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<mods:title>The woodwasp genus Tremex (Hymenoptera, Siricidae) of Japan</mods:title>
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<taxonomicName authority="Konow, 1896" authorityName="Konow" authorityYear="1896" box="[151,545,319,346]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="longicollis">
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<emphasis bold="true" box="[151,376,319,346]" italics="true" pageId="21" pageNumber="22">Tremex longicollis</emphasis>
<bibRefCitation author="Konow, F. W." box="[383,545,320,346]" pageId="21" pageNumber="22" pagination="41 - 59" refId="ref25417" refString="Konow, F. W. (1896) Verschiedenes aus der Hymenopteren-Gruppe der Tenthrediniden. Wiener Entomologische Zeitung, 15, 41 - 59." type="journal article" year="1896">Konow, 1896</bibRefCitation>
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(
<figureCitation box="[159,287,356,383]" captionStart="FIGURE 2" captionStartId="2.[151,250,1940,1964]" captionTargetBox="[163,1424,188,1911]" captionTargetId="figure-20@2.[151,1435,181,1917]" captionTargetPageId="2" captionText="FIGURE 2. Tremex spp., female, with parts of measurements (white lines in A, B, D–F) and the names of cells in forewing (C).—A, D, T. longicollis, Okazaki, Aichi; B, F, T. fuscicornis, Inchon, Korea; C, T. nakanei, Tokyo; E, T. katayamai, holotype. OOCL: ocellar-occipital (carina) line. MPL: median length of pronotum seen from above." figureDoi="http://doi.org/10.5281/zenodo.7621755" httpUri="https://zenodo.org/record/7621755/files/figure.png" pageId="21" pageNumber="22">Figs 2A, D</figureCitation>
, 
<figureCitation box="[298,333,356,382]" captionStart="FIGURE 3" captionStartId="10.[151,250,1870,1894]" captionTargetBox="[157,1430,190,1838]" captionTargetId="figure-20@10.[151,1435,181,1846]" captionTargetPageId="10" captionText="FIGURE 3. Head, thorax and base of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, T. fuscicornis, Schwedt, Germany, F, G, T. fuscicornis, Inchon, Korea; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621757" httpUri="https://zenodo.org/record/7621757/files/figure.png" pageId="21" pageNumber="22">3C</figureCitation>
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<figureCitation box="[345,379,356,382]" captionStart="FIGURE 4" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[158,1429,188,1197]" captionTargetId="figure-15@11.[151,1435,181,1203]" captionTargetPageId="11" captionText="FIGURE 4. Head, thorax and base of abdomen, male.—A, B, T. apicalis, Minamichitose, Hokkaido; C, D, T. fuscicornis, Shikotsu-ko, Hokkaido; E, T. longicollis, Muroto, Kochi; F, T. nakanei, Noda, Chiba; G, T. contractus, Okayama-shi, Okayama; H, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621759" httpUri="https://zenodo.org/record/7621759/files/figure.png" pageId="21" pageNumber="22">4E</figureCitation>
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<figureCitation box="[390,450,356,383]" captionStart="FIGURE 5" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[155,1432,565,1761]" captionTargetId="figure-186@12.[151,1435,561,1764]" captionTargetPageId="12" captionText="FIGURE 5. Antenna, female (A–G, I) and male (H, J–N).—A. T. apicalis, Minamichitose, Hokkaido; B, T. bicinctus, holotype; C, T. longicollis, Toyota, Aichi; D, T. fuscicornis, Inchon, Korea; E, T. nakanei, Ôginosen, Tottori; F, T. contractus, Okayamashi, Okayama; G, T. okinawensis, holotype; H. T. apicalis, Tsukuba, Ibaraki; I, T. katayamai, holotype; J, T. longicollis, Muroto, Kochi; K, T. fuscicornis, Shikotsu-ko, Hokkaido; L, T. nakanei, Noda, Chiba; M, T. contractus, Tatsuno, Hyogo; N, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621761" httpUri="https://zenodo.org/record/7621761/files/figure.png" pageId="21" pageNumber="22">5C, J</figureCitation>
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<figureCitation box="[462,499,356,382]" captionStart="FIGURE 6" captionStartId="13.[151,250,1061,1085]" captionTargetBox="[155,1432,184,1034]" captionTargetId="figure-15@13.[151,1435,181,1037]" captionTargetPageId="13" captionText="FIGURE 6. Hind leg, female.—A, T. longicollis, Tsushima, Nagasaki; B, T. katayamai, holotype; C, T. fuscicornis, Inchon, Korea; D, T. nakanei, holotype of T. kaedei; E, T. contractus, Okayama-shi, Okayama; F, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621763" httpUri="https://zenodo.org/record/7621763/files/figure.png" pageId="21" pageNumber="22">6A</figureCitation>
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<figureCitation box="[510,571,357,383]" captionStart="FIGURE 7" captionStartId="14.[151,250,1592,1616]" captionTargetBox="[158,1430,190,1546]" captionTargetId="figure-20@14.[151,1435,181,1553]" captionTargetPageId="14" captionText="FIGURE 7. Wing, female.—A, T. apicalis, Kisofukushima, Nagano; B, T. bicinctus, holotype; C, T. bicinctus, Hôfu, Yamauchi; D, T. bicinctus, Fukuoka-shi, Fukuoka; E, T. longicollis, Okazaki, Aichi; F, T. longicollis, Osaka; G, H, T. katayamai, paratype; I, T. fuscicornis, Akigase, Saitama; J, T. fuscicornis, Kyoto." figureDoi="http://doi.org/10.5281/zenodo.7621766" httpUri="https://zenodo.org/record/7621766/files/figure.png" pageId="21" pageNumber="22">7E, F</figureCitation>
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<figureCitation box="[582,617,356,382]" captionStart="FIGURE 9" captionStartId="16.[151,250,1266,1290]" captionTargetBox="[158,1429,276,1239]" captionTargetId="figure-20@16.[151,1435,271,1243]" captionTargetPageId="16" captionText="FIGURE 9. Wing, male.—A, T. apicalis, Tsukuba, Ibaraki; B, T. longicollis, Muroto, Kochi; C, T. fuscicornis, Chitose, Hokkaido; D, T. nakanei, Noda, Chiba; E, T. contractus, Tatsuno, Hyogo; F, T. okinawensis, Amami-ôshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621770" httpUri="https://zenodo.org/record/7621770/files/figure.png" pageId="21" pageNumber="22">9B</figureCitation>
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<figureCitation box="[629,678,356,382]" captionStart="FIGURE 10" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[180,1408,190,1875]" captionTargetId="figure-15@17.[173,1414,181,1884]" captionTargetPageId="17" captionText="FIGURE 10. Posterior part of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, F, T. fuscicornis Inchon, Korea; G, T. fuscicornis, Akigase, Saitama; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621772" httpUri="https://zenodo.org/record/7621772/files/figure.png" pageId="21" pageNumber="22">10C</figureCitation>
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<figureCitation box="[690,737,356,382]" captionStart="FIGURE 11" captionStartId="18.[151,250,1565,1589]" captionTargetBox="[175,1383,189,1534]" captionTargetId="figure-20@18.[151,1435,181,1542]" captionTargetPageId="18" captionText="FIGURE 11. Posterior part of abdomen, female.—A, T. bicinctus, holotype; B, T. longicollis, Okazaki, Aichi; C, T. katayamai, holotype; D, T. fuscicornis, Inchon, Korea; E, T. contractus, Okayama-shi, Okayama; F, T. okinawensis, holotype." figureDoi="http://doi.org/10.5281/zenodo.7621776" httpUri="https://zenodo.org/record/7621776/files/figure.png" pageId="21" pageNumber="22">11B</figureCitation>
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<figureCitation box="[749,797,356,382]" captionStart="FIGURE 12" captionStartId="19.[151,250,1268,1292]" captionTargetBox="[159,1428,186,1239]" captionTargetId="figure-15@19.[151,1435,181,1244]" captionTargetPageId="19" captionText="FIGURE 12. Abdomen, male.—A, F, T. nakanei, Noda, Chiba; B, G, T. contractus, Tatsuno, Hyogo; C, T. apicalis, Tsukuba, Ibaraki; D, T. fuscicornis, Chitose, Hokkaido; E, T. longicollis, Muroto, Kochi; H, T. okinawensis, Amami-ôshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621778" httpUri="https://zenodo.org/record/7621778/files/figure.png" pageId="21" pageNumber="22">12E</figureCitation>
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<figureCitation box="[808,839,356,382]" captionStart="FIGURE 14" captionStartId="29.[151,250,1904,1928]" captionTargetBox="[170,1415,636,1877]" captionTargetId="figure-225@29.[170,1417,633,1880]" captionTargetPageId="29" captionText="FIGURE 14. Distribution of T. longicollis and T. fuscicornis in Japan based on the specimens examined (see Appendix for the collection records)." figureDoi="http://doi.org/10.5281/zenodo.7621785" httpUri="https://zenodo.org/record/7621785/files/figure.png" pageId="21" pageNumber="22">14</figureCitation>
; 
<tableCitation box="[851,973,356,382]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStartId-0="6.[151,239,980,1004]" captionStartId-1="7.[151,239,469,493]" captionStartId-2="7.[151,239,1276,1300]" captionStartId-3="20.[150,238,639,665]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="7" captionTargetPageId-3="20" captionText-0="TABLE 1. Variation of structural characters, female, 1." captionText-1="TABLE 2. Variation of structural characters, female, 2." captionText-2="TABLE 3. Variation of structural characters, male." captionText-3="TABLE 4. Number of specimens collected in each 10–11 days of April to November." httpUri-0="http://table.plazi.org/id/1FB4E1BC120442039FC7FC3BFD4C9581" httpUri-1="http://table.plazi.org/id/1FB4E1BC120542029FC7FE3AFD4D9781" httpUri-2="http://table.plazi.org/id/1FB4E1BC120542029FC7FB13FD1F9378" httpUri-3="http://table.plazi.org/id/1FB4E1BC121642119FC6FD90FB8E94F5" pageId="21" pageNumber="22" tableUuid-0="1FB4E1BC120442039FC7FC3BFD4C9581" tableUuid-1="1FB4E1BC120542029FC7FE3AFD4D9781" tableUuid-2="1FB4E1BC120542029FC7FB13FD1F9378" tableUuid-3="1FB4E1BC121642119FC6FD90FB8E94F5">Tables 1–4</tableCitation>
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<emphasis box="[151,340,424,448]" italics="true" pageId="21" pageNumber="22">Tremex longicollis</emphasis>
<bibRefCitation author="Konow, F. W." box="[347,528,424,448]" pageId="21" pageNumber="22" pagination="41 - 59" refId="ref25417" refString="Konow, F. W. (1896) Verschiedenes aus der Hymenopteren-Gruppe der Tenthrediniden. Wiener Entomologische Zeitung, 15, 41 - 59." type="journal article" year="1896">Konow, 1896: 45</bibRefCitation>
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<treatmentCitation author="Yano, M." box="[539,709,425,449]" page="120" pageId="21" pageNumber="22" year="1917">
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(“ 
<emphasis box="[736,850,424,448]" italics="true" pageId="21" pageNumber="22">longicolis”</emphasis>
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<treatmentCitation author="Yano, M." box="[873,1046,425,449]" page="473" pageId="21" pageNumber="22" year="1932">
<bibRefCitation author="Yano, M." box="[873,1046,425,449]" pageId="21" pageNumber="22" pagination="471 - 475" refId="ref29882" refString="Yano, M. (1932) Siricidae. In: Issiki, S., Ishii, T., Niijima, Y., Hori, H., Hozawa, S., Okamoto, H., Oguma, M., Kawada, A., Kawamura, T., Yokoyama, K., Takahashi, R., Uchida, S., Ueno, M., Kuwana, I., Kuwayama, S., Yano, M., Yamada, S., Maruge, N., Koizumi, M., Kojima, T., Esaki, T., Kinoshita, S., Kishida, K., Yuasa, H., Shiraki, T. &amp; Shinji, O., Iconographia Insectorum Japonicorum Editio Prima. Hokuryukan, Tokyo, pp. 471 - 475. [in Japanese]" type="book chapter" year="1932">Yano, 1932: 473</bibRefCitation>
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<treatmentCitation author="Gussakovskij, V. V." box="[1058,1302,424,448]" page="67" pageId="21" pageNumber="22" year="1935">
<bibRefCitation author="Gussakovskij, V. V." box="[1058,1302,424,448]" pageId="21" pageNumber="22" refId="ref24744" refString="Gussakovskij, V. V. (1935) Insectes Hymenopteres, Chalastogastra 1, Fauna SSSR. 2 (1). Izdatel'stvo Akademii Nauk SSSR, Moskva, Leningrad, 453 pp. [in Russian and German]" type="book" year="1935">Gussakovskij, 1935: 67</bibRefCitation>
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<treatmentCitation author="Esaki, T. &amp; Hori, H. &amp; Yasumatsu, K." page="330" pageId="21" pageNumber="22" year="1939">
<bibRefCitation author="Esaki, T. &amp; Hori, H. &amp; Yasumatsu, K." pageId="21" pageNumber="22" refId="ref24625" refString="Esaki, T., Hori, H. &amp; Yasumatsu, K. (1939) Insectorum Japonicorum Illustratio Iconographica Coloribus Ad Naturam Depicta. Sanseido, Tokyo, 485 pp. [in Japanese]" type="book" year="1939">
Esaki 
<emphasis box="[1378,1424,424,448]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
., 1939: 330
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; 
<treatmentCitation author="Takeuchi, K." box="[317,529,455,479]" page="193" pageId="21" pageNumber="22" year="1938">
<bibRefCitation author="Takeuchi, K." box="[317,529,455,479]" pageId="21" pageNumber="22" pagination="173 - 229" refId="ref28712" refString="Takeuchi, K. (1938) A systematic study on the suborder Symphyta (Hymenoptera) of the Japanese Empire (1). Tenthredo, 2, 173 - 229." type="journal article" year="1938">Takeuchi, 1938: 193</bibRefCitation>
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; 
<treatmentCitation author="Benson, R. B." box="[541,724,455,479]" page="30" pageId="21" pageNumber="22" year="1943">
<bibRefCitation author="Benson, R. B." box="[541,724,455,479]" pageId="21" pageNumber="22" pagination="27 - 51" refId="ref24461" refString="Benson, R. B. (1943) Studies in Siricidae, especially of Europe and southern Asia (Hymenoptera; Symphyta). Bulletin of Entomological Research, 34 (1), 27 - 51. https: // doi. org / 10.1016 / 0165 - 3806 (90) 90255 - W" type="journal article" year="1943">Benson, 1943: 30</bibRefCitation>
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; 
<treatmentCitation author="Maa, T. - C." box="[736,901,455,479]" page="159" pageId="21" pageNumber="22" year="1949">
<bibRefCitation author="Maa, T. - C." box="[736,901,455,479]" pageId="21" pageNumber="22" pagination="11 - 189" refId="ref25790" refString="Maa, T. - C. (1949) A synopsis of Asiatic Siricoidea with notes on certain exotic and fossil forms. Notes d'Entomologie Chinoise, 13, 11 - 189." type="journal article" year="1949">Maa, 1949: 159</bibRefCitation>
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; 
<treatmentCitation author="Takeuchi, K." box="[912,1111,455,479]" page="47" pageId="21" pageNumber="22" year="1949">
<bibRefCitation author="Takeuchi, K." box="[912,1111,455,479]" pageId="21" pageNumber="22" pagination="47 - 50" refId="ref28745" refString="Takeuchi, K. (1949) A list of the food-plants of Japanese sawflies. Transactions of the Kansai Entomological Society, Osaka, 14, 47 - 50." type="journal article" year="1949">Takeuchi, 1949: 47</bibRefCitation>
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<treatmentCitation author="Miyatake, M." box="[1123,1326,455,479]" page="38" pageId="21" pageNumber="22" year="1950">
<bibRefCitation author="Miyatake, M." box="[1123,1326,455,479]" pageId="21" pageNumber="22" pagination="38 - 39" refId="ref26137" refString="Miyatake, M. (1950) A contribution to the Siricoidea of Shikoku (Hymenoptera). Transactions of the Shikoku Entomological Society, Matsuyama, 1, 38 - 39." type="journal article" year="1950">Miyatake, 1950: 38</bibRefCitation>
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<treatmentCitation author="Takeuchi, K." page="1333" pageId="21" pageNumber="22" year="1950">
<bibRefCitation author="Takeuchi, K." pageId="21" pageNumber="22" pagination="1332 - 1334" refId="ref28776" refString="Takeuchi, K. (1950) Siricidae. In: Ishii, T., Esaki, T., Kinoshita, S., Shiraki, T., Uchida, S., Kawamura, T., Kuwayama, S. &amp; Yuasa, H. (Eds.), Iconographia Insectorum Japonicorum Editio Secunda Reformata. Hokuryukan, Tokyo, pp. 1332 - 1334. [in Japanese]" type="book chapter" year="1950">Takeuchi, 1950: 1333</bibRefCitation>
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<treatmentCitation author="Takeuchi, K." box="[335,539,486,510]" page="2" pageId="21" pageNumber="22" year="1955">
<bibRefCitation author="Takeuchi, K." box="[335,539,486,510]" pageId="21" pageNumber="22" pagination="1 - 9" refId="ref28852" refString="Takeuchi, K. (1955 a) Siricidae of Japan (Hymenoptera). Akitu, Kyoto, 4, 1 - 9. [in Japanese with English abstract]" type="journal article" year="1955">Takeuchi, 1955a: 2</bibRefCitation>
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, 7; 
<treatmentCitation author="Takeuchi, K." box="[584,815,486,510]" page="115" pageId="21" pageNumber="22" year="1955">
<bibRefCitation author="Takeuchi, K." box="[584,815,486,510]" pageId="21" pageNumber="22" refId="ref28884" refString="Takeuchi, K. (1955 b) Coloured Illustrations of the Insects of Japan. Vol. II. Hoikusha, Osaka, 190 pp., 68 pls. [in Japanese]" type="book" year="1955">Takeuchi, 1955b: 115</bibRefCitation>
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; 
<treatmentCitation author="Okutani, T." box="[829,1031,486,510]" page="554" pageId="21" pageNumber="22" year="1959">
<bibRefCitation author="Okutani, T." box="[829,1031,486,510]" pageId="21" pageNumber="22" pagination="548 - 582" refId="ref26904" refString="Okutani, T. (1959) [Symphyta]. In: Esaki, T., Ishii, T., Kawada, A., Shiraki T. &amp; Yuasa, H. (Eds.), Illustrated Insect Larvae of Japan. Hokuryukan, Tokyo, pp. 548 - 582. [in Japanese]" type="book chapter" year="1959">Okutani 1959: 554</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Kondo, T." box="[1046,1215,486,510]" page="4" pageId="21" pageNumber="22" year="1961">
<bibRefCitation author="Kondo, T." box="[1046,1215,486,510]" pageId="21" pageNumber="22" pagination="4 - 5" refId="ref25239" refString="Kondo, T. (1961) [A record of Tremex longicollis with a fragmentary note on its biology]. Suzumushi, Kurashiki, 11 (2), 4 - 5. [in Japanese]" type="journal article" year="1961">Kondo, 1961: 4</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Okutani, T." box="[1231,1432,486,510]" page="343" pageId="21" pageNumber="22" year="1962">
<bibRefCitation author="Okutani, T." box="[1231,1432,486,510]" pageId="21" pageNumber="22" pagination="342 - 343" refId="ref26965" refString="Okutani, T. (1962) [On the larvae of Siricidae]. Transactions of the 72 nd Meeting of the Japanese Forestry Society, 1962, 342 - 343. [in Japanese]" type="journal article" year="1962">Okutani 1962: 343</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Takeuchi, K." box="[199,383,516,541]" page="3" pageId="21" pageNumber="22" year="1962">
<bibRefCitation author="Takeuchi, K." box="[199,383,516,541]" pageId="21" pageNumber="22" refId="ref28919" refString="Takeuchi, K. (1962) Insecta Japonica. Series 2. Part 4. Hymenoptera: Siricidae. Hokuryukan, Tokyo, 2 + 12 pp., 1 pl. [in Japanese with English summary]" type="book" year="1962">Takeuchi, 1962: 3</bibRefCitation>
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, 10; 
<treatmentCitation author="Kim, C. - W." box="[434,597,516,540]" page="295" pageId="21" pageNumber="22" year="1963">
<bibRefCitation author="Kim, C. - W." box="[434,597,516,540]" pageId="21" pageNumber="22" pagination="243 - 374" refId="ref25110" refString="Kim, C. - W. (1963) Hymenoptera of Korea. Humanities and Sciences, Natural Science, Korea University, 6, 243 - 374. [in Korean]" type="journal article" year="1963">Kim, 1963: 295</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Okutani, T." box="[608,789,516,540]" page="25" pageId="21" pageNumber="22" year="1963">
<bibRefCitation author="Okutani, T." box="[608,789,516,540]" pageId="21" pageNumber="22" pagination="23 - 27" refId="ref27002" refString="Okutani, T. (1963) Description of the larvae of horntails. Science Reports of the Hyogo University of Agriculture, Series Agricultural Biology, 6 (1), 23 - 27." type="journal article" year="1963">Okutani 1963: 25</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Okutani, T." box="[800,987,516,541]" page="44" pageId="21" pageNumber="22" year="1967">
<bibRefCitation author="Okutani, T." box="[800,987,516,541]" pageId="21" pageNumber="22" pagination="43 - 49" refId="ref27093" refString="Okutani, T. (1967) Food plants of Japanese Symphyta (I). Japanese Journal of Applied Entomology and Zoology, 11, 43 - 49. [in Japanese with English abstract] https: // doi. org / 10.1303 / jjaez. 11.90" type="journal article" year="1967">Okutani, 1967: 44</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Kim, C. - W." box="[998,1148,516,541]" page="85" pageId="21" pageNumber="22" year="1971">
<bibRefCitation author="Kim, C. - W." box="[998,1148,516,541]" pageId="21" pageNumber="22" pagination="85 - 86" refId="ref25144" refString="Kim, C. - W. (1971) Tremex apicalis Matsumura as a host of an Ichneumon-fly, Megarhyssa jezoensis Matsumura. Korean Journal of Zoology, 14, 85 - 86. [in Korean with English summary]" type="journal article" year="1971">Kim, 1971: 85</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Okutani, T." box="[1159,1346,516,541]" page="18" pageId="21" pageNumber="22" year="1971">
<bibRefCitation author="Okutani, T." box="[1159,1346,516,541]" pageId="21" pageNumber="22" pagination="17 - 19" refId="ref27190" refString="Okutani, T. (1971) [Symphyta of Japan II]. Nature and Insects, Tokyo, 6 (12), 17 - 19. [in Japanese]" type="journal article" year="1971">Okutani, 1971: 18</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Tanaka, C." page="202" pageId="21" pageNumber="22" year="1971">
<bibRefCitation author="Tanaka, C." pageId="21" pageNumber="22" refId="ref28959" refString="Tanaka, C. (1971) A List of Insects of Toyama Prefecture. Privately published, Toyama, 242 pp. [in Japanese]" type="book" year="1971">Tanaka, 1971: 202</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Kondo, T. &amp; Miyake, M." box="[316,604,547,571]" page="15" pageId="21" pageNumber="22" year="1974">
<bibRefCitation author="Kondo, T. &amp; Miyake, M." box="[316,604,547,571]" pageId="21" pageNumber="22" pagination="15 - 18" refId="ref25278" refString="Kondo, T. &amp; Miyake, M. (1974) [A fragmentary note on the biology of Tremex longicollis]. Nature and Insects, Tokyo, 9 (14), 15 - 18. [in Japanese]" type="journal article" year="1974">Kondo &amp; Miyake, 1974: 15</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Kondo, T. &amp; Miyake, M." box="[615,889,547,571]" page="3" pageId="21" pageNumber="22" year="1976">
<bibRefCitation author="Kondo, T. &amp; Miyake, M." box="[615,889,547,571]" pageId="21" pageNumber="22" pagination="1 - 13" refId="ref25321" refString="Kondo, T. &amp; Miyake, M. (1976) [Symphyta (Hymenoptera) of Okayama Prefecture, 2]. Suzumushi, Kurashiki, 113, 1 - 13. [in Japanese]" type="journal article" year="1976">Kondo &amp; Miyake, 1976: 3</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Okutani, T." box="[901,1103,547,571]" page="286" pageId="21" pageNumber="22" year="1977">
<bibRefCitation author="Okutani, T." box="[901,1103,547,571]" pageId="21" pageNumber="22" pagination="285 - 286" refId="ref27260" refString="Okutani, T. (1977) Siricidae. In: Ito, S., Okutani, T. &amp; Hiura, I. (Eds.), Colored Illustrations of the Insects of Japan. Vol. II. Hoikusha, Osaka, pp. 285 - 286, pl. 54. [in Japanese]" type="journal article" year="1977">Okutani, 1977: 286</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Smith, D. R." box="[1114,1281,547,571]" page="99" pageId="21" pageNumber="22" year="1978">
<bibRefCitation author="Smith, D. R." box="[1114,1281,547,571]" pageId="21" pageNumber="22" pagination="43 - 128" refId="ref28065" refString="Smith, D. R. (1978) Suborder Symphyta (Xyelidae, Pararchexyelidae, Parapamphiliidae, Xyelydidae, Karatavitidae, Gigasiricidae, Sepulcidae, Pseudosiricidae, Anaxyelidae, Siricidae, Xiphydriidae, Paroryssidae, Xyelotomidae, Blasticotomidae, Pergidae). In: Vecht, J. &amp; Shenefelt, R. D. (Eds.), Hymenopterorum Catalogues Part 14. Dr. W. Junk B. V. Publishers, Hague, pp. 43 - 128." type="book chapter" year="1978">Smith, 1978: 99</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Tanaka, C." page="459" pageId="21" pageNumber="22" year="1979">
<bibRefCitation author="Tanaka, C." pageId="21" pageNumber="22" pagination="459 - 489" refId="ref28986" refString="Tanaka, C. (1979) Hymenoptera. In: Toyama-ken Konchu Kenkyu-kai (Ed.), [Insects of Toyama Prefecture]. Toyama Prefectural Government, Toyama, pp. 459 - 489. [in Japanese]" type="book chapter" year="1979">Tanaka, 1979: 459</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Shimoyama, K." box="[252,466,578,602]" page="7" pageId="21" pageNumber="22" year="1986">
<bibRefCitation author="Shimoyama, K." box="[252,466,578,602]" pageId="21" pageNumber="22" pagination="7 - 8" refId="ref27731" refString="Shimoyama, K. (1986) Siricoidea (Hymenoptera) of Aomori Prefecture. Journal of Aomori-ken Biological Society, 23, 7 - 8. [in Japanese]" type="journal article" year="1986">Shimoyama, 1986: 7</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Abe, M. &amp; Togashi, I." box="[477,753,578,602]" page="558" pageId="21" pageNumber="22" year="1989">
<bibRefCitation author="Abe, M. &amp; Togashi, I." box="[477,753,578,602]" pageId="21" pageNumber="22" pagination="558 - 559" refId="ref24237" refString="Abe, M. &amp; Togashi, I. (1989) Siricidae. In: Hirashima, Y. (Ed.), A Check List of Japanese Insects. Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka, pp. 558 - 559. [in Japanese]" type="book chapter" year="1989">Abe &amp; Togashi, 1989: 558</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Xiao, G. &amp; Huang, X. &amp; Zhou, S. &amp; Wu, J. &amp; Zhang, P." box="[764,979,578,602]" page="56" pageId="21" pageNumber="22" year="1992">
<bibRefCitation author="Xiao, G. &amp; Huang, X. &amp; Zhou, S. &amp; Wu, J. &amp; Zhang, P." box="[764,979,578,602]" pageId="21" pageNumber="22" refId="ref29568" refString="Xiao, G., Huang, X., Zhou, S., Wu, J. &amp; Zhang, P. (1992) Economic Sawfly Fauna of China. Tianze Eldonejo, Beijing, 4 + 8 + 221 pp. [in Chinese]" type="book" year="1992">
Xiao 
<emphasis box="[821,866,578,602]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
., 1992: 56
</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Tabata, M. &amp; Abe, Y." box="[990,1252,578,602]" page="447" pageId="21" pageNumber="22" year="1995">
<bibRefCitation author="Tabata, M. &amp; Abe, Y." box="[990,1252,578,602]" pageId="21" pageNumber="22" pagination="447 - 450" refId="ref28433" refString="Tabata, M. &amp; Abe, Y. (1995) Cerrena unicolor isolated from the mycangia of a horntail, Tremex longicollis, in Kochi Prefecture, Japan. Mycoscience, 36, 447 - 450. https: // doi. org / 10.1007 / BF 02268630" type="journal article" year="1995">Tabata &amp; Abe, 1995: 447</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Sakamoto, M." page="25" pageId="21" pageNumber="22" year="1997">
<bibRefCitation author="Sakamoto, M." pageId="21" pageNumber="22" pagination="23 - 34" refId="ref27594" refString="Sakamoto, M. (1997) The siricid woodwasps (Hymenoptera, Symphyta, Siricidae) of Hiroshima Prefecture. Journal of the Hiba Society of Natural History, 177, 23 - 34. [in Japanese with English abstract]" type="journal article" year="1997">Sakamoto, 1997: 25</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Lee, J. - W. &amp; Ryu, S. - M. &amp; Chung, D. - K." box="[238,449,608,633]" page="299" pageId="21" pageNumber="22" year="1998">
<bibRefCitation author="Lee, J. - W. &amp; Ryu, S. - M. &amp; Chung, D. - K." box="[238,449,608,633]" pageId="21" pageNumber="22" pagination="297 - 302" refId="ref25717" refString="Lee, J. - W., Ryu, S. - M. &amp; Chung, D. - K. (1998) Taxonomy of the genus Tremex Jurine (Hymenoptera: Siricidae) from Korea, with descriptions of one new species and one unrecorded species. Korean Journal of Biological Sciences, 2, 297 - 302. https: // doi. org / 10.1080 / 12265071.1998.9647422" type="journal article" year="1998">
Lee 
<emphasis box="[282,326,608,632]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
., 1998: 299
</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Nagase, H." box="[459,664,608,633]" page="1254" pageId="21" pageNumber="22" year="2004">
<bibRefCitation author="Nagase, H." box="[459,664,608,633]" pageId="21" pageNumber="22" pagination="1241 - 1326" refId="ref26245" refString="Nagase, H. (2004) Hymenoptera (excl. Formicidae). In: Insect Fauna of Kanagawa. Kanagawa Konchu Danwakai, Odawara, pp. 1241 - 1326. [in Japanese]" type="book chapter" year="2004">Nagase, 2004: 1254</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Naito, T. &amp; Yoshida, H. &amp; Nakamine, H. &amp; Morita, T. &amp; Ikeda, T. &amp; Suzuki, H. &amp; Nakanishi, A." box="[674,891,608,633]" page="68" pageId="21" pageNumber="22" year="2004">
<bibRefCitation author="Naito, T. &amp; Yoshida, H. &amp; Nakamine, H. &amp; Morita, T. &amp; Ikeda, T. &amp; Suzuki, H. &amp; Nakanishi, A." box="[674,891,608,633]" pageId="21" pageNumber="22" pagination="1 - 2" refId="ref26562" refString="Naito, T., Yoshida, H., Nakamine, H., Morita, T., Ikeda, T., Suzuki, H. &amp; Nakanishi, A. (2004) Species diversity of sawflies in Hyogo Prefecture, central Japan. Museum of Nature and Human Activities, Hyogo, Monograph of Natural History and Environmental Science, 1, 1 - 2, pls. 1 - 10 + 1 - 85. [in Japanese]" type="journal article" year="2004">
Naito 
<emphasis box="[736,787,608,632]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
, 2004: 68
</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Yoshida, H." box="[900,1099,608,632]" page="105" pageId="21" pageNumber="22" year="2006">
<bibRefCitation author="Yoshida, H." box="[900,1099,608,632]" pageId="21" pageNumber="22" refId="ref30082" refString="Yoshida, H. (2006) Symphyta (Hymenoptera) of Osaka Prefecture, Japan. West Japan Hymenopterists' Club, Kakogawa, 24 pls., 128 pp. [in Japanese]" type="book" year="2006">Yoshida, 2006: 105</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Nagase, H." box="[1109,1301,609,633]" page="291" pageId="21" pageNumber="22" year="2007">
<bibRefCitation author="Nagase, H." box="[1109,1301,609,633]" pageId="21" pageNumber="22" pagination="286 - 310" refId="ref26282" refString="Nagase, H. (2007) Hymenoptera. In: Research Group of the Tanzawa Mountains (Ed.), Results of the Scientific Research on the Tanzawa Mountains, Appendix: A List of Plants and Animals. Hiraoka Environmental Science Laboratory, Sagamihara, pp. 286 - 310. [in Japanese]" type="book chapter" year="2007">Nagase, 2007: 291</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Yamazaki, K. &amp; Matsumoto, R." page="313" pageId="21" pageNumber="22" year="2009">
<bibRefCitation author="Yamazaki, K. &amp; Matsumoto, R." pageId="21" pageNumber="22" pagination="313 - 315" refId="ref29741" refString="Yamazaki, K. &amp; Matsumoto, R. (2009) Predation on the woodwasp Tremex longicollis Konow (Hymenoptera: Siricidae) and its parasitoid Megarhyssa jezoensis (Matsukura) (Hymenoptera: Ichneumonidae) adults during oviposition. Journal of Asia- Pacific Entomology, 12, 313 - 315. https: // doi. org / 10.1016 / j. aspen. 2009.05.003" type="journal article" year="2009">Yamazaki &amp; Matsumoto, 2009: 313</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Taeger, A. &amp; Blank, S. M. &amp; Liston, A. D." box="[443,688,639,664]" page="108" pageId="21" pageNumber="22" year="2010">
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Taeger 
<emphasis box="[519,564,639,663]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
., 2010: 108
</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Funamoto, D." box="[699,907,639,663]" page="10" pageId="21" pageNumber="22" year="2010">
<bibRefCitation author="Funamoto, D." box="[699,907,639,663]" pageId="21" pageNumber="22" pagination="10" refId="ref24715" refString="Funamoto, D. (2010) [Records of two Tremex woodwasps collected in Kanagawa Prefecture]. Kanagawa-Chuho, 171, 10. [in Japanese]" type="journal article" year="2010">Funamoto, 2010: 10</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Tanaka, S." box="[918,1096,639,663]" page="38" pageId="21" pageNumber="22" year="2015">
<bibRefCitation author="Tanaka, S." box="[918,1096,639,663]" pageId="21" pageNumber="22" pagination="27 - 44" refId="ref29028" refString="Tanaka, S. (2015) [Catalogue of sawflies and woodwasps of Yamaguchi Prefecture]. Bulletin of the Firefly Museum of Toyata Town, (7), 27 - 44. [in Japanese]" type="journal article" year="2015">Tanaka, 2015: 38</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Nagase, H. &amp; Watanabe, K." box="[1107,1432,639,664]" page="950" pageId="21" pageNumber="22" year="2018">
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</treatmentCitation>
; 
<treatmentCitation author="Higurashi, T." box="[199,404,670,694]" page="93" pageId="21" pageNumber="22" year="2019">
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</treatmentCitation>
; 
<treatmentCitation author="Lee, J. - W. &amp; Choi, J. - K. &amp; Park, B." box="[415,614,670,694]" page="14" pageId="21" pageNumber="22" year="2019">
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Lee 
<emphasis box="[459,503,670,694]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
., 2019: 14
</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Naito, T." box="[625,786,670,694]" page="20" pageId="21" pageNumber="22" year="2019">
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</treatmentCitation>
; 
<treatmentCitation author="Naito, T." box="[797,971,670,694]" page="467" pageId="21" pageNumber="22" year="2020">
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</treatmentCitation>
; 
<treatmentCitation author="Ogata, K." box="[981,1159,670,694]" page="186" pageId="21" pageNumber="22" year="2021">
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</treatmentCitation>
.
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<paragraph blockId="21.[151,1437,424,817]" pageId="21" pageNumber="22">
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<treatmentCitation author="Marlatt, C. L." box="[151,493,700,725]" page="49" pageId="21" pageNumber="22" year="1898">
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<emphasis box="[151,300,700,724]" italics="true" pageId="21" pageNumber="22">Tremex similis</emphasis>
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(synonym of 
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<emphasis box="[836,943,700,724]" italics="true" pageId="21" pageNumber="22">T. apicalis</emphasis>
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); 
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; 
<treatmentCitation author="Matsumura, S." box="[1207,1432,700,725]" page="59" pageId="21" pageNumber="22" year="1930">
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; 
<treatmentCitation author="Matsumura, S." box="[199,419,731,755]" page="79" pageId="21" pageNumber="22" year="1931">
<bibRefCitation author="Matsumura, S." box="[199,419,731,755]" pageId="21" pageNumber="22" refId="ref26031" refString="Matsumura, S. (1931) 6000 Illustrated Insects of Japan-Empire. Tokoshoin, Tokyo, 1497 + 23 pp. [in Japanese]" type="book" year="1931">Matsumura, 1931: 79</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Matsumura, S." box="[430,676,731,755]" page="952" pageId="21" pageNumber="22" year="1932">
<bibRefCitation author="Matsumura, S." box="[430,676,731,755]" pageId="21" pageNumber="22" refId="ref26102" refString="Matsumura, S. (1932 b) Conspectus of Japanese Injurious Insects. Meijitosho, Tokyo, 62 pls., 3 + 51 + 971 + 115 pp. [in Japanese]" type="book" year="1932">Matsumura, 1932b: 952</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation author="Smith, D. R." box="[686,851,731,755]" page="99" pageId="21" pageNumber="22" year="1978">
<bibRefCitation author="Smith, D. R." box="[686,851,731,755]" pageId="21" pageNumber="22" pagination="43 - 128" refId="ref28065" refString="Smith, D. R. (1978) Suborder Symphyta (Xyelidae, Pararchexyelidae, Parapamphiliidae, Xyelydidae, Karatavitidae, Gigasiricidae, Sepulcidae, Pseudosiricidae, Anaxyelidae, Siricidae, Xiphydriidae, Paroryssidae, Xyelotomidae, Blasticotomidae, Pergidae). In: Vecht, J. &amp; Shenefelt, R. D. (Eds.), Hymenopterorum Catalogues Part 14. Dr. W. Junk B. V. Publishers, Hague, pp. 43 - 128." type="book chapter" year="1978">Smith, 1978: 99</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="21.[151,1437,424,817]" box="[151,957,762,786]" pageId="21" pageNumber="22">
<treatmentCitationGroup box="[151,957,762,786]" pageId="21" pageNumber="22">
<treatmentCitation author="Togashi, I." box="[151,552,762,786]" page="244" pageId="21" pageNumber="22" year="1965">
<taxonomicName authority=": Togashi, 1965: 244" authorityName=": Togashi" authorityPageNumber="244" authorityYear="1965" box="[151,552,762,786]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fuscicornis">
<emphasis box="[151,342,762,786]" italics="true" pageId="21" pageNumber="22">Tremex fuscicornis</emphasis>
: 
<bibRefCitation author="Togashi, I." box="[355,552,762,786]" pageId="21" pageNumber="22" pagination="122" refId="ref29136" refString="Togashi, I. (1965) Siricidae. In: Asahina, S., Ishihara, T. &amp; Yasumatsu, K. (Eds.), Iconographia Insectorum Japonicorum Colore naturali edita. Vol. III. Hokuryukan, Tokyo, pp. 244, pl. 122. [in Japanese]" type="book chapter" year="1965">Togashi, 1965: 244</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, plate 122, fig. 15. Not 
<bibRefCitation author="Fabricius, J. C." box="[792,953,762,786]" pageId="21" pageNumber="22" refId="ref24662" refString="Fabricius, J. C. (1787) Mantissa Insectorum sistens eorum species nuper detectas adiectis characteribus genericis differentiis specificis emendationibus observationibus. Vol. 1. C. G. Proft, Hafniae, 348 pp. https: // doi. org / 10.5962 / bhl. title. 11657" type="book" year="1787">Fabricius, 1787</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="description">
<paragraph blockId="21.[151,1437,424,817]" box="[151,911,792,817]" pageId="21" pageNumber="22">
See 
<bibRefCitation author="Smith, D. R." box="[194,328,792,817]" pageId="21" pageNumber="22" pagination="43 - 128" refId="ref28065" refString="Smith, D. R. (1978) Suborder Symphyta (Xyelidae, Pararchexyelidae, Parapamphiliidae, Xyelydidae, Karatavitidae, Gigasiricidae, Sepulcidae, Pseudosiricidae, Anaxyelidae, Siricidae, Xiphydriidae, Paroryssidae, Xyelotomidae, Blasticotomidae, Pergidae). In: Vecht, J. &amp; Shenefelt, R. D. (Eds.), Hymenopterorum Catalogues Part 14. Dr. W. Junk B. V. Publishers, Hague, pp. 43 - 128." type="book chapter" year="1978">Smith (1978)</bibRefCitation>
and 
<bibRefCitation author="Lee, J. - W. &amp; Choi, J. - K. &amp; Park, B." box="[378,546,792,817]" pageId="21" pageNumber="22" pagination="1 - 96" refId="ref25673" refString="Lee, J. - W., Choi, J. - K. &amp; Park, B. (2019) Synoptic list of Symphyta (Hymenoptera) in Korea. Journal of Species Research, 8, 1 - 96." type="journal article" year="2019">
Lee 
<emphasis box="[422,467,792,816]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
. (2019)
</bibRefCitation>
for more synonyms and references.
</paragraph>
<paragraph blockId="21.[151,1437,862,2041]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[151,420,862,889]" pageId="21" pageNumber="22">Diagnostic characters.</emphasis>
<emphasis box="[430,514,863,889]" italics="true" pageId="21" pageNumber="22">Female</emphasis>
. Length without ovipositor 
<quantity box="[839,965,862,889]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.5" metricValueMin="2.2" pageId="21" pageNumber="22" unit="mm" value="28.5" valueMax="35.0" valueMin="22.0">22–35 mm</quantity>
. Head pale brown, usually with blackish mark across ocellar area. Antenna blackish brown, with two to four basal and one apical antennomeres pale brown (
<figureCitation box="[159,250,934,961]" captionStart="FIGURE 5" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[155,1432,565,1761]" captionTargetId="figure-186@12.[151,1435,561,1764]" captionTargetPageId="12" captionText="FIGURE 5. Antenna, female (A–G, I) and male (H, J–N).—A. T. apicalis, Minamichitose, Hokkaido; B, T. bicinctus, holotype; C, T. longicollis, Toyota, Aichi; D, T. fuscicornis, Inchon, Korea; E, T. nakanei, Ôginosen, Tottori; F, T. contractus, Okayamashi, Okayama; G, T. okinawensis, holotype; H. T. apicalis, Tsukuba, Ibaraki; I, T. katayamai, holotype; J, T. longicollis, Muroto, Kochi; K, T. fuscicornis, Shikotsu-ko, Hokkaido; L, T. nakanei, Noda, Chiba; M, T. contractus, Tatsuno, Hyogo; N, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621761" httpUri="https://zenodo.org/record/7621761/files/figure.png" pageId="21" pageNumber="22">Fig. 5C</figureCitation>
). Thorax black, at least dorsal half of pronotum (usually without blackish line medially) and sometimes propleuron, parascutal lobe and mesoscutellum marked with pale brown. Legs brown, most of coxae, trochanters, anterior and dorsal surfaces of mid femur, hind femur, dorsal parts of hind tibia and hind tarsomere 1 blackish brown to black (
<figureCitation box="[255,345,1043,1069]" captionStart="FIGURE 6" captionStartId="13.[151,250,1061,1085]" captionTargetBox="[155,1432,184,1034]" captionTargetId="figure-15@13.[151,1435,181,1037]" captionTargetPageId="13" captionText="FIGURE 6. Hind leg, female.—A, T. longicollis, Tsushima, Nagasaki; B, T. katayamai, holotype; C, T. fuscicornis, Inchon, Korea; D, T. nakanei, holotype of T. kaedei; E, T. contractus, Okayama-shi, Okayama; F, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621763" httpUri="https://zenodo.org/record/7621763/files/figure.png" pageId="21" pageNumber="22">Fig. 6A</figureCitation>
). Forewing pale brownish hyaline, without distinct infuscation (
<figureCitation box="[1061,1177,1043,1069]" captionStart="FIGURE 7" captionStartId="14.[151,250,1592,1616]" captionTargetBox="[158,1430,190,1546]" captionTargetId="figure-20@14.[151,1435,181,1553]" captionTargetPageId="14" captionText="FIGURE 7. Wing, female.—A, T. apicalis, Kisofukushima, Nagano; B, T. bicinctus, holotype; C, T. bicinctus, Hôfu, Yamauchi; D, T. bicinctus, Fukuoka-shi, Fukuoka; E, T. longicollis, Okazaki, Aichi; F, T. longicollis, Osaka; G, H, T. katayamai, paratype; I, T. fuscicornis, Akigase, Saitama; J, T. fuscicornis, Kyoto." figureDoi="http://doi.org/10.5281/zenodo.7621766" httpUri="https://zenodo.org/record/7621766/files/figure.png" pageId="21" pageNumber="22">Fig. 7E, F</figureCitation>
). Abdomen black, with following pale brown: Most of tergum 2, anterior and often posterior margins of terga 3–6 (tergum 3 often almost entirely), tergum 8 (except anterolateral corner and large roughly heart-shaped black mark at middle), posterior half of tergum 9 (without black area along posterior margin), including entire precornal basin, and cornus. Antenna with 14–16 antennomeres. Pronotum (
<figureCitation box="[525,611,1187,1213]" captionStart="FIGURE 3" captionStartId="10.[151,250,1870,1894]" captionTargetBox="[157,1430,190,1838]" captionTargetId="figure-20@10.[151,1435,181,1846]" captionTargetPageId="10" captionText="FIGURE 3. Head, thorax and base of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, T. fuscicornis, Schwedt, Germany, F, G, T. fuscicornis, Inchon, Korea; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621757" httpUri="https://zenodo.org/record/7621757/files/figure.png" pageId="21" pageNumber="22">Fig. 3C</figureCitation>
) long, MPL 0.74–0.95 (average 0.85) times as long as OOCL. Hind tibia 1.00–1.12 (average 1.07) times as long as hind tarsomere 1; dorsal margin of hind tibia straight or very shallowly concave (
<figureCitation box="[262,356,1259,1285]" captionStart="FIGURE 6" captionStartId="13.[151,250,1061,1085]" captionTargetBox="[155,1432,184,1034]" captionTargetId="figure-15@13.[151,1435,181,1037]" captionTargetPageId="13" captionText="FIGURE 6. Hind leg, female.—A, T. longicollis, Tsushima, Nagasaki; B, T. katayamai, holotype; C, T. fuscicornis, Inchon, Korea; D, T. nakanei, holotype of T. kaedei; E, T. contractus, Okayama-shi, Okayama; F, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621763" httpUri="https://zenodo.org/record/7621763/files/figure.png" pageId="21" pageNumber="22">Fig. 6A</figureCitation>
). Forewing 1.10–1.38 (average 1.2) times as long as ovipositor sheath. Abdominal tergum 8 0.94–1.29 (average 1.10) times as long as terga 5–7 combined; precornal basin long, its length 0.74–0.98 (average 0.86) times as long as wide; ovipositor sheath 2.24–2.74 (average 2.55) times as long as apical sheath.
</paragraph>
<paragraph blockId="21.[151,1437,862,2041]" pageId="21" pageNumber="22">
<emphasis box="[199,257,1367,1393]" italics="true" pageId="21" pageNumber="22">Male</emphasis>
. Length 
<quantity box="[354,435,1367,1393]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="21" pageNumber="22" unit="mm" value="21.0">21 mm</quantity>
. Head, antenna, thorax and legs black except as follows: postocular area and temple (gena) entirely pale brown. Antenna with scape and pedicel pale brown (
<figureCitation box="[900,982,1402,1429]" captionStart="FIGURE 5" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[155,1432,565,1761]" captionTargetId="figure-186@12.[151,1435,561,1764]" captionTargetPageId="12" captionText="FIGURE 5. Antenna, female (A–G, I) and male (H, J–N).—A. T. apicalis, Minamichitose, Hokkaido; B, T. bicinctus, holotype; C, T. longicollis, Toyota, Aichi; D, T. fuscicornis, Inchon, Korea; E, T. nakanei, Ôginosen, Tottori; F, T. contractus, Okayamashi, Okayama; G, T. okinawensis, holotype; H. T. apicalis, Tsukuba, Ibaraki; I, T. katayamai, holotype; J, T. longicollis, Muroto, Kochi; K, T. fuscicornis, Shikotsu-ko, Hokkaido; L, T. nakanei, Noda, Chiba; M, T. contractus, Tatsuno, Hyogo; N, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621761" httpUri="https://zenodo.org/record/7621761/files/figure.png" pageId="21" pageNumber="22">Fig. 5J</figureCitation>
). Lateral sunken area of parascutal lobe pale brown. Fore leg with posterior half of coxa, femur, tibia and tarsus pale brown; mid leg with part of femur, ventral surface of tibia and apical tarsomere pale brown; hind leg with most of femur and apical tarsomere pale brown. Wings yellowish hyaline, with brownish veins (
<figureCitation box="[782,873,1511,1537]" captionStart="FIGURE 9" captionStartId="16.[151,250,1266,1290]" captionTargetBox="[158,1429,276,1239]" captionTargetId="figure-20@16.[151,1435,271,1243]" captionTargetPageId="16" captionText="FIGURE 9. Wing, male.—A, T. apicalis, Tsukuba, Ibaraki; B, T. longicollis, Muroto, Kochi; C, T. fuscicornis, Chitose, Hokkaido; D, T. nakanei, Noda, Chiba; E, T. contractus, Tatsuno, Hyogo; F, T. okinawensis, Amami-ôshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621770" httpUri="https://zenodo.org/record/7621770/files/figure.png" pageId="21" pageNumber="22">Fig. 9B</figureCitation>
). Abdomen pale brown, with entire tergum 1 and narrow anterior lateral margins of each segment blackish. Antenna with 14 antennomeres. Pronotum (
<figureCitation box="[1281,1365,1547,1573]" captionStart="FIGURE 4" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[158,1429,188,1197]" captionTargetId="figure-15@11.[151,1435,181,1203]" captionTargetPageId="11" captionText="FIGURE 4. Head, thorax and base of abdomen, male.—A, B, T. apicalis, Minamichitose, Hokkaido; C, D, T. fuscicornis, Shikotsu-ko, Hokkaido; E, T. longicollis, Muroto, Kochi; F, T. nakanei, Noda, Chiba; G, T. contractus, Okayama-shi, Okayama; H, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621759" httpUri="https://zenodo.org/record/7621759/files/figure.png" pageId="21" pageNumber="22">Fig. 4E</figureCitation>
) long, MPL about 0.91 times as long as OOCL. Hind tibia short, 1.19 times as long as hind tarsomere 1.
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="materials_examined">
<paragraph blockId="21.[151,1437,862,2041]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[199,434,1618,1644]" pageId="21" pageNumber="22">Material examined.</emphasis>
<materialsCitation box="[442,737,1619,1647]" country="Taiwan" location="Taiwan" pageId="21" pageNumber="22" specimenCount="138" specimenCount-female="138">
<specimenCount box="[442,502,1620,1647]" pageId="21" pageNumber="22" type="female">69♀</specimenCount>
1³ (
<specimenCount box="[546,578,1619,1645]" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
from 
<collectingCountry box="[648,733,1619,1645]" name="Taiwan" pageId="21" pageNumber="22">Taiwan</collectingCountry>
,
</materialsCitation>
<materialsCitation box="[745,959,1619,1645]" country="Taiwan" location="Taiwan" pageId="21" pageNumber="22" specimenCount="138" specimenCount-female="138">
<specimenCount box="[745,795,1619,1645]" pageId="21" pageNumber="22" type="female">34♀</specimenCount>
from Honshu,
</materialsCitation>
<materialsCitation box="[967,1221,1619,1647]" country="Taiwan" location="Taiwan" pageId="21" pageNumber="22" specimenCount="138" specimenCount-female="138">
<specimenCount box="[967,1024,1620,1647]" pageId="21" pageNumber="22" type="female">11♀</specimenCount>
1³ from Shikoku,
</materialsCitation>
<materialsCitation country="Taiwan" location="Taiwan" pageId="21" pageNumber="22" specimenCount="138" specimenCount-female="138">
<specimenCount box="[1229,1278,1619,1645]" pageId="21" pageNumber="22" type="female">21♀</specimenCount>
from Kyushu and
</materialsCitation>
<materialsCitation box="[199,868,1655,1681]" country="Taiwan" location="Taiwan" pageId="21" pageNumber="22" specimenCount="138" specimenCount-female="138">
<specimenCount box="[199,233,1655,1681]" pageId="21" pageNumber="22" type="female">2♀</specimenCount>
with no locality data). See Appendix for collection data.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="distribution">
<paragraph blockId="21.[151,1437,862,2041]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[199,345,1690,1716]" pageId="21" pageNumber="22">Distribution</emphasis>
(
<figureCitation box="[363,449,1691,1717]" captionStart="FIGURE 14" captionStartId="29.[151,250,1904,1928]" captionTargetBox="[170,1415,636,1877]" captionTargetId="figure-225@29.[170,1417,633,1880]" captionTargetPageId="29" captionText="FIGURE 14. Distribution of T. longicollis and T. fuscicornis in Japan based on the specimens examined (see Appendix for the collection records)." figureDoi="http://doi.org/10.5281/zenodo.7621785" httpUri="https://zenodo.org/record/7621785/files/figure.png" pageId="21" pageNumber="22">Fig. 14</figureCitation>
). 
<collectingCountry box="[472,536,1691,1717]" name="Japan" pageId="21" pageNumber="22">Japan</collectingCountry>
(Honshu, Shikoku, Kyushu, Tsushima Island),? 
<collectingCountry box="[1091,1162,1691,1717]" name="South Korea" pageId="21" pageNumber="22">Korea</collectingCountry>
(see discussion below), 
<collectingCountry box="[151,236,1727,1753]" name="Taiwan" pageId="21" pageNumber="22">Taiwan</collectingCountry>
, 
<collectingCountry box="[247,314,1727,1753]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
(
<collectingRegion box="[328,402,1727,1753]" country="China" name="Fujian" pageId="21" pageNumber="22">Fujian</collectingRegion>
(
<bibRefCitation author="Xiao, G. &amp; Huang, X. &amp; Zhou, S. &amp; Wu, J. &amp; Zhang, P." box="[417,602,1727,1753]" pageId="21" pageNumber="22" refId="ref29568" refString="Xiao, G., Huang, X., Zhou, S., Wu, J. &amp; Zhang, P. (1992) Economic Sawfly Fauna of China. Tianze Eldonejo, Beijing, 4 + 8 + 221 pp. [in Chinese]" type="book" year="1992">
Xiao 
<emphasis box="[480,529,1727,1753]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
. 1992
</bibRefCitation>
)).
</paragraph>
</subSubSection>
<subSubSection box="[199,1090,1763,1789]" pageId="21" pageNumber="22" type="biology_ecology">
<paragraph blockId="21.[151,1437,862,2041]" box="[199,1090,1763,1789]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[199,324,1763,1789]" pageId="21" pageNumber="22">Host plant</emphasis>
. 
<taxonomicName box="[338,490,1763,1789]" class="Magnoliopsida" family="Cannabaceae" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="family">Cannabaceae</taxonomicName>
: 
<taxonomicName authority="Pers." authorityName="Pers." box="[501,725,1763,1789]" class="Magnoliopsida" family="Cannabaceae" genus="Celtis" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="sinensis">
<emphasis box="[501,662,1763,1789]" italics="true" pageId="21" pageNumber="22">Celtis sinensis</emphasis>
Pers.
</taxonomicName>
(
<bibRefCitation author="Takeuchi, K." box="[738,909,1763,1789]" pageId="21" pageNumber="22" pagination="47 - 50" refId="ref28745" refString="Takeuchi, K. (1949) A list of the food-plants of Japanese sawflies. Transactions of the Kansai Entomological Society, Osaka, 14, 47 - 50." type="journal article" year="1949">Takeuchi 1949</bibRefCitation>
; 
<bibRefCitation author="Okutani, T." box="[920,1078,1763,1789]" pageId="21" pageNumber="22" pagination="43 - 49" refId="ref27093" refString="Okutani, T. (1967) Food plants of Japanese Symphyta (I). Japanese Journal of Applied Entomology and Zoology, 11, 43 - 49. [in Japanese with English abstract] https: // doi. org / 10.1303 / jjaez. 11.90" type="journal article" year="1967">Okutani 1967</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="discussion">
<paragraph blockId="21.[151,1437,862,2041]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[199,308,1799,1825]" pageId="21" pageNumber="22">Remarks</emphasis>
. This species occurs in autumn (
<tableCitation box="[673,759,1799,1825]" captionStart="TABLE 4" captionStartId="20.[150,238,639,665]" captionTargetPageId="20" captionText="TABLE 4. Number of specimens collected in each 10–11 days of April to November." httpUri="http://table.plazi.org/id/1FB4E1BC121642119FC6FD90FB8E94F5" pageId="21" pageNumber="22" tableUuid="1FB4E1BC121642119FC6FD90FB8E94F5">Table 4</tableCitation>
) and the female is well characterized by the largely blackish brown antenna (
<figureCitation box="[331,418,1834,1861]" captionStart="FIGURE 5" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[155,1432,565,1761]" captionTargetId="figure-186@12.[151,1435,561,1764]" captionTargetPageId="12" captionText="FIGURE 5. Antenna, female (A–G, I) and male (H, J–N).—A. T. apicalis, Minamichitose, Hokkaido; B, T. bicinctus, holotype; C, T. longicollis, Toyota, Aichi; D, T. fuscicornis, Inchon, Korea; E, T. nakanei, Ôginosen, Tottori; F, T. contractus, Okayamashi, Okayama; G, T. okinawensis, holotype; H. T. apicalis, Tsukuba, Ibaraki; I, T. katayamai, holotype; J, T. longicollis, Muroto, Kochi; K, T. fuscicornis, Shikotsu-ko, Hokkaido; L, T. nakanei, Noda, Chiba; M, T. contractus, Tatsuno, Hyogo; N, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621761" httpUri="https://zenodo.org/record/7621761/files/figure.png" pageId="21" pageNumber="22">Fig. 5C</figureCitation>
), the long pronotum (
<figureCitation box="[660,749,1835,1861]" captionStart="FIGURE 2" captionStartId="2.[151,250,1940,1964]" captionTargetBox="[163,1424,188,1911]" captionTargetId="figure-20@2.[151,1435,181,1917]" captionTargetPageId="2" captionText="FIGURE 2. Tremex spp., female, with parts of measurements (white lines in A, B, D–F) and the names of cells in forewing (C).—A, D, T. longicollis, Okazaki, Aichi; B, F, T. fuscicornis, Inchon, Korea; C, T. nakanei, Tokyo; E, T. katayamai, holotype. OOCL: ocellar-occipital (carina) line. MPL: median length of pronotum seen from above." figureDoi="http://doi.org/10.5281/zenodo.7621755" httpUri="https://zenodo.org/record/7621755/files/figure.png" pageId="21" pageNumber="22">Fig. 2A</figureCitation>
), the dorsally blackish and ventrally brownish hind tibia (
<figureCitation captionStart="FIGURE 6" captionStartId="13.[151,250,1061,1085]" captionTargetBox="[155,1432,184,1034]" captionTargetId="figure-15@13.[151,1435,181,1037]" captionTargetPageId="13" captionText="FIGURE 6. Hind leg, female.—A, T. longicollis, Tsushima, Nagasaki; B, T. katayamai, holotype; C, T. fuscicornis, Inchon, Korea; D, T. nakanei, holotype of T. kaedei; E, T. contractus, Okayama-shi, Okayama; F, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621763" httpUri="https://zenodo.org/record/7621763/files/figure.png" pageId="21" pageNumber="22">Fig. 6A</figureCitation>
), the pale brownish wings without distinct infuscation (
<figureCitation box="[818,934,1871,1897]" captionStart="FIGURE 7" captionStartId="14.[151,250,1592,1616]" captionTargetBox="[158,1430,190,1546]" captionTargetId="figure-20@14.[151,1435,181,1553]" captionTargetPageId="14" captionText="FIGURE 7. Wing, female.—A, T. apicalis, Kisofukushima, Nagano; B, T. bicinctus, holotype; C, T. bicinctus, Hôfu, Yamauchi; D, T. bicinctus, Fukuoka-shi, Fukuoka; E, T. longicollis, Okazaki, Aichi; F, T. longicollis, Osaka; G, H, T. katayamai, paratype; I, T. fuscicornis, Akigase, Saitama; J, T. fuscicornis, Kyoto." figureDoi="http://doi.org/10.5281/zenodo.7621766" httpUri="https://zenodo.org/record/7621766/files/figure.png" pageId="21" pageNumber="22">Fig. 7E, F</figureCitation>
) and the elongate precornal basin (
<figureCitation box="[1332,1424,1871,1897]" captionStart="FIGURE 2" captionStartId="2.[151,250,1940,1964]" captionTargetBox="[163,1424,188,1911]" captionTargetId="figure-20@2.[151,1435,181,1917]" captionTargetPageId="2" captionText="FIGURE 2. Tremex spp., female, with parts of measurements (white lines in A, B, D–F) and the names of cells in forewing (C).—A, D, T. longicollis, Okazaki, Aichi; B, F, T. fuscicornis, Inchon, Korea; C, T. nakanei, Tokyo; E, T. katayamai, holotype. OOCL: ocellar-occipital (carina) line. MPL: median length of pronotum seen from above." figureDoi="http://doi.org/10.5281/zenodo.7621755" httpUri="https://zenodo.org/record/7621755/files/figure.png" pageId="21" pageNumber="22">Fig. 2D</figureCitation>
). The male of this species is very rare. It was described by 
<bibRefCitation author="Yano, M." box="[815,959,1907,1933]" pageId="21" pageNumber="22" pagination="471 - 475" refId="ref29882" refString="Yano, M. (1932) Siricidae. In: Issiki, S., Ishii, T., Niijima, Y., Hori, H., Hozawa, S., Okamoto, H., Oguma, M., Kawada, A., Kawamura, T., Yokoyama, K., Takahashi, R., Uchida, S., Ueno, M., Kuwana, I., Kuwayama, S., Yano, M., Yamada, S., Maruge, N., Koizumi, M., Kojima, T., Esaki, T., Kinoshita, S., Kishida, K., Yuasa, H., Shiraki, T. &amp; Shinji, O., Iconographia Insectorum Japonicorum Editio Prima. Hokuryukan, Tokyo, pp. 471 - 475. [in Japanese]" type="book chapter" year="1932">Yano (1932)</bibRefCitation>
, 
<bibRefCitation author="Esaki, T. &amp; Hori, H. &amp; Yasumatsu, K." box="[973,1188,1907,1933]" pageId="21" pageNumber="22" refId="ref24625" refString="Esaki, T., Hori, H. &amp; Yasumatsu, K. (1939) Insectorum Japonicorum Illustratio Iconographica Coloribus Ad Naturam Depicta. Sanseido, Tokyo, 485 pp. [in Japanese]" type="book" year="1939">
Esaki 
<emphasis box="[1044,1095,1907,1933]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
. (1939)
</bibRefCitation>
, 
<bibRefCitation author="Takeuchi, K." box="[1201,1387,1907,1933]" pageId="21" pageNumber="22" refId="ref28919" refString="Takeuchi, K. (1962) Insecta Japonica. Series 2. Part 4. Hymenoptera: Siricidae. Hokuryukan, Tokyo, 2 + 12 pp., 1 pl. [in Japanese with English summary]" type="book" year="1962">Takeuchi (1962)</bibRefCitation>
and 
<bibRefCitation author="Naito, T." box="[151,297,1943,1969]" pageId="21" pageNumber="22" pagination="462 - 468" refId="ref26504" refString="Naito, T. (2020) Family Siricidae. In: Naito, T., Shinohara, A., Hara, H. &amp; Ito, F., Sawflies and Woodwasps of Japan. Hokkaido University Press, Sapporo, pp. 179 - 182 + 462 - 468. [in Japanese]" type="book chapter" year="2020">Naito (2020)</bibRefCitation>
but only 
<specimenCount box="[408,563,1943,1969]" pageId="21" pageNumber="22" type="generic">one specimen</specimenCount>
listed above was available for the present study. It differs from the males of other Japanese congeners by the pale brown temple, gena, antennal scape and pedicel (
<figureCitation box="[1120,1211,1979,2005]" captionStart="FIGURE 4" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[158,1429,188,1197]" captionTargetId="figure-15@11.[151,1435,181,1203]" captionTargetPageId="11" captionText="FIGURE 4. Head, thorax and base of abdomen, male.—A, B, T. apicalis, Minamichitose, Hokkaido; C, D, T. fuscicornis, Shikotsu-ko, Hokkaido; E, T. longicollis, Muroto, Kochi; F, T. nakanei, Noda, Chiba; G, T. contractus, Okayama-shi, Okayama; H, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621759" httpUri="https://zenodo.org/record/7621759/files/figure.png" pageId="21" pageNumber="22">Figs 4E</figureCitation>
, 
<figureCitation box="[1222,1249,1978,2005]" captionStart="FIGURE 5" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[155,1432,565,1761]" captionTargetId="figure-186@12.[151,1435,561,1764]" captionTargetPageId="12" captionText="FIGURE 5. Antenna, female (A–G, I) and male (H, J–N).—A. T. apicalis, Minamichitose, Hokkaido; B, T. bicinctus, holotype; C, T. longicollis, Toyota, Aichi; D, T. fuscicornis, Inchon, Korea; E, T. nakanei, Ôginosen, Tottori; F, T. contractus, Okayamashi, Okayama; G, T. okinawensis, holotype; H. T. apicalis, Tsukuba, Ibaraki; I, T. katayamai, holotype; J, T. longicollis, Muroto, Kochi; K, T. fuscicornis, Shikotsu-ko, Hokkaido; L, T. nakanei, Noda, Chiba; M, T. contractus, Tatsuno, Hyogo; N, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621761" httpUri="https://zenodo.org/record/7621761/files/figure.png" pageId="21" pageNumber="22">5J</figureCitation>
), the pale brown wings without dark cloud (
<figureCitation box="[452,538,2015,2041]" captionStart="FIGURE 9" captionStartId="16.[151,250,1266,1290]" captionTargetBox="[158,1429,276,1239]" captionTargetId="figure-20@16.[151,1435,271,1243]" captionTargetPageId="16" captionText="FIGURE 9. Wing, male.—A, T. apicalis, Tsukuba, Ibaraki; B, T. longicollis, Muroto, Kochi; C, T. fuscicornis, Chitose, Hokkaido; D, T. nakanei, Noda, Chiba; E, T. contractus, Tatsuno, Hyogo; F, T. okinawensis, Amami-ôshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621770" httpUri="https://zenodo.org/record/7621770/files/figure.png" pageId="21" pageNumber="22">Fig. 9B</figureCitation>
) and mostly pale brown abdomen (
<figureCitation box="[934,1036,2015,2041]" captionStart="FIGURE 12" captionStartId="19.[151,250,1268,1292]" captionTargetBox="[159,1428,186,1239]" captionTargetId="figure-15@19.[151,1435,181,1244]" captionTargetPageId="19" captionText="FIGURE 12. Abdomen, male.—A, F, T. nakanei, Noda, Chiba; B, G, T. contractus, Tatsuno, Hyogo; C, T. apicalis, Tsukuba, Ibaraki; D, T. fuscicornis, Chitose, Hokkaido; E, T. longicollis, Muroto, Kochi; H, T. okinawensis, Amami-ôshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621778" httpUri="https://zenodo.org/record/7621778/files/figure.png" pageId="21" pageNumber="22">Fig. 12E</figureCitation>
).
</paragraph>
<paragraph blockId="22.[151,1437,150,681]" pageId="22" pageNumber="23">
<bibRefCitation author="Gussakovskij, V. V." box="[199,431,150,177]" pageId="22" pageNumber="23" refId="ref24744" refString="Gussakovskij, V. V. (1935) Insectes Hymenopteres, Chalastogastra 1, Fauna SSSR. 2 (1). Izdatel'stvo Akademii Nauk SSSR, Moskva, Leningrad, 453 pp. [in Russian and German]" type="book" year="1935">Gussakovskij (1935)</bibRefCitation>
noted “ 
<collectingCountry box="[518,589,151,177]" name="South Korea" pageId="22" pageNumber="23">Korea</collectingCountry>
(?)” for the distribution of this species and, probably after him, 
<bibRefCitation author="Kim, C. - W." box="[1305,1436,151,177]" pageId="22" pageNumber="23" pagination="243 - 374" refId="ref25110" refString="Kim, C. - W. (1963) Hymenoptera of Korea. Humanities and Sciences, Natural Science, Korea University, 6, 243 - 374. [in Korean]" type="journal article" year="1963">Kim (1963)</bibRefCitation>
also included it in the Korean fauna without giving any collection data. 
<bibRefCitation author="Lee, J. - W. &amp; Ryu, S. - M. &amp; Chung, D. - K." box="[946,1131,187,213]" pageId="22" pageNumber="23" pagination="297 - 302" refId="ref25717" refString="Lee, J. - W., Ryu, S. - M. &amp; Chung, D. - K. (1998) Taxonomy of the genus Tremex Jurine (Hymenoptera: Siricidae) from Korea, with descriptions of one new species and one unrecorded species. Korean Journal of Biological Sciences, 2, 297 - 302. https: // doi. org / 10.1080 / 12265071.1998.9647422" type="journal article" year="1998">
Lee 
<emphasis box="[995,1043,187,213]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. (1998)
</bibRefCitation>
gave the collection records of 
<specimenCount box="[183,312,223,249]" pageId="22" pageNumber="23" type="male">three males</specimenCount>
collected in April and 
<specimenCount box="[576,691,223,249]" pageId="22" pageNumber="23" type="male">two males</specimenCount>
collected in August but no females. The identity of these Korean specimens, particularly those collected in April, needs confirmation, because the Japanese specimens examined are all females except for 
<specimenCount box="[456,561,295,321]" pageId="22" pageNumber="23" type="male">one male</specimenCount>
as noted above and all were collected in September to October with a few exceptional collection data in July, August and November (
<tableCitation box="[812,900,331,357]" captionStart="TABLE 4" captionStartId="20.[150,238,639,665]" captionTargetPageId="20" captionText="TABLE 4. Number of specimens collected in each 10–11 days of April to November." httpUri="http://table.plazi.org/id/1FB4E1BC121642119FC6FD90FB8E94F5" pageId="22" pageNumber="23" tableUuid="1FB4E1BC121642119FC6FD90FB8E94F5">Table 4</tableCitation>
).
</paragraph>
<paragraph blockId="22.[151,1437,150,681]" pageId="22" pageNumber="23">
The specimens listed in Appendix (also 
<figureCitation box="[642,721,367,393]" captionStart="FIGURE 14" captionStartId="29.[151,250,1904,1928]" captionTargetBox="[170,1415,636,1877]" captionTargetId="figure-225@29.[170,1417,633,1880]" captionTargetPageId="29" captionText="FIGURE 14. Distribution of T. longicollis and T. fuscicornis in Japan based on the specimens examined (see Appendix for the collection records)." figureDoi="http://doi.org/10.5281/zenodo.7621785" httpUri="https://zenodo.org/record/7621785/files/figure.png" pageId="22" pageNumber="23">Fig. 14</figureCitation>
) from 
<collectingRegion isEnumeration="true" pageId="22" pageNumber="23">Fukushima, Ibaraki, Tochigi, Saitama, Aichi, Mie, Kyoto, Kagawa, Fukuoka, Nagasaki</collectingRegion>
(
<location LSID="urn:lsid:plazi:treatment:C3620022121742139FC7FEAFFCD994C5:4E14E7EF121442139EBFFE7CFD0597C1" box="[495,684,403,429]" name="Tsushima Island" pageId="22" pageNumber="23" stateProvince="Fukushima, Ibaraki, Tochigi, Saitama, Aichi, Mie, Kyoto, Kagawa, Fukuoka, Nagasaki">Tsushima Island</location>
) and 
<collectingRegion box="[752,880,403,429]" country="Japan" name="Kagoshima" pageId="22" pageNumber="23">Kagoshima</collectingRegion>
prefectures represent the first collection records from these prefectures.
</paragraph>
<paragraph blockId="22.[151,1437,150,681]" pageId="22" pageNumber="23">
<bibRefCitation author="Matsumura, S." box="[199,419,475,501]" pageId="22" pageNumber="23" refId="ref25923" refString="Matsumura, S. (1906) Nihon Gaichu Mokuroku, Zen. Rokumeikan, Tokyo, 154 pp. [in Japanese]" type="book" year="1906">Matsumura’s (1906</bibRefCitation>
, 
<bibRefCitation author="Matsumura, S." box="[429,500,475,501]" pageId="22" pageNumber="23" refId="ref26102" refString="Matsumura, S. (1932 b) Conspectus of Japanese Injurious Insects. Meijitosho, Tokyo, 62 pls., 3 + 51 + 971 + 115 pp. [in Japanese]" type="book" year="1932">1932b</bibRefCitation>
) host records of conifers and 
<bibRefCitation author="Okutani, T." box="[825,1014,474,501]" pageId="22" pageNumber="23" pagination="548 - 582" refId="ref26904" refString="Okutani, T. (1959) [Symphyta]. In: Esaki, T., Ishii, T., Kawada, A., Shiraki T. &amp; Yuasa, H. (Eds.), Illustrated Insect Larvae of Japan. Hokuryukan, Tokyo, pp. 548 - 582. [in Japanese]" type="book chapter" year="1959">Okutani’s (1959)</bibRefCitation>
host record of “Muku” (= 
<taxonomicName authority="(Thunb.) Planch." authorityName="Planch." baseAuthorityName="Thunb." class="Magnoliopsida" family="Cannabaceae" genus="Aphananthe" kingdom="Plantae" order="Rosales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis italics="true" pageId="22" pageNumber="23">Aphananthe aspera</emphasis>
(Thunb.) Planch.
</taxonomicName>
(
<taxonomicName box="[448,599,511,537]" class="Magnoliopsida" family="Cannabaceae" kingdom="Plantae" order="Rosales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="family">Cannabaceae</taxonomicName>
)) are probably erroneous. 
<bibRefCitation author="Ogata, K." box="[908,1060,511,537]" pageId="22" pageNumber="23" pagination="186" refId="ref26830" refString="Ogata, K. (2021) [Tremex longicollis ovipositing on Podocarpus macrophyllus f. angustifolius]. Satsuma, 167, 186. [in Japanese]" type="journal article" year="2021">Ogata (2021)</bibRefCitation>
observed the oviposition of this woodwasp on 
<taxonomicName authority="(Thunb.) Sweet f." authorityName="Sweet f." baseAuthorityName="Thunb." box="[318,827,547,573]" class="Magnoliopsida" family="Sapotaceae" genus="Podocarpus" kingdom="Plantae" order="Ericales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="macrophyllus">
<emphasis box="[318,614,547,573]" italics="true" pageId="22" pageNumber="23">Podocarpus macrophyllus</emphasis>
(Thunb.) Sweet f.
</taxonomicName>
<emphasis box="[837,982,547,573]" italics="true" pageId="22" pageNumber="23">angustifolius</emphasis>
(Blume) Pilg. (
<taxonomicName box="[1163,1338,547,573]" class="Pinopsida" family="Podocarpaceae" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="family">Podocarpaceae</taxonomicName>
). This is an interesting observation, because all the known host plants of 
<taxonomicName box="[873,1043,583,609]" pageId="22" pageNumber="23">
<emphasis box="[873,954,583,608]" italics="true" pageId="22" pageNumber="23">Tremex</emphasis>
species
</taxonomicName>
are angiosperms and 
<taxonomicName authorityName="Konow" authorityYear="1896" box="[1290,1436,583,609]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis box="[1290,1436,583,609]" italics="true" pageId="22" pageNumber="23">T. longicollis</emphasis>
</taxonomicName>
is thought to be monophagous on 
<taxonomicName authority="(Sakamoto 1997)" baseAuthorityName="Sakamoto" baseAuthorityYear="1997" box="[543,914,619,645]" class="Magnoliopsida" family="Cannabaceae" genus="Celtis" kingdom="Plantae" order="Rosales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="sinensis">
<emphasis box="[543,706,619,645]" italics="true" pageId="22" pageNumber="23">Celtis sinensis</emphasis>
(
<bibRefCitation author="Sakamoto, M." box="[722,905,619,645]" pageId="22" pageNumber="23" pagination="23 - 34" refId="ref27594" refString="Sakamoto, M. (1997) The siricid woodwasps (Hymenoptera, Symphyta, Siricidae) of Hiroshima Prefecture. Journal of the Hiba Society of Natural History, 177, 23 - 34. [in Japanese with English abstract]" type="journal article" year="1997">Sakamoto 1997</bibRefCitation>
)
</taxonomicName>
. The host association of 
<taxonomicName authorityName="Konow" authorityYear="1896" box="[1200,1348,619,645]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis box="[1200,1348,619,645]" italics="true" pageId="22" pageNumber="23">T. longicollis</emphasis>
</taxonomicName>
with 
<taxonomicName class="Magnoliopsida" family="Sapotaceae" genus="Podocarpus" kingdom="Plantae" order="Ericales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="macrophyllus">
<emphasis italics="true" pageId="22" pageNumber="23">P. macrophyllus</emphasis>
</taxonomicName>
f. 
<emphasis box="[333,478,655,681]" italics="true" pageId="22" pageNumber="23">angustifolius</emphasis>
is unlikely and needs confirmation.
</paragraph>
</subSubSection>
</treatment>
</document>