291 lines
38 KiB
XML
291 lines
38 KiB
XML
<document id="C4CDA43F2B130D16FD78B66BC7A5B01C" ID-CLB-Dataset="7784" ID-DOI="10.11646/zootaxa.4933.4.1" ID-GBIF-Dataset="2ffdac24-b2ae-448c-b88e-b4f0dd1a9547" ID-ISSN="1175-5326" ID-Zenodo-Dep="4556546" ID-ZooBank="1FC0D2E0-888E-4F79-ABFE-BC7E91ADEECE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614066053014" checkinUser="plazi" docAuthor="Wood, Thomas James" docDate="2021" docId="E52A87BCFF86FF8BFF79FA6889496F24" docLanguage="en" docName="zootaxa.4933.4.1.pdf" docOrigin="Zootaxa 4933 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andrena (Graecandrena) virguladivina Wood 2021, spec. nov." docType="treatment" docUuid="5124F4A3-EA43-40E9-A8FE-E0180CEF4CF1" docUuidSource="ZooBank" docVersion="6" lastPageNumber="469" masterDocId="1913FFC4FF89FF99FFEEFFE18A626C79" masterDocTitle="Fifteen new Andrena species from little-visited arid, Mediterranean, and mountainous parts of the Old World (Hymenoptera: Andrenidae)" masterLastPageNumber="492" masterPageNumber="451" pageNumber="466" updateTime="1698930763140" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="EAFBD477834C707766C1418F5D1D1B3D">
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<mods:title id="E901BC1997D8AC14855DA227289BDB18">Fifteen new Andrena species from little-visited arid, Mediterranean, and mountainous parts of the Old World (Hymenoptera: Andrenidae)</mods:title>
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<mods:name id="2802328E1C95FE44FF4DA94DB094FBF3" type="personal">
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<mods:roleTerm id="0A9242782E2398F7B6394011363CBC85">Author</mods:roleTerm>
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<mods:namePart id="802706575A1B3B85B4563B32E551B28C">Wood, Thomas James</mods:namePart>
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<mods:affiliation id="EFC888FCBAE223B0A6171DDA03F32DFE">Laboratoire de Zoologie, Université de Mons, Mons, 7000, Belgium</mods:affiliation>
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<mods:typeOfResource id="955F369FFBF58C496A1EDE812E8B6978">text</mods:typeOfResource>
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<mods:title id="6F223FCFFD450CA39345B3C3006121CB">Zootaxa</mods:title>
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<mods:part id="E535F73B8B0CAFCEEE212193C5DA49A3">
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<mods:date id="05DD5263FC8D9F98E4A63E6464B386C6">2021</mods:date>
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<mods:number id="62CC5AFFD230DCDAE2E80612FCA5DAB6">2021-02-23</mods:number>
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<mods:number id="2E8949CED68C3A7874E83A62C306EB7D">4933</mods:number>
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<mods:identifier id="1DDCAF90CBAA7A2A12C0FFF1CE9305D9" type="DOI">10.11646/zootaxa.4933.4.1</mods:identifier>
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<treatment id="E52A87BCFF86FF8BFF79FA6889496F24" ID-DOI="http://doi.org/10.5281/zenodo.4556556" ID-GBIF-Taxon="180326871" ID-Zenodo-Dep="4556556" LSID="urn:lsid:zoobank.org:act:5124F4A3-EA43-40E9-A8FE-E0180CEF4CF1" httpUri="http://treatment.plazi.org/id/E52A87BCFF86FF8BFF79FA6889496F24" lastPageId="18" lastPageNumber="469" pageId="15" pageNumber="466">
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<subSubSection id="25996521FF86FF96FF79FA6889FD69BD" pageId="15" pageNumber="466" type="nomenclature">
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<paragraph id="6D3C36AAFF86FF96FF79FA68889069DA" blockId="15.[151,927,1417,1476]" box="[151,754,1417,1444]" pageId="15" pageNumber="466">
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<heading id="367481C6FF86FF96FF79FA68889069DA" bold="true" box="[151,754,1417,1444]" fontSize="11" level="1" pageId="15" pageNumber="466" reason="1">
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<emphasis id="5FF7EAB8FF86FF96FF79FA68889069DA" bold="true" box="[151,754,1417,1444]" pageId="15" pageNumber="466">
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<taxonomicName id="AA834D29FF86FF96FF79FA68881169DD" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[151,627,1417,1444]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="virguladivina" status="sp. nov." subGenus="Graecandrena">
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<emphasis id="5FF7EAB8FF86FF96FF79FA688B6369DD" bold="true" box="[151,257,1417,1444]" italics="true" pageId="15" pageNumber="466">Andrena</emphasis>
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||
(
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<emphasis id="5FF7EAB8FF86FF96FEFCFA688BA269DD" bold="true" box="[274,448,1417,1444]" italics="true" pageId="15" pageNumber="466">Graecandrena</emphasis>
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||
)
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<emphasis id="5FF7EAB8FF86FF96FE3CFA68881169DD" bold="true" box="[466,627,1417,1444]" italics="true" pageId="15" pageNumber="466">virguladivina</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="44C457C3FF86FF96FD94FA68889069DA" box="[634,754,1417,1443]" pageId="15" pageNumber="466" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6D3C36AAFF86FF96FF79FA4D89FD69BD" blockId="15.[151,927,1417,1476]" box="[151,927,1452,1476]" pageId="15" pageNumber="466">
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||
<uri id="19123AA8FF86FF96FF79FA4D89FD69BD" box="[151,927,1452,1476]" pageId="15" pageNumber="466">
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||
urn:lsid:zoobank.org:act:
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||
<uuid id="19250C7FFF86FF96FE75FA4D89FD69BD" box="[411,927,1452,1476]" pageId="15" pageNumber="466">5124F4A3-EA43-40E9-A8FE-E0180CEF4CF1</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="25996521FF86FF96FF79FA0C8E336A50" pageId="15" pageNumber="466" type="materials_examined">
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||
<paragraph id="6D3C36AAFF86FF96FF79FA0C8F4B6A73" blockId="15.[151,1437,1517,2010]" box="[151,1321,1517,1546]" pageId="15" pageNumber="466">
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||
<materialsCitation id="DDEB3CF7FF86FF96FF79FA0C8F4B6A73" ID-GBIF-Occurrence="3050682370" box="[151,1321,1517,1546]" collectingDate="1996-05-26" collectionCode="OOLM" collectorName="Marek Halada" country="Syria" location="Jisr" municipality="Shughur" pageId="15" pageNumber="466" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="5FF7EAB8FF86FF96FF79FA0C8BC56A7E" bold="true" box="[151,423,1517,1543]" pageId="15" pageNumber="466">
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||
<typeStatus id="B2388808FF86FF96FF79FA0C8B586A7E" box="[151,314,1517,1543]" pageId="15" pageNumber="466" type="holotype">HOLOTYPE</typeStatus>
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||
:
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||
<collectingCountry id="1594763AFF86FF96FEA8FA0C8BC36A7E" box="[326,417,1517,1543]" name="Syria" pageId="15" pageNumber="466">SYRIA</collectingCountry>
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||
:
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||
</emphasis>
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||
<location id="685C6071FF86FF96FE40FA0C8BB76A7E" LSID="urn:lsid:plazi:treatment:E52A87BCFF86FF8BFF79FA6889496F24:685C6071FF86FF96FE40FA0C8BB76A7E" box="[430,469,1517,1543]" country="Syria" municipality="Shughur" name="Jisr" pageId="15" pageNumber="466">Jisr</location>
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ash
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<collectingMunicipality id="8D58ACD0FF86FF96FDE6FA0C88056A7E" box="[520,615,1517,1543]" pageId="15" pageNumber="466">Shughur</collectingMunicipality>
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||
[Jisr al-Shughur],
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||
<date id="193D106AFF86FF96FCD7FA0C89CB6A7E" box="[825,937,1517,1543]" pageId="15" pageNumber="466" value="1996-05-26">
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||
<collectingDate id="0979E982FF86FF96FCD7FA0C89CB6A7E" box="[825,937,1517,1543]" pageId="15" pageNumber="466" value="1996-05-26">26.v.1996</collectingDate>
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</date>
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, 1³, leg.
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<collectorName id="C076537CFF86FF96FBF8FA0C8ED76A7E" box="[1046,1205,1517,1543]" pageId="15" pageNumber="466">Marek Halada</collectorName>
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||
(
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||
<collectionCode id="0B92AE6FFF86FF96FB2AFA0E8F7F6A73" box="[1220,1309,1519,1546]" pageId="15" pageNumber="466">OÖLM</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="6D3C36AAFF86FF96FF29F9EF8E336A50" blockId="15.[151,1437,1517,2010]" box="[199,1105,1550,1579]" pageId="15" pageNumber="466">
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<materialsCitation id="DDEB3CF7FF86FF96FF29F9EF8E2F6A50" ID-GBIF-Occurrence="3050682362" box="[199,1101,1550,1579]" collectionCode="OOLM, SMNHTAU" country="Syria" location="Same" pageId="15" pageNumber="466" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
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||
<emphasis id="5FF7EAB8FF86FF96FF29F9EF8BBE6A51" bold="true" box="[199,476,1550,1576]" pageId="15" pageNumber="466">
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||
<typeStatus id="B2388808FF86FF96FF29F9EF8B0D6A51" box="[199,367,1550,1576]" pageId="15" pageNumber="466" type="paratype">PARATYPES</typeStatus>
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||
:
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||
<collectingCountry id="1594763AFF86FF96FE95F9EF8BB46A51" box="[379,470,1550,1576]" name="Syria" pageId="15" pageNumber="466">SYRIA</collectingCountry>
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||
:
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||
</emphasis>
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||
<location id="685C6071FF86FF96FE0DF9EE88436A50" LSID="urn:lsid:plazi:treatment:E52A87BCFF86FF8BFF79FA6889496F24:685C6071FF86FF96FE0DF9EE88436A50" box="[483,545,1551,1577]" country="Syria" name="Same" pageId="15" pageNumber="466">Same</location>
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as
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||
<typeStatus id="B2388808FF86FF96FDA8F9EE88C96A50" box="[582,683,1551,1577]" pageId="15" pageNumber="466" type="holotype">holotype</typeStatus>
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||
, 1³,
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<specimenCount id="7B85FD23FF86FF96FD09F9EE89686A50" box="[743,778,1551,1577]" pageId="15" pageNumber="466" type="female">2♀</specimenCount>
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||
(
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||
<collectionCode id="0B92AE6FFF86FF96FCF7F9F189136A52" box="[793,881,1552,1579]" pageId="15" pageNumber="466">OÖLM</collectionCode>
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||
),
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||
<specimenCount id="7B85FD23FF86FF96FC6AF9EE89C56A50" box="[900,935,1551,1577]" pageId="15" pageNumber="466" type="female">1♀</specimenCount>
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||
(
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||
<collectionCode id="0B92AE6FFF86FF96FC58F9EE8E276A50" box="[950,1093,1551,1577]" pageId="15" pageNumber="466">SMNHTAU</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="25996521FF86FF88FF29F9D18FEC6DAB" lastPageId="17" lastPageNumber="468" pageId="15" pageNumber="466" type="description">
|
||
<paragraph id="6D3C36AAFF86FF89FF29F9D188366DE4" blockId="15.[151,1437,1517,2010]" lastBlockId="16.[151,1437,150,413]" lastPageId="16" lastPageNumber="467" pageId="15" pageNumber="466">
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<emphasis id="5FF7EAB8FF86FF96FF29F9D18BA36A33" bold="true" box="[199,449,1584,1610]" pageId="15" pageNumber="466">Description: Female:</emphasis>
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Body length
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<quantity id="AA7B9B4FFF86FF96FDB8F9D088F46A32" box="[598,662,1585,1611]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="466" unit="mm" value="7.0">7 mm</quantity>
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||
(
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<figureCitation id="F5B82A2FFF86FF96FD4DF9D189716A32" box="[675,787,1584,1611]" captionStart="FIGURES 51–56" captionStartId="16.[151,264,1907,1931]" captionTargetBox="[151,1436,437,1883]" captionTargetId="figure-140@16.[151,1436,437,1883]" captionTargetPageId="16" captionText="FIGURES 51–56. Andrena virguladivina spec. nov. 51. female profile, 52. female face, 53. female scutum, 54. female propodeum, 55. female terga, 56. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556572" httpUri="https://zenodo.org/record/4556572/files/figure.png" pageId="15" pageNumber="466">Figure 51</figureCitation>
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).
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<emphasis id="5FF7EAB8FF86FF96FCC8F9D1890E6A33" box="[806,876,1584,1610]" italics="true" pageId="15" pageNumber="466">Head:</emphasis>
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Black, 1.2 times wider than long. Clypeus domed, centrally broadly flattened, strongly punctate, punctures separated by 0.5–1 puncture diameter basally, becoming more densely punctate apically, strength of punctures increasing; underlying surface deformed, irregular, uneven (
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<figureCitation id="F5B82A2FFF86FF96FF71F9758B706AD6" box="[159,274,1684,1711]" captionStart="FIGURES 51–56" captionStartId="16.[151,264,1907,1931]" captionTargetBox="[151,1436,437,1883]" captionTargetId="figure-140@16.[151,1436,437,1883]" captionTargetPageId="16" captionText="FIGURES 51–56. Andrena virguladivina spec. nov. 51. female profile, 52. female face, 53. female scutum, 54. female propodeum, 55. female terga, 56. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556572" httpUri="https://zenodo.org/record/4556572/files/figure.png" pageId="15" pageNumber="466">Figure 52</figureCitation>
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). Clypeus in basal half with central, longitudinal, raised impunctate line, apically disappearing; underlying surface strongly shagreened in basal 2/3rds, dull, becoming weaker in apical 1/3
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<superScript id="9AF69BE2FF86FF96FB80F9548E1E6ABA" attach="right" box="[1134,1148,1717,1731]" fontSize="6" pageId="15" pageNumber="466">rd</superScript>
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, weakly shining. Process of labrum narrowly trapezoidal, apical margin straight, with slight raised shiny ridge. Gena equalling width of compound eye; ocelloccipital distance broad, 1.5 times width of lateral ocellus. Fovea narrow, subequal to width of flagellum, extending ventrally to level of antennal insertion, here separated from inner margin of compound eye by distance subequal to own width. Gena, face, and scape with short white hairs, vertex with longer brown hairs, not exceeding length of scape. Antenna dark, A5–12 lightened orange below, A3 exceeding A4, shorter than A4+5.
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<taxonomicName id="AA834D29FF86FF96FF79F89F8B1A6BEE" authority=": Scutum" authorityName="Scutum" box="[151,376,1917,1943]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="466" phylum="Arthropoda" rank="genus">
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<emphasis id="5FF7EAB8FF86FF96FF79F89F8B7A6BEE" box="[151,280,1918,1943]" italics="true" pageId="15" pageNumber="466">Mesosoma:</emphasis>
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Scutum
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</taxonomicName>
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and scutellum regularly and evenly punctate, punctures separated by 2 puncture diameters, underlying surface shagreened, weakly shining (
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<figureCitation id="F5B82A2FFF86FF96FD5FF87E89436BC0" box="[689,801,1950,1977]" captionStart="FIGURES 51–56" captionStartId="16.[151,264,1907,1931]" captionTargetBox="[151,1436,437,1883]" captionTargetId="figure-140@16.[151,1436,437,1883]" captionTargetPageId="16" captionText="FIGURES 51–56. Andrena virguladivina spec. nov. 51. female profile, 52. female face, 53. female scutum, 54. female propodeum, 55. female terga, 56. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556572" httpUri="https://zenodo.org/record/4556572/files/figure.png" pageId="15" pageNumber="466">Figure 53</figureCitation>
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). Pronotum rounded, without humeral angle. Episternum and propodeum finely microreticulate, weakly shining, propodeal triangle clearly marked by change in surface sculpture, finely shagreened with weak basal rugosity (
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<figureCitation id="F5B82A2FFF99FF89FCFCFF7689E46CC8" box="[786,902,150,177]" captionStart="FIGURES 51–56" captionStartId="16.[151,264,1907,1931]" captionTargetBox="[151,1436,437,1883]" captionTargetId="figure-140@16.[151,1436,437,1883]" captionTargetPageId="16" captionText="FIGURES 51–56. Andrena virguladivina spec. nov. 51. female profile, 52. female face, 53. female scutum, 54. female propodeum, 55. female terga, 56. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556572" httpUri="https://zenodo.org/record/4556572/files/figure.png" pageId="16" pageNumber="467">Figure 54</figureCitation>
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). Episternum and propodeum with moderately long white hairs, scutum and scutellum with short brown to light brown hairs. Legs dark, apical tarsal segments and hind basitarsi lightened brown to dark orange, pubescence whitish to light brown, scopa white. Hind tarsal claws without inner teeth. Wings hyaline, venation dark brown, stigma dark orange, nervulus interstitial.
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<emphasis id="5FF7EAB8FF99FF89FB34FF1A8F3A6D6D" box="[1242,1368,251,276]" italics="true" pageId="16" pageNumber="467">Metasoma:</emphasis>
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Terga dark, apical margins slightly depressed, occupying 2/5ths of segment, faintly lightened dark brown (
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<figureCitation id="F5B82A2FFF99FF89FAF5FEFD8FF26D4E" box="[1307,1424,284,311]" captionStart="FIGURES 51–56" captionStartId="16.[151,264,1907,1931]" captionTargetBox="[151,1436,437,1883]" captionTargetId="figure-140@16.[151,1436,437,1883]" captionTargetPageId="16" captionText="FIGURES 51–56. Andrena virguladivina spec. nov. 51. female profile, 52. female face, 53. female scutum, 54. female propodeum, 55. female terga, 56. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556572" httpUri="https://zenodo.org/record/4556572/files/figure.png" pageId="16" pageNumber="467">Figure 55</figureCitation>
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). Tergal discs finely microreticulate, essentially impunctate, weakly shining; T2–3 with very weak hair fringes laterally (
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<figureCitation id="F5B82A2FFF99FF89FF3EFEBE8B236D00" box="[208,321,350,377]" captionStart="FIGURES 51–56" captionStartId="16.[151,264,1907,1931]" captionTargetBox="[151,1436,437,1883]" captionTargetId="figure-140@16.[151,1436,437,1883]" captionTargetPageId="16" captionText="FIGURES 51–56. Andrena virguladivina spec. nov. 51. female profile, 52. female face, 53. female scutum, 54. female propodeum, 55. female terga, 56. female terga detail." figureDoi="http://doi.org/10.5281/zenodo.4556572" httpUri="https://zenodo.org/record/4556572/files/figure.png" pageId="16" pageNumber="467">Figure 56</figureCitation>
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). Apical fringe of T5 and hairs flanking pygidial plate light to dark brown; pygidial plate triangular, apically weakly pointed, flattened, dull.
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</paragraph>
|
||
<caption id="39FC6622FF99FF89FF79F89288146BD6" ID-DOI="http://doi.org/10.5281/zenodo.4556572" ID-Zenodo-Dep="4556572" httpUri="https://zenodo.org/record/4556572/files/figure.png" pageId="16" pageNumber="467" startId="16.[151,264,1907,1931]" targetBox="[151,1436,437,1883]" targetPageId="16">
|
||
<paragraph id="6D3C36AAFF99FF89FF79F89288146BD6" blockId="16.[151,1437,1907,1968]" pageId="16" pageNumber="467">
|
||
<emphasis id="5FF7EAB8FF99FF89FF79F8928B366BF2" bold="true" box="[151,340,1907,1931]" pageId="16" pageNumber="467">FIGURES 51–56.</emphasis>
|
||
<taxonomicName id="AA834D29FF99FF89FEB5F89288226BF2" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[347,576,1907,1931]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="virguladivina" status="sp. nov.">
|
||
<emphasis id="5FF7EAB8FF99FF89FEB5F89288226BF2" box="[347,576,1907,1931]" italics="true" pageId="16" pageNumber="467">Andrena virguladivina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5FF7EAB8FF99FF89FDA8F89588CF6BF5" bold="true" box="[582,685,1908,1932]" pageId="16" pageNumber="467">
|
||
<taxonomicNameLabel id="44C457C3FF99FF89FDA8F89588CF6BF5" box="[582,685,1908,1932]" pageId="16" pageNumber="467" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
51. female profile, 52. female face, 53. female scutum, 54. female propodeum, 55. female terga, 56. female terga detail.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6D3C36AAFF98FF88FF29FF768FEC6DAB" blockId="17.[151,1437,150,466]" pageId="17" pageNumber="468">
|
||
<emphasis id="5FF7EAB8FF98FF88FF29FF768B6F6CC8" bold="true" box="[199,269,151,177]" pageId="17" pageNumber="468">Male:</emphasis>
|
||
Body length
|
||
<quantity id="AA7B9B4FFF98FF88FE42FF768B8D6CC8" box="[428,495,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="468" unit="mm" value="6.0">6 mm</quantity>
|
||
(
|
||
<figureCitation id="F5B82A2FFF98FF88FDEEFF7688166CC8" box="[512,628,150,177]" captionStart="FIGURES 57–62" captionStartId="17.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-162@17.[151,1436,489,1935]" captionTargetPageId="17" captionText="FIGURES 57–62. Andrena virguladivina spec. nov. 57. male profile, 58. male face, 59. male terga, 60. male genitalia dorsal view, 61. male genitalia profile, 62. male genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.4556574" httpUri="https://zenodo.org/record/4556574/files/figure.png" pageId="17" pageNumber="468">Figure 57</figureCitation>
|
||
).
|
||
<emphasis id="5FF7EAB8FF98FF88FD64FF7688B26CC8" box="[650,720,151,177]" italics="true" pageId="17" pageNumber="468">Head:</emphasis>
|
||
Black, 1.2 times wider than long (
|
||
<figureCitation id="F5B82A2FFF98FF88FB8CFF768EB56CC8" box="[1122,1239,150,177]" captionStart="FIGURES 57–62" captionStartId="17.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-162@17.[151,1436,489,1935]" captionTargetPageId="17" captionText="FIGURES 57–62. Andrena virguladivina spec. nov. 57. male profile, 58. male face, 59. male terga, 60. male genitalia dorsal view, 61. male genitalia profile, 62. male genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.4556574" httpUri="https://zenodo.org/record/4556574/files/figure.png" pageId="17" pageNumber="468">Figure 58</figureCitation>
|
||
). Clypeus evenly domed, regularly punctate, punctures separated by 1 puncture diameter, underlying surface shiny. Process of labrum narrowly rounded trapezoidal. Gena subequal to width of compound eye; ocelloccipital distance broad, 1.5 times width of lateral ocellus. Gena, face, and scape with long white hairs, vertex with long light brown hairs, exceeding length of scape. Antenna dark, A4–13 brown below, A3 equals A4.
|
||
<taxonomicName id="AA834D29FF98FF88FC73FEC68EC96D38" authority=": Structurally" authorityName="Structurally" box="[925,1195,295,321]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="468" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FF7EAB8FF98FF88FC73FEC68E7C6D39" box="[925,1054,295,320]" italics="true" pageId="17" pageNumber="468">Mesosoma:</emphasis>
|
||
Structurally
|
||
</taxonomicName>
|
||
as in female, though propodeal triangle with more extensive basal rugosity, all basitarsi more clearly lightened orange.
|
||
<emphasis id="5FF7EAB8FF98FF88FB35FEAA8F356D1D" box="[1243,1367,331,356]" italics="true" pageId="17" pageNumber="468">Metasoma:</emphasis>
|
||
Terga structurally as in female (
|
||
<figureCitation id="F5B82A2FFF98FF88FE5FFE8E88436DF0" box="[433,545,366,393]" captionStart="FIGURES 57–62" captionStartId="17.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-162@17.[151,1436,489,1935]" captionTargetPageId="17" captionText="FIGURES 57–62. Andrena virguladivina spec. nov. 57. male profile, 58. male face, 59. male terga, 60. male genitalia dorsal view, 61. male genitalia profile, 62. male genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.4556574" httpUri="https://zenodo.org/record/4556574/files/figure.png" pageId="17" pageNumber="468">Figure 59</figureCitation>
|
||
). Genitalia rounded (
|
||
<figureCitation id="F5B82A2FFF98FF88FCE4FE8E891B6DF3" box="[778,889,367,394]" captionStart="FIGURES 57–62" captionStartId="17.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-162@17.[151,1436,489,1935]" captionTargetPageId="17" captionText="FIGURES 57–62. Andrena virguladivina spec. nov. 57. male profile, 58. male face, 59. male terga, 60. male genitalia dorsal view, 61. male genitalia profile, 62. male genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.4556574" httpUri="https://zenodo.org/record/4556574/files/figure.png" pageId="17" pageNumber="468">Figure 60</figureCitation>
|
||
), in profile compact (
|
||
<figureCitation id="F5B82A2FFF98FF88FB86FE8E8EB56DF3" box="[1128,1239,367,394]" captionStart="FIGURES 57–62" captionStartId="17.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-162@17.[151,1436,489,1935]" captionTargetPageId="17" captionText="FIGURES 57–62. Andrena virguladivina spec. nov. 57. male profile, 58. male face, 59. male terga, 60. male genitalia dorsal view, 61. male genitalia profile, 62. male genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.4556574" httpUri="https://zenodo.org/record/4556574/files/figure.png" pageId="17" pageNumber="468">Figure 61</figureCitation>
|
||
), gonocoxae with small, weakly produced apical teeth, gonostyli evenly arched, apically truncate, inner margin slightly thickened; penis valve with base narrowly triangular in dorsal view, extending ventrally, long, apically strongly bifurcate (
|
||
<figureCitation id="F5B82A2FFF98FF88FAFCFE568FE36DAB" box="[1298,1409,439,466]" captionStart="FIGURES 57–62" captionStartId="17.[151,264,1958,1982]" captionTargetBox="[151,1436,489,1935]" captionTargetId="figure-162@17.[151,1436,489,1935]" captionTargetPageId="17" captionText="FIGURES 57–62. Andrena virguladivina spec. nov. 57. male profile, 58. male face, 59. male terga, 60. male genitalia dorsal view, 61. male genitalia profile, 62. male genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.4556574" httpUri="https://zenodo.org/record/4556574/files/figure.png" pageId="17" pageNumber="468">Figure 62</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="39FC6622FF98FF88FF79F847897F6B9B" ID-DOI="http://doi.org/10.5281/zenodo.4556574" ID-Zenodo-Dep="4556574" httpUri="https://zenodo.org/record/4556574/files/figure.png" pageId="17" pageNumber="468" startId="17.[151,264,1958,1982]" targetBox="[151,1436,489,1935]" targetPageId="17">
|
||
<paragraph id="6D3C36AAFF98FF88FF79F847897F6B9B" blockId="17.[151,1436,1958,2018]" pageId="17" pageNumber="468">
|
||
<emphasis id="5FF7EAB8FF98FF88FF79F8478B376BC7" bold="true" box="[151,341,1958,1982]" pageId="17" pageNumber="468">FIGURES 57–62.</emphasis>
|
||
<taxonomicName id="AA834D29FF98FF88FEB2F84788206BC7" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[348,578,1958,1982]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="virguladivina" status="sp. nov.">
|
||
<emphasis id="5FF7EAB8FF98FF88FEB2F84788206BC7" box="[348,578,1958,1982]" italics="true" pageId="17" pageNumber="468">Andrena virguladivina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5FF7EAB8FF98FF88FDA7F84688D36BC6" bold="true" box="[585,689,1959,1983]" pageId="17" pageNumber="468">
|
||
<taxonomicNameLabel id="44C457C3FF98FF88FDA7F84688D36BC6" box="[585,689,1959,1983]" pageId="17" pageNumber="468" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
57. male profile, 58. male face, 59. male terga, 60. male genitalia dorsal view, 61. male genitalia profile, 62. male genitalia ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="25996521FF9BFF8BFF29FF778F0C6F6C" pageId="18" pageNumber="469" type="diagnosis">
|
||
<paragraph id="6D3C36AAFF9BFF8BFF29FF778A896E18" blockId="18.[151,1437,150,861]" pageId="18" pageNumber="469">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFF29FF778B266CC9" bold="true" box="[199,324,150,176]" pageId="18" pageNumber="469">Diagnosis:</emphasis>
|
||
Subgeneric placement of
|
||
<taxonomicName id="AA834D29FF9BFF8BFD9CFF7689486CC8" authorityName="Wood" authorityYear="2021" box="[626,810,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="virguladivina">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFD9CFF7689486CC8" box="[626,810,151,177]" italics="true" pageId="18" pageNumber="469">A. virguladivina</emphasis>
|
||
</taxonomicName>
|
||
faces the same challenges as the previously described
|
||
<taxonomicName id="AA834D29FF9BFF8BFF79FF5A8B7B6CAC" box="[151,281,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="456" phylum="Arthropoda" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFF79FF5A8B7B6CAC" box="[151,281,187,213]" italics="true" pageId="18" pageNumber="469">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
species because of deep problems in subgeneric classification for small black Old World
|
||
<taxonomicName id="AA834D29FF9BFF8BFAD3FF5A8FFE6CAC" box="[1341,1436,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFAD3FF5A8FFE6CAC" box="[1341,1436,187,213]" italics="true" pageId="18" pageNumber="469">Andrena</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation id="09124B5BFF9BFF8BFF16FF3E8BB66C80" author="Pisanty, G. & Richter, R. & Martin, T. & Dettman, J. & Cardinal, S." box="[248,468,223,249]" pageId="18" pageNumber="469" refId="ref23835" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. & Cardinal, S. (2020) Molecular phylogeny and historical biogeography of an- drenine bees (Hymenoptera: Andrenidae). bioRxiv. [published online] https: // doi. org / 10.1101 / 2020.06.09.103556" type="book" year="2020">
|
||
Pisanty
|
||
<emphasis id="5FF7EAB8FF9BFF8BFEBBFF3E8BEA6C80" box="[341,392,223,249]" italics="true" pageId="18" pageNumber="469">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
). In the female sex, the combination of small body size, finely shagreened propodeal triangle (though with basal rugosity more typically found in
|
||
<emphasis id="5FF7EAB8FF9BFF8BFCDDFEE289B76D64" box="[819,981,259,285]" italics="true" pageId="18" pageNumber="469">Graecandrena</emphasis>
|
||
), and narrow facial foveae could suggest
|
||
<taxonomicName id="AA834D29FF9BFF8BFF79FEC68B7B6D38" box="[151,281,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="456" phylum="Arthropoda" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFF79FEC68B7B6D38" box="[151,281,295,321]" italics="true" pageId="18" pageNumber="469">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
, but the structure of the male genitalia combined with a black clypeus (typically yellow in
|
||
<taxonomicName id="AA834D29FF9BFF8BFAF5FEC68FFF6D38" box="[1307,1437,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="456" phylum="Arthropoda" rank="subGenus" subGenus="Aciandrena">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFAF5FEC68FFF6D38" box="[1307,1437,295,321]" italics="true" pageId="18" pageNumber="469">Aciandrena</emphasis>
|
||
</taxonomicName>
|
||
but this can be intra- and interspecifically variable) suggests a much stronger affinity with the true
|
||
<emphasis id="5FF7EAB8FF9BFF8BFB14FEAA8FFE6D1C" box="[1274,1436,331,357]" italics="true" pageId="18" pageNumber="469">Graecandrena</emphasis>
|
||
(species around
|
||
<taxonomicName id="AA834D29FF9BFF8BFEA3FE8E88EB6DF0" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[333,649,366,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="graecella">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFEA3FE8E8BB56DF0" box="[333,471,367,393]" italics="true" pageId="18" pageNumber="469">A. graecella</emphasis>
|
||
<bibRefCitation id="09124B5BFF9BFF8BFE33FE8E88EB6DF0" author="Warncke, K." box="[477,649,366,393]" pageId="18" pageNumber="469" pagination="27 - 76" refId="ref24173" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
|
||
</taxonomicName>
|
||
), which have a genital capsule that is rounded, the inner margin of the gonostyli creating a broadly circular area, without gonocoxal teeth, and with a penis valve that is comparatively broad basally, narrowing apically (e.g.
|
||
<taxonomicName id="AA834D29FF9BFF8BFDB8FE56890E6DA8" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[598,876,438,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="butea">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFDB8FE5688D76DA8" box="[598,693,439,465]" italics="true" pageId="18" pageNumber="469">A. butea</emphasis>
|
||
<bibRefCitation id="09124B5BFF9BFF8BFD50FE56890E6DA8" author="Warncke, K." box="[702,876,438,465]" pageId="18" pageNumber="469" pagination="27 - 76" refId="ref24173" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="AA834D29FF9BFF8BFC96FE568ECD6DA8" authority="Warncke, 1973" authorityName="Warncke" authorityYear="1973" box="[888,1199,439,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="hyemala">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFC96FE56899B6DA8" box="[888,1017,439,465]" italics="true" pageId="18" pageNumber="469">A. hyemala</emphasis>
|
||
Warncke, 1973
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="AA834D29FF9BFF8BFB53FE568AB06D8C" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="pelopa">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFB53FE568F486DA8" box="[1213,1322,439,465]" italics="true" pageId="18" pageNumber="469">A. pelopa</emphasis>
|
||
<bibRefCitation id="09124B5BFF9BFF8BFADCFE568AB06D8C" author="Warncke, K." pageId="18" pageNumber="469" pagination="29 - 102" refId="ref24254" refString="Warncke, K. (1975) Die Sandbienen der Turkei (Hymenoptera, Apoidea, Andrena), Teil B. Mitteilungen der Munchner Entomologischen Gesellschaft, 65, 29 - 102." type="journal article" year="1975">Warncke, 1975</bibRefCitation>
|
||
</taxonomicName>
|
||
, excluding the clearly divergent structure seen in
|
||
<taxonomicName id="AA834D29FF9BFF8BFCE0FE3A89E16D8D" authorityName="Schmiedeknecht" authorityYear="1900" box="[782,899,475,501]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="arsinoe">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFCE0FE3A89E16D8D" box="[782,899,475,501]" italics="true" pageId="18" pageNumber="469">A. arsinoe</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AA834D29FF9BFF8BFC53FE3A8E5B6D8C" authorityName="Warncke" authorityYear="1967" box="[957,1081,475,501]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="amicula">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFC53FE3A8E5B6D8C" box="[957,1081,475,501]" italics="true" pageId="18" pageNumber="469">A. amicula</emphasis>
|
||
</taxonomicName>
|
||
that do not fall within the true
|
||
<emphasis id="5FF7EAB8FF9BFF8BFF79FE1E8B5B6E60" box="[151,313,511,537]" italics="true" pageId="18" pageNumber="469">Graecandrena</emphasis>
|
||
,
|
||
<bibRefCitation id="09124B5BFF9BFF8BFEA6FE1E887F6E60" author="Pisanty, G. & Richter, R. & Martin, T. & Dettman, J. & Cardinal, S." box="[328,541,511,537]" pageId="18" pageNumber="469" refId="ref23835" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. & Cardinal, S. (2020) Molecular phylogeny and historical biogeography of an- drenine bees (Hymenoptera: Andrenidae). bioRxiv. [published online] https: // doi. org / 10.1101 / 2020.06.09.103556" type="book" year="2020">
|
||
Pisanty
|
||
<emphasis id="5FF7EAB8FF9BFF8BFE4CFE1E8BB66E60" box="[418,468,511,537]" italics="true" pageId="18" pageNumber="469">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
, see above).
|
||
<taxonomicName id="AA834D29FF9BFF8BFD40FE1E89C96E60" authorityName="Wood" authorityYear="2021" box="[686,939,511,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="virguladivina">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFD40FE1E89C96E60" box="[686,939,511,537]" italics="true" pageId="18" pageNumber="469">Andrena virguladivina</emphasis>
|
||
</taxonomicName>
|
||
deviates from this structure by the presence of very slight gonocoxal teeth (though these are also present in
|
||
<taxonomicName id="AA834D29FF9BFF8BFCB2FDC28E086E44" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[860,1130,546,573]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="butea">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFCB2FDC289DB6E44" box="[860,953,547,573]" italics="true" pageId="18" pageNumber="469">A. butea</emphasis>
|
||
<bibRefCitation id="09124B5BFF9BFF8BFC2EFDC28E086E44" author="Warncke, K." box="[960,1130,546,573]" pageId="18" pageNumber="469" pagination="27 - 76" refId="ref24173" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
|
||
</taxonomicName>
|
||
) but overall construction is similar.
|
||
</paragraph>
|
||
<paragraph id="6D3C36AAFF9BFF8BFF29FD8A8F0C6F6C" blockId="18.[151,1437,150,861]" pageId="18" pageNumber="469">
|
||
Males can be instantly diagnosed from all other
|
||
<emphasis id="5FF7EAB8FF9BFF8BFD0BFD8A8E736EFC" box="[741,1041,619,645]" italics="true" pageId="18" pageNumber="469">
|
||
<taxonomicName id="AA834D29FF9BFF8BFD0BFD8A89046EFC" box="[741,870,619,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="456" phylum="Arthropoda" rank="subGenus" subGenus="Aciandrena">Aciandrena</taxonomicName>
|
||
/Graecandrena
|
||
</emphasis>
|
||
species by the genital constriction, specifically the structure of the penis valve which is apically bifurcate, a unique character. Females can be diagnosed by the combination of a finely shagreened propodeal triangle, narrow facial foveae, and the sculpturing of the clypeus which is domed, slightly flattened centrally, and strongly and coarsely punctate, leading to an uneven, misshapen, and partially deformed apical third, with punctures separated by less than half a puncture diameter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25996521FF9BFF8BFF29FCFE89496F24" pageId="18" pageNumber="469" type="etymology">
|
||
<paragraph id="6D3C36AAFF9BFF8BFF29FCFE89496F24" blockId="18.[151,1437,150,861]" pageId="18" pageNumber="469">
|
||
<emphasis id="5FF7EAB8FF9BFF8BFF29FCFE8B306F40" bold="true" box="[199,338,799,825]" pageId="18" pageNumber="469">Etymology:</emphasis>
|
||
The name is Latin for divining rod (or dowsing rod) because of the shape of the male penis valve which is apically bifurcate, thus resembling a divining rod.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |