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<document id="7F66D061BBFED4F46D919FC73B7063A9" ID-DOI="10.11646/zootaxa.5443.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="11045403" ID-ZooBank="C64715E1-18DC-40E3-AED1-86F5A63E6AD6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1713857862667" checkinUser="plazi" docAuthor="Bollino, Maurizio &amp; Anderson, Robert S." docDate="2024" docId="DF6187DAFFD87C0B07DF96876240840E" docLanguage="en" docName="zootaxa.5443.3.7.pdf" docOrigin="Zootaxa 5443 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5443.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eupholus sulisi Bollino and Anderson 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="435" masterDocId="2358FFA2FFD97C0F0748955561518200" masterDocTitle="A new species of Eupholus Boisduval, 1835 from Indonesian New Guinea with notes on the taxonomy of the genus (Coleoptera, Curculionidae)" masterLastPageNumber="445" masterPageNumber="431" pageNumber="432" updateTime="1714075692918" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="09E95D6BE7815FE3FE84FA8A016CFC01" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F9EBE0CFC36C62D6677E1AF651723A2D">A new species of Eupholus Boisduval, 1835 from Indonesian New Guinea with notes on the taxonomy of the genus (Coleoptera, Curculionidae)</mods:title>
</mods:titleInfo>
<mods:name id="07B6CA57A34583EEAD3BDB79D8CCA4F5" type="personal">
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<mods:namePart id="CB8B213376549483BDE02B5B053CB806">Bollino, Maurizio</mods:namePart>
<mods:affiliation id="3340EA559021E5E8DDC48A56758C3458">Museo di Storia naturale del Salento, 73021 Calimera (Lecce), Italy.</mods:affiliation>
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<mods:namePart id="F8FCCC31674E7F9165BC066CF0EE41EA">Anderson, Robert S.</mods:namePart>
<mods:affiliation id="DF7DBF72464DAEECE75136AE68C564AB">Beaty Centre for Species Discovery, Canadian Museum of Nature, 1740 Chemin Pink, Gatineau, Quebec, J 9 J 3 N 7, Canada.</mods:affiliation>
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<mods:title id="D5B325C0926ED05C6C5B8BD6D0137442">Zootaxa</mods:title>
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<mods:date id="C1FDCDDF63B1978DA5B83E09A290C3C9">2024</mods:date>
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<mods:number id="F03B2F34990375D981B6FFD298279CAA">2024-04-23</mods:number>
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<mods:number id="11CFDAB994B8CA6C96AB4E2CAFA079E1">5443</mods:number>
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<treatment id="DF6187DAFFD87C0B07DF96876240840E" ID-DOI="http://doi.org/10.5281/zenodo.11045385" ID-Zenodo-Dep="11045385" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E" httpUri="http://treatment.plazi.org/id/DF6187DAFFD87C0B07DF96876240840E" lastPageId="4" lastPageNumber="435" pageId="1" pageNumber="432">
<subSubSection id="1FD26547FFD87C0E07DF9687638681ED" box="[151,727,978,1005]" pageId="1" pageNumber="432" type="nomenclature">
<paragraph id="577736CCFFD87C0E07DF9687638681ED" blockId="1.[151,727,978,1041]" box="[151,727,978,1005]" pageId="1" pageNumber="432">
<heading id="0C3F81A0FFD87C0E07DF9687638681ED" bold="true" box="[151,727,978,1005]" fontSize="11" level="1" pageId="1" pageNumber="432" reason="1">
<emphasis id="65BCEADEFFD87C0E07DF9687638681ED" bold="true" box="[151,727,978,1005]" pageId="1" pageNumber="432">
<taxonomicName id="90C84D4FFFD87C0E07DF9687633C81EC" authority="Bollino and Anderson" authorityName="Bollino and Anderson" authorityYear="2024" box="[151,621,978,1005]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="sulisi" status="sp. nov.">
<emphasis id="65BCEADEFFD87C0E07DF9687600581ED" bold="true" box="[151,340,978,1005]" italics="true" pageId="1" pageNumber="432">Eupholus sulisi</emphasis>
Bollino and Anderson
</taxonomicName>
,
<taxonomicNameLabel id="7E8F57A5FFD87C0E05329686638681ED" box="[634,727,979,1005]" pageId="1" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1FD26547FFD87C0E07DF96A2607D8611" box="[151,300,1015,1041]" pageId="1" pageNumber="432" type="description">
<paragraph id="577736CCFFD87C0E07DF96A2607D8611" blockId="1.[151,727,978,1041]" box="[151,300,1015,1041]" pageId="1" pageNumber="432">
(
<figureCitation id="CFF32A49FFD87C0E07D796A260758611" box="[159,292,1015,1041]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[284,1308,192,1905]" captionTargetId="figure-29@2.[272,1316,179,1917]" captionTargetPageId="2" captionText="FIGURE 1: A: Eupholus sulisi Holotype female, dorsal view; B: same, lateral view; C: E. sulisi male, Cyclops Mt.; D–E: E. sulisi females, Cyclops Mt.; F: E. sulisi male, Arfak Mts.; G: E. sulisi male, Star Mts." figureDoi="http://doi.org/10.5281/zenodo.11045405" httpUri="https://zenodo.org/record/11045405/files/figure.png" pageId="1" pageNumber="432">Figs. 1A–G</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1FD26547FFD87C0E07DF916B60A68755" pageId="1" pageNumber="432" type="diagnosis">
<paragraph id="577736CCFFD87C0E07DF916B60A68755" blockId="1.[151,1437,1086,2013]" pageId="1" pageNumber="432">
<emphasis id="65BCEADEFFD87C0E07DF916B605B8658" bold="true" box="[151,266,1086,1112]" pageId="1" pageNumber="432">Diagnosis</emphasis>
. A brilliant blue-green
<taxonomicName id="90C84D4FFFD87C0E056D916A63BD8659" box="[549,748,1087,1113]" pageId="1" pageNumber="432">
<emphasis id="65BCEADEFFD87C0E056D916A63DE8659" box="[549,655,1087,1113]" italics="true" pageId="1" pageNumber="432">Eupholus</emphasis>
species
</taxonomicName>
with two glabrous transverse black bands towards elytral declivity (one just in front of declivity and one just behind) and black glabrous, swollen humeri; scales of body and appendages a metallic blue-green but more vividly blue (but sparse) anterolaterally on pronotum, in posthumeral area and surrounding second transverse black band on elytra. It is very similar in pattern to
<taxonomicName id="90C84D4FFFD87C0E03C991FE619E86E9" authority="Heller 1910" authorityName="Heller" authorityYear="1910" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="65BCEADEFFD87C0E03C991FE640186C4" box="[1153,1360,1195,1221]" italics="true" pageId="1" pageNumber="432">Eupholus prasinus</emphasis>
<bibRefCitation id="33594B3DFFD87C0E021E91FE619E86E9" author="Heller, K. M." pageId="1" pageNumber="432" pagination="1 - 44" refId="ref4860" refString="Heller, K. M. (1910) Funfter Beitrage zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausheute von Dr. Schlaginhaufen. Mit 1 Tafel und 2 Figuren im Texte. Abhandlungen und Berichte des Koniglichen Zoologischen Anthropologischen- Ethnographischen Museums zu Dresden, 13 (3), 1 - 44." type="book chapter" year="1910">Heller 1910</bibRefCitation>
</taxonomicName>
from which it is immediately distinguished by the shape of the rostrum, furrowed in
<taxonomicName id="90C84D4FFFD87C0E033C919A659C86E9" authorityName="Bollino and Anderson" authorityYear="2024" box="[1140,1229,1231,1257]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFD87C0E033C919A65D486E8" box="[1140,1157,1231,1256]" italics="true" pageId="1" pageNumber="432">E</emphasis>
.
<emphasis id="65BCEADEFFD87C0E03DA919A659C86E9" box="[1170,1229,1231,1257]" italics="true" pageId="1" pageNumber="432">sulisi</emphasis>
</taxonomicName>
and with a median ridge in
<taxonomicName id="90C84D4FFFD87C0E07B291A6602A870C" authorityName="Heller" authorityYear="1910" box="[250,379,1267,1293]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="65BCEADEFFD87C0E07B291A6605A870C" box="[250,267,1267,1292]" italics="true" pageId="1" pageNumber="432">E</emphasis>
.
<emphasis id="65BCEADEFFD87C0E065391A6602A870C" box="[283,379,1267,1292]" italics="true" pageId="1" pageNumber="432">prasinus</emphasis>
</taxonomicName>
, and to
<taxonomicName id="90C84D4FFFD87C0E069691A66252870D" authority="Heller 1908" authorityName="Heller" authorityYear="1908" box="[478,771,1267,1293]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="65BCEADEFFD87C0E069691A660BE870C" box="[478,495,1267,1292]" italics="true" pageId="1" pageNumber="432">E</emphasis>
.
<emphasis id="65BCEADEFFD87C0E06B791A6633E870D" box="[511,623,1267,1293]" italics="true" pageId="1" pageNumber="432">humeralis</emphasis>
<bibRefCitation id="33594B3DFFD87C0E053191A66252870D" author="Heller, K. M." box="[633,771,1267,1293]" pageId="1" pageNumber="432" pagination="1 - 34" refId="ref4821" refString="Heller, K. M. (1908) Vierter Beitrag zur Papuanischen Kaferfauna. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch-Etnographischen Museums zu Dresden, 12 (1), 1 - 34, 1 pl." type="journal article" year="1908">Heller 1908</bibRefCitation>
</taxonomicName>
, from which it is distinguished by the different pattern of elytral scales, the glabrous black, although also swollen, humerus in
<taxonomicName id="90C84D4FFFD87C0E04C3904262B28731" authorityName="Bollino and Anderson" authorityYear="2024" box="[907,995,1303,1329]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFD87C0E04C3904262B28731" box="[907,995,1303,1329]" italics="true" pageId="1" pageNumber="432">E. sulisi</emphasis>
</taxonomicName>
, and the different shape of the aedeagus and of the everted endophallus.
</paragraph>
</subSubSection>
<subSubSection id="1FD26547FFD87C0E078F900A633A8499" pageId="1" pageNumber="432" type="materials_examined">
<paragraph id="577736CCFFD87C0E078F900A622587C1" blockId="1.[151,1437,1086,2013]" pageId="1" pageNumber="432">
<emphasis id="65BCEADEFFD87C0E078F900A60238778" bold="true" box="[199,370,1374,1401]" pageId="1" pageNumber="432">Type material</emphasis>
.
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<typeStatus id="8873886EFFD87C0E06CC900A60BE8779" box="[388,495,1375,1401]" pageId="1" pageNumber="432" type="holotype">Holotype</typeStatus>
, female (
<figureCitation id="CFF32A49FFD87C0E0515900A63B68779" box="[605,743,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[284,1308,192,1905]" captionTargetId="figure-29@2.[272,1316,179,1917]" captionTargetPageId="2" captionText="FIGURE 1: A: Eupholus sulisi Holotype female, dorsal view; B: same, lateral view; C: E. sulisi male, Cyclops Mt.; D–E: E. sulisi females, Cyclops Mt.; F: E. sulisi male, Arfak Mts.; G: E. sulisi male, Star Mts." figureDoi="http://doi.org/10.5281/zenodo.11045405" httpUri="https://zenodo.org/record/11045405/files/figure.png" pageId="1" pageNumber="432">Figs. 1A–B</figureCitation>
):
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—
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/
<collectingCounty id="BE164E40FFD87C0E0367900A65C68779" box="[1071,1175,1375,1401]" pageId="1" pageNumber="432">Dormena</collectingCounty>
vill. (
<collectingMunicipality id="B713ACB6FFD87C0E03A9900A64188779" box="[1249,1353,1375,1401]" pageId="1" pageNumber="432">Jayapura</collectingMunicipality>
,
<collectingRegion id="950CF82EFFD87C0E0210900A64CD8779" box="[1368,1436,1375,1401]" country="Indonesia" name="Papua Barat" pageId="1" pageNumber="432">Papua</collectingRegion>
<location id="52176017FFD87C0E07DF90D661AA879D" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E:52176017FFD87C0E07DF90D661AA879D" box="[151,251,1411,1437]" country="Indonesia" county="Dormena" municipality="Jayapura" name="Regency" pageId="1" pageNumber="432" stateProvince="Papua Barat">Regency</location>
) /
<location id="52176017FFD87C0E065690D660AD879D" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E:52176017FFD87C0E065690D660AD879D" box="[286,508,1411,1437]" country="Indonesia" county="Dormena" municipality="Jayapura" name="Cyclops Mountains" pageId="1" pageNumber="432" stateProvince="Papua Barat">Cyclops Mountains</location>
/
<date id="2376100CFFD87C0E055090D6632F879D" box="[536,638,1411,1437]" pageId="1" pageNumber="432" value="2023-07">
<collectingDate id="3332E9E4FFD87C0E055090D6632F879D" box="[536,638,1411,1437]" pageId="1" pageNumber="432" value="2023-07">VII.2023</collectingDate>
</date>
/
<collectorName id="FA3D531AFFD87C0E05D290D66399879D" box="[666,712,1411,1437]" pageId="1" pageNumber="432">Lgt.</collectorName>
local people—coll.
<collectorName id="FA3D531AFFD87C0E04FC90D66559879D" box="[948,1032,1411,1437]" pageId="1" pageNumber="432">Bollino</collectorName>
//
</materialsCitation>
<materialsCitation id="E7A03C91FFD87C0E036490D6622187C1" collectionCode="CMNC" country="Indonesia" pageId="1" pageNumber="432" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<typeStatus id="8873886EFFD87C0E036490D66591879D" box="[1068,1216,1411,1437]" pageId="1" pageNumber="432" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="90C84D4FFFD87C0E039390D664DB879D" authorityName="Bollino and Anderson" authorityYear="2024" box="[1243,1418,1411,1437]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFD87C0E039390D664DB879D" box="[1243,1418,1411,1437]" italics="true" pageId="1" pageNumber="432">Eupholus sulisi</emphasis>
</taxonomicName>
/ BOLLINO &amp; ANDERSON, 2024 (typed on red card), in
<collectionCode id="31D9AE09FFD87C0E045290F2622187C1" box="[794,880,1447,1473]" pageId="1" pageNumber="432">CMNC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="577736CCFFD87C0E078F909E633A8499" blockId="1.[151,1437,1086,2013]" pageId="1" pageNumber="432">
<typeStatus id="8873886EFFD87C0E078F909E606587E5" box="[199,308,1483,1509]" pageId="1" pageNumber="432" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="41CEFD45FFD87C0E060E909E60D087E5" box="[326,385,1483,1509]" count="6" pageId="1" pageNumber="432" type="male">6♂♂</specimenCount>
,
<specimenCount id="41CEFD45FFD87C0E06D8909F609887E5" box="[400,457,1482,1509]" count="5" pageId="1" pageNumber="432" type="female">5♀♀</specimenCount>
):
<materialsCitation id="E7A03C91FFD87C0E06AA909E62D58409" collectingDate="2023-07" collectorName="Lgt. &amp; Bollino" country="Indonesia" county="Dormena" location="Regency" municipality="Jayapura" pageId="1" pageNumber="432" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="41CEFD45FFD87C0E06AA909E637787E5" box="[482,550,1483,1509]" count="2" pageId="1" pageNumber="432" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="41CEFD45FFD87C0E057D909E632787E5" box="[565,630,1483,1509]" count="2" pageId="1" pageNumber="432" type="female">2 ♀♀</specimenCount>
<collectingCountry id="2FDF765CFFD87C0E05C8909E63BD87E5" box="[640,748,1483,1509]" name="Indonesia" pageId="1" pageNumber="432">Indonesia</collectingCountry>
—
<collectingRegion id="950CF82EFFD87C0E0441909E62C387E5" box="[777,914,1483,1509]" country="Indonesia" name="Papua Barat" pageId="1" pageNumber="432">Papua Prov.</collectingRegion>
/
<collectingCounty id="BE164E40FFD87C0E04E4909E654587E5" box="[940,1044,1483,1509]" pageId="1" pageNumber="432">Dormena</collectingCounty>
vill. (
<collectingMunicipality id="B713ACB6FFD87C0E0314909E659587E5" box="[1116,1220,1483,1509]" pageId="1" pageNumber="432">Jayapura</collectingMunicipality>
,
<collectingRegion id="950CF82EFFD87C0E0399909E644487E5" box="[1233,1301,1483,1509]" country="Indonesia" name="Papua Barat" pageId="1" pageNumber="432">Papua</collectingRegion>
<location id="52176017FFD87C0E0268909E64D587E5" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E:52176017FFD87C0E0268909E64D587E5" box="[1312,1412,1483,1509]" country="Indonesia" county="Dormena" municipality="Jayapura" name="Regency" pageId="1" pageNumber="432" stateProvince="Papua Barat">Regency</location>
) /
<location id="52176017FFD87C0E07DF90BA60248409" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E:52176017FFD87C0E07DF90BA60248409" box="[151,373,1519,1545]" country="Indonesia" county="Dormena" municipality="Jayapura" name="Cyclops Mountains" pageId="1" pageNumber="432" stateProvince="Papua Barat">Cyclops Mountains</location>
/
<date id="2376100CFFD87C0E06D890BA60A78409" box="[400,502,1519,1545]" pageId="1" pageNumber="432" value="2023-07">
<collectingDate id="3332E9E4FFD87C0E06D890BA60A78409" box="[400,502,1519,1545]" pageId="1" pageNumber="432" value="2023-07">VII.2023</collectingDate>
</date>
/
<collectorName id="FA3D531AFFD87C0E055A90BA63118409" box="[530,576,1519,1545]" pageId="1" pageNumber="432">Lgt.</collectorName>
local people—coll.
<collectorName id="FA3D531AFFD87C0E046490BA62D58409" box="[812,900,1519,1545]" pageId="1" pageNumber="432">Bollino</collectorName>
</materialsCitation>
;
<materialsCitation id="E7A03C91FFD87C0E04DA90BA64C9842D" collectingDate="2023-07" collectorName="Lgt. &amp; Bollino" country="Indonesia" county="Highland" location="Bintang Mountains Regency" municipality="Weime District" pageId="1" pageNumber="432" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="41CEFD45FFD87C0E04DA90BA62908409" box="[914,961,1519,1545]" count="1" pageId="1" pageNumber="432" type="male">1 ♂</specimenCount>
,
<specimenCount id="41CEFD45FFD87C0E049890BA62AC8409" box="[976,1021,1519,1545]" count="1" pageId="1" pageNumber="432" type="female">1 ♀</specimenCount>
<collectingCountry id="2FDF765CFFD87C0E034F90BA65258409" box="[1031,1140,1519,1545]" name="Indonesia" pageId="1" pageNumber="432">Indonesia</collectingCountry>
—
<collectingCounty id="BE164E40FFD87C0E03D890BA65A98409" box="[1168,1272,1519,1545]" pageId="1" pageNumber="432">Highland</collectingCounty>
<collectingRegion id="950CF82EFFD87C0E024A90BA64DA8409" box="[1282,1419,1519,1545]" country="Indonesia" name="Papua Barat" pageId="1" pageNumber="432">Papua Prov.</collectingRegion>
/
<collectingMunicipality id="B713ACB6FFD87C0E07DF93466010842D" box="[151,321,1555,1581]" pageId="1" pageNumber="432">Weime District</collectingMunicipality>
/
<location id="52176017FFD87C0E061F934663C6842D" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E:52176017FFD87C0E061F934663C6842D" box="[343,663,1555,1581]" country="Indonesia" county="Highland" municipality="Weime District" name="Bintang Mountains Regency" pageId="1" pageNumber="432" stateProvince="Papua Barat">
<location id="52176017FFD87C0E061F9346637F842D" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E:52176017FFD87C0E061F9346637F842D" box="[343,558,1555,1581]" country="Indonesia" county="Highland" municipality="Weime District" name="Bintang Mountains" pageId="1" pageNumber="432" stateProvince="Papua Barat">
Bintang
<location id="52176017FFD87C0E06FE9346637F842D" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E:52176017FFD87C0E06FE9346637F842D" box="[438,558,1555,1581]" country="Indonesia" county="Highland" municipality="Weime District" name="Mountains Regency" pageId="1" pageNumber="432" stateProvince="Papua Barat">Mountains</location>
</location>
Regency
</location>
/ (
<location id="52176017FFD87C0E05FD9346624D842D" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E:52176017FFD87C0E05FD9346624D842D" box="[693,796,1555,1581]" country="Indonesia" county="Highland" municipality="Weime District" name="Star Mts." pageId="1" pageNumber="432" stateProvince="Papua Barat">Star Mts.</location>
)—m 2000—
<date id="2376100CFFD87C0E04FF93466548842D" box="[951,1049,1555,1581]" pageId="1" pageNumber="432" value="2023-07">
<collectingDate id="3332E9E4FFD87C0E04FF93466548842D" box="[951,1049,1555,1581]" pageId="1" pageNumber="432" value="2023-07">VII.2023</collectingDate>
</date>
/
<collectorName id="FA3D531AFFD87C0E03679346650F842D" box="[1071,1118,1555,1581]" pageId="1" pageNumber="432">Lgt.</collectorName>
local people—coll.
<collectorName id="FA3D531AFFD87C0E0208934664C9842D" box="[1344,1432,1555,1581]" pageId="1" pageNumber="432">Bollino</collectorName>
</materialsCitation>
;
<materialsCitation id="E7A03C91FFD87C0E07DF936263678475" collectingDate="2022-03" collectorName="Lgt. &amp; M. Bollino" country="Indonesia" location="Manokwari province" municipality="Neney village - Ransiki" pageId="1" pageNumber="432" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="41CEFD45FFD87C0E07DF936261868451" box="[151,215,1591,1617]" count="3" pageId="1" pageNumber="432" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="41CEFD45FFD87C0E07AB936260708451" box="[227,289,1591,1617]" count="2" pageId="1" pageNumber="432" type="female">2 ♀♀</specimenCount>
<collectingCountry id="2FDF765CFFD87C0E066F936260C38451" box="[295,402,1591,1617]" name="Indonesia" pageId="1" pageNumber="432">Indonesia</collectingCountry>
—
<collectingRegion id="950CF82EFFD87C0E06E6936263638451" box="[430,562,1591,1617]" country="Indonesia" name="Papua Barat" pageId="1" pageNumber="432">West Papua</collectingRegion>
/
<collectingMunicipality id="B713ACB6FFD87C0E050E936262028451" box="[582,851,1591,1617]" pageId="1" pageNumber="432">Neney village—Ransiki</collectingMunicipality>
distrik /
<location id="52176017FFD87C0E04FD936265CE8451" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E:52176017FFD87C0E04FD936265CE8451" box="[949,1183,1591,1617]" country="Indonesia" municipality="Neney village - Ransiki" name="Manokwari province" pageId="1" pageNumber="432" stateProvince="Papua Barat">Manokwari province</location>
/
<date id="2376100CFFD87C0E03FB9362646B8451" box="[1203,1338,1591,1617]" pageId="1" pageNumber="432" value="2022-03">
<collectingDate id="3332E9E4FFD87C0E03FB9362646B8451" box="[1203,1338,1591,1617]" pageId="1" pageNumber="432" value="2022-03">March 2022</collectingDate>
</date>
—m 900 /
<collectorName id="FA3D531AFFD87C0E07EE930E61848475" box="[166,213,1627,1653]" pageId="1" pageNumber="432">Lgt.</collectorName>
local people—coll.
<collectorName id="FA3D531AFFD87C0E06F0930E63678475" box="[440,566,1627,1653]" pageId="1" pageNumber="432">M. Bollino</collectorName>
</materialsCitation>
, all in
<collectionCode id="31D9AE09FFD87C0E05C9930E639A8475" box="[641,715,1627,1653]" pageId="1" pageNumber="432">MBLI</collectionCode>
.
<location id="52176017FFD87C0E059C930E63A98475" LSID="urn:lsid:plazi:treatment:DF6187DAFFD87C0B07DF96876240840E:52176017FFD87C0E059C930E63A98475" box="[724,760,1627,1653]" country="Indonesia" name="All" pageId="1" pageNumber="432" stateProvince="Papua Barat">All</location>
<typeStatus id="8873886EFFD87C0E05B6930E62388475" box="[766,873,1627,1653]" pageId="1" pageNumber="432" type="paratype">paratypes</typeStatus>
with additional red label:
<typeStatus id="8873886EFFD87C0E03D9930E644D8475" box="[1169,1308,1627,1653]" pageId="1" pageNumber="432" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="90C84D4FFFD87C0E027A930E61838499" authorityName="Bollino and Anderson" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFD87C0E027A930E61838499" italics="true" pageId="1" pageNumber="432">Eupholus sulisi</emphasis>
</taxonomicName>
/ BOLLINO &amp; ANDERSON, 2024.
</paragraph>
</subSubSection>
<subSubSection id="1FD26547FFD87C0C078F93F7657687FE" lastPageId="3" lastPageNumber="434" pageId="1" pageNumber="432" type="description">
<paragraph id="577736CCFFD87C0C078F93F762BD81A5" blockId="1.[151,1437,1086,2013]" lastBlockId="3.[151,1437,151,1221]" lastPageId="3" lastPageNumber="434" pageId="1" pageNumber="432">
<emphasis id="65BCEADEFFD87C0E078F93F7600084BC" bold="true" box="[199,337,1698,1724]" pageId="1" pageNumber="432">Description</emphasis>
. LB
<quantity id="90309B29FFD87C0E06C793F660A984BD" box="[399,504,1699,1725]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="1" pageNumber="432" unit="mm" value="18.9">18.9 mm</quantity>
; LE
<quantity id="90309B29FFD87C0E057993F663CA84BD" box="[561,667,1699,1725]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="1" pageNumber="432" unit="mm" value="14.2">14.2 mm</quantity>
; LP
<quantity id="90309B29FFD87C0E059993F6627D84BD" box="[721,812,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="432" unit="mm" value="4.2">4.2 mm</quantity>
; LR
<quantity id="90309B29FFD87C0E042F93F6629384BD" box="[871,962,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="432" unit="mm" value="3.7">3.7 mm</quantity>
; WE
<quantity id="90309B29FFD87C0E034C93F6650E84BD" box="[1028,1119,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="1" pageNumber="432" unit="mm" value="8.7">8.7 mm</quantity>
; WP
<quantity id="90309B29FFD87C0E03D793F765AB84BD" box="[1183,1274,1698,1725]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="432" unit="mm" value="5.2">5.2 mm</quantity>
; WR
<quantity id="90309B29FFD87C0E027593F664C884BD" box="[1341,1433,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="432" unit="mm" value="2.4">2.4 mm</quantity>
.
<typeStatus id="8873886EFFD87C0E07DF939261AE84E1" box="[151,255,1735,1761]" pageId="1" pageNumber="432" type="holotype">Holotype</typeStatus>
female. Length. Total length
<quantity id="90309B29FFD87C0E05149392639784E1" box="[604,710,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="1" pageNumber="432" unit="mm" value="23.9">23.9 mm</quantity>
; pronotum + elytron
<quantity id="90309B29FFD87C0E04889392657B84E1" box="[960,1066,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="1" pageNumber="432" unit="mm" value="18.9">18.9 mm</quantity>
. Coloration brilliant blue-green, elytra with two glabrous transverse black bands towards elytral declivity (one just in front of declivity and one just behind it) and humeri also black, glabrous, humeri swollen; scales of body and appendages metallic blue-green but more vividly blue anterolaterally on pronotum, in posthumeral area and surrounding second transverse black band on elytra. Head with pale translucent, recumbent to suberect apically truncate scales in addition to appressed metallic scales, both types of scales more dense bordering eye; surface dull, micro-reticulate, crenulate behind eye. Eye height approximately 1.3 × the width. Rostrum 1.83 × longer than wide at base; maximum width in front of antennal insertion, there 1.46 × width at base; dorsum trisulcate, median furrow terminating well in front of eyes, with low glabrous median carina terminating at indistinct linear impression between front of eyes, and with pair of sublateral furrows in front of eyes; in addition to metallic scales with pale translucent, recumbent to suberect apically subtruncate scales; epistome V-shaped, sunken posteriorly, ridged; scrobes short, descending, not reaching eyes; venter densely covered with adpressed metallic blue-green scales interspersed with scattered long suberect setae, with median furrow and pair of sublateral furrows, anteriorly behind prementum with 2 stiff erect setae, half the length of prementum. Antenna: funicle + club about 2 × longer than scape; scape with metallic scales and sparse elongate erect seta-like scales; funicle covered with appressed pale green scales and dense long erect seta-like scales; club about 3 × as long as wide. Pronotum 1.23 × wider than long; sulcate and glabrous medially, especially so just in front of midlength; with dense metallic green scales not fully concealing surface, scales less dense and more vividly blue (although sparse) in lateral band starting behind eyes and extended posteriorly to hind margin pronotum; surface texture irregular, weakly punctate-rugose. Scutellum circular, covered with erect setiform pale translucent scales. Thoracic venter densely squamose with blue-green, round to lanceolate, recumbent scales; posterior process between fore coxae glabrous. Elytron 1.89 × longer than width at humerus; 1.42 × wider at humerus than maximum width of the pronotum, greatest width near middle, there 1.1 × wider than at humerus; humerus evenly rounded, with protruding knoblike swelling; all intervals flat or very slightly convex; surface with dense appressed round metallic scales almost fully covering elytral surface except for two glabrous transverse black bands towards elytral declivity (one just in front of declivity and one just behind it) and humeri; striae distinct, punctures small but deep; epipleural edge with fringe of pale, lanceolate scales. Legs with uniform vestiture of metallic green rounded scales, interspersed with dense suberect yellow setalike scales, especially on tibia; front and middle tibia with inner margins toothed distally, teeth small, widely separated; front tibia with apical one-third rather strongly curved inwardly. Abdomen densely squamose with blue-green, round to lanceolate, subrecumbent scales. Genitalia. Hemisternite (9E) similar to those of the other species of the
<emphasis id="65BCEADEFFDA7C0C05E396DE62FA81A5" box="[683,939,907,933]" italics="true" pageId="3" pageNumber="434">
<taxonomicName id="90C84D4FFFDA7C0C05E396DE624481A5" authorityName="Boisduval" authorityYear="1835" box="[683,789,907,933]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">Eupholus</taxonomicName>
schoenherri-
</emphasis>
group
</paragraph>
<caption id="03B76644FFDB7C0D07D092C2655485D3" ID-DOI="http://doi.org/10.5281/zenodo.11045405" ID-Zenodo-Dep="11045405" httpUri="https://zenodo.org/record/11045405/files/figure.png" pageId="2" pageNumber="433" startId="2.[152,255,1943,1967]" targetBox="[284,1308,192,1905]" targetPageId="2" targetType="figure">
<paragraph id="577736CCFFDB7C0D07D092C2655485D3" blockId="2.[151,1436,1943,2003]" pageId="2" pageNumber="433">
<emphasis id="65BCEADEFFDB7C0D07D092C2604A85AF" bold="true" box="[152,283,1943,1967]" pageId="2" pageNumber="433">FIGURE 1:</emphasis>
A:
<taxonomicName id="90C84D4FFFDB7C0D060992C2608C85AF" authorityName="Bollino and Anderson" authorityYear="2024" box="[321,477,1943,1967]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFDB7C0D060992C2608C85AF" box="[321,477,1943,1967]" italics="true" pageId="2" pageNumber="433">Eupholus sulisi</emphasis>
</taxonomicName>
Holotype female, dorsal view; B: same, lateral view; C:
<taxonomicName id="90C84D4FFFDB7C0D036292CD652D85AF" authorityName="Bollino and Anderson" authorityYear="2024" box="[1066,1148,1943,1967]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFDB7C0D036292CD656885AF" box="[1066,1081,1944,1967]" italics="true" pageId="2" pageNumber="433">E</emphasis>
.
<emphasis id="65BCEADEFFDB7C0D030E92C2652D85AF" box="[1094,1148,1943,1967]" italics="true" pageId="2" pageNumber="433">sulisi</emphasis>
</taxonomicName>
male, Cyclops Mt.; D–E:
<taxonomicName id="90C84D4FFFDB7C0D02CE92CD619C85D3" authorityName="Bollino and Anderson" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFDB7C0D02CE92CD64C485AF" box="[1414,1429,1944,1967]" italics="true" pageId="2" pageNumber="433">E</emphasis>
.
<emphasis id="65BCEADEFFDB7C0D07DF92EE619C85D3" box="[151,205,1979,2003]" italics="true" pageId="2" pageNumber="433">sulisi</emphasis>
</taxonomicName>
females, Cyclops Mt.; F:
<taxonomicName id="90C84D4FFFDB7C0D069392E9637C85D3" authorityName="Bollino and Anderson" authorityYear="2024" box="[475,557,1979,2003]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFDB7C0D069392E960BB85D3" box="[475,490,1980,2003]" italics="true" pageId="2" pageNumber="433">E</emphasis>
.
<emphasis id="65BCEADEFFDB7C0D06BF92EE637C85D3" box="[503,557,1979,2003]" italics="true" pageId="2" pageNumber="433">sulisi</emphasis>
</taxonomicName>
male, Arfak Mts.; G:
<taxonomicName id="90C84D4FFFDB7C0D045A92E9623585D3" authorityName="Bollino and Anderson" authorityYear="2024" box="[786,868,1979,2003]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFDB7C0D045A92E9627085D3" box="[786,801,1980,2003]" italics="true" pageId="2" pageNumber="433">E</emphasis>
.
<emphasis id="65BCEADEFFDB7C0D046692EE623585D3" box="[814,868,1979,2003]" italics="true" pageId="2" pageNumber="433">sulisi</emphasis>
</taxonomicName>
male, Star Mts.
</paragraph>
</caption>
<paragraph id="577736CCFFDA7C0C078F96FA62F086C5" blockId="3.[151,1437,151,1221]" pageId="3" pageNumber="434">
Male (
<figureCitation id="CFF32A49FFDA7C0C065D96FA609581C9" box="[277,452,943,969]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[284,1308,192,1905]" captionTargetId="figure-29@2.[272,1316,179,1917]" captionTargetPageId="2" captionText="FIGURE 1: A: Eupholus sulisi Holotype female, dorsal view; B: same, lateral view; C: E. sulisi male, Cyclops Mt.; D–E: E. sulisi females, Cyclops Mt.; F: E. sulisi male, Arfak Mts.; G: E. sulisi male, Star Mts." figureDoi="http://doi.org/10.5281/zenodo.11045405" httpUri="https://zenodo.org/record/11045405/files/figure.png" pageId="3" pageNumber="434">Figs. 1C, F–G</figureCitation>
). Like female, but slightly smaller (LB: 17.2–18.8); 1.25–1.27 × wider at humerus than maximum width of the pronotum, greatest width near middle, there only slightly (1.03–1.04 ×) wider than at humerus; humerus uniformly rounded, with barely noticeable swelling. Genitalia. Aedeagus (
<figureCitation id="CFF32A49FFDA7C0C025E96A264C28611" box="[1302,1427,1015,1041]" captionStart="FIGURE 9" captionStartId="13.[152,255,1874,1898]" captionTargetBox="[236,1353,211,1824]" captionTargetId="figure-24@13.[206,1381,181,1850]" captionTargetPageId="13" captionText="FIGURE 9: Female left hemisternites of Eupholus schoenherri-group and Eupholus alternans-group. A: Eupholus azureus; B: E. dhuyi; C: E. amalulu; D: E. browni; E: E. sulisi; F: E. alternans; G: E. amaliae; H: E. antonkozlovi. Male genital of Eupholus spp., aedeagus in dorsal (I, N) and lateral view (L, O); sternite IX in dorsal view (M, P): E. sulisi sp. n. (I, L, M), E. humeralis (N,O, P)" figureDoi="http://doi.org/10.5281/zenodo.11045421" httpUri="https://zenodo.org/record/11045421/files/figure.png" pageId="3" pageNumber="434">Figs. 9I–L</figureCitation>
) and everted endophallus (
<figureCitation id="CFF32A49FFDA7C0C06F7914E636A8635" box="[447,571,1051,1077]" captionStart="FIGURE 3" captionStartId="6.[152,255,1807,1831]" captionTargetBox="[281,1307,181,1783]" captionTargetId="figure-29@6.[281,1307,181,1783]" captionTargetPageId="6" captionText="FIGURE 3: Lateral view of Eupholus loriae-group male A: E. cuvierii Female left hemisternites of Eupholus loriae-group B: E. loriae; C: E. prasinus; D: E. euphrosyne; E: E. bruijni; F: E. cuvierii, Lateral view of endophallus of Eupholus schoenherrigroup males G: E. sulisi Cyclops Mts.; H: E. sulisi Arfak Mts.; I: E. sulisi Star Mts.; L: E. humeralis Kutubu Lake; M: E. humeralis Bosavi Mt.; N: E. humeralis Merauke Prov." figureDoi="http://doi.org/10.5281/zenodo.11045409" httpUri="https://zenodo.org/record/11045409/files/figure.png" pageId="3" pageNumber="434">Figs. 3G–I</figureCitation>
) as illustrated. Apart from the evident difference in the morphology of the aedeagus compared to the externally most similar species (
<taxonomicName id="90C84D4FFFDA7C0C046F916A62E78659" authorityName="Heller" authorityYear="1908" box="[807,950,1087,1113]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="65BCEADEFFDA7C0C046F916A62E78659" box="[807,950,1087,1113]" italics="true" pageId="3" pageNumber="434">E. humeralis</emphasis>
</taxonomicName>
), it is worth noting that
<taxonomicName id="90C84D4FFFDA7C0C038A916A644D8659" authorityName="Bollino and Anderson" authorityYear="2024" box="[1218,1308,1087,1113]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFDA7C0C038A916A644D8659" box="[1218,1308,1087,1113]" italics="true" pageId="3" pageNumber="434">E. sulisi</emphasis>
</taxonomicName>
exhibits an unusual morphology of the subapical diverticulum of the everted endophallus (see red arrows in
<figureCitation id="CFF32A49FFDA7C0C039391366406867D" box="[1243,1367,1123,1149]" captionStart="FIGURE 3" captionStartId="6.[152,255,1807,1831]" captionTargetBox="[281,1307,181,1783]" captionTargetId="figure-29@6.[281,1307,181,1783]" captionTargetPageId="6" captionText="FIGURE 3: Lateral view of Eupholus loriae-group male A: E. cuvierii Female left hemisternites of Eupholus loriae-group B: E. loriae; C: E. prasinus; D: E. euphrosyne; E: E. bruijni; F: E. cuvierii, Lateral view of endophallus of Eupholus schoenherrigroup males G: E. sulisi Cyclops Mts.; H: E. sulisi Arfak Mts.; I: E. sulisi Star Mts.; L: E. humeralis Kutubu Lake; M: E. humeralis Bosavi Mt.; N: E. humeralis Merauke Prov." figureDoi="http://doi.org/10.5281/zenodo.11045409" httpUri="https://zenodo.org/record/11045409/files/figure.png" pageId="3" pageNumber="434">Figs. 3G–I</figureCitation>
). This diverticulum, in fact, has a subquadrangular shape in lateral view, whereas all other species we studied within the
<emphasis id="65BCEADEFFDA7C0C07DF91FE60C686C5" box="[151,407,1195,1221]" italics="true" pageId="3" pageNumber="434">
<taxonomicName id="90C84D4FFFDA7C0C07DF91FE605086C5" authorityName="Boisduval" authorityYear="1835" box="[151,257,1195,1221]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="genus">Eupholus</taxonomicName>
schoenherri-
</emphasis>
group have it triangular and oriented ventrally.
</paragraph>
<paragraph id="577736CCFFDA7C0C07DF905960D38726" blockId="3.[151,386,1292,1318]" box="[151,386,1292,1318]" pageId="3" pageNumber="434">
<heading id="0C3F81A0FFDA7C0C07DF905960D38726" bold="true" box="[151,386,1292,1318]" fontSize="11" level="1" pageId="3" pageNumber="434" reason="1">
<emphasis id="65BCEADEFFDA7C0C07DF905960D38726" bold="true" box="[151,386,1292,1318]" pageId="3" pageNumber="434">Variability (1C–G)</emphasis>
</heading>
</paragraph>
<paragraph id="577736CCFFDA7C0C07DF9001657687FE" blockId="3.[151,1437,1364,1535]" pageId="3" pageNumber="434">
<taxonomicName id="90C84D4FFFDA7C0C07DF90016012876E" authorityName="Bollino and Anderson" authorityYear="2024" box="[151,323,1364,1390]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFDA7C0C07DF90016012876E" box="[151,323,1364,1390]" italics="true" pageId="3" pageNumber="434">Eupholus sulisi</emphasis>
</taxonomicName>
is a species with considerable variability, both among populations and within the same population. Individuals from the Arfak Mountains lack glabrous areas to trace lines, while those from the typical locality vary greatly both in terms of glabrous areas and the extent of the vivid blue posthumeral bands of elytra. Finally, the pair from the Star Mountains exhibit an unusual gray coloration of the scales, a hue shared by the majority of specimens of
<taxonomicName id="90C84D4FFFDA7C0C07FE90B1637487FE" authority="Boisduval" authorityName="Boisduval" box="[182,549,1508,1534]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="schoenherri">
<emphasis id="65BCEADEFFDA7C0C07FE90B160FD87FE" box="[182,428,1508,1534]" italics="true" pageId="3" pageNumber="434">Eupholus schoenherri</emphasis>
Boisduval
</taxonomicName>
that we have received from the same locality.
</paragraph>
</subSubSection>
<subSubSection id="1FD26547FFDA7C0C07DF930561A384FA" pageId="3" pageNumber="434" type="distribution">
<paragraph id="577736CCFFDA7C0C07DF93056069846A" blockId="3.[151,312,1616,1642]" box="[151,312,1616,1642]" pageId="3" pageNumber="434">
<heading id="0C3F81A0FFDA7C0C07DF93056069846A" bold="true" box="[151,312,1616,1642]" fontSize="11" level="1" pageId="3" pageNumber="434" reason="1">
<emphasis id="65BCEADEFFDA7C0C07DF93056069846A" bold="true" box="[151,312,1616,1642]" pageId="3" pageNumber="434">Distribution.</emphasis>
</heading>
</paragraph>
<paragraph id="577736CCFFDA7C0C07DF93CD61A384FA" blockId="3.[151,1436,1688,1786]" pageId="3" pageNumber="434">
The new species is distributed to the north of the highlands running west to east along the entire island of New
<collectingCountry id="2FDF765CFFDA7C0C07DF93E961BD84D6" box="[151,236,1724,1750]" name="Guinea-Bissau" pageId="3" pageNumber="434">Guinea</collectingCountry>
. South of this mountain range,
<taxonomicName id="90C84D4FFFDA7C0C057693E9624C84D6" authorityName="Heller" authorityYear="1908" box="[574,797,1724,1750]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="65BCEADEFFDA7C0C057693E9624C84D6" box="[574,797,1724,1750]" italics="true" pageId="3" pageNumber="434">Eupholus humeralis</emphasis>
</taxonomicName>
, the most closely related species, appears to be distributed (
<figureCitation id="CFF32A49FFDA7C0C07D793B561B684FA" box="[159,231,1760,1786]" captionStart="MAP 1" captionStartId="4.[152,210,877,901]" captionTargetBox="[152,1436,181,854]" captionTargetId="figure-29@4.[152,1436,181,854]" captionTargetPageId="4" captionText="MAP 1. Known distribution of Eupholus sulisi (red asterisk: type locality; red squares: other localities) and Eupholus humeralis (blue asterisk: type locality; blue squares: other localities)." figureDoi="http://doi.org/10.5281/zenodo.11068902" httpUri="https://zenodo.org/record/11068902/files/figure.png" pageId="3" pageNumber="434">Map 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1FD26547FFDA7C0C07DF921960E485F7" pageId="3" pageNumber="434" type="etymology">
<paragraph id="577736CCFFDA7C0C07DF921960758566" blockId="3.[151,292,1868,1894]" box="[151,292,1868,1894]" pageId="3" pageNumber="434">
<heading id="0C3F81A0FFDA7C0C07DF921960758566" bold="true" box="[151,292,1868,1894]" fontSize="11" level="1" pageId="3" pageNumber="434" reason="1">
<emphasis id="65BCEADEFFDA7C0C07DF921960758566" bold="true" box="[151,292,1868,1894]" pageId="3" pageNumber="434">Etymology.</emphasis>
</heading>
</paragraph>
<paragraph id="577736CCFFDA7C0C07DF92C160E485F7" blockId="3.[151,1437,1940,2039]" pageId="3" pageNumber="434">
This species is dedicated to the friend of the first author, Sulistiningsih (Sulis for friends) (Sentani,
<collectingCountry id="2FDF765CFFDA7C0C03BF92C1643985AE" box="[1271,1384,1940,1966]" name="Indonesia" pageId="3" pageNumber="434">Indonesia</collectingCountry>
), for the generous assistance provided, enabling him to obtain interesting specimens. The species name is a feminine noun in the genitive case.
</paragraph>
</subSubSection>
<caption id="03B76644FFDD7C0B07D0963863B881AA" ID-DOI="http://doi.org/10.5281/zenodo.11068902" ID-Zenodo-Dep="11068902" httpUri="https://zenodo.org/record/11068902/files/figure.png" pageId="4" pageNumber="435" startId="4.[152,210,877,901]" targetBox="[152,1436,181,854]" targetPageId="4">
<paragraph id="577736CCFFDD7C0B07D0963863B881AA" blockId="4.[151,1436,877,938]" pageId="4" pageNumber="435">
<emphasis id="65BCEADEFFDD7C0B07D0963861BA8185" bold="true" box="[152,235,877,901]" pageId="4" pageNumber="435">MAP 1.</emphasis>
Known distribution of
<taxonomicName id="90C84D4FFFDD7C0B069D963B63218186" authorityName="Bollino and Anderson" authorityYear="2024" box="[469,624,878,902]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="sulisi">
<emphasis id="65BCEADEFFDD7C0B069D963B63218186" box="[469,624,878,902]" italics="true" pageId="4" pageNumber="435">Eupholus sulisi</emphasis>
</taxonomicName>
(red asterisk:
<typeStatus id="8873886EFFDD7C0B05B5963B62788186" box="[765,809,878,902]" pageId="4" pageNumber="435">type</typeStatus>
locality; red squares: other localities) and
<taxonomicName id="90C84D4FFFDD7C0B039A963B64CD8186" authorityName="Heller" authorityYear="1908" box="[1234,1436,878,902]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="65BCEADEFFDD7C0B039A963B64CD8186" box="[1234,1436,878,902]" italics="true" pageId="4" pageNumber="435">Eupholus humeralis</emphasis>
</taxonomicName>
(blue asterisk:
<typeStatus id="8873886EFFDD7C0B066496C7600981AA" box="[300,344,914,938]" pageId="4" pageNumber="435">type</typeStatus>
locality; blue squares: other localities).
</paragraph>
</caption>
<subSubSection id="1FD26547FFDD7C0B07DF96A96240840E" pageId="4" pageNumber="435" type="discussion">
<paragraph id="577736CCFFDD7C0B07DF96A960388616" blockId="4.[151,361,1020,1046]" box="[151,361,1020,1046]" pageId="4" pageNumber="435">
<heading id="0C3F81A0FFDD7C0B07DF96A960388616" bold="true" box="[151,361,1020,1046]" fontSize="11" level="1" pageId="4" pageNumber="435" reason="1">
<emphasis id="65BCEADEFFDD7C0B07DF96A960388616" bold="true" box="[151,361,1020,1046]" pageId="4" pageNumber="435">Taxonomic notes</emphasis>
</heading>
</paragraph>
<paragraph id="577736CCFFDD7C0B07DF911163868736" blockId="4.[151,1436,1092,1550]" pageId="4" pageNumber="435">
<bibRefCitation id="33594B3DFFDD7C0B07DF91116063865E" author="Riedel, A." box="[151,306,1092,1118]" pageId="4" pageNumber="435" pagination="1 - 16" refId="ref5199" refString="Riedel, A. (2002) Two new species of Eupholus Boisduval (Coleoptera, Curculionidae, Entiminae) from West New Guinea, a discussion of their taxonomic characters, and notes on nomenclature. Zootaxa, 90 (1), 1 - 16. https: // doi. org / 10.11646 / zootaxa. 90.1.1" type="journal article" year="2002">Riedel (2002)</bibRefCitation>
, identified two distinct
<typeStatus id="8873886EFFDD7C0B057A9111633C865E" box="[562,621,1092,1118]" pageId="4" pageNumber="435">types</typeStatus>
of transfer apparatus occurring within the genus
<taxonomicName id="90C84D4FFFDD7C0B03CC911165BF865E" authorityName="Boisduval" authorityYear="1835" box="[1156,1262,1092,1118]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="65BCEADEFFDD7C0B03CC911165BF865E" box="[1156,1262,1092,1118]" italics="true" pageId="4" pageNumber="435">Eupholus</emphasis>
</taxonomicName>
. This character, combined with the structure of the dorsal surface of the rostrum, whose differences in various species were already recognized by
<bibRefCitation id="33594B3DFFDD7C0B067191D9608386A6" author="Heller, K. M." box="[313,466,1164,1190]" pageId="4" pageNumber="435" pagination="146 - 154" refId="ref4914" refString="Heller, K. M. (1923) Bestimmungsschlussel aussereuropaischer Kafer. Curculionidae: Genus Eupholus Guer. Koleopterologische Rundschau, 10 (4), 146 - 154." type="journal article" year="1923">Heller (1923)</bibRefCitation>
, allowed our contemporary colleague to identify two groups of species: 1) the
<emphasis id="65BCEADEFFDD7C0B027A91D961B286CA" italics="true" pageId="4" pageNumber="435">
<taxonomicName id="90C84D4FFFDD7C0B027A91D964CD86A6" authorityName="Boisduval" authorityYear="1835" box="[1330,1436,1164,1190]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">Eupholus</taxonomicName>
loriae-
</emphasis>
group, characterized by a rather compact transfer apparatus (if observed with the inflexed endophallus), and a median ridge on the rostrum in most species, even if some species have a medially furrowed rostrum, and 2) the
<emphasis id="65BCEADEFFDD7C0B07DF91AD60CB8712" box="[151,410,1272,1298]" italics="true" pageId="4" pageNumber="435">
<taxonomicName id="90C84D4FFFDD7C0B07DF91AD60508712" authorityName="Boisduval" authorityYear="1835" box="[151,257,1272,1298]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">Eupholus</taxonomicName>
schoenherri-
</emphasis>
group, characterized by an elongate sting-like transfer apparatus when observed with the inflexed endophallus, and with a furrowed rostrum.
</paragraph>
<paragraph id="577736CCFFDD7C0B078F90156240840E" blockId="4.[151,1436,1092,1550]" pageId="4" pageNumber="435">
Having the opportunity to study the everted endophallus and/or the female genitalia of 38 of 71 known species recognized as valid at the moment (
<bibRefCitation id="33594B3DFFDD7C0B056190316205877E" author="Porion, T. &amp; Audibert, C." box="[553,852,1380,1406]" pageId="4" pageNumber="435" refId="ref5171" refString="Porion, T. &amp; Audibert, C. (2020) Le Genre Eupholus. Taita Publishers, Hradec Kralove, Republique Tcheque, 171 pp." type="book" year="2020">Porion and Audibert 2020</bibRefCitation>
;
<bibRefCitation id="33594B3DFFDD7C0B0417903165C2877E" author="Limoges, R. &amp; Le Tirant, S." box="[863,1171,1380,1406]" pageId="4" pageNumber="435" pagination="1 - 7" refId="ref5079" refString="Limoges, R. &amp; Le Tirant, S. (2022) Nouvelle espece du genre Eupholus Boisduval, 1835 de la Papouasie, Indonesie (Coleoptera, Curculionidae, Entiminae). Faunitaxys, 10 (63), 1 - 7." type="journal article" year="2022">Limoges &amp; Le Tirant 2022</bibRefCitation>
; present paper), we can confirm the existence of the two species group (
<emphasis id="65BCEADEFFDD7C0B059990DC63A287A2" box="[721,755,1417,1442]" italics="true" pageId="4" pageNumber="435">i.e.</emphasis>
the
<emphasis id="65BCEADEFFDD7C0B046390DD62BD87A2" box="[811,1004,1416,1442]" italics="true" pageId="4" pageNumber="435">
<taxonomicName id="90C84D4FFFDD7C0B046390DD62C487A2" authorityName="Boisduval" authorityYear="1835" box="[811,917,1416,1442]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">Eupholus</taxonomicName>
loriae-
</emphasis>
group and the
<taxonomicName id="90C84D4FFFDD7C0B03D090DD618B87C6" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="schoenherri-group">
<emphasis id="65BCEADEFFDD7C0B03D090DD618B87C6" italics="true" pageId="4" pageNumber="435">Eupholus schoenherri-group</emphasis>
</taxonomicName>
) plus another species group to which we will refer to as the
<emphasis id="65BCEADEFFDD7C0B04D490F965D287C6" box="[924,1155,1452,1478]" italics="true" pageId="4" pageNumber="435">
<taxonomicName id="90C84D4FFFDD7C0B04D490F9655787C6" authorityName="Boisduval" authorityYear="1835" box="[924,1030,1452,1478]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="genus">Eupholus</taxonomicName>
alternans-
</emphasis>
group. Finally,
<taxonomicName id="90C84D4FFFDD7C0B027A90F9634487EA" authority="Porion and Audibert" authorityName="Porion and Audibert" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="antonkozlovi">
<emphasis id="65BCEADEFFDD7C0B027A90F9607787EA" italics="true" pageId="4" pageNumber="435">Eupholus antonkozlovi</emphasis>
Porion and Audibert
</taxonomicName>
, due to the morphology of the everted endophallus, appears not to belong to any of the three identified groups and will be discussed later.
</paragraph>
</subSubSection>
</treatment>
</document>
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