257 lines
32 KiB
XML
257 lines
32 KiB
XML
<document id="78DF3A9EAAB875880E56A82961AFA33B" ID-CLB-Dataset="301504" ID-DOI="10.1093/zooldnnean/zlad146" ID-GBIF-Dataset="5e66aeaf-44c5-4be9-924d-cc13680ace6a" ID-ISSN="0024-4082" ID-Zenodo-Dep="13315269" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1722845279467" checkinUser="plazi" docAuthor="Lumen, Ryan & Kamdńskd, Marcdn J." docDate="2024" docId="C27387A7FFE6FFC99ABD0612FB0A73AD" docLanguage="en" docName="zlad146.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (201)" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Biolus granulatus, comb. nov." docType="treatment" docVersion="4" lastPageNumber="378" masterDocId="3E4AFFDFFFF2FFDD9A72053DFF83770F" masterDocTitle="evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae" masterLastPageNumber="386" masterPageNumber="358" pageNumber="378" updateTime="1723663925820" updateUser="ExternalLinkService">
|
||
<mods:mods id="BF1B8596C812231B42256FE8B2F18C79" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="F2C83E8D05A83993BEC6447ADD93362B">
|
||
<mods:title id="4B6E133B6E99074B87F8A11FFEE46FA6">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="F9AFCDDBBD99412E9A48C844E5AF7E77" type="personal">
|
||
<mods:role id="A2F053A93DC4E6DE6CFE28D70788CDB4">
|
||
<mods:roleTerm id="1425FD93807239467BD0321750C5885D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1254D6AF8121B6674E8D088A90E11DCD">Lumen, Ryan</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A8B06C4D0BF1C7994246730DF459CD72" type="personal">
|
||
<mods:role id="B55CC3B49E5A91868CA8832E5449FD68">
|
||
<mods:roleTerm id="BF39B9D684F41FA28F83F048235F4D94">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5050899DAC9AF6E597B2D90E66D80F6B">Kamdńskd, Marcdn J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="29C275937835F5F48FEEA37579A7087F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A73E99FDEFE74B24052FFF374F022C75" type="host">
|
||
<mods:titleInfo id="9177BCBAC98332EB17A9A35688EB490A">
|
||
<mods:title id="701588801D6F23A398963B5FFE6C7547">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="2512013A166805D0A2491D4D3CA57303">
|
||
<mods:date id="3C9127A68DC81A306D40CD5324EE1858">2024</mods:date>
|
||
<mods:detail id="E58EB27B51403957023308EA098A44B1" type="volume">
|
||
<mods:number id="F8FC7BB6C2272A9AAE610EE791D1DB85">201</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CE33C0D199CFEA84CC7F1E7E3DBEBCFA" type="issue">
|
||
<mods:number id="DCC29CFFE188DB298A1FB36D983EBAED">201</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="54FA34BF983C02F7CE73D9B706C0172E" unit="page">
|
||
<mods:start id="E9DCE25512CCB0F84CB30FB0ABDA2DE8">358</mods:start>
|
||
<mods:end id="003752FADF9217921996913702018D76">386</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="580CECD9D8CAB136D0A5936DAAA5E32E">journal article</mods:classification>
|
||
<mods:identifier id="0EA6062A1CA48C947351EC366D3BD2E1" type="CLB-Dataset">301504</mods:identifier>
|
||
<mods:identifier id="88EB6DFCD556F047B11ADD5A88FE0513" type="DOI">10.1093/zooldnnean/zlad146</mods:identifier>
|
||
<mods:identifier id="6EC6877C166F2005352A3F234B6F7622" type="GBIF-Dataset">5e66aeaf-44c5-4be9-924d-cc13680ace6a</mods:identifier>
|
||
<mods:identifier id="61D1C0D498376FCCEB3512A954E256F7" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="82847EEDC4EC3B95EA8B984F13A09DE3" type="Zenodo-Dep">13315269</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="C27387A7FFE6FFC99ABD0612FB0A73AD" ID-DOI="http://doi.org/10.5281/zenodo.13323866" ID-Zenodo-Dep="13323866" LSID="urn:lsid:plazi:treatment:C27387A7FFE6FFC99ABD0612FB0A73AD" httpUri="http://treatment.plazi.org/id/C27387A7FFE6FFC99ABD0612FB0A73AD" lastPageNumber="378" pageId="20" pageNumber="378">
|
||
<subSubSection id="02C0653AFFE6FFC99ABD0612FD1E7445" box="[207,669,815,842]" pageId="20" pageNumber="378" type="nomenclature">
|
||
<paragraph id="4A6536B1FFE6FFC99ABD0612FD1E7445" blockId="20.[207,669,815,842]" box="[207,669,815,842]" pageId="20" pageNumber="378">
|
||
<heading id="112D81DDFFE6FFC99ABD0612FD1E7445" box="[207,669,815,842]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="378" reason="4">
|
||
<emphasis id="78AEEAA3FFE6FFC99ABD0612FD1E7445" bold="true" box="[207,669,815,842]" pageId="20" pageNumber="378">
|
||
6.
|
||
<taxonomicName id="8DDA4D32FFE6FFC99A940612FDAB7446" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[230,552,815,842]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="granulatus" status="comb. nov.">
|
||
<emphasis id="78AEEAA3FFE6FFC99A940612FE2E7445" bold="true" box="[230,429,815,842]" italics="true" pageId="20" pageNumber="378">Biolus granulatus</emphasis>
|
||
(Fabricius)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="639D57D8FFE6FFC9985E060DFD1E7445" box="[556,669,816,842]" pageId="20" pageNumber="378" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="02C0653AFFE6FFC99B5C0653FDBD7487" box="[302,574,878,904]" pageId="20" pageNumber="378" type="description">
|
||
<paragraph id="4A6536B1FFE6FFC99B5C0653FDBD7487" blockId="20.[302,574,878,904]" box="[302,574,878,904]" pageId="20" pageNumber="378">
|
||
<heading id="112D81DDFFE6FFC99B5C0653FDBD7487" box="[302,574,878,904]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="378" reason="3">
|
||
<emphasis id="78AEEAA3FFE6FFC99B5C0653FDBD7487" bold="true" box="[302,574,878,904]" pageId="20" pageNumber="378">
|
||
(
|
||
<figureCitation id="D2E12A34FFE6FFC99B4B0653FE047487" box="[313,391,878,904]" captionStart="Figure1" captionStartId="1.[129,191,1217,1241]" captionTargetBox="[131,1459,153,1189]" captionTargetId="figure-312@1.[128,1475,144,1190]" captionTargetPageId="1" captionText="Figure1. Ovovdvdparydn ºenebrdonddae and dts geographdcal ddstrdbutdon.A, sympatrdcPlatynotdnd specdes ddsplaydng ddfferent reproductdonstrategdes: Biolusgranulatus–ovovdvdparous,Crypticanus sp.–ovdparous.B, female gendtal tract ofBiolus brunoi sp.nov.wdth developed 1stdnstar larva.Scale" figureDoi="http://doi.org/10.5281/zenodo.13315275" httpUri="https://zenodo.org/record/13315275/files/figure.png" pageId="20" pageNumber="378">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="D2E12A34FFE6FFC99BE00653FE3C7487" box="[402,447,878,904]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." figureDoi="http://doi.org/10.5281/zenodo.13315277" httpUri="https://zenodo.org/record/13315277/files/figure.png" pageId="20" pageNumber="378">2F, I</figureCitation>
|
||
,
|
||
<figureCitation id="D2E12A34FFE6FFC99BBB0653FE647487" box="[457,487,878,904]" captionStart="Figure 5" captionStartId="6.[113,178,1366,1390]" captionTargetBox="[117,1454,147,1335]" captionTargetId="figure-238@6.[114,1458,144,1338]" captionTargetPageId="6" captionText="Figure 5. Phylogeny of the subtrdbe Eurynotdna based on morphology. The most parsdmondous cladogram obtadned dn ºNº analysds" figureDoi="http://doi.org/10.5281/zenodo.13315287" httpUri="https://zenodo.org/record/13315287/files/figure.png" pageId="20" pageNumber="378">5A</figureCitation>
|
||
,
|
||
<figureCitation id="D2E12A34FFE6FFC99B810653FD8D7487" box="[499,526,878,904]" captionStart="Figure 7" captionStartId="12.[113,178,1803,1827]" captionTargetBox="[118,1458,150,1772]" captionTargetId="figure-7@12.[114,1458,144,1775]" captionTargetPageId="12" captionText="Figure 7. Phylogeny of morphology, ovovdvdpary, and female reproductdve structures of Eurynotdna.A, Eurynotus barbosai, 1st dnstar larva ddssected from bursa copulatrdx. B, Biolus omeri, gendtal tubes. C, B. brunoi, spermatheca and ovdposdtor.D, E. capensis, gendtal tubes of two dnddvdduals.E, B.analisetosus, sclerdte and egg ddssected from bursa copulatrdx. F, B. granulatus, sclerdte from bursa copulatrdx.G, E. asperatus, padred sclerdtes on ddstal part of bursa copulatrdx. H, Oncotus cedrimontis, bursa copulatrdx. Abbrevdatdons: c1–c4, lobes of coxdtdes. Numbers dn parentheses represent characters and states used dn analysds." figureDoi="http://doi.org/10.5281/zenodo.13323862" httpUri="https://zenodo.org/record/13323862/files/figure.png" pageId="20" pageNumber="378">7F</figureCitation>
|
||
,
|
||
<figureCitation id="D2E12A34FFE6FFC998650653FDB17487" box="[535,562,878,904]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." figureDoi="http://doi.org/10.5281/zenodo.13315291" httpUri="https://zenodo.org/record/13315291/files/figure.png" pageId="20" pageNumber="378">8F</figureCitation>
|
||
)
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="02C0653AFFE6FFC99A0306A8FD5D7387" pageId="20" pageNumber="378" type="reference_group">
|
||
<paragraph id="4A6536B1FFE6FFC99A0306A8FE7574A2" blockId="20.[113,762,917,1160]" box="[113,502,917,941]" pageId="20" pageNumber="378">
|
||
<treatmentCitationGroup id="6ACA119FFFE6FFC99A0306A8FE7574A2" box="[113,502,917,941]" pageId="20" pageNumber="378">
|
||
<treatmentCitation id="CB7B10A0FFE6FFC99A0306A8FE7174A2" author="Fabrdcdus JC" box="[113,498,917,941]" page="214" pageId="20" pageNumber="378" year="1787">
|
||
<taxonomicName id="8DDA4D32FFE6FFC99A0306A8FE7174A2" authority="Fabrdcdus 1787: 214" authorityName="Fabrdcdus" authorityPageNumber="214" authorityYear="1787" box="[113,498,917,941]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="78AEEAA3FFE6FFC99A0306A8FEA374A2" box="[113,288,917,941]" italics="true" pageId="20" pageNumber="378">Helops granulatus</emphasis>
|
||
<bibRefCitation id="2E4B4B40FFE6FFC99B5706A8FE7174A2" author="Fabrdcdus JC" box="[293,498,917,941]" pageId="20" pageNumber="378" refId="ref25568" refString="Fabrdcdus JC. Mantissa Insectorum Sistens Eorum Species Nuper Detectas Adiectis Characteribus Genericis, Differentiis Specificis, Emendationibus, Observationibus. 1. Copenhagen: Impensds C. G. Proft, 1787, XX + 348." type="journal volume" year="1787">Fabrdcdus 1787: 214</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE6FFC99AFF0689FD4274C3" blockId="20.[113,762,917,1160]" box="[141,705,948,972]" pageId="20" pageNumber="378">
|
||
<treatmentCitationGroup id="6ACA119FFFE6FFC99AFF0689FD4274C3" box="[141,705,948,972]" pageId="20" pageNumber="378">
|
||
<taxonomicName id="8DDA4D32FFE6FFC99AFF0689FEA574C3" box="[141,294,948,972]" class="Insecta" family="Tenebrionidae" genus="Blaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="granulata">
|
||
<emphasis id="78AEEAA3FFE6FFC99AFF0689FEA574C3" box="[141,294,948,972]" italics="true" pageId="20" pageNumber="378">Blaps granulata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="2E4B4B40FFE6FFC99B440689FE4E74C3" author="Fabrdcdus JC" box="[310,461,948,972]" pageId="20" pageNumber="378" refId="ref25568" refString="Fabrdcdus JC. Mantissa Insectorum Sistens Eorum Species Nuper Detectas Adiectis Characteribus Genericis, Differentiis Specificis, Emendationibus, Observationibus. 1. Copenhagen: Impensds C. G. Proft, 1787, XX + 348." type="journal volume" year="1787">Fabrdcdus 1787</bibRefCitation>
|
||
%.–
|
||
<treatmentCitation id="CB7B10A0FFE6FFC99B820689FD3E74C3" author="Fabrdcdus JC" box="[496,701,948,972]" page="107" pageId="20" pageNumber="378" year="1792">
|
||
<bibRefCitation id="2E4B4B40FFE6FFC99B820689FD3E74C3" author="Fabrdcdus JC" box="[496,701,948,972]" pageId="20" pageNumber="378" refId="ref25606" refString="Fabrdcdus JC. Entomologia Systematica Emendata et Aucta: Secundun Classes, Ordines, Genera, Species, Adjectis Synonimis, Locis, Observationibus, Descriptionibus. Hafndae: Proft, 1792, 330." type="journal volume" year="1792">Fabrdcdus 1792: 107</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE6FFC99AFF06E9FE2D7325" blockId="20.[113,762,917,1160]" pageId="20" pageNumber="378">
|
||
<treatmentCitationGroup id="6ACA119FFFE6FFC99AFF06E9FE2D7325" pageId="20" pageNumber="378">
|
||
<taxonomicName id="8DDA4D32FFE6FFC99AFF06E9FEE374E4" authorityName="Muir" authorityYear="1921" box="[141,352,979,1003]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="78AEEAA3FFE6FFC99AFF06E9FEE374E4" box="[141,352,979,1003]" italics="true" pageId="20" pageNumber="378">Eurynotus granulatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="2E4B4B40FFE6FFC99B0606EEFD8D74E4" author="Fabrdcdus JC" box="[372,526,979,1003]" pageId="20" pageNumber="378" refId="ref25568" refString="Fabrdcdus JC. Mantissa Insectorum Sistens Eorum Species Nuper Detectas Adiectis Characteribus Genericis, Differentiis Specificis, Emendationibus, Observationibus. 1. Copenhagen: Impensds C. G. Proft, 1787, XX + 348." type="journal volume" year="1787">Fabrdcdus 1787</bibRefCitation>
|
||
%.–
|
||
<treatmentCitation id="CB7B10A0FFE6FFC9984706EEFD7474E4" author="Gebden H" box="[565,759,979,1003]" page="230" pageId="20" pageNumber="378" year="1906">
|
||
<bibRefCitation id="2E4B4B40FFE6FFC9984706EEFD7474E4" author="Gebden H" box="[565,759,979,1003]" pageId="20" pageNumber="378" pagination="209 - 37" refId="ref25970" refString="Gebden H. Uber dde von Fabrdcdus beschrdebenen ºypen von ºenebrdondden dn den Museen von Kopenhagen und Kdel. Deutsche Entomologische Zeitschrift 1906; 1906: 209 - 37." type="journal article" year="1906">Gebden 1906: 230</bibRefCitation>
|
||
</treatmentCitation>
|
||
,
|
||
<treatmentCitation id="CB7B10A0FFE6FFC99A0306CEFF587304" author="Gebden H" box="[113,219,1011,1035]" page="414" pageId="20" pageNumber="378" year="1938">
|
||
<bibRefCitation id="2E4B4B40FFE6FFC99A0306CEFF587304" author="Gebden H" box="[113,219,1011,1035]" pageId="20" pageNumber="378" pagination="49 - 80" refId="ref26000" refString="Gebden H. Katalog der Tenebrioniden. Teil II. Munchen: Mdttedlungen der Muncher Entomologddche Gesellschaft, 1938, 28, 49 - 80, 283 - 428 [370 - 465]." type="journal article" year="1938">1938: 414</bibRefCitation>
|
||
</treatmentCitation>
|
||
; Koch 1954a: 280; 1954b: 2; 1956: 25;
|
||
<treatmentCitation id="CB7B10A0FFE6FFC9981406CEFF4C7325" author="Banaszkdewdcz M" page="63" pageId="20" pageNumber="378" year="2006">
|
||
<bibRefCitation id="2E4B4B40FFE6FFC9981406CEFF4C7325" author="Banaszkdewdcz M" pageId="20" pageNumber="378" pagination="59 - 77" refId="ref25294" refString="Banaszkdewdcz M. Comparatdve study of female gendtalda dn Peddndnd sens Iwan 2004 (Coleoptera: ºenebrdonddae: Peddndnd %, wdth notes on the classdficatdon. Annales Zoologici 2006; 56: 59 - 77." type="journal article" year="2006">Banaszkdewdcz 2006: 63</bibRefCitation>
|
||
</treatmentCitation>
|
||
; Kamdńskd 2016: 235.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE6FFC99AFF010FFD5D7387" blockId="20.[113,762,917,1160]" pageId="20" pageNumber="378">
|
||
<treatmentCitationGroup id="6ACA119FFFE6FFC99AFF010FFD5D7387" pageId="20" pageNumber="378">
|
||
=
|
||
<taxonomicName id="8DDA4D32FFE6FFC99AEE010FFD007346" authority="Mulsant and Rey 1854a: 27" authorityName="Mulsant and Rey" authorityPageNumber="27" authorityYear="1854" box="[156,643,1073,1098]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="norrisi" status="syn. nov.">
|
||
<emphasis id="78AEEAA3FFE6FFC99AEE010FFEC97346" box="[156,330,1074,1097]" italics="true" pageId="20" pageNumber="378">Eurynotus norrisi</emphasis>
|
||
Mulsant and Rey 1854a: 27
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="639D57D8FFE6FFC998FC010FFD6E7346" box="[654,749,1074,1098]" pageId="20" pageNumber="378">syn. nov.</taxonomicNameLabel>
|
||
– Mulsant and Rey 1854b: 171; Lacordadre 1859: 239;
|
||
<treatmentCitation id="CB7B10A0FFE6FFC998DD016CFF5F7387" author="Gebden H" page="414" pageId="20" pageNumber="378" year="1938">
|
||
<bibRefCitation id="2E4B4B40FFE6FFC998DD016CFF5F7387" author="Gebden H" pageId="20" pageNumber="378" pagination="49 - 80" refId="ref26000" refString="Gebden H. Katalog der Tenebrioniden. Teil II. Munchen: Mdttedlungen der Muncher Entomologddche Gesellschaft, 1938, 28, 49 - 80, 283 - 428 [370 - 465]." type="journal article" year="1938">Gebden 1938: 414</bibRefCitation>
|
||
</treatmentCitation>
|
||
; Koch 1954a: 277; 1954b: 2; Kamdńskd 2016: 235.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="02C0653AFFE6FFC99A030197FB0A73AD" pageId="20" pageNumber="378" type="diagnosis">
|
||
<paragraph id="4A6536B1FFE6FFC99A030197FF4871FA" blockId="20.[113,764,1193,1781]" pageId="20" pageNumber="378">
|
||
<emphasis id="78AEEAA3FFE6FFC99A030197FF5473CE" box="[113,215,1194,1217]" italics="true" pageId="20" pageNumber="378">Diagnosis:</emphasis>
|
||
Can be ddfficult to ddfferentdate from other specdes, but ds most sdmdlar to
|
||
<taxonomicName id="8DDA4D32FFE6FFC99B3C01F4FE1C73EF" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[334,415,1225,1248]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="omeri">
|
||
<emphasis id="78AEEAA3FFE6FFC99B3C01F4FE1C73EF" box="[334,415,1225,1248]" italics="true" pageId="20" pageNumber="378">B. omeri</emphasis>
|
||
</taxonomicName>
|
||
(e.g. both have sdmple punctatdon of abdomdnal ventrdte V, and are sdmdlar dn sdze, wdth tubercles/ radsed callosdtdes on elytral ddsc%. The two can be ddfferentdated based on setatdon of the tarsd (
|
||
<taxonomicName id="8DDA4D32FFE6FFC99BD9001AFD867231" box="[427,517,1319,1342]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="norrisi">
|
||
<emphasis id="78AEEAA3FFE6FFC99BD9001AFD867231" box="[427,517,1319,1342]" italics="true" pageId="20" pageNumber="378">B. norrisi</emphasis>
|
||
</taxonomicName>
|
||
golden;
|
||
<taxonomicName id="8DDA4D32FFE6FFC9982D001AFD327231" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[607,689,1319,1342]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="omeri">
|
||
<emphasis id="78AEEAA3FFE6FFC9982D001AFD327231" box="[607,689,1319,1342]" italics="true" pageId="20" pageNumber="378">B. omeri</emphasis>
|
||
</taxonomicName>
|
||
black/ dark brown%, the sclerdte wdthdn the bursa copulatrdx (present dn
|
||
<taxonomicName id="8DDA4D32FFE6FFC99AFE005BFE8D7272" authorityName="Muir" authorityYear="1921" box="[140,270,1381,1405]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="78AEEAA3FFE6FFC99AFE005BFE8D7272" box="[140,270,1381,1405]" italics="true" pageId="20" pageNumber="378">B. granulatus</emphasis>
|
||
</taxonomicName>
|
||
; absent dn
|
||
<emphasis id="78AEEAA3FFE6FFC99B0D005BFE587272" box="[383,475,1382,1405]" italics="true" pageId="20" pageNumber="378">
|
||
<taxonomicName id="8DDA4D32FFE6FFC99B0D005BFE577272" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[383,468,1382,1405]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="omeri">B. omeri</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
, and characterdzatdon of the dorsal surface (
|
||
<taxonomicName id="8DDA4D32FFE6FFC99B6800B8FEFB7293" box="[282,376,1413,1436]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="norrisi">
|
||
<emphasis id="78AEEAA3FFE6FFC99B6800B8FEFB7293" box="[282,376,1413,1436]" italics="true" pageId="20" pageNumber="378">B. norrisi</emphasis>
|
||
</taxonomicName>
|
||
blunter/shorter and more crowded tubercles; whdle
|
||
<taxonomicName id="8DDA4D32FFE6FFC99B500098FEF472B3" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[290,375,1445,1468]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="omeri">
|
||
<emphasis id="78AEEAA3FFE6FFC99B500098FEF472B3" box="[290,375,1445,1468]" italics="true" pageId="20" pageNumber="378">B. omeri</emphasis>
|
||
</taxonomicName>
|
||
has sharper/taller and more wddely spaced tuberculatdon wdth a sldghtly more lustrous surface%. Insofar as other specdes are concerned, the abdomdnal ventrdte V punctatdon (sdmple% separates dt from
|
||
<taxonomicName id="8DDA4D32FFE6FFC9984F033EFD507115" authorityName="Koch" authorityYear="1954" box="[573,723,1538,1562]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="analisetosus">
|
||
<emphasis id="78AEEAA3FFE6FFC9984F033EFD507115" box="[573,723,1538,1562]" italics="true" pageId="20" pageNumber="378">B. analisetosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8DDA4D32FFE6FFC99894033EFF6B7136" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="asperipennis">
|
||
<emphasis id="78AEEAA3FFE6FFC99894033EFF6B7136" italics="true" pageId="20" pageNumber="378">B. asperipennis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8DDA4D32FFE6FFC99A87031FFECB7136" box="[245,328,1569,1593]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="graafi">
|
||
<emphasis id="78AEEAA3FFE6FFC99A87031FFECB7136" box="[245,328,1569,1593]" italics="true" pageId="20" pageNumber="378">B. graafi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="8DDA4D32FFE6FFC99BF3031FFE757136" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[385,502,1569,1593]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="sauroides">
|
||
<emphasis id="78AEEAA3FFE6FFC99BF3031FFE757136" box="[385,502,1569,1593]" italics="true" pageId="20" pageNumber="378">B. sauroides</emphasis>
|
||
</taxonomicName>
|
||
(wdth two or more
|
||
<typeStatus id="95618813FFE6FFC998B1031FFD797135" box="[707,762,1570,1594]" pageId="20" pageNumber="378">types</typeStatus>
|
||
of punctatdon%. The tubercles/radsed callosdtdes present on the elytral ddsc further separate dt from
|
||
<taxonomicName id="8DDA4D32FFE6FFC99BA1035CFDE17177" box="[467,610,1633,1656]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="asperipennis">
|
||
<emphasis id="78AEEAA3FFE6FFC99BA1035CFDE17177" box="[467,610,1633,1656]" italics="true" pageId="20" pageNumber="378">B. asperipennis</emphasis>
|
||
</taxonomicName>
|
||
(lackdng ddscal tubercles%. The presence of a sclerdte dn the bursa copulatrdx sep* arates thds specdes from all but
|
||
<taxonomicName id="8DDA4D32FFE6FFC99BDA03A2FDB771B9" authorityName="Koch" authorityYear="1954" box="[424,564,1694,1718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="analisetosus">
|
||
<emphasis id="78AEEAA3FFE6FFC99BDA03A2FDB771B9" box="[424,564,1694,1718]" italics="true" pageId="20" pageNumber="378">B. analisetosus</emphasis>
|
||
</taxonomicName>
|
||
(whdch, dn contrast to
|
||
<taxonomicName id="8DDA4D32FFE6FFC99AF90382FE8971D9" authorityName="Muir" authorityYear="1921" box="[139,266,1726,1750]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="78AEEAA3FFE6FFC99AF90382FE8971D9" box="[139,266,1726,1750]" italics="true" pageId="20" pageNumber="378">B. granulatus</emphasis>
|
||
</taxonomicName>
|
||
’ axe*shaped sclerdte, has a flatter, chevron*shaped sclerdte%.
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE6FFC99A03022BFD7970A3" blockId="20.[113,763,1814,1964]" pageId="20" pageNumber="378">
|
||
<emphasis id="78AEEAA3FFE6FFC99A03022BFE817021" box="[113,258,1814,1838]" italics="true" pageId="20" pageNumber="378">Redescription:</emphasis>
|
||
Length
|
||
<quantity id="8D229B54FFE6FFC99B18022BFE6B7021" box="[362,488,1814,1838]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="20" pageNumber="378" unit="mm" value="15.0" valueMax="16.0" valueMin="14.0">14–16 mm</quantity>
|
||
; wddth
|
||
<quantity id="8D229B54FFE6FFC99834022AFD2B7021" box="[582,680,1815,1839]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="20" pageNumber="378" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
|
||
. Head: Coarsely punctate, punctures closely spaced, separated by ≤1 puncture ddameter to confluent. Mentum wdth apdcal notch. Eye wdth dmpressed sulcus around margdn. Prothorax: Pronotum coarsely punctate, punctures separated by ≤1
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE6FFC9995805ADFB997530" blockId="20.[810,1459,144,575]" pageId="20" pageNumber="378">puncture ddameter to confluent. Hypomeron rugose, wdth large and shallow punctures. Pterothorax: Elytron not costate, wdth macrotubercles and wdth ddstdnct or fadnt mdcrotuberculatdon dn dnterstdces on ddsc. At most, a few wddely spaced, small mdcrotubercles dn basal regdon of epdpleura. Abdomen: Punctate. Abdomdnal ventrdte V punctures separated by ≥1 puncture ddameter wdthout larger, setdgerous punctures dnter* mdxed. Females wdthout meddan apdcal notch. Legs: Male and Female legs unmoddfied. ºarsd wdth golden setae. Males: pro/meso tarsd robust wdth large, ventral, tomentose pads. Male termdnalda: Parameres weakly taperdng wdth tdps sharply taperdng and ddrected meddally. Female termdnalda: Bursa copulatrdx wdth accessory pouch and axe*shaped sclerdte near spermathecal opendng.</paragraph>
|
||
<paragraph id="4A6536B1FFE6FFC99958075DFB11741A" blockId="20.[810,1460,608,977]" pageId="20" pageNumber="378">
|
||
<emphasis id="78AEEAA3FFE6FFC99958075DFB397577" box="[810,1210,608,632]" italics="true" pageId="20" pageNumber="378">
|
||
Material examined:
|
||
<taxonomicName id="8DDA4D32FFE6FFC99E70075DFB397577" authorityName="Fabrdcdus" authorityYear="1787" box="[1026,1210,608,632]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="granulatus">Helops granulatus</taxonomicName>
|
||
</emphasis>
|
||
:
|
||
<materialsCitation id="FAB23CECFFE6FFC99EA3075CFB11741A" collectionCode="ZMUK" pageId="20" pageNumber="378" specimenCount="3" typeStatus="lectotype">
|
||
<typeStatus id="95618813FFE6FFC99EA3075CFAB87576" box="[1233,1339,609,633]" pageId="20" pageNumber="378" type="lectotype">Lectotype</typeStatus>
|
||
, here des* dgnated (
|
||
<collectionCode id="2CCBAE74FFE6FFC999FA07BDFC567597" box="[904,981,640,664]" pageId="20" pageNumber="378">ZMUK</collectionCode>
|
||
%: QR code label ‘ZMUKFabrdcdus 004512’.
|
||
<emphasis id="78AEEAA3FFE6FFC99958079DFCE275B8" box="[810,865,672,695]" italics="true" pageId="20" pageNumber="378">Note:</emphasis>
|
||
As the ordgdnal descrdptdons lack dnformatdon defindng the number or specdficdty of specdmens used, a specdmen from the presumed syntype serdes ds here desdgnated as lectotype to fix the taxonomdc status of the specdes.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE6FFC999370620FC50749D" blockId="20.[810,1460,608,977]" pageId="20" pageNumber="378">
|
||
<taxonomicName id="8DDA4D32FFE6FFC999370620FC70743B" box="[837,1011,797,820]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="378" phylum="Arthropoda" rank="species" species="norrisi">
|
||
<emphasis id="78AEEAA3FFE6FFC999370620FC70743B" box="[837,1011,797,820]" italics="true" pageId="20" pageNumber="378">Eurynotus norrisi</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="95618813FFE6FFC9998C0620FBE4743A" box="[1022,1127,797,821]" pageId="20" pageNumber="378" type="lectotype">Lectotype</typeStatus>
|
||
(MNHN%:
|
||
<emphasis id="78AEEAA3FFE6FFC99E9E0621FB70743B" bold="true" box="[1260,1267,796,820]" pageId="20" pageNumber="378">‘</emphasis>
|
||
<taxonomicName id="8DDA4D32FFE6FFC99E800620FADD743A" box="[1266,1374,797,821]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="378" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
|
||
norrdsdd buq? spdnola dn ldtt. cap.bon.sp./ small blue cdrcular label:
|
||
<taxonomicName id="8DDA4D32FFE6FFC999580666FC15747C" box="[810,918,859,883]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="378" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
|
||
norrdsd Mls Cap. B. Sp. 2 º?/ ºype Mulsant/ Norrdsdd/ Red label: ºype’.
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE6FFC9993706A7FC5B74DE" blockId="20.[810,1460,608,977]" pageId="20" pageNumber="378">(See Supportdng Informatdon, Appenddx S1 for adddtdonal ma* terdal examdned.%</paragraph>
|
||
<paragraph id="4A6536B1FFE6FFC9995806CEFB337367" blockId="20.[810,1459,1010,1128]" pageId="20" pageNumber="378">
|
||
<emphasis id="78AEEAA3FFE6FFC9995806CEFCE27305" box="[810,865,1011,1034]" italics="true" pageId="20" pageNumber="378">Note:</emphasis>
|
||
As the ordgdnal descrdptdons lack dnformatdon defindng the number or specdficdty of specdmens used, a specdmen from the presumed
|
||
<typeStatus id="95618813FFE6FFC999B2010CFB927346" box="[960,1041,1073,1097]" pageId="20" pageNumber="378" type="syntype">syntype</typeStatus>
|
||
serdes ds here desdgnated as
|
||
<typeStatus id="95618813FFE6FFC99F45010CFA147346" box="[1335,1431,1073,1097]" pageId="20" pageNumber="378" type="lectotype">lectotype</typeStatus>
|
||
to fix the taxonomdc status of the specdes.
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE6FFC9995801B4FB0A73AD" blockId="20.[810,1161,1161,1186]" box="[810,1161,1161,1186]" pageId="20" pageNumber="378">
|
||
<emphasis id="78AEEAA3FFE6FFC9995801B4FC2B73AE" box="[810,936,1161,1185]" italics="true" pageId="20" pageNumber="378">Distribution:</emphasis>
|
||
South Afrdca (Fdg. 9%.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |