17" pageNumber="18">P. t h e a e</emphasis>
and sometimes is about the same length as the spine. Specimens from
<collectingCountry id="F3295C9FFF84FFAAC741FD67404C6BCE" box="[283,353,712,738]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
collected on the same host genus as
<emphasis id="B94AC01DFF84FFAAC556FD6742556BCD" box="[780,888,712,737]" italics="true" pageId="17" pageNumber="18">P. m i t h i l a</emphasis>
, i.e., on
<taxonomicName id="4C3E678CFF84FFAAC585FD6745C96BCD" box="[991,1252,712,737]" class="Magnoliopsida" family="Rosaceae" genus="Photinia" kingdom="Plantae" order="Rosales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="benthamiana">
<emphasis id="B94AC01DFF84FFAAC585FD6745C96BCD" box="[991,1252,712,737]" italics="true" pageId="17" pageNumber="18">Photinia benthamiana</emphasis>
</taxonomicName>
, were available for study (see 'Other material studied' under
<emphasis id="B94AC01DFF84FFAAC4F9FD5F43D16A25" box="[675,764,752,777]" italics="true" pageId="17" pageNumber="18">P. t h e a e</emphasis>
), but these could not be separated morphologically from
<emphasis id="B94AC01DFF84FFAAC6CDFCB741DD6A1D" box="[151,240,792,817]" italics="true" pageId="17" pageNumber="18">P. t h e a e</emphasis>
. Thus adult females of
<taxonomicName id="4C3E678CFF84FFAAC455FCB743A06A1D" box="[527,653,792,817]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="mithilae">
<emphasis id="B94AC01DFF84FFAAC455FCB743A06A1D" box="[527,653,792,817]" italics="true" pageId="17" pageNumber="18">P. mithilae</emphasis>
</taxonomicName>
and
<emphasis id="B94AC01DFF84FFAAC492FCB7420F6A1D" box="[712,802,792,817]" italics="true" pageId="17" pageNumber="18">P. t h e a e</emphasis>
appear similar in all features considered and the two species cannot be separated with the available information (see also 'Diagnosis' of
<taxonomicName id="4C3E678CFF84FFAAC204FCEF45FD6A75" box="[1118,1232,832,857]" class="Insecta" family="Kerriidae" genus="Paratachardina" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ternata">
<emphasis id="B94AC01DFF84FFAAC204FCEF45FD6A75" box="[1118,1232,832,857]" italics="true" pageId="17" pageNumber="18">P. ternata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>