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<document id="F8C17BF4E86DD0F6A3F08420B9C5572B" ID-CLB-Dataset="6363" ID-DOI="10.11646/zootaxa.4974.2.1" ID-GBIF-Dataset="16a1742e-cafe-4c2e-a8c3-2e1b3b57d456" ID-ISSN="1175-5326" ID-Zenodo-Dep="4775440" ID-ZooBank="0CCA430E-5601-42CB-847F-87B22BFD3112" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621509898047" checkinUser="plazi" docAuthor="Schools, Molly &amp; Hedges, S. Blair" docDate="2021" docId="DA66BA10FFC1FFD80DF10D9D04CDD321" docLanguage="en" docName="zootaxa.4974.2.1.pdf" docOrigin="Zootaxa 4974 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wetmorena Cochran 1927" docType="treatment" docVersion="8" lastPageNumber="234" masterDocId="265FC268FFDEFFF90D6608570654D778" masterDocTitle="Phylogenetics, classification, and biogeography of the Neotropical forest lizards (Squamata, Diploglossidae)" masterLastPageNumber="257" masterPageNumber="201" pageNumber="232" updateTime="1698940127518" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3540B8FA6C00FB476BF988D0E6CBBDAF">Phylogenetics, classification, and biogeography of the Neotropical forest lizards (Squamata, Diploglossidae)</mods:title>
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<mods:namePart id="FE54B5B015775C26B3EAC302295D9D1C">Schools, Molly</mods:namePart>
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<mods:namePart id="C742D5D52FDB4399702664A32A84ECAE">Hedges, S. Blair</mods:namePart>
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<mods:affiliation id="884D677717353843700BD2475A64AD09">sbh @ temple. edu; https: // orcid. org / 0000 - 0002 - 0652 - 2411</mods:affiliation>
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<mods:date id="CA5FE3DFB2418BCD1A744FF565CBD004">2021</mods:date>
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<treatment id="DA66BA10FFC1FFD80DF10D9D04CDD321" ID-DOI="http://doi.org/10.5281/zenodo.4891080" ID-GBIF-Taxon="182482706" ID-Zenodo-Dep="4891080" LSID="urn:lsid:plazi:treatment:DA66BA10FFC1FFD80DF10D9D04CDD321" httpUri="http://treatment.plazi.org/id/DA66BA10FFC1FFD80DF10D9D04CDD321" lastPageId="33" lastPageNumber="234" pageId="31" pageNumber="232">
<subSubSection id="1AD5588DFFC1FFE60DF10D9D041DD171" pageId="31" pageNumber="232" type="nomenclature">
<paragraph id="52700B06FFC1FFE60DF10D9D0463D29C" blockId="31.[151,585,1482,1581]" box="[151,567,1482,1509]" pageId="31" pageNumber="232">
<heading id="0938BC6AFFC1FFE60DF10D9D0463D29C" bold="true" box="[151,567,1482,1509]" fontSize="11" level="1" pageId="31" pageNumber="232" reason="1">
<emphasis id="60BBD714FFC1FFE60DF10D9D0463D29C" bold="true" box="[151,567,1482,1509]" pageId="31" pageNumber="232">
Genus
<taxonomicName id="95CF7085FFC1FFE60D880D9C0463D29C" authority="Cochran, 1927" authorityName="Cochran" authorityYear="1927" box="[238,567,1482,1509]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFC1FFE60D880D9C072CD29D" bold="true" box="[238,376,1483,1509]" italics="true" pageId="31" pageNumber="232">Wetmorena</emphasis>
<bibRefCitation id="365E76F7FFC1FFE60CE60D9D0463D29C" author="Cochran, D. M." box="[384,567,1482,1508]" pageId="31" pageNumber="232" pagination="91 - 92" refId="ref21513" refString="Cochran, D. M. (1927) A new genus of anguid lizards from Haiti. Proceedings of the Biological Society of Washington, 40, 91 - 92." type="journal article" year="1927">Cochran, 1927</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="52700B06FFC1FFE60DF10DB8041DD171" blockId="31.[151,585,1482,1581]" box="[151,585,1519,1545]" pageId="31" pageNumber="232">Short-headed Four-toed Forest Lizards</paragraph>
</subSubSection>
<subSubSection id="1AD5588DFFC1FFE60DF10E4406B3D155" box="[151,231,1555,1581]" pageId="31" pageNumber="232" type="description">
<paragraph id="52700B06FFC1FFE60DF10E4406B3D155" blockId="31.[151,585,1482,1581]" box="[151,231,1555,1581]" pageId="31" pageNumber="232">
<figureCitation id="CAF41783FFC1FFE60DF10E4406B3D155" box="[151,231,1555,1581]" captionStart="FIGURE 20" captionStartId="32.[151,250,1943,1967]" captionTargetBox="[184,1405,187,1920]" captionTargetId="figure-26@32.[183,1405,181,1920]" captionTargetPageId="32" captionText="FIGURE 20. In life images of (A) Wetmorena agasepsoides (ANSP 38713, SVL 57.6 mm) and (B) Wetmorena haetiana surda (USNM 328897). Photographs by S. B. Hedges. (C) The distribution of Wetmorena on Hispaniola." figureDoi="http://doi.org/10.5281/zenodo.4775499" httpUri="https://zenodo.org/record/4775499/files/figure.png" pageId="31" pageNumber="232">Fig. 20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="1AD5588DFFC1FFE60DF10E0F02BDD117" box="[151,1257,1623,1648]" pageId="31" pageNumber="232" type="reference_group">
<paragraph id="52700B06FFC1FFE60DF10E0F02BDD117" blockId="31.[151,1257,1623,1648]" box="[151,1257,1623,1648]" pageId="31" pageNumber="232">
<treatmentCitationGroup id="72DF2C28FFC1FFE60DF10E0F02BDD117" box="[151,1257,1623,1648]" pageId="31" pageNumber="232">
<treatmentCitation id="D36E2D17FFC1FFE60DF10E0F0793D117" author="Cochran, D. M." box="[151,455,1623,1647]" page="91" pageId="31" pageNumber="232" year="1927">
<taxonomicName id="95CF7085FFC1FFE60DF10E0F0793D117" authority="Cochran, 1927: 91" authorityName="Cochran" authorityPageNumber="91" authorityYear="1927" box="[151,455,1623,1647]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFC1FFE60DF10E0F075DD117" box="[151,265,1624,1647]" italics="true" pageId="31" pageNumber="232">Wetmorena</emphasis>
<bibRefCitation id="365E76F7FFC1FFE60C690E000793D117" author="Cochran, D. M." box="[271,455,1623,1647]" pageId="31" pageNumber="232" pagination="91 - 92" refId="ref21513" refString="Cochran, D. M. (1927) A new genus of anguid lizards from Haiti. Proceedings of the Biological Society of Washington, 40, 91 - 92." type="journal article" year="1927">Cochran, 1927:91</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="8D74B5A4FFC1FFE60CB70E0F0457D108" box="[465,515,1624,1648]" pageId="31" pageNumber="232">Type</typeStatus>
species.
<treatmentCitation id="D36E2D17FFC1FFE60F390E0F05BAD117" author="Cochran, D. M." box="[607,1006,1623,1647]" page="91" pageId="31" pageNumber="232" year="1927">
<taxonomicName id="95CF7085FFC1FFE60F390E0F05BAD117" ID-CoL="5BXYT" authority="Cochran, 1927: 91" authorityName="Cochran" authorityPageNumber="91" authorityYear="1927" box="[607,1006,1623,1647]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="species" species="haetiana">
<emphasis id="60BBD714FFC1FFE60F390E0F0564D117" box="[607,816,1623,1647]" italics="true" pageId="31" pageNumber="232">Wetmorena haetiana</emphasis>
<bibRefCitation id="365E76F7FFC1FFE60E500E0005BAD117" author="Cochran, D. M." box="[822,1006,1623,1647]" pageId="31" pageNumber="232" pagination="91 - 92" refId="ref21513" refString="Cochran, D. M. (1927) A new genus of anguid lizards from Haiti. Proceedings of the Biological Society of Washington, 40, 91 - 92." type="journal article" year="1927">Cochran, 1927:91</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, by original designation.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1AD5588DFFC1FFD80DF10EC102F8D68D" lastPageId="33" lastPageNumber="234" pageId="31" pageNumber="232" type="diagnosis">
<paragraph id="52700B06FFC1FFE60DF10EC10775D01D" blockId="31.[151,1437,1686,2038]" pageId="31" pageNumber="232">
<emphasis id="60BBD714FFC1FFE60DF10EC10746D1C8" bold="true" box="[151,274,1686,1712]" pageId="31" pageNumber="232">Diagnosis.</emphasis>
Species of
<taxonomicName id="95CF7085FFC1FFE60CFE0EC00441D1C8" authorityName="Cochran" authorityYear="1927" box="[408,533,1687,1712]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFC1FFE60CFE0EC00441D1C8" box="[408,533,1687,1712]" italics="true" pageId="31" pageNumber="232">Wetmorena</emphasis>
</taxonomicName>
have (1) claw sheath, present, (2) contact between the nasal and rostral scales, absent, (3) scales in contact with the nasal scale, four, (4) postnasal scales, one, (5) position of the nostril in the nasal scale, central, (6) keels on dorsal body scales, absent, (7) digits per limb, four, (8) longest toe lamellae, 812, (9) dorsal scale rows, 98117, (10) relative head width, 8.5214.5, (11) relative rostral height, 38.761.8, (12) relative frontonasal length, 1.472.69, (13) relative interparietal distance, 0.4471.03, (14) relative axilla-groin distance (59.971.4).
</paragraph>
<paragraph id="52700B06FFC1FFD80DA10F38055DD6A9" blockId="31.[151,1437,1686,2038]" lastBlockId="33.[151,1437,151,1113]" lastPageId="33" lastPageNumber="234" pageId="31" pageNumber="232">
From
<taxonomicName id="95CF7085FFC1FFE60C6B0F380738D0F1" authority="Schools &amp; Hedges, 2021" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[269,364,1903,1929]" class="Reptilia" family="Diploglossidae" genus="Advenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="60BBD714FFC1FFE60C6B0F380738D0F1" box="[269,364,1903,1929]" italics="true" pageId="31" pageNumber="232">Advenus</emphasis>
</taxonomicName>
<emphasis id="60BBD714FFC1FFE60C120F38078BD0F1" bold="true" box="[372,479,1903,1929]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="7B886A6FFFC1FFE60C120F38078BD0F1" box="[372,479,1903,1929]" pageId="31" pageNumber="232" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, we distinguish
<taxonomicName id="95CF7085FFC1FFE60FFD0F38054CD0F0" authorityName="Cochran" authorityYear="1927" box="[667,792,1903,1928]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFC1FFE60FFD0F38054CD0F0" box="[667,792,1903,1928]" italics="true" pageId="31" pageNumber="232">Wetmorena</emphasis>
</taxonomicName>
by the digits per limb (four versus five in
<taxonomicName id="95CF7085FFC1FFE608620F380337D0F1" authority="Schools &amp; Hedges, 2021" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[1284,1379,1903,1929]" class="Reptilia" family="Diploglossidae" genus="Advenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="60BBD714FFC1FFE608620F380337D0F1" box="[1284,1379,1903,1929]" italics="true" pageId="31" pageNumber="232">Advenus</emphasis>
</taxonomicName>
<emphasis id="60BBD714FFC1FFE6080D0F38069CD0D5" bold="true" pageId="31" pageNumber="232">
<taxonomicNameLabel id="7B886A6FFFC1FFE6080D0F38069CD0D5" pageId="31" pageNumber="232" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
), longest toe lamellae (812 versus 1617), and dorsal scale rows (98117 versus 96). From
<taxonomicName id="95CF7085FFC1FFE609920FC40336D0D5" authority="Schools &amp; Hedges, 2021" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[1268,1378,1939,1965]" class="Reptilia" family="Diploglossidae" genus="Caribicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="60BBD714FFC1FFE609920FC40336D0D5" box="[1268,1378,1939,1965]" italics="true" pageId="31" pageNumber="232">Caribicus</emphasis>
</taxonomicName>
<emphasis id="60BBD714FFC1FFE6080D0FC4069CD0A9" bold="true" pageId="31" pageNumber="232">
<taxonomicNameLabel id="7B886A6FFFC1FFE6080D0FC4069CD0A9" pageId="31" pageNumber="232" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, we distinguish
<taxonomicName id="95CF7085FFC1FFE60CE00FE00457D0A8" authorityName="Cochran" authorityYear="1927" box="[390,515,1975,2000]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFC1FFE60CE00FE00457D0A8" box="[390,515,1975,2000]" italics="true" pageId="31" pageNumber="232">Wetmorena</emphasis>
</taxonomicName>
by the claw sheath (present versus its absence in
<taxonomicName id="95CF7085FFC1FFE609240FE002E4D0A9" authority="Schools &amp; Hedges, 2021" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[1090,1200,1975,2001]" class="Reptilia" family="Diploglossidae" genus="Caribicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="60BBD714FFC1FFE609240FE002E4D0A9" box="[1090,1200,1975,2001]" italics="true" pageId="31" pageNumber="232">Caribicus</emphasis>
</taxonomicName>
<emphasis id="60BBD714FFC1FFE609DF0FE00370D0A9" bold="true" box="[1209,1316,1975,2001]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="7B886A6FFFC1FFE609DF0FE00370D0A9" box="[1209,1316,1975,2001]" pageId="31" pageNumber="232" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
), keels on the dorsal scales (absent versus their presence in
<taxonomicName id="95CF7085FFC1FFE60FB80F8C0518D08D" authority="Schools &amp; Hedges, 2021" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[734,844,2011,2037]" class="Reptilia" family="Diploglossidae" genus="Caribicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="60BBD714FFC1FFE60FB80F8C0518D08D" box="[734,844,2011,2037]" italics="true" pageId="31" pageNumber="232">Caribicus</emphasis>
</taxonomicName>
<emphasis id="60BBD714FFC1FFE60E310F8C0591D08D" bold="true" box="[855,965,2011,2037]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="7B886A6FFFC1FFE60E310F8C0591D08D" box="[855,965,2011,2037]" pageId="31" pageNumber="232" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
), digits per limb (four versus five), and the relative frontonasal length (1.472.69 versus 2.983.32). From
<emphasis id="60BBD714FFFFFFD80ECE08C00259D7C9" box="[936,1037,151,177]" italics="true" pageId="33" pageNumber="234">
<taxonomicName id="95CF7085FFFFFFD80ECE08C0025DD7C9" authorityName="Gray" authorityYear="1839" box="[936,1033,151,177]" class="Reptilia" family="Anguidae" genus="Celestus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">Celestus</taxonomicName>
,
</emphasis>
we distinguish
<taxonomicName id="95CF7085FFFFFFD809AC08C00313D7C8" authorityName="Cochran" authorityYear="1927" box="[1226,1351,151,176]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD809AC08C00313D7C8" box="[1226,1351,151,176]" italics="true" pageId="33" pageNumber="234">Wetmorena</emphasis>
</taxonomicName>
by the claw sheath (present versus its absence in
<taxonomicName id="95CF7085FFFFFFD80F1208EC0487D7AD" authorityName="Gray" authorityYear="1839" box="[628,723,187,213]" class="Reptilia" family="Anguidae" genus="Celestus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80F1208EC0487D7AD" box="[628,723,187,213]" italics="true" pageId="33" pageNumber="234">Celestus</emphasis>
</taxonomicName>
) and the digits per limb (four versus five). From
<taxonomicName id="95CF7085FFFFFFD8086608EC0330D7AD" authority="Schools &amp; Hedges, 2021" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[1280,1380,187,213]" class="Reptilia" family="Diploglossidae" genus="Comptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="60BBD714FFFFFFD8086608EC0330D7AD" box="[1280,1380,187,213]" italics="true" pageId="33" pageNumber="234">Comptus</emphasis>
</taxonomicName>
<emphasis id="60BBD714FFFFFFD8080D08EC069CD781" bold="true" pageId="33" pageNumber="234">
<taxonomicNameLabel id="7B886A6FFFFFFFD8080D08EC069CD781" pageId="33" pageNumber="234" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, we distinguish
<taxonomicName id="95CF7085FFFFFFD80CE6088807A9D780" authorityName="Cochran" authorityYear="1927" box="[384,509,223,248]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80CE6088807A9D780" box="[384,509,223,248]" italics="true" pageId="33" pageNumber="234">Wetmorena</emphasis>
</taxonomicName>
by the claw sheath (present versus its absence in
<taxonomicName id="95CF7085FFFFFFD8094D088802DBD781" authority="Schools &amp; Hedges, 2021" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[1067,1167,223,249]" class="Reptilia" family="Diploglossidae" genus="Comptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="60BBD714FFFFFFD8094D088802DBD781" box="[1067,1167,223,249]" italics="true" pageId="33" pageNumber="234">Comptus</emphasis>
</taxonomicName>
<emphasis id="60BBD714FFFFFFD809F3088802ABD781" bold="true" box="[1173,1279,223,249]" pageId="33" pageNumber="234">
<taxonomicNameLabel id="7B886A6FFFFFFFD809F3088802ABD781" box="[1173,1279,223,249]" pageId="33" pageNumber="234" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
), keels on the dorsal body scales (absent versus their presence in
<taxonomicName id="95CF7085FFFFFFD80FBD0954056BD665" authority="Schools &amp; Hedges, 2021" authorityName="Schools &amp; Hedges" authorityYear="2021" box="[731,831,259,285]" class="Reptilia" family="Diploglossidae" genus="Comptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="60BBD714FFFFFFD80FBD0954056BD665" box="[731,831,259,285]" italics="true" pageId="33" pageNumber="234">Comptus</emphasis>
</taxonomicName>
<emphasis id="60BBD714FFFFFFD80E21095405E5D665" bold="true" box="[839,945,259,285]" pageId="33" pageNumber="234">
<taxonomicNameLabel id="7B886A6FFFFFFFD80E21095405E5D665" box="[839,945,259,285]" pageId="33" pageNumber="234" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
), digits per limb (four versus five), longest toe lamellae (812 versus 1323), the relative frontonasal length (1.472.69 versus 2.953.65), and the distance between the parietal scales (0.4471.03 versus 00.435). From
<emphasis id="60BBD714FFFFFFD80E05091C05B5D61D" box="[867,993,331,357]" italics="true" pageId="33" pageNumber="234">
<taxonomicName id="95CF7085FFFFFFD80E05091C058AD61D" authorityName="Cope" authorityYear="1862" box="[867,990,331,357]" class="Squamata" family="Diploglossidae" genus="Panolopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">Panolopus</taxonomicName>
,
</emphasis>
we distinguish
<taxonomicName id="95CF7085FFFFFFD809F2091C0345D61C" authorityName="Cochran" authorityYear="1927" box="[1172,1297,331,356]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD809F2091C0345D61C" box="[1172,1297,331,356]" italics="true" pageId="33" pageNumber="234">Wetmorena</emphasis>
</taxonomicName>
by the claw sheath (present versus its absence in
<taxonomicName id="95CF7085FFFFFFD80F55093804FFD6F1" authorityName="Cope" authorityYear="1862" box="[563,683,367,393]" class="Squamata" family="Diploglossidae" genus="Panolopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80F55093804FFD6F1" box="[563,683,367,393]" italics="true" pageId="33" pageNumber="234">Panolopus</emphasis>
</taxonomicName>
), the digits per limb (four versus five), and the relative axilla-groin distance (59.971.4 versus 49.759.6). From
<emphasis id="60BBD714FFFFFFD80FCF09C40544D6D5" box="[681,784,403,429]" italics="true" pageId="33" pageNumber="234">
<taxonomicName id="95CF7085FFFFFFD80FCF09C40559D6D5" authorityName="Gray" authorityYear="1852" box="[681,781,403,429]" class="Squamata" family="Diploglossidae" genus="Sauresia" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">Sauresia</taxonomicName>
,
</emphasis>
we distinguish
<taxonomicName id="95CF7085FFFFFFD80EA809C4021FD6D4" authorityName="Cochran" authorityYear="1927" box="[974,1099,403,428]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80EA809C4021FD6D4" box="[974,1099,403,428]" italics="true" pageId="33" pageNumber="234">Wetmorena</emphasis>
</taxonomicName>
by the distance between the parietal scales (0.4471.03 versus
<quantity id="9537A6E3FFFFFFD80F7F09E004C5D6A9" box="[537,657,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="5.4737" metricValueMax="10.9474" metricValueMin="0.0" pageId="33" pageNumber="234" unit="in" value="0.2155" valueMax="0.431" valueMin="0.0">00.431 in</quantity>
<taxonomicName id="95CF7085FFFFFFD80FFE09E004ADD6A9" authorityName="Gray" authorityYear="1852" box="[664,761,439,465]" class="Squamata" family="Diploglossidae" genus="Sauresia" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80FFE09E004ADD6A9" box="[664,761,439,465]" italics="true" pageId="33" pageNumber="234">Sauresia</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="06B05B8EFFFEFFD90DF10FC0022AD0AC" ID-DOI="http://doi.org/10.5281/zenodo.4775499" ID-Zenodo-Dep="4775499" httpUri="https://zenodo.org/record/4775499/files/figure.png" pageId="32" pageNumber="233" startId="32.[151,250,1943,1967]" targetBox="[184,1405,187,1920]" targetPageId="32">
<paragraph id="52700B06FFFEFFD90DF10FC0022AD0AC" blockId="32.[151,1436,1943,2004]" pageId="32" pageNumber="233">
<emphasis id="60BBD714FFFEFFD90DF10FC00774D0D7" bold="true" box="[151,288,1943,1967]" pageId="32" pageNumber="233">FIGURE 20.</emphasis>
In life images of (A)
<taxonomicName id="95CF7085FFFEFFD90C9A0FCF04AFD0C8" baseAuthorityName="Thomas" baseAuthorityYear="1971" box="[508,763,1944,1968]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="233" phylum="Chordata" rank="species" species="agasepsoides">
<emphasis id="60BBD714FFFEFFD90C9A0FCF04AFD0C8" box="[508,763,1944,1968]" italics="true" pageId="32" pageNumber="233">Wetmorena agasepsoides</emphasis>
</taxonomicName>
(ANSP 38713, SVL 57.6 mm) and (B)
<taxonomicName id="95CF7085FFFEFFD909EB0FCF03C8D0C8" box="[1165,1436,1944,1968]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="233" phylum="Chordata" rank="subSpecies" species="haetiana" subSpecies="surda">
<emphasis id="60BBD714FFFEFFD909EB0FCF03C8D0C8" box="[1165,1436,1944,1968]" italics="true" pageId="32" pageNumber="233">Wetmorena haetiana surda</emphasis>
</taxonomicName>
(USNM 328897). Photographs by S. B. Hedges. (C) The distribution of
<taxonomicName id="95CF7085FFFEFFD90E140FEB05B0D0AB" authorityName="Cochran" authorityYear="1927" box="[882,996,1980,2003]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="233" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFEFFD90E140FEB05B0D0AB" box="[882,996,1980,2003]" italics="true" pageId="32" pageNumber="233">Wetmorena</emphasis>
</taxonomicName>
on Hispaniola.
</paragraph>
</caption>
<paragraph id="52700B06FFFFFFD80DA1098D02F8D68D" blockId="33.[151,1437,151,1113]" box="[199,1196,474,501]" pageId="33" pageNumber="234">
<emphasis id="60BBD714FFFFFFD80DA1098D077BD68C" bold="true" box="[199,303,474,500]" pageId="33" pageNumber="234">Content.</emphasis>
Two species (
<tableCitation id="1F4D3EBDFFFFFFD80CAB098C0470D68D" box="[461,548,475,501]" captionStart="TABLE 3" captionStartId="14.[151,239,1291,1315]" captionTargetBox="[163,1425,1404,2011]" captionTargetPageId="14" captionText="TABLE 3. Classification and distribution of lizards of the family Diploglossidae (55 species). For common names we use modifying words from literature sources, in some cases modified for consistency following guidelines (Hedges et al. 2019). The IUCN Redlist threat status is indicated in parentheses (NA = not assessed)." httpUri="http://table.plazi.org/id/06B05B8EFFD0FFF70DF10D5C0264D214" pageId="33" pageNumber="234" tableUuid="06B05B8EFFD0FFF70DF10D5C0264D214">Table 3</tableCitation>
):
<taxonomicName id="95CF7085FFFFFFD80F5C098C0500D68D" baseAuthorityName="Thomas" baseAuthorityYear="1971" box="[570,852,475,501]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="agasepsoides">
<emphasis id="60BBD714FFFFFFD80F5C098C0500D68D" box="[570,852,475,501]" italics="true" pageId="33" pageNumber="234">Wetmorena agasepsoides</emphasis>
</taxonomicName>
(comb. nov.) and
<emphasis id="60BBD714FFFFFFD80979098C02F8D68D" box="[1055,1196,475,501]" italics="true" pageId="33" pageNumber="234">
<taxonomicName id="95CF7085FFFFFFD80979098C02FCD68D" authorityName="Cochran" authorityYear="1927" box="[1055,1192,475,501]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="haetiana">W. haetiana</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="1AD5588DFFFFFFD80DA109A90457D545" pageId="33" pageNumber="234" type="distribution">
<paragraph id="52700B06FFFFFFD80DA109A90457D545" blockId="33.[151,1437,151,1113]" pageId="33" pageNumber="234">
<emphasis id="60BBD714FFFFFFD80DA109A90735D560" bold="true" box="[199,353,510,536]" pageId="33" pageNumber="234">Distribution.</emphasis>
<taxonomicName id="95CF7085FFFFFFD80C0E09A807B1D560" authorityName="Cochran" authorityYear="1927" box="[360,485,511,536]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80C0E09A807B1D560" box="[360,485,511,536]" italics="true" pageId="33" pageNumber="234">Wetmorena</emphasis>
</taxonomicName>
occurs only on Hispaniola, in the Massif de la Selle (
<collectingCountry id="2AD84B96FFFFFFD8092109A802D6D562" box="[1095,1154,511,538]" name="Haiti" pageId="33" pageNumber="234">Haiti</collectingCountry>
) and Sierra de
<collectingRegion id="900BC5E4FFFFFFD8084909A803C8D561" box="[1327,1436,511,537]" country="Dominican Republic" name="Baoruco" pageId="33" pageNumber="234">Bahoruco</collectingRegion>
(
<collectingCountry id="2AD84B96FFFFFFD80DF90A7407DFD546" box="[159,395,547,574]" name="Dominican Republic" pageId="33" pageNumber="234">Dominican Republic</collectingCountry>
) (
<figureCitation id="CAF41783FFFFFFD80CC40A7407A2D545" box="[418,502,547,573]" captionStart="FIGURE 20" captionStartId="32.[151,250,1943,1967]" captionTargetBox="[184,1405,187,1920]" captionTargetId="figure-26@32.[183,1405,181,1920]" captionTargetPageId="32" captionText="FIGURE 20. In life images of (A) Wetmorena agasepsoides (ANSP 38713, SVL 57.6 mm) and (B) Wetmorena haetiana surda (USNM 328897). Photographs by S. B. Hedges. (C) The distribution of Wetmorena on Hispaniola." figureDoi="http://doi.org/10.5281/zenodo.4775499" httpUri="https://zenodo.org/record/4775499/files/figure.png" pageId="33" pageNumber="234">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1AD5588DFFFFFFD80DA10A1005CBD5FD" pageId="33" pageNumber="234" type="etymology">
<paragraph id="52700B06FFFFFFD80DA10A1005CBD5FD" blockId="33.[151,1437,151,1113]" pageId="33" pageNumber="234">
<emphasis id="60BBD714FFFFFFD80DA10A10071AD519" bold="true" box="[199,334,583,609]" pageId="33" pageNumber="234">Etymology.</emphasis>
The generic name references Dr. Alexander Wetmore, who was Assistant Secretary of the Smithsonian Institution (
<collectingRegion id="900BC5E4FFFFFFD80CCA0A3C043BD5FD" box="[428,623,619,645]" country="United States of America" name="District of Columbia" pageId="33" pageNumber="234">Washington D.C.</collectingRegion>
) at the time of description.
</paragraph>
</subSubSection>
<subSubSection id="1AD5588DFFFFFFD80DA10AD804CDD321" pageId="33" pageNumber="234" type="discussion">
<paragraph id="52700B06FFFFFFD80DA10AD805BCD425" blockId="33.[151,1437,151,1113]" pageId="33" pageNumber="234">
<emphasis id="60BBD714FFFFFFD80DA10AD8076FD5D1" bold="true" box="[199,315,655,681]" pageId="33" pageNumber="234">Remarks.</emphasis>
<taxonomicName id="95CF7085FFFFFFD80C240AD807EBD5D0" authorityName="Cochran" authorityYear="1927" box="[322,447,655,680]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80C240AD807EBD5D0" box="[322,447,655,680]" italics="true" pageId="33" pageNumber="234">Wetmorena</emphasis>
</taxonomicName>
is a monophyletic clade that has a support value of 100% in Bayesian and ML analyses (
<figureCitation id="CAF41783FFFFFFD80DC60AE406BED5B5" box="[160,234,691,717]" captionStart="FIGURE 2" captionStartId="9.[151,250,1748,1772]" captionTargetBox="[175,1412,181,1725]" captionTargetId="figure-17@9.[175,1412,181,1725]" captionTargetPageId="9" captionText="FIGURE 2. Phylogenetic tree of diploglossid lizards based on sequences of nine genes: four mitochondrial genes (CytB, ND2, 12S rRNA, and 16S rRNA) and five nuclear genes (AMEL, BDNF, PLPR, RAG1, and ZFP36). Maximum likelihood tree obtained from the nine-gene dataset (59 individuals; 6,949 sites).A scale bar indicates 5% sequence divergence. The numbers at nodes are ML bootstrap values, followed by Bayesian posterior probabilities; asterisks indicate significant (≥ 95%) support, and a dash or zero value indicates weak (&lt;50%) support. The tree is rooted with Pseudopus apodus (Anguidae)." figureDoi="http://doi.org/10.5281/zenodo.4775446" httpUri="https://zenodo.org/record/4775446/files/figure.png" pageId="33" pageNumber="234">Fig. 2</figureCitation>
). Our phylogenies include all three subspecies of
<taxonomicName id="95CF7085FFFFFFD80E580AE4027DD5B5" authorityName="Cochran" authorityYear="1927" box="[830,1065,691,717]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="haetiana">
<emphasis id="60BBD714FFFFFFD80E580AE4027DD5B5" box="[830,1065,691,717]" italics="true" pageId="33" pageNumber="234">Wetmorena haetiana</emphasis>
</taxonomicName>
. Previously, authors considered
<taxonomicName id="95CF7085FFFFFFD80DF10A8007E1D589" baseAuthorityName="Thomas" baseAuthorityYear="1971" box="[151,437,727,753]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="agasepsoides">
<emphasis id="60BBD714FFFFFFD80DF10A8007E1D589" box="[151,437,727,753]" italics="true" pageId="33" pageNumber="234">Wetmorena agasepsoides</emphasis>
</taxonomicName>
to be a species of
<taxonomicName id="95CF7085FFFFFFD80FFA0A8004A9D589" authorityName="Gray" authorityYear="1852" box="[668,765,727,753]" class="Squamata" family="Diploglossidae" genus="Sauresia" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80FFA0A8004A9D589" box="[668,765,727,753]" italics="true" pageId="33" pageNumber="234">Sauresia</emphasis>
</taxonomicName>
because of similarities in their appearance, including the presence of an ear opening. However, molecular data support the clustering of
<taxonomicName id="95CF7085FFFFFFD809470AAC0368D46D" box="[1057,1340,763,789]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="agasepsiodes">
<emphasis id="60BBD714FFFFFFD809470AAC0368D46D" box="[1057,1340,763,789]" italics="true" pageId="33" pageNumber="234">Wetmorena agasepsiodes</emphasis>
</taxonomicName>
with
<taxonomicName id="95CF7085FFFFFFD808E60AAC06ADD441" authorityName="Cochran" authorityYear="1927" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="haetiana">
<emphasis id="60BBD714FFFFFFD808E60AAC06ADD441" italics="true" pageId="33" pageNumber="234">W. haetiana</emphasis>
</taxonomicName>
with a support value of 100% in Bayesian and ML analyses, which indicates that the loss of the auricular opening occurred within the clade recognized here as the genus
<taxonomicName id="95CF7085FFFFFFD80E020B1405B5D424" authorityName="Cochran" authorityYear="1927" box="[868,993,835,860]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80E020B1405B5D424" box="[868,993,835,860]" italics="true" pageId="33" pageNumber="234">Wetmorena</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="52700B06FFFFFFD80DA10B3004CDD321" blockId="33.[151,1437,151,1113]" pageId="33" pageNumber="234">
For most of the time since it was named in 1927,
<taxonomicName id="95CF7085FFFFFFD80F800B300537D4F8" authorityName="Cochran" authorityYear="1927" box="[742,867,871,896]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80F800B300537D4F8" box="[742,867,871,896]" italics="true" pageId="33" pageNumber="234">Wetmorena</emphasis>
</taxonomicName>
has been recognized as a valid genus, distinct from
<taxonomicName id="95CF7085FFFFFFD80DF10BDC06A2D4DD" authorityName="Gray" authorityYear="1839" box="[151,246,907,933]" class="Reptilia" family="Anguidae" genus="Celestus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80DF10BDC06A2D4DD" box="[151,246,907,933]" italics="true" pageId="33" pageNumber="234">Celestus</emphasis>
</taxonomicName>
. The genus was synonomized with
<taxonomicName id="95CF7085FFFFFFD80FE40BDC04B5D4DD" authorityName="Gray" authorityYear="1839" box="[642,737,907,933]" class="Reptilia" family="Anguidae" genus="Celestus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80FE40BDC04B5D4DD" box="[642,737,907,933]" italics="true" pageId="33" pageNumber="234">Celestus</emphasis>
</taxonomicName>
by
<bibRefCitation id="365E76F7FFFFFFD80E6F0BDC059AD4DD" author="Hass, C. A. &amp; Maxson, L. R. &amp; Hedges, S. B." box="[777,974,907,933]" pageId="33" pageNumber="234" pagination="15 - 33" refId="ref22877" refString="Hass, C. A., Maxson, L. R. &amp; Hedges, S. B. (2001) Relationships and divergence times of West Indian amphibians and reptiles: Insights from albumin immunology. In: Woods, C. A. &amp; Sergile, F. E. (Eds.), Biogeography of the West Indies: Patterns and Perspectives. 2 nd Edition. CRC Press, Boca Raton, Florida, pp. 15 - 33. https: // doi. org / 10.1201 / 9781420039481 - 11" type="book chapter" year="2001">
Hass
<emphasis id="60BBD714FFFFFFD80E200BDC0529D4DD" box="[838,893,907,933]" italics="true" pageId="33" pageNumber="234">et al.</emphasis>
(2001)
</bibRefCitation>
because their study using immunological data found relationships that differed from previous studies (
<bibRefCitation id="365E76F7FFFFFFD80E220BF80265D4B1" author="Savage, J. M. &amp; Lips, K. R." box="[836,1073,943,969]" pageId="33" pageNumber="234" pagination="817 - 842" refId="ref24772" refString="Savage, J. M. &amp; Lips, K. R. (1993) A review of the status and biogeography of the lizard genera Celestus and Diploglossus (Squamata: Anguidae), with description of two new species from Costa Rica. Revista de Biologia Tropical, 41, 817 - 842." type="journal article" year="1993">Savage &amp; Lips 1993</bibRefCitation>
). Our study shows that the stem divergence time of
<taxonomicName id="95CF7085FFFFFFD80C0B0B8407BED494" authorityName="Cochran" authorityYear="1927" box="[365,490,979,1004]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD80C0B0B8407BED494" box="[365,490,979,1004]" italics="true" pageId="33" pageNumber="234">Wetmorena</emphasis>
</taxonomicName>
is comparable to the stem times of other genera of celestines (
<figureCitation id="CAF41783FFFFFFD809FC0B840289D495" box="[1178,1245,979,1005]" captionStart="FIGURE 3" captionStartId="10.[151,250,1680,1704]" captionTargetBox="[152,1436,181,1655]" captionTargetId="figure-26@10.[152,1436,181,1656]" captionTargetPageId="10" captionText="FIGURE 3. Timetree of Diploglossidae based on sequences of nine genes: four mitochondrial genes (CytB, ND2, 12S rRNA, and 16S rRNA) and five nuclear genes (AMEL, BDNF, PLPR, RAG1, and ZFP36). Nodes show divergence times in millions of years. A scale bar indicates time (6 million years ago, Mya). We show Bayesian credibility ranges as gray bars at nodes." figureDoi="http://doi.org/10.5281/zenodo.4775450" httpUri="https://zenodo.org/record/4775450/files/figure.png" pageId="33" pageNumber="234">Fig. 3</figureCitation>
). The high levels of divergence (46 Mya) among subspecies of
<taxonomicName id="95CF7085FFFFFFD80FC60BA005D2D369" authorityName="Cochran" authorityYear="1927" box="[672,902,1015,1041]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="haetiana">
<emphasis id="60BBD714FFFFFFD80FC60BA005D2D369" box="[672,902,1015,1041]" italics="true" pageId="33" pageNumber="234">Wetmorena haetiana</emphasis>
</taxonomicName>
(
<figureCitation id="CAF41783FFFFFFD80EF30BA00583D369" box="[917,983,1015,1041]" captionStart="FIGURE 3" captionStartId="10.[151,250,1680,1704]" captionTargetBox="[152,1436,181,1655]" captionTargetId="figure-26@10.[152,1436,181,1656]" captionTargetPageId="10" captionText="FIGURE 3. Timetree of Diploglossidae based on sequences of nine genes: four mitochondrial genes (CytB, ND2, 12S rRNA, and 16S rRNA) and five nuclear genes (AMEL, BDNF, PLPR, RAG1, and ZFP36). Nodes show divergence times in millions of years. A scale bar indicates time (6 million years ago, Mya). We show Bayesian credibility ranges as gray bars at nodes." figureDoi="http://doi.org/10.5281/zenodo.4775450" httpUri="https://zenodo.org/record/4775450/files/figure.png" pageId="33" pageNumber="234">Fig. 3</figureCitation>
), together with additional molecular and morphological data (Schools &amp; Hedges, unpubl.), and greater sampling of populations, suggest that
<taxonomicName id="95CF7085FFFFFFD809940C4C033BD34C" authorityName="Cochran" authorityYear="1927" box="[1266,1391,1051,1076]" class="Squamata" family="Diploglossidae" genus="Wetmorena" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="234" phylum="Chordata" rank="genus">
<emphasis id="60BBD714FFFFFFD809940C4C033BD34C" box="[1266,1391,1051,1076]" italics="true" pageId="33" pageNumber="234">Wetmorena</emphasis>
</taxonomicName>
is a complex that includes four additional species.
</paragraph>
</subSubSection>
</treatment>
</document>