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<document id="BC1ECAAC9DB83267E12D8AFBA954131F" ID-DOI="10.21248/contrib.entomol.69.1.091-173" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742724" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638269330072" checkinUser="felipe" docAuthor="Assing, Volker &amp; Schülke, Michael" docDate="2019" docId="F078561B7269710BFC846232FBC1FE20" docLanguage="en" docName="ContribEntomol.69.1.91-173.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 69 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Bryaxis meghruicus BRACHAT 2019, spec. nov." docType="treatment" docUuid="AA6E7E2A-4450-48E8-8048-8061C3C6DF5C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="155" masterDocId="0C412E637256714BFFD26338F915FFC6" masterDocTitle="The Staphylinidae of Armenia and Nagorno-Karabakh (Coleoptera)" masterLastPageNumber="173" masterPageNumber="91" pageNumber="154" updateTime="1699437206044" updateUser="plazi">
<mods:mods id="958BB61180610FCFA790BEF77D5A5F5F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7B9B4DA87B6A6C350FEE1676CF26FF34">
<mods:title id="902A0DF293E4363513A97DD78E374FB4">The Staphylinidae of Armenia and Nagorno-Karabakh (Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name id="065E9BA1EFF5DE8AFD8D944C269A4687" type="personal">
<mods:role id="0FA1A38CC9AF50CD0A3F2D1BC047849E">
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<mods:namePart id="D64FFBAAF4FE7A81CCAE292A02381B80">Assing, Volker</mods:namePart>
</mods:name>
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<mods:namePart id="E096229923BEAFE602E9B315CA44866F">Schülke, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource id="9498AEAA7F43C0E9AA6083A6AFCEBF1C">text</mods:typeOfResource>
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<mods:title id="05ED7699BEF1EECF124D3707FCBA49CB">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="DFC26A7BF332B82D2B1308EF8B8030FF">
<mods:date id="2C585E8EA9D7241E0FF1C8C9107DF7E4">2019</mods:date>
<mods:detail id="C6390AE278D080FA553667DDDA51C630" type="pubDate">
<mods:number id="93CA908128637D76D7313CE183FD5CF7">2019-06-24</mods:number>
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<mods:detail id="506DAB7BEF17CECCF188C1423862F638" type="volume">
<mods:number id="E8F4BF62D7299E0D6DAE62384D5667F6">69</mods:number>
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<mods:number id="71E6B3C7437020118C283E29F428DE46">1</mods:number>
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<mods:classification id="CF03A912E49DC5F940E8504721BF28B9">journal article</mods:classification>
<mods:identifier id="B166D6A63E559654A9325C567DDE815B" type="DOI">10.21248/contrib.entomol.69.1.091-173</mods:identifier>
<mods:identifier id="315764821C0B00A9C3F9E003678818FE" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="0DF2FF35977CF6024E7E8BE9B706F420" type="Zenodo-Dep">5742724</mods:identifier>
<mods:identifier id="56AEEDD6722213D46A2C73BDF95F96D1" type="ZooBank">9B8BCFC0-C491-45EF-9BC8-04A6B6B7DB8C</mods:identifier>
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<treatment id="F078561B7269710BFC846232FBC1FE20" LSID="urn:lsid:zoobank.org:act:AA6E7E2A-4450-48E8-8048-8061C3C6DF5C" httpUri="http://treatment.plazi.org/id/F078561B7269710BFC846232FBC1FE20" lastPageId="64" lastPageNumber="155" pageId="63" pageNumber="154">
<subSubSection id="30CBB48672697174FC846232FC4BFE83" pageId="63" pageNumber="154" type="nomenclature">
<paragraph id="786EE70D72697174FC846232FDFAFEE0" blockId="63.[854,1374,265,357]" box="[854,1263,265,294]" pageId="63" pageNumber="154">
<heading id="2326506172697174FC846232FDFAFEE0" box="[854,1263,265,294]" fontSize="12" level="2" pageId="63" pageNumber="154" reason="2">
<taxonomicName id="BFD19C8E72697174FC846232FD95FEE3" ID-CoL="8XW3B" authority="BRACHAT" authorityName="BRACHAT" authorityYear="2019" box="[854,1152,265,294]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="154" phylum="Arthropoda" rank="species" species="meghruicus" status="sp. nov.">
<emphasis id="4AA53B1F72697174FC846232FD08FEE0" box="[854,1053,266,294]" italics="true" pageId="63" pageNumber="154">Bryaxis meghruicus</emphasis>
BRACHAT
</taxonomicName>
<taxonomicNameLabel id="5196866472697174FB546232FDFAFEE0" box="[1158,1263,266,294]" pageId="63" pageNumber="154" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="786EE70D72697174FC846209FC4BFE83" blockId="63.[854,1374,265,357]" box="[854,1374,305,325]" pageId="63" pageNumber="154">
<uri id="0C40EB0F72697174FC846209FC4BFE83" box="[854,1374,305,325]" pageId="63" pageNumber="154">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="30CBB48672697174FC846269FAA4FEA3" box="[854,945,337,357]" pageId="63" pageNumber="154" type="description">
<paragraph id="786EE70D72697174FC846269FAA4FEA3" blockId="63.[854,1374,265,357]" box="[854,945,337,357]" pageId="63" pageNumber="154">
(
<figureCitation id="E0EAFB8872697174FC886269FAB8FEA3" box="[858,941,337,357]" captionStart="Figs 7–23" captionStartId="57.[178,218,1988,2010]" captionTargetBox="[184,1416,256,1980]" captionTargetId="figure-27@57.[177,1417,254,1983]" captionTargetPageId="57" captionText="Figs 7–23:Omalium kociani (7–10), Proteinus baculatus (11–14), Bryaxis armeniacus (15, 17), B. meghruicus (16, 18), Aleochara fugax (19–21), and Atheta meghruica (22–23): (median lobe of) aedeagus in lateral and in ventral view (7–8, 13–16); female accessory sclerites (9); spermatheca (10, 21); habitus (11, 19, 22); thorax in ventral view (12); male antennomeres I–III (17–18); posterior margin of female sternite VIII (20); forebody (23). Scale bars: 11, 19, 22: 1.0 mm; 12, 23: 0.5 mm; 7–10, 13–16, 20–21: 0.1 mm; 17–18: without scale." figureDoi="http://doi.org/10.5281/zenodo.5742742" httpUri="https://zenodo.org/record/5742742/files/figure.png" pageId="63" pageNumber="154">Figs 16, 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="30CBB48672697174FCE262B8FC0DFDDE" pageId="63" pageNumber="154" type="materials_examined">
<paragraph id="786EE70D72697174FCE262B8FC81FE31" blockId="63.[816,1428,383,536]" pageId="63" pageNumber="154">
<emphasis id="4AA53B1F72697174FCE262B8FADCFE51" bold="true" box="[816,969,383,408]" pageId="63" pageNumber="154">Type material</emphasis>
:
<materialsCitation id="C8B9ED5072697174FC0462B8FC81FE31" collectingDate="2018-07-11" collectorName="V. Assing" country="Armenia" elevation="1780" latitude="39.244446" location="Kapan" longLatPrecision="19" longitude="46.106667" pageId="63" pageNumber="154" specimenCount="1" typeStatus="holotype">
<typeStatus id="A76A59AF72697174FC0462B8FD54FE5E" box="[982,1089,384,408]" pageId="63" pageNumber="154" type="holotype">Holotype</typeStatus>
[left antennomeres X and XI and right hind leg missing]: “
<collectingCountry id="00C6A79D72697174FBB76298FDC8FE7E" box="[1125,1245,416,440]" name="Armenia" pageId="63" pageNumber="154">ARMENIA</collectingCountry>
[29] – ca.
<quantity id="BF294AE872697174FA9B6298FC86FE7E" box="[1353,1427,416,440]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="154" unit="km" value="30.0">30 km</quantity>
W
<location id="7D0EB1D672697174FC8362F8FA8FFE11" LSID="urn:lsid:plazi:treatment:F078561B7269710BFC846232FBC1FE20:7D0EB1D672697174FC8362F8FA8FFE11" box="[849,922,448,471]" country="Armenia" latitude="39.244446" longLatPrecision="19" longitude="46.106667" name="Kapan" pageId="63" pageNumber="154">Kapan</location>
,
<geoCoordinate id="1DE581CA72697174FC7562F8FD37FE1E" box="[935,1058,448,472]" degrees="39" direction="north" minutes="14" orientation="latitude" pageId="63" pageNumber="154" precision="15" seconds="40" value="39.244446">39°14'40&quot;N</geoCoordinate>
,
<geoCoordinate id="1DE581CA72697174FBE262F8FDB2FE1E" box="[1072,1191,448,472]" degrees="46" direction="east" minutes="06" orientation="longitude" pageId="63" pageNumber="154" precision="15" seconds="24" value="46.106667">46°06'24&quot;E</geoCoordinate>
,
<quantity id="BF294AE872697174FB6662F8FC13FE11" box="[1204,1286,448,472]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="63" pageNumber="154" unit="m" value="1780.0">
<elevation id="F3FC003E72697174FB6662F8FC13FE11" box="[1204,1286,448,472]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="63" pageNumber="154" unit="m" value="1780.0">1780 m</elevation>
</quantity>
, river bank,
<date id="0C6FC1CD72697174FCE262D8FAA6FE3E" box="[816,947,480,504]" pageId="63" pageNumber="154" value="2018-07-11">
<collectingDate id="1C2B382572697174FCE262D8FAA6FE3E" box="[816,947,480,504]" pageId="63" pageNumber="154" value="2018-07-11">11.VII.2018</collectingDate>
</date>
,
<collectorName id="D52482DB72697174FC1062D8FD32FE3E" box="[962,1063,480,504]" pageId="63" pageNumber="154">V. Assing</collectorName>
/
<taxonomicName id="BFD19C8E72697174FB9662D8FC0FFE3E" authority="Assing &amp; Schülke, 2019" authorityName="Assing &amp; Schülke" authorityYear="2019" box="[1092,1306,480,504]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="154" phylum="Arthropoda" rank="species" species="meghruicus" status="sp. nov.">
<emphasis id="4AA53B1F72697174FB9662D8FC0FFE3E" box="[1092,1306,480,504]" italics="true" pageId="63" pageNumber="154">Bryaxis meghruicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5196866472697174FAF662D8FC81FE31" box="[1316,1428,480,503]" pageId="63" pageNumber="154" rank="species">spec. nov.</taxonomicNameLabel>
</materialsCitation>
</paragraph>
<paragraph id="786EE70D72697174FC976138FC0DFDDE" blockId="63.[816,1428,383,536]" box="[837,1304,512,536]" pageId="63" pageNumber="154">
, det. Brachat 10.2018 /
<typeStatus id="A76A59AF72697174FB976138FDA2FDDE" box="[1093,1207,512,536]" pageId="63" pageNumber="154" type="holotype">Holotypus</typeStatus>
” (cBra).
</paragraph>
</subSubSection>
<subSubSection id="30CBB48672697174FCE26178FD3CFD5E" pageId="63" pageNumber="154" type="etymology">
<paragraph id="786EE70D72697174FCE26178FD3CFD5E" blockId="63.[816,1427,576,664]" pageId="63" pageNumber="154">
<emphasis id="4AA53B1F72697174FCE26178FAB2FD9E" bold="true" box="[816,935,576,600]" pageId="63" pageNumber="154">Etymology</emphasis>
: The specific epithet is an adjective derived from Meghru, the name of the mountain range where the
<typeStatus id="A76A59AF72697174FCE261B8FA4BFD5E" box="[816,862,640,664]" pageId="63" pageNumber="154">type</typeStatus>
locality is situated.
</paragraph>
</subSubSection>
<subSubSection id="30CBB48672697174FCE26187FDC6F83E" pageId="63" pageNumber="154" type="description">
<paragraph id="786EE70D72697174FCE26187FC4CFCF1" blockId="63.[816,1428,703,1880]" pageId="63" pageNumber="154">
<emphasis id="4AA53B1F72697174FCE26187FAA0FD11" bold="true" box="[816,949,703,727]" pageId="63" pageNumber="154">Description</emphasis>
: Body length
<quantity id="BF294AE872697174FBB561F8FDA8FD11" box="[1127,1213,704,728]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="63" pageNumber="154" unit="mm" value="1.6">1.6 mm</quantity>
. Coloration: dark reddish-brown with the antennae, the maxillary palpi, and the legs slightly paler. Pubescence rather long and sub-erect; head with scattered long and erect setae.
</paragraph>
<paragraph id="786EE70D72697174FCE26078FA62FADE" blockId="63.[816,1428,703,1880]" pageId="63" pageNumber="154">
Head distinctly transverse,
<quantity id="BF294AE872697174FB8F6078FDA8FC91" box="[1117,1213,832,856]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="63" pageNumber="154" unit="mm" value="0.26">0.26 mm</quantity>
long and
<quantity id="BF294AE872697174FAFF6078FC86FC91" box="[1325,1427,832,856]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="63" pageNumber="154" unit="mm" value="0.34">0.34 mm</quantity>
broad, weakly punctate in anterior half; frontal lobe
<quantity id="BF294AE872697174FCE260B8FA85FC51" box="[816,912,896,920]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="63" pageNumber="154" unit="mm" value="0.17">0.17 mm</quantity>
broad; vertex with unmodified median keel. Eyes large and distinctly projecting, composed of approximately 25 ommatidia, slightly longer than the weakly convex postocular region in dorsal view. Antenna rather short; antennomeres III–XI
<quantity id="BF294AE872697174FAFF6738FC86FBD1" box="[1325,1427,1024,1048]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="63" pageNumber="154" unit="mm" value="0.46">0.46 mm</quantity>
long; antennomeres III weakly oblong, IV–VII globulous and approximately as long as broad, VIII weakly transverse, IX–XI forming a distinct club, IX and X distinctly transverse, and XI slightly broader than X and longer than combined length of VII–X. Maxillary palpi rather short; palpomeres II and III with few tubercles; palpomere IV petiolate,
<quantity id="BF294AE872697174FB9F67D8FDB8FB31" box="[1101,1197,1248,1272]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="63" pageNumber="154" unit="mm" value="0.22">0.22 mm</quantity>
long and
<quantity id="BF294AE872697174FAFF67D8FC86FB31" box="[1325,1427,1248,1272]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="63" pageNumber="154" unit="mm" value="0.09">0.09 mm</quantity>
broad.
</paragraph>
<paragraph id="786EE70D72697174FCE26618FDC8FA9E" blockId="63.[816,1428,703,1880]" pageId="63" pageNumber="154">
Pronotum
<quantity id="BF294AE872697174FC746618FD11FAF1" box="[934,1028,1312,1336]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="63" pageNumber="154" unit="mm" value="0.36">0.36 mm</quantity>
long and
<quantity id="BF294AE872697174FBA16618FDC4FAF1" box="[1139,1233,1312,1336]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="63" pageNumber="154" unit="mm" value="0.38">0.38 mm</quantity>
broad, convex in cross-section, broadest in anterior third.
</paragraph>
<paragraph id="786EE70D72697174FCE26658FA9AFA7E" blockId="63.[816,1428,703,1880]" pageId="63" pageNumber="154">
Elytra
<quantity id="BF294AE872697174FCAB6658FAC2FAB1" box="[889,983,1376,1400]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="63" pageNumber="154" unit="mm" value="0.6">0.60 mm</quantity>
long and
<quantity id="BF294AE872697174FB9A6658FDB3FAB1" box="[1096,1190,1376,1400]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="63" pageNumber="154" unit="mm" value="0.64">0.64 mm</quantity>
broad, with fine and posteriorly more distinct punctation; humeral angles marked.
</paragraph>
<paragraph id="786EE70D72697174FC9766F8FD64F89E" blockId="63.[816,1428,703,1880]" pageId="63" pageNumber="154">
: head ventrally with short transverse gular impression, anterior margin of this impression elevated; antenna: scapus
<quantity id="BF294AE872697174FC266538FD46F9D1" box="[1012,1107,1536,1560]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="63" pageNumber="154" unit="mm" value="0.15">0.15 mm</quantity>
long and
<quantity id="BF294AE872697174FB0B6538FC2CF9D1" box="[1241,1337,1536,1560]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="63" pageNumber="154" unit="mm" value="0.12">0.12 mm</quantity>
broad, dorsally slightly dilated, inner margin with a small flat tubercle (
<figureCitation id="E0EAFB8872697174FC496578FAFDF99E" box="[923,1000,1600,1624]" captionStart="Figs 7–23" captionStartId="57.[178,218,1988,2010]" captionTargetBox="[184,1416,256,1980]" captionTargetId="figure-27@57.[177,1417,254,1983]" captionTargetPageId="57" captionText="Figs 7–23:Omalium kociani (7–10), Proteinus baculatus (11–14), Bryaxis armeniacus (15, 17), B. meghruicus (16, 18), Aleochara fugax (19–21), and Atheta meghruica (22–23): (median lobe of) aedeagus in lateral and in ventral view (7–8, 13–16); female accessory sclerites (9); spermatheca (10, 21); habitus (11, 19, 22); thorax in ventral view (12); male antennomeres I–III (17–18); posterior margin of female sternite VIII (20); forebody (23). Scale bars: 11, 19, 22: 1.0 mm; 12, 23: 0.5 mm; 7–10, 13–16, 20–21: 0.1 mm; 17–18: without scale." figureDoi="http://doi.org/10.5281/zenodo.5742742" httpUri="https://zenodo.org/record/5742742/files/figure.png" pageId="63" pageNumber="154">Fig. 18</figureCitation>
); pedicel (
<figureCitation id="E0EAFB8872697174FBB36578FDB8F99E" box="[1121,1197,1600,1624]" captionStart="Figs 7–23" captionStartId="57.[178,218,1988,2010]" captionTargetBox="[184,1416,256,1980]" captionTargetId="figure-27@57.[177,1417,254,1983]" captionTargetPageId="57" captionText="Figs 7–23:Omalium kociani (7–10), Proteinus baculatus (11–14), Bryaxis armeniacus (15, 17), B. meghruicus (16, 18), Aleochara fugax (19–21), and Atheta meghruica (22–23): (median lobe of) aedeagus in lateral and in ventral view (7–8, 13–16); female accessory sclerites (9); spermatheca (10, 21); habitus (11, 19, 22); thorax in ventral view (12); male antennomeres I–III (17–18); posterior margin of female sternite VIII (20); forebody (23). Scale bars: 11, 19, 22: 1.0 mm; 12, 23: 0.5 mm; 7–10, 13–16, 20–21: 0.1 mm; 17–18: without scale." figureDoi="http://doi.org/10.5281/zenodo.5742742" httpUri="https://zenodo.org/record/5742742/files/figure.png" pageId="63" pageNumber="154">Fig. 18</figureCitation>
) oval and unmodified,
<quantity id="BF294AE872697174FCB76558FAC7F9B1" box="[869,978,1632,1656]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="63" pageNumber="154" unit="mm" value="0.06">0.060 mm</quantity>
long and
<quantity id="BF294AE872697174FBE86558FDB2F9B1" box="[1082,1191,1632,1656]" metricMagnitude="-5" metricUnit="m" metricValue="5.5" pageId="63" pageNumber="154" unit="mm" value="0.055">0.055 mm</quantity>
broad in dorsal view; metaventrite with median sulcus posteriorly; legs robust; pro- and metafemora dilated; profemur with depression near base; protibia dilated and with deep emargination in apical third; metatibia distinctly dilated, in apical half with projecting tooth, a deep emargination, and a pronounced spine at apical margin; aedeagus
<quantity id="BF294AE872697174FAFF6418FC98F8F1" box="[1325,1421,1824,1848]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="63" pageNumber="154" unit="mm" value="0.41">0.41 mm</quantity>
long and shaped as in
<figureCitation id="E0EAFB8872697174FBF36478FD7BF89E" box="[1057,1134,1856,1880]" captionStart="Figs 7–23" captionStartId="57.[178,218,1988,2010]" captionTargetBox="[184,1416,256,1980]" captionTargetId="figure-27@57.[177,1417,254,1983]" captionTargetPageId="57" captionText="Figs 7–23:Omalium kociani (7–10), Proteinus baculatus (11–14), Bryaxis armeniacus (15, 17), B. meghruicus (16, 18), Aleochara fugax (19–21), and Atheta meghruica (22–23): (median lobe of) aedeagus in lateral and in ventral view (7–8, 13–16); female accessory sclerites (9); spermatheca (10, 21); habitus (11, 19, 22); thorax in ventral view (12); male antennomeres I–III (17–18); posterior margin of female sternite VIII (20); forebody (23). Scale bars: 11, 19, 22: 1.0 mm; 12, 23: 0.5 mm; 7–10, 13–16, 20–21: 0.1 mm; 17–18: without scale." figureDoi="http://doi.org/10.5281/zenodo.5742742" httpUri="https://zenodo.org/record/5742742/files/figure.png" pageId="63" pageNumber="154">Fig. 16</figureCitation>
.
</paragraph>
<paragraph id="786EE70D72697174FCE26447FDC6F83E" blockId="63.[816,1427,1919,2040]" pageId="63" pageNumber="154">
<emphasis id="4AA53B1F72697174FCE26447FD11F85E" bold="true" box="[816,1028,1919,1944]" pageId="63" pageNumber="154">Comparative notes</emphasis>
:
<taxonomicName id="BFD19C8E72697174FBC664B8FDF0F85E" authorityName="Assing &amp; Schülke" authorityYear="2019" box="[1044,1253,1920,1944]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="154" phylum="Arthropoda" rank="species" species="meghruicus">
<emphasis id="4AA53B1F72697174FBC664B8FDF0F85E" box="[1044,1253,1920,1944]" italics="true" pageId="63" pageNumber="154">Bryaxis meghruicus</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="BFD19C8E72697174FCE26498FA89F87E" baseAuthorityName="MOTSCHULSKY" baseAuthorityYear="1851" box="[816,924,1952,1976]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="154" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis id="4AA53B1F72697174FCE26498FA89F87E" box="[816,924,1952,1976]" italics="true" pageId="63" pageNumber="154">B. clavipes</emphasis>
</taxonomicName>
group. It is distinguished from other species of this group by the internal structures of the aedeagus and by the combination of other characters.
</paragraph>
</subSubSection>
<subSubSection id="30CBB4867216710BFF0663C7FBC1FE20" pageId="64" pageNumber="155" type="distribution">
<paragraph id="786EE70D7216710BFF0663C7F8AEFEB1" blockId="64.[212,823,255,376]" pageId="64" pageNumber="155">
<emphasis id="4AA53B1F7216710BFF0663C7FB53FED1" bold="true" box="[212,582,255,280]" pageId="64" pageNumber="155">Distribution and natural history</emphasis>
: The type locality is situated some
<quantity id="BF294AE87216710BFEB56218F8BCFEFE" box="[359,425,288,312]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="64" pageNumber="155" unit="km" value="30.0">30 km</quantity>
to the west of Kapan, South
<collectingCountry id="00C6A79D7216710BFD196218FA26FEFE" box="[715,819,288,312]" name="Armenia" pageId="64" pageNumber="155">Armenia</collectingCountry>
. The
<typeStatus id="A76A59AF7216710BFED76278F876FE9E" box="[261,355,320,344]" pageId="64" pageNumber="155" type="holotype">holotype</typeStatus>
was floated from gravel on a river bank at an altitude of
<quantity id="BF294AE87216710BFEB46258F8A2FEB1" box="[358,439,352,376]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="64" pageNumber="155" unit="m" value="1780.0">1780 m</quantity>
.
</paragraph>
<paragraph id="786EE70D7216710BFF2B62F0FBC1FE20" blockId="64.[249,724,451,487]" box="[249,724,451,487]" pageId="64" pageNumber="155">
3.6.4
<taxonomicName id="BFD19C8E7216710BFEEA62F0F8C5FE20" authorityName="MacLeay" authorityYear="1825" box="[312,464,456,486]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="155" phylum="Arthropoda" rank="subFamily" subFamily="Tachyporinae">Tachyporinae</taxonomicName>
(by MICHAEL SCHÜLKE)
</paragraph>
</subSubSection>
</treatment>
</document>
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