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<document id="8458E9AAACB3BC8DCAA98BA74400CA0C" ID-DOI="10.5281/zenodo.178539" ID-GBIF-Dataset="dda53384-9974-4b83-b780-a9cd51fffacf" ID-ISSN="1175-5326" ID-Zenodo-Dep="178539" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459899930198" checkinUser="plazi" docAuthor="Freitag, Hendrik &amp; Jäch, Manfred A." docDate="2007" docId="03939707FFAC2553FF2CFE70E2BDFE35" docLanguage="en" docName="zt01590p059.pdf" docOrigin="Zootaxa 1590" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ancyronyx minutulus Freitag &amp; Jach, sp. n." docType="treatment" docVersion="10" lastPageNumber="55" masterDocId="FFAAEF7FFFBC2541FFBBFFBDE027FF97" masterDocTitle="The genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) in Palawan and Busuanga, (Philippines) with descriptions of six new species" masterLastPageNumber="59" masterPageNumber="37" pageNumber="53" updateTime="1698228715028" updateUser="plazi">
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<mods:title id="DCFC4B95EA2A650D4BB9A508E146A89D">The genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) in Palawan and Busuanga, (Philippines) with descriptions of six new species</mods:title>
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<mods:namePart id="EFA6F9324C38128CCC7C7A74DE63076C">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="4DE049104D3CA0C4D6C4C2E21204F12C">Jäch, Manfred A.</mods:namePart>
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<mods:date id="C0A8318DACF4B581A85B1A71941F9429">2007</mods:date>
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<treatment id="03939707FFAC2553FF2CFE70E2BDFE35" ID-DOI="http://doi.org/10.5281/zenodo.6243457" ID-GBIF-Taxon="119372909" ID-Zenodo-Dep="6243457" LSID="urn:lsid:zoobank.org:act:F84835DC-D96D-4A08-A65F-9BDF597B8C09" httpUri="http://treatment.plazi.org/id/03939707FFAC2553FF2CFE70E2BDFE35" lastPageId="18" lastPageNumber="55" pageId="16" pageNumber="53">
<subSubSection id="C320759AFFAC2551FF2CFE70E16CFD9D" pageId="16" pageNumber="53" type="nomenclature">
<paragraph id="8B852611FFAC2551FF2CFE70E28CFE70" blockId="16.[151,683,461,522]" box="[151,683,461,487]" pageId="16" pageNumber="53">
<heading id="D0CD917DFFAC2551FF2CFE70E28CFE70" bold="true" box="[151,683,461,487]" fontSize="11" level="1" pageId="16" pageNumber="53" reason="1">
<emphasis id="B94EFA03FFAC2551FF2CFE70E28CFE70" bold="true" box="[151,683,461,487]" pageId="16" pageNumber="53">
<taxonomicName id="4C3A5D92FFAC2551FF2CFE70E245FE70" ID-CoL="DPSS" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[151,610,461,487]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="53" phylum="Arthropoda" rank="species" species="minutulus" status="sp. nov.">
<emphasis id="B94EFA03FFAC2551FF2CFE70E1B9FE70" bold="true" box="[151,414,461,487]" italics="true" pageId="16" pageNumber="53">Ancyronyx minutulus</emphasis>
Freitag &amp; Jäch
</taxonomicName>
<taxonomicNameLabel id="A27D4778FFAC2551FDD2FE70E28CFE70" box="[617,683,461,487]" pageId="16" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B852611FFAC2551FF2CFE4DE16CFD9D" blockId="16.[151,683,461,522]" box="[151,331,496,522]" pageId="16" pageNumber="53">
(
<figureCitation id="13013A94FFAC2551FF24FE4DE0D4FD9D" box="[159,243,496,522]" captionStart="FIGURES 7 10" captionStartId="6.[151,269,1956,1980]" captionTargetBox="[156,1436,194,1932]" captionTargetId="figure@6.[151,1436,194,1932]" captionTargetPageId="6" captionText="FIGURES 7 10. Habitus of 7) Ancyronyx minerva sp. n., paratype Ψ; 8) A. minerva sp. n., paratype Ψ, winged, partly completed; 9) A. minutulus sp. n., holotype ♂, partly completed; 10) A. helgeschneideri sp. n., paratype Ψ." httpUri="https://zenodo.org/record/178543/files/figure.png" pageId="16" pageNumber="53">Figs. 9</figureCitation>
,
<figureCitation id="13013A94FFAC2551FF44FE4DE13DFD9D" box="[255,282,496,522]" captionStart="FIGURE 15" captionStartId="17.[151,255,1743,1767]" captionTargetBox="[151,1436,183,1683]" captionTargetId="figure@17.[151,1436,183,1683]" captionTargetPageId="17" captionText="FIGURE 15. Ancyronyx minutulus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male sternite VIII (strioles omitted); e) male tergite VIII (setae and structures of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178548/files/figure.png" pageId="16" pageNumber="53">15</figureCitation>
af)
</paragraph>
</subSubSection>
<subSubSection id="C320759AFFAC2551FF2CFD87E280FD5B" pageId="16" pageNumber="53" type="materials_examined">
<paragraph id="8B852611FFAC2551FF2CFD87E498FDC3" blockId="16.[151,1437,570,2036]" box="[151,1215,570,596]" pageId="16" pageNumber="53">
<emphasis id="B94EFA03FFAC2551FF2CFD87E11AFDC3" bold="true" box="[151,317,570,596]" pageId="16" pageNumber="53">
<typeStatus id="548198B3FFAC2551FF2CFD87E0F3FDC3" box="[151,212,570,596]" pageId="16" pageNumber="53">Type</typeStatus>
locality.
</emphasis>
Balsahan River, Iwahig District, Puerto Princesa City, Palawan,
<collectingCountry id="F32D6681FFAC2551FB8FFD87E49BFDC3" box="[1076,1212,570,596]" name="Philippines" pageId="16" pageNumber="53">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8B852611FFAC2551FF7DFDDFE280FD5B" blockId="16.[151,1437,570,2036]" pageId="16" pageNumber="53">
<emphasis id="B94EFA03FFAC2551FF7DFDDFE1D3FDEB" bold="true" box="[198,500,610,636]" pageId="16" pageNumber="53">
<typeStatus id="548198B3FFAC2551FF7DFDDFE126FDEB" box="[198,257,610,636]" pageId="16" pageNumber="53">Type</typeStatus>
material.
<typeStatus id="548198B3FFAC2551FE39FDDFE1D3FDEB" box="[386,500,610,636]" pageId="16" pageNumber="53" type="holotype">Holotype</typeStatus>
</emphasis>
ď (
<collectionCode id="ED2BBED4FFAC2551FDA6FDDFE248FDEB" box="[541,623,610,636]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="53">NMW</collectionCode>
): “
<collectingCountry id="F32D6681FFAC2551FD37FDDFE36EFDEB" box="[652,841,610,636]" name="Philippines" pageId="16" pageNumber="53">PHILIPPINEN</collectingCountry>
.: Palawan
<quantity id="4CC28BF4FFAC2551FC7AFDDFE3DBFDEB" box="[961,1020,610,636]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="53" unit="km" value="9.0">9 km</quantity>
W P. Princesa Iwahig,Balsahan riv. lg. Zettel,
<date id="FF8400D1FFAC2551FEA3FD37E1B1FD33" box="[280,406,650,676]" pageId="16" pageNumber="53" value="1994-03-24">24.3.1994</date>
(48)”, dissected (terminal parts of abdomen, aedeagus and one leg glued separately), major parts of abdomen and one leg missing.
</paragraph>
</subSubSection>
<subSubSection id="C320759AFFAC2553FF7DFD67E22CFEBD" lastPageId="18" lastPageNumber="55" pageId="16" pageNumber="53" type="diagnosis">
<paragraph id="8B852611FFAC2551FF7DFD67E273FC8B" blockId="16.[151,1437,570,2036]" pageId="16" pageNumber="53">
<emphasis id="B94EFA03FFAC2551FF7DFD67E161FD63" bold="true" box="[198,326,730,756]" pageId="16" pageNumber="53">Diagnosis.</emphasis>
1.0 mm long (CL),
<quantity id="4CC28BF4FFAC2551FD86FD67E28FFD63" box="[573,680,730,756]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="16" pageNumber="53" unit="mm" value="0.38">0.38 mm</quantity>
broad (EW), 2.6 times as broad as wide (CL/EW). Body form elongate, moderately convex dorsally.
</paragraph>
<paragraph id="8B852611FFAC2551FF2CFC97E48DFC2B" blockId="16.[151,1437,570,2036]" pageId="16" pageNumber="53">
Colouration (
<figureCitation id="13013A94FFAC2551FE89FC97E15FFCD3" box="[306,376,810,836]" captionStart="FIGURES 7 10" captionStartId="6.[151,269,1956,1980]" captionTargetBox="[156,1436,194,1932]" captionTargetId="figure@6.[151,1436,194,1932]" captionTargetPageId="6" captionText="FIGURES 7 10. Habitus of 7) Ancyronyx minerva sp. n., paratype Ψ; 8) A. minerva sp. n., paratype Ψ, winged, partly completed; 9) A. minutulus sp. n., holotype ♂, partly completed; 10) A. helgeschneideri sp. n., paratype Ψ." httpUri="https://zenodo.org/record/178543/files/figure.png" pageId="16" pageNumber="53">Fig. 9</figureCitation>
) predominantly dark brown (labrum, maxillary palpi, clypeus, frons, pronotum, scutellum, parts of elytra); elytra with two pairs of elongately oval yellowish patches; ventral side distinctly paler brown than dark dorsal parts; legs predominantly yellowish, but base and apex of femur, base and tip of tibia, and apex of terminal tarsal segment brownish; antennae yellowish, distally slightly darkened.
</paragraph>
<paragraph id="8B852611FFAC2551FF7DFC77E2D7FBCB" blockId="16.[151,1437,570,2036]" pageId="16" pageNumber="53">
Head
<quantity id="4CC28BF4FFAC2551FEAAFC77E158FC73" box="[273,383,970,996]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="16" pageNumber="53" unit="mm" value="0.24">0.24 mm</quantity>
broad (HW); ID
<quantity id="4CC28BF4FFAC2551FDE4FC77E2F7FC73" box="[607,720,970,996]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="53" unit="mm" value="0.15">0.15 mm</quantity>
; labrum slightly pubescent, appearing glabrous; frons and clypeus finely punctate, moderately densely pubescent; frontoclypeal suture hardly discernible. Eyes moderately protruding. Antennae 11-segmented, slender, approximately as long as HW. Gula microreticulate; gular sutures narrowly separated mesally, inconspicuous.
</paragraph>
<paragraph id="8B852611FFAC2551FF7DFBD7E252FAB3" blockId="16.[151,1437,570,2036]" pageId="16" pageNumber="53">
Pronotum
<quantity id="4CC28BF4FFAC2551FEF9FBD7E18BFB13" box="[322,428,1130,1156]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="53" unit="mm" value="0.3">0.30 mm</quantity>
long (PL),
<quantity id="4CC28BF4FFAC2551FD8CFBD7E286FB13" box="[567,673,1130,1156]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="53" unit="mm" value="0.32">0.32 mm</quantity>
broad (MW), ca. 0.9 times as long as wide (PL/MW), widest at ca. posterior 0.4, anteriorly attenuate; anterior margin nearly straight; margin between pronotum and hypomeron conspiciuous; anterior transverse groove distinct; posterolateral oblique grooves inconspicuous; pronotal surface microreticulate and pubescent, a small part close to scutellum almost glabrous. Prosternum distinctly punctate, interstices granulose.
</paragraph>
<paragraph id="8B852611FFAC2551FF7DFA8FE19CF983" blockId="16.[151,1437,570,2036]" pageId="16" pageNumber="53">
Elytra elongately oval,
<quantity id="4CC28BF4FFAC2551FE6CFA8FE267FADB" box="[471,576,1330,1356]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="16" pageNumber="53" unit="mm" value="0.74">0.74 mm</quantity>
long (EL), ca. 1.8 times as long as wide (EL/EW), with ca. eight longitudinal, moderately regular, slightly impressed rows of punctures (five strial rows between suture and shoulder); punctures moderately large and moderately deeply impressed, including those at yellowish patches; interstices and intervals convex, glabrous; lateral elytral gutter narrow; elytra apically conjointly rounded. Meso- and metaventrite distinctly punctate; interstices glabrous; metaventral longitudinal groove distinct and moderately long.
<collectingCountry id="F32D6681FFAC2551FF61FA47E132F983" box="[218,277,1530,1556]" name="India" pageId="16" pageNumber="53">Hind</collectingCountry>
wings absent.
</paragraph>
<paragraph id="8B852611FFAC2551FF7DF99FE4E0F9F3" blockId="16.[151,1437,570,2036]" pageId="16" pageNumber="53">Legs slightly shorter than body; setae rather small and inconspicuous; claws well developed, strongly curved, base of each claw with projection carrying one very small and inconspicuous tooth.</paragraph>
<paragraph id="8B852611FFAC2551FF7DF9CFE284F923" blockId="16.[151,1437,570,2036]" pageId="16" pageNumber="53">
Sternite IX (
<figureCitation id="13013A94FFAC2551FEE6F9CFE193F91B" box="[349,436,1650,1676]" captionStart="FIGURE 15" captionStartId="17.[151,255,1743,1767]" captionTargetBox="[151,1436,183,1683]" captionTargetId="figure@17.[151,1436,183,1683]" captionTargetPageId="17" captionText="FIGURE 15. Ancyronyx minutulus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male sternite VIII (strioles omitted); e) male tergite VIII (setae and structures of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178548/files/figure.png" pageId="16" pageNumber="53">Fig. 15</figureCitation>
c) ca.
<quantity id="4CC28BF4FFAC2551FDBAF9CFE269F91B" box="[513,590,1650,1676]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="16" pageNumber="53" unit="m" value="275.0">275 m</quantity>
long; apical margin almost straight, very slightly emarginate, without setae; paraprocts not reaching apical margin.
</paragraph>
<paragraph id="8B852611FFAC2551FF7DF97FE19DF863" blockId="16.[151,1437,570,2036]" pageId="16" pageNumber="53">
Aedeagus (
<figureCitation id="13013A94FFAC2551FEF1F97FE1B8F94B" box="[330,415,1730,1756]" captionStart="FIGURE 15" captionStartId="17.[151,255,1743,1767]" captionTargetBox="[151,1436,183,1683]" captionTargetId="figure@17.[151,1436,183,1683]" captionTargetPageId="17" captionText="FIGURE 15. Ancyronyx minutulus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male sternite VIII (strioles omitted); e) male tergite VIII (setae and structures of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178548/files/figure.png" pageId="16" pageNumber="53">Fig. 15</figureCitation>
a, b) ca.
<quantity id="4CC28BF4FFAC2551FDB9F97FE26BF94B" box="[514,588,1730,1756]" metricMagnitude="2" metricUnit="m" metricValue="3.25" pageId="16" pageNumber="53" unit="m" value="325.0">325 m</quantity>
long. Median lobe rather long and slender (ca.
<quantity id="4CC28BF4FFAC2551FBC3F97FE493F94B" box="[1144,1204,1730,1756]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="16" pageNumber="53" unit="m" value="65.0">65 m</quantity>
broad), subapically evenly acuminate, slightly curved ventrad (lateral view), with numerous distinct microtube-like structures mainly ending lateroapically; ventral sac plicate, with moderately sclerotized, conspicuous fibula; ejaculatory duct distinctly speckled from level of parameral base to parameral apex; corona weekly developed. Phallobase asymmetrical, basally slightly curved ventrad (lateral view), with conspicuous, strongly sclerotized margin and short medial longitudinal ridge; basolateral (penile) apophyses rather long. Parameres elongately subtriangular (lateral view), reaching ca. basal 0.6 of aedeagus; basal margin sinuately emarginate (lateral view), not contiguous ventrally.
</paragraph>
<caption id="DF457699FFAD2550FF2CF972E2FBF8BB" httpUri="https://zenodo.org/record/178548/files/figure.png" pageId="17" pageNumber="54" targetBox="[151,1436,183,1683]" targetPageId="17">
<paragraph id="8B852611FFAD2550FF2CF972E2FBF8BB" blockId="17.[151,1436,1743,1836]" pageId="17" pageNumber="54">
<emphasis id="B94EFA03FFAD2550FF2CF972E100F970" bold="true" box="[151,295,1743,1767]" pageId="17" pageNumber="54">FIGURE 15.</emphasis>
<taxonomicName id="4C3A5D92FFAD2550FE95F96DE237F970" box="[302,528,1744,1767]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="minutulus" status="sp. nov.">
<emphasis id="B94EFA03FFAD2550FE95F96DE237F970" box="[302,528,1744,1767]" italics="true" pageId="17" pageNumber="54">Ancyronyx minutulus</emphasis>
</taxonomicName>
<emphasis id="B94EFA03FFAD2550FDACF972E273F970" bold="true" box="[535,596,1743,1767]" pageId="17" pageNumber="54">
<taxonomicNameLabel id="A27D4778FFAD2550FDACF972E273F970" box="[535,596,1743,1767]" pageId="17" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male sternite VIII (strioles omitted); e) male tergite VIII (setae and structures of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted).
</paragraph>
</caption>
<paragraph id="8B852611FFAD2550FF7DF8DDE33AF865" blockId="17.[151,1436,1888,2034]" pageId="17" pageNumber="54">
Secondary sexual characters. Male sternite VIII incomplete (
<figureCitation id="13013A94FFAD2550FC20F8DDE3D5F8ED" box="[923,1010,1888,1914]" captionStart="FIGURE 15" captionStartId="17.[151,255,1743,1767]" captionTargetBox="[151,1436,183,1683]" captionTargetId="figure@17.[151,1436,183,1683]" captionTargetPageId="17" captionText="FIGURE 15. Ancyronyx minutulus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male sternite VIII (strioles omitted); e) male tergite VIII (setae and structures of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178548/files/figure.png" pageId="17" pageNumber="54">Fig. 15</figureCitation>
d), weekly sclerotized; median strut short, apically rounded. Male tergite VIII (
<figureCitation id="13013A94FFAD2550FD31F835E2F9F835" box="[650,734,1928,1954]" captionStart="FIGURE 15" captionStartId="17.[151,255,1743,1767]" captionTargetBox="[151,1436,183,1683]" captionTargetId="figure@17.[151,1436,183,1683]" captionTargetPageId="17" captionText="FIGURE 15. Ancyronyx minutulus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male sternite VIII (strioles omitted); e) male tergite VIII (setae and structures of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178548/files/figure.png" pageId="17" pageNumber="54">Fig. 15</figureCitation>
e) slightly broader than long, condyles inconspicuous, basal half with microstrial pattern. Male ventrite 5 (
<figureCitation id="13013A94FFAD2550FD15F80DE326F85D" box="[686,769,1968,1994]" captionStart="FIGURE 15" captionStartId="17.[151,255,1743,1767]" captionTargetBox="[151,1436,183,1683]" captionTargetId="figure@17.[151,1436,183,1683]" captionTargetPageId="17" captionText="FIGURE 15. Ancyronyx minutulus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male sternite VIII (strioles omitted); e) male tergite VIII (setae and structures of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted)." httpUri="https://zenodo.org/record/178548/files/figure.png" pageId="17" pageNumber="54">Fig. 15</figureCitation>
f) subtrapezoidal, apically slightly microreticulate, medially rather glabrous, basal half with microstrial pattern.
</paragraph>
<paragraph id="8B852611FFAE2553FF7DFF25E1BEFF25" blockId="18.[151,1436,152,418]" box="[198,409,152,178]" pageId="18" pageNumber="55">Female unknown.</paragraph>
<paragraph id="8B852611FFAE2553FF7DFF02E22CFEBD" blockId="18.[151,1436,152,418]" pageId="18" pageNumber="55">
<emphasis id="B94EFA03FFAE2553FF7DFF02E1FEFF4E" bold="true" box="[198,473,191,217]" pageId="18" pageNumber="55">Differential diagnosis.</emphasis>
In this species the elytral colour patches are more elongate than in the other species described herein. Additionally,
<taxonomicName id="4C3A5D92FFAE2553FDFBFF55E2F2FE96" box="[576,725,232,258]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="55" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis id="B94EFA03FFAE2553FDFBFF55E275FE96" box="[576,594,232,257]" italics="true" pageId="18" pageNumber="55">A</emphasis>
.
<emphasis id="B94EFA03FFAE2553FDDAFF55E2F2FE96" box="[609,725,232,257]" italics="true" pageId="18" pageNumber="55">minutulus</emphasis>
</taxonomicName>
can easily be distinguished by its small size in combination with its entirely dark pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C320759AFFAE2553FF7DFE8AE35FFEC5" box="[198,888,311,338]" pageId="18" pageNumber="55" type="distribution">
<paragraph id="8B852611FFAE2553FF7DFE8AE35FFEC5" blockId="18.[151,1436,152,418]" box="[198,888,311,338]" pageId="18" pageNumber="55">
<emphasis id="B94EFA03FFAE2553FF7DFE8AE178FEC6" bold="true" box="[198,351,311,337]" pageId="18" pageNumber="55">Distribution</emphasis>
(Fig. 17). Only known from the
<typeStatus id="548198B3FFAE2553FD5BFE85E335FEC5" box="[736,786,312,338]" pageId="18" pageNumber="55">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C320759AFFAE2553FF7DFEE2E2BDFE35" pageId="18" pageNumber="55" type="etymology">
<paragraph id="8B852611FFAE2553FF7DFEE2E2BDFE35" blockId="18.[151,1436,152,418]" pageId="18" pageNumber="55">
<emphasis id="B94EFA03FFAE2553FF7DFEE2E174FEEE" bold="true" box="[198,339,351,377]" pageId="18" pageNumber="55">Etymology.</emphasis>
<taxonomicName id="4C3A5D92FFAE2553FEE6FEDDE1F2FEED" box="[349,469,352,378]" class="Insecta" family="Elmidae" genus="Ancyronyx" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="58" phylum="Arthropoda" rank="species" species="minutulus">Minutulus</taxonomicName>
(lat.) very small. Named in reference to the fact that this species is the smallest representative of the genus so far described.
</paragraph>
</subSubSection>
</treatment>
</document>