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<document id="55BAC743F5FEA7B7324124C017A8525B" ID-CLB-Dataset="9161" ID-DOI="10.11646/zootaxa.4822.2.7" ID-GBIF-Dataset="1cd743eb-44c1-4a65-aa50-2ee46f963270" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401527" ID-ZooBank="BB252D1B-239A-4DC4-B538-99E313259B9F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609336462336" checkinUser="plazi" docAuthor="Garcia, Lluc, Miralles-Núñez, Adrià &amp; Pérez-Fernández, Toni" docDate="2020" docId="B3661632714B1579FF54F9BCFAF4FB72" docLanguage="en" docName="zootaxa.4822.2.7.pdf" docOrigin="Zootaxa 4822 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Baeticoniscus Garcia &amp; Miralles-Núñez &amp; Pérez-Fernández 2020, n. gen." docType="treatment" docUuid="9AE11B33-FA0D-4BCD-9061-BB3EB2CD9205" docUuidSource="ZooBank" docVersion="6" lastPageNumber="260" masterDocId="4F5F6E4A714A157AFFC3FFC3FFBBFF9B" masterDocTitle="A new genus and species of cave-dwelling terrestrial isopod (Crustacea: Oniscidea Trichoniscidae) from Southern Spain" masterLastPageNumber="268" masterPageNumber="257" pageNumber="258" updateTime="1698849674458" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FAAB2AF5F1ADC7938CB398CC76B64BEF">A new genus and species of cave-dwelling terrestrial isopod (Crustacea: Oniscidea Trichoniscidae) from Southern Spain</mods:title>
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<mods:namePart id="0C6E9125C40B9982D38D563AA8E0C995">Garcia, Lluc</mods:namePart>
<mods:affiliation id="88249BC4F08A35BA7B81CE6CF2A8F754">Museu Balear de Ciències Naturals-Balearic Museum of Natural Sciences. Sóller (Majorca) Balearic Islands. Interdisciplinary Ecology Group University of the Balearic Islands (UIB), Spain.</mods:affiliation>
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<mods:namePart id="A1587F2D7C1687E23ACA780500DB07AD">Miralles-Núñez, Adrià</mods:namePart>
<mods:affiliation id="58990C3FECC873645ABFDB9D8848A3FC">Institució Catalana dHistòria Natural, Barcelona. Spain.</mods:affiliation>
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<mods:namePart id="01F676F0DF8189879AE5272B409F376D">Pérez-Fernández, Toni</mods:namePart>
<mods:affiliation id="D1A1B713B31FC02B9AA03DA93A2C6E62">Grupo Espeleológico de Villacarrillo-G. E. V. - Villacarrillo, Jaén. Spain.</mods:affiliation>
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<treatment id="B3661632714B1579FF54F9BCFAF4FB72" ID-DOI="http://doi.org/10.5281/zenodo.4401519" ID-GBIF-Taxon="176347548" ID-Zenodo-Dep="4401519" LSID="urn:lsid:zoobank.org:act:9AE11B33-FA0D-4BCD-9061-BB3EB2CD9205" httpUri="http://treatment.plazi.org/id/B3661632714B1579FF54F9BCFAF4FB72" lastPageId="3" lastPageNumber="260" pageId="1" pageNumber="258">
<subSubSection id="73D5F4AF714B157BFF54F9BCFC09F927" pageId="1" pageNumber="258" type="nomenclature">
<paragraph id="3B70A724714B157BFF54F9BCFE46F902" blockId="1.[151,946,1662,1724]" box="[151,509,1662,1689]" pageId="1" pageNumber="258">
<heading id="60381048714B157BFF54F9BCFE46F902" bold="true" box="[151,509,1662,1689]" fontSize="11" level="1" pageId="1" pageNumber="258" reason="1">
<emphasis id="09BB7B36714B157BFF54F9BCFE46F902" bold="true" box="[151,509,1662,1689]" pageId="1" pageNumber="258">
<taxonomicName id="FCCFDCA7714B157BFF54F9BCFE25F903" ID-CoL="845FG" authority="Garcia" authorityName="Garcia &amp; Miralles-Núñez &amp; Pérez-Fernández" authorityYear="2020" box="[151,414,1662,1689]" class="Malacostraca" family="Trichoniscidae" genus="Baeticoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="258" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="09BB7B36714B157BFF54F9BCFE84F902" bold="true" box="[151,319,1663,1689]" italics="true" pageId="1" pageNumber="258">Baeticoniscus</emphasis>
Garcia
</taxonomicName>
,
<taxonomicNameLabel id="1288C64D714B157BFE68F9BCFE46F902" box="[427,509,1663,1689]" pageId="1" pageNumber="258" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
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<paragraph id="3B70A724714B157BFF54F967FC09F927" blockId="1.[151,946,1662,1724]" box="[151,946,1700,1724]" pageId="1" pageNumber="258">
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urn:lsid:zoobank.org:act:
<uuid id="4F699DF1714B157BFE5EF967FC09F927" box="[413,946,1700,1724]" pageId="1" pageNumber="258">9AE11B33-FA0D-4BCD-9061-BB3EB2CD9205</uuid>
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</paragraph>
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<subSubSection id="73D5F4AF714B157BFF54F929FDC0F846" pageId="1" pageNumber="258" type="diagnosis">
<paragraph id="3B70A724714B157BFF54F929FDC0F846" blockId="1.[151,1436,1770,2013]" pageId="1" pageNumber="258">
<emphasis id="09BB7B36714B157BFF54F929FEB1F89F" bold="true" box="[151,266,1770,1796]" pageId="1" pageNumber="258">Diagnosis</emphasis>
.
<taxonomicName id="FCCFDCA7714B157BFED9F928FD11F89D" authority="(Haplophthalminae)" baseAuthorityName="Haplophthalminae" box="[282,682,1771,1798]" class="Malacostraca" family="Trichoniscidae" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="258" phylum="Arthropoda" rank="family">Trichoniscidae (Haplophthalminae)</taxonomicName>
with strong tubercles in the cephalon and pereonites, and smooth third pleonite. Head (cephalothorax) with two small ribs on the front; vertex raised, with a prominent tubercle divided at the apex and two blunt protuberances on the sides; central part of the cephalon with a smaller bilobed tubercle; lateral parts of the cephalon with two tubercles on each side. Pereonites 16 with four raised tubercles (2 + 2), the central ones bigger than the external ones, and accessory rugosities on each side; pereonite 7 with two central large and closely tubercles, separated at the base, and two accessory rugosities on the sides. First male pleopod with thick endopod, with striated subconical tip.
</paragraph>
</subSubSection>
<caption id="6FB0F7AC71481578FF54F85CFDD4F840" ID-DOI="http://doi.org/10.5281/zenodo.4401529" ID-Zenodo-Dep="4401529" httpUri="https://zenodo.org/record/4401529/files/figure.png" pageId="2" pageNumber="259" startId="2.[151,250,1951,1975]" targetBox="[163,1423,188,1923]" targetPageId="2">
<paragraph id="3B70A72471481578FF54F85CFDD4F840" blockId="2.[151,1436,1951,2011]" pageId="2" pageNumber="259">
<emphasis id="09BB7B3671481578FF54F85CFEAFF82C" bold="true" box="[151,276,1951,1975]" pageId="2" pageNumber="259">FIGURE 1.</emphasis>
Cueva de la Pileta of Benaoján, Málaga, Spain. A, entrance to the cave; B, view of one of the halls inside (photos provided by
<taxonomicName id="FCCFDCA771481578FED4F800FEE7F840" box="[279,348,1987,2011]" family="Bullon" pageId="2" pageNumber="266" rank="family">Bullón</taxonomicName>
family and Juan Mayoral).
</paragraph>
</caption>
<subSubSection id="73D5F4AF71491579FF04FF54FC63FF2A" box="[199,984,150,177]" pageId="3" pageNumber="260" type="type_taxon">
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<emphasis id="09BB7B3671491579FF04FF54FED9FF2B" bold="true" box="[199,354,150,177]" pageId="3" pageNumber="260">
<typeStatus id="E474198671491579FF04FF54FEB9FF2A" box="[199,258,151,177]" pageId="3" pageNumber="260">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="FCCFDCA771491579FEAAFF54FD59FF2A" ID-CoL="846YD" authority="Garcia" authorityName="Garcia &amp; Miralles-Núñez &amp; Pérez-Fernández" authorityYear="2020" box="[361,738,151,177]" class="Malacostraca" family="Trichoniscidae" genus="Baeticoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="species" species="bullonorum" status="sp. nov.">
<emphasis id="09BB7B3671491579FEAAFF54FD34FF2A" box="[361,655,151,177]" italics="true" pageId="3" pageNumber="260">Baeticoniscus bullonorum</emphasis>
Garcia
</taxonomicName>
,
<taxonomicNameLabel id="1288C64D71491579FD2DFF54FC91FF2A" box="[750,810,151,177]" pageId="3" pageNumber="260" rank="species">n. sp.</taxonomicNameLabel>
(by monotypy)
</paragraph>
</subSubSection>
<subSubSection id="73D5F4AF71491579FF04FF78FE35FF62" pageId="3" pageNumber="260" type="etymology">
<paragraph id="3B70A72471491579FF04FF78FE35FF62" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="260">
<emphasis id="09BB7B3671491579FF04FF78FEF5FF4E" bold="true" box="[199,334,187,213]" pageId="3" pageNumber="260">Etymology.</emphasis>
From Baetica, Roman province of Hispania, in the south of the Iberian Peninsula, and oνίσκος (onískos), woodlouse.
</paragraph>
</subSubSection>
<subSubSection id="73D5F4AF71491579FF04FEC0FAF4FB72" pageId="3" pageNumber="260" type="discussion">
<paragraph id="3B70A72471491579FF04FEC0FE69FD82" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="260">
<emphasis id="09BB7B3671491579FF04FEC0FE80FE86" bold="true" box="[199,315,259,285]" pageId="3" pageNumber="260">Remarks.</emphasis>
Within the
<taxonomicName id="FCCFDCA771491579FE7EFEC0FD34FE86" authorityName="Verhoeff. A" authorityYear="1908" box="[445,655,259,285]" class="Malacostraca" family="Trichoniscidae" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="subFamily" subFamily="Haplophthalminae">Haplophthalminae</taxonomicName>
, only three known genera
<taxonomicName id="FCCFDCA771491579FC09FEC0FB56FE86" authority="Vandel, 1955" authorityName="Vandel" authorityYear="1955" box="[970,1261,258,285]" class="Malacostraca" family="Trichoniscidae" genus="Acteoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FC09FEC0FBEBFE86" box="[970,1104,259,285]" italics="true" pageId="3" pageNumber="260">Acteoniscus</emphasis>
<bibRefCitation id="5F5EDAD571491579FB95FEC0FB56FE86" author="Vandel, A." box="[1110,1261,258,285]" pageId="3" pageNumber="260" refId="ref5372" refString="Vandel, A. (1955) Isopodes terrestres recoltes dans les grottes de l'Attique par M. Pierre Strinati. Notes biospeleologiques, 10," type="book" year="1955">Vandel, 1955</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FCCFDCA771491579FB34FEC0FEF6FEDA" authority="Brian, 1958" authorityName="Brian" authorityYear="1958" class="Malacostraca" family="Trichoniscidae" genus="Paracyphoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FB34FEC0FF07FEDB" italics="true" pageId="3" pageNumber="260">Paracyphoniscus</emphasis>
<bibRefCitation id="5F5EDAD571491579FF05FEE4FEF6FEDA" author="Brian, A." box="[198,333,294,321]" pageId="3" pageNumber="260" refId="ref3878" refString="Brian, A. (1958) Intorno ad alcuni trichoniscidi raccolti in grotte vicentine e trevigiane dai signori Prof. C. Conci, L. Tamanini e G. Meggiolaro, con descrizione di un nuovo genere (isopodi terrestri). Tial, Genoa, 17 pp." type="book" year="1958">Brian, 1958</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="FCCFDCA771491579FE4BFEE4FCB4FEDA" authority="Tabacaru, 1993" authorityName="Tabacaru" authorityYear="1993" box="[392,783,295,321]" class="Malacostraca" family="Trichoniscidae" genus="Strouhaloniscellus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FE4BFEE4FDECFEDA" box="[392,599,295,321]" italics="true" pageId="3" pageNumber="260">Strouhaloniscellus</emphasis>
<bibRefCitation id="5F5EDAD571491579FDA2FEE4FCB4FEDA" author="Tabacaru, I." box="[609,783,295,321]" pageId="3" pageNumber="260" pagination="43 - 85" refId="ref5173" refString="Tabacaru, I. (1993) Sur la classification des Trichoniscidae et la position systematique de Thaumatoniscellus orghidani Tabacaru, 1973 (Crustacea, Isopoda, Oniscidea). Travaux de l'Institut de Speologie &quot; Emile Racovitza &quot;, 32, 43 - 85." type="journal article" year="1993">Tabacaru, 1993</bibRefCitation>
</taxonomicName>
have the third pleonite completely smooth, which differentiates them from the rest of the genera of this subfamily (
<bibRefCitation id="5F5EDAD571491579FCA7FE88FC43FEFE" author="Vandel, A." box="[868,1016,330,357]" pageId="3" pageNumber="260" refId="ref5372" refString="Vandel, A. (1955) Isopodes terrestres recoltes dans les grottes de l'Attique par M. Pierre Strinati. Notes biospeleologiques, 10," type="book" year="1955">Vandel 1955</bibRefCitation>
,
<bibRefCitation id="5F5EDAD571491579FBC6FE88FBFBFEFE" author="Vandel, A." box="[1029,1088,331,357]" pageId="3" pageNumber="260" refId="ref5398" refString="51 - 61. Vandel, A. (1960) Isopodes terrestres (premiere partie). Faune de France. Vol. 64. Editions Paul Lechevalier, Paris, 416 pp. Wallace, L. &amp; Berrocal, J. A. (2002) s. n. Guia de las cuevas de Malaga. Biblioteca Popular Malaguena, 90, Diputacion de" type="book" year="1960">1960</bibRefCitation>
;
<bibRefCitation id="5F5EDAD571491579FB8DFE88FB6BFEFE" author="Brian, A." box="[1102,1232,330,357]" pageId="3" pageNumber="260" refId="ref3878" refString="Brian, A. (1958) Intorno ad alcuni trichoniscidi raccolti in grotte vicentine e trevigiane dai signori Prof. C. Conci, L. Tamanini e G. Meggiolaro, con descrizione di un nuovo genere (isopodi terrestri). Tial, Genoa, 17 pp." type="book" year="1958">Brian 1958</bibRefCitation>
;
<bibRefCitation id="5F5EDAD571491579FB1DFE88FA36FEFE" author="Tabacaru, I." box="[1246,1421,331,357]" pageId="3" pageNumber="260" pagination="43 - 85" refId="ref5173" refString="Tabacaru, I. (1993) Sur la classification des Trichoniscidae et la position systematique de Thaumatoniscellus orghidani Tabacaru, 1973 (Crustacea, Isopoda, Oniscidea). Travaux de l'Institut de Speologie &quot; Emile Racovitza &quot;, 32, 43 - 85." type="journal article" year="1993">Tabacaru, 1993</bibRefCitation>
).
<taxonomicName id="FCCFDCA771491579FF54FEACFE88FE13" authorityName="Garcia &amp; Miralles-Núñez &amp; Pérez-Fernández" authorityYear="2020" box="[151,307,367,392]" class="Malacostraca" family="Trichoniscidae" genus="Baeticoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FF54FEACFE88FE13" box="[151,307,367,392]" italics="true" pageId="3" pageNumber="260">Baeticoniscus</emphasis>
</taxonomicName>
is morphologically very close to
<taxonomicName id="FCCFDCA771491579FD02FEACFC52FE12" authority="Vandel, 1955" authorityName="Vandel" authorityYear="1955" box="[705,1001,366,393]" class="Malacostraca" family="Trichoniscidae" genus="Acteoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FD02FEACFCFCFE12" box="[705,839,367,393]" italics="true" pageId="3" pageNumber="260">Acteoniscus</emphasis>
<bibRefCitation id="5F5EDAD571491579FC92FEACFC52FE12" author="Vandel, A." box="[849,1001,366,393]" pageId="3" pageNumber="260" refId="ref5372" refString="Vandel, A. (1955) Isopodes terrestres recoltes dans les grottes de l'Attique par M. Pierre Strinati. Notes biospeleologiques, 10," type="book" year="1955">Vandel, 1955</bibRefCitation>
</taxonomicName>
and to
<taxonomicName id="FCCFDCA771491579FB8BFEACFA23FE12" authority="Brian 1958" authorityName="Brian" authorityYear="1958" box="[1096,1432,366,393]" class="Malacostraca" family="Trichoniscidae" genus="Paracyphoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FB8BFEACFAB2FE12" box="[1096,1289,367,393]" italics="true" pageId="3" pageNumber="260">Paracyphoniscus</emphasis>
<bibRefCitation id="5F5EDAD571491579FAD7FEACFA23FE12" author="Brian, A." box="[1300,1432,366,393]" pageId="3" pageNumber="260" refId="ref3878" refString="Brian, A. (1958) Intorno ad alcuni trichoniscidi raccolti in grotte vicentine e trevigiane dai signori Prof. C. Conci, L. Tamanini e G. Meggiolaro, con descrizione di un nuovo genere (isopodi terrestri). Tial, Genoa, 17 pp." type="book" year="1958">Brian 1958</bibRefCitation>
</taxonomicName>
, both monotypic and only known from
<collectingCountry id="43D8E7B471491579FD9CFE50FD15FE36" box="[607,686,403,429]" name="Greece" pageId="3" pageNumber="260">Greece</collectingCountry>
and northeast
<collectingCountry id="43D8E7B471491579FC9DFE50FC2AFE36" box="[862,913,403,429]" name="Italy" pageId="3" pageNumber="260">Italy</collectingCountry>
respectively (
<bibRefCitation id="5F5EDAD571491579FBF6FE50FABBFE36" author="Schmalfuss, H." box="[1077,1280,403,429]" pageId="3" pageNumber="260" pagination="1 - 341" refId="ref4865" refString="Schmalfuss, H. (2003) World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde, Series A, 654, 1 - 341." type="journal article" year="2003">Schmalfuss 2003</bibRefCitation>
).
<taxonomicName id="FCCFDCA771491579FAD5FE50FE00FE4A" authority="Vandel, 1955" authorityName="Vandel" authorityYear="1955" class="Malacostraca" family="Trichoniscidae" genus="Acteoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="species" species="petrochilosi">
<emphasis id="09BB7B3671491579FAD5FE50FEA7FE4A" italics="true" pageId="3" pageNumber="260">Acteoniscus petrochilosi</emphasis>
<bibRefCitation id="5F5EDAD571491579FEE6FE74FE00FE4A" author="Vandel, A." box="[293,443,438,465]" pageId="3" pageNumber="260" refId="ref5372" refString="Vandel, A. (1955) Isopodes terrestres recoltes dans les grottes de l'Attique par M. Pierre Strinati. Notes biospeleologiques, 10," type="book" year="1955">Vandel, 1955</bibRefCitation>
</taxonomicName>
is only known from the Keratea cave, in Attiki (
<bibRefCitation id="5F5EDAD571491579FC32FE74FAE5FE4A" author="Alexiou, S. &amp; Sfenthourakis, S." box="[1009,1374,439,465]" pageId="3" pageNumber="260" pagination="3 - 50" refId="ref3698" refString="Alexiou, S. &amp; Sfenthourakis, S. (2013) The terrestrial Isopods (Isopoda: Oniscidea) of Greece. Parnassia Archives, 1, 3 - 50." type="journal article" year="2013">Alexiou &amp; Sfenthourakis, 2013</bibRefCitation>
), and
<taxonomicName id="FCCFDCA771491579FF54FE18FDCDFE6E" authority="Brian, 1958" authorityName="Brian" authorityYear="1958" box="[151,630,474,501]" class="Malacostraca" family="Trichoniscidae" genus="Paracyphoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="species" species="meggiolaroi">
<emphasis id="09BB7B3671491579FF54FE18FE53FE6E" box="[151,488,475,501]" italics="true" pageId="3" pageNumber="260">Paracyphoniscus meggiolaroi</emphasis>
<bibRefCitation id="5F5EDAD571491579FE32FE18FDCDFE6E" author="Brian, A." box="[497,630,474,501]" pageId="3" pageNumber="260" refId="ref3878" refString="Brian, A. (1958) Intorno ad alcuni trichoniscidi raccolti in grotte vicentine e trevigiane dai signori Prof. C. Conci, L. Tamanini e G. Meggiolaro, con descrizione di un nuovo genere (isopodi terrestri). Tial, Genoa, 17 pp." type="book" year="1958">Brian, 1958</bibRefCitation>
</taxonomicName>
has been collected only from a few caves in the Eastern Alps (
<bibRefCitation id="5F5EDAD571491579FA86FE18FF69FD82" author="Paoletti, M. G." pageId="3" pageNumber="260" pagination="546 - 577" refId="ref4527" refString="Paoletti, M. G. (1982) L'artropodofauna ipogea delle Alpi Orientali dal confine orientale al Garda. Con particolare riguardo a Bathysciinae ed Isopodi terrestri. Lavori della Societa' Italiana di biogeografia, New Series, 1978, 546 - 577. https: // doi. org / 10.21426 / B 67110180" type="journal article" year="1982">Paoletti 1982</bibRefCitation>
;
<bibRefCitation id="5F5EDAD571491579FF1DFE3CFE7DFD82" author="Ruffo, S. &amp; Stoch, F." box="[222,454,510,537]" pageId="3" pageNumber="260" pagination="1 - 307" refId="ref4822" refString="Ruffo, S. &amp; Stoch, F. (2005) Checklist e distribuzione della fauna italiana. Memorie del Museo Civico di Storia Naturale di Verona, Serie 2, Sezione Scienze della Vita, 16, 1 - 307." type="journal article" year="2005">Ruffo &amp; Stoch 2005</bibRefCitation>
).
</paragraph>
<paragraph id="3B70A72471491579FF04FDE0FD9EFD55" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="260">
However, the first pleopods of the males of their respective
<typeStatus id="E474198671491579FCADFDE0FC25FDA6" box="[878,926,547,573]" pageId="3" pageNumber="260">type</typeStatus>
species are very different.
<taxonomicName id="FCCFDCA771491579FB11FDE0FAD5FDA7" authorityName="Garcia &amp; Miralles-Núñez &amp; Pérez-Fernández" authorityYear="2020" box="[1234,1390,547,572]" class="Malacostraca" family="Trichoniscidae" genus="Baeticoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FB11FDE0FAD5FDA7" box="[1234,1390,547,572]" italics="true" pageId="3" pageNumber="260">Baeticoniscus</emphasis>
</taxonomicName>
can be distinguished morphologically from both genera by the cephalic structure and the shape and arrangement of the tubercles of the pereon, and resembles them by presenting the third pleonite completely smooth, a rare morphological feature in the
<taxonomicName id="FCCFDCA771491579FE9AFD4CFD93FD32" authorityName="Verhoeff. A" authorityYear="1908" box="[345,552,655,681]" class="Malacostraca" family="Trichoniscidae" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="subFamily" subFamily="Haplophthalminae">Haplophthalminae</taxonomicName>
subfamily since, in all other known genera, there exist ribs or protuberances in this body part (
<bibRefCitation id="5F5EDAD571491579FEFCFD70FE6AFD56" author="Vandel, A." box="[319,465,690,717]" pageId="3" pageNumber="260" refId="ref5372" refString="Vandel, A. (1955) Isopodes terrestres recoltes dans les grottes de l'Attique par M. Pierre Strinati. Notes biospeleologiques, 10," type="book" year="1955">Vandel 1955</bibRefCitation>
,
<bibRefCitation id="5F5EDAD571491579FE1FFD70FDA3FD55" author="Vandel, A." box="[476,536,691,718]" pageId="3" pageNumber="260" refId="ref5398" refString="51 - 61. Vandel, A. (1960) Isopodes terrestres (premiere partie). Faune de France. Vol. 64. Editions Paul Lechevalier, Paris, 416 pp. Wallace, L. &amp; Berrocal, J. A. (2002) s. n. Guia de las cuevas de Malaga. Biblioteca Popular Malaguena, 90, Diputacion de" type="book" year="1960">1960</bibRefCitation>
).
</paragraph>
<paragraph id="3B70A72471491579FF04FD14FAF2FC1A" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="260">
The first pleopods of the male of
<taxonomicName id="FCCFDCA771491579FDF3FD14FD71FD6B" authorityName="Garcia &amp; Miralles-Núñez &amp; Pérez-Fernández" authorityYear="2020" box="[560,714,727,752]" class="Malacostraca" family="Trichoniscidae" genus="Baeticoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FDF3FD14FD71FD6B" box="[560,714,727,752]" italics="true" pageId="3" pageNumber="260">Baeticoniscus</emphasis>
</taxonomicName>
are of the same
<typeStatus id="E474198671491579FCB9FD14FC11FD6A" box="[890,938,727,753]" pageId="3" pageNumber="260">type</typeStatus>
as those of the major part of
<taxonomicName id="FCCFDCA771491579FB20FD14FA27FD6A" box="[1251,1436,727,753]" class="Malacostraca" family="Trichoniscidae" genus="Haplophthalmus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FB20FD14FA27FD6A" box="[1251,1436,727,753]" italics="true" pageId="3" pageNumber="260">Haplophthalmus</emphasis>
</taxonomicName>
species (i.e. a thick endopod, with striated subconical, or more or less dilated, tip) and different from those of
<taxonomicName id="FCCFDCA771491579FA97FD38FF67FCA3" authorityName="Vandel" authorityYear="1955" class="Malacostraca" family="Trichoniscidae" genus="Acteoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FA97FD38FF67FCA3" italics="true" pageId="3" pageNumber="260">Acteoniscus</emphasis>
</taxonomicName>
and
<taxonomicName id="FCCFDCA771491579FED3FCDCFE74FCA2" authorityName="Brian" authorityYear="1958" box="[272,463,799,825]" class="Malacostraca" family="Trichoniscidae" genus="Paracyphoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FED3FCDCFE74FCA2" box="[272,463,799,825]" italics="true" pageId="3" pageNumber="260">Paracyphoniscus</emphasis>
</taxonomicName>
.
<taxonomicName id="FCCFDCA771491579FE1FFCDCFDDBFCA2" authorityName="Vandel" authorityYear="1955" box="[476,608,799,825]" class="Malacostraca" family="Trichoniscidae" genus="Acteoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FE1FFCDCFDDBFCA2" box="[476,608,799,825]" italics="true" pageId="3" pageNumber="260">Acteoniscus</emphasis>
</taxonomicName>
has the endopod of the first male pleopod showing a striated distal part, but it is narrow and ends with an acute tip (
<bibRefCitation id="5F5EDAD571491579FD94FC80FD50FCC6" author="Vandel, A." box="[599,747,834,861]" pageId="3" pageNumber="260" refId="ref5372" refString="Vandel, A. (1955) Isopodes terrestres recoltes dans les grottes de l'Attique par M. Pierre Strinati. Notes biospeleologiques, 10," type="book" year="1955">Vandel 1955</bibRefCitation>
).
<taxonomicName id="FCCFDCA771491579FD3CFC80FC05FCC6" authorityName="Brian" authorityYear="1958" box="[767,958,835,861]" class="Malacostraca" family="Trichoniscidae" genus="Paracyphoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FD3CFC80FC05FCC6" box="[767,958,835,861]" italics="true" pageId="3" pageNumber="260">Paracyphoniscus</emphasis>
</taxonomicName>
, according to the illustrations provided by
<bibRefCitation id="5F5EDAD571491579FF54FCA4FE93FC1A" author="Brian, A." box="[151,296,870,897]" pageId="3" pageNumber="260" refId="ref3878" refString="Brian, A. (1958) Intorno ad alcuni trichoniscidi raccolti in grotte vicentine e trevigiane dai signori Prof. C. Conci, L. Tamanini e G. Meggiolaro, con descrizione di un nuovo genere (isopodi terrestri). Tial, Genoa, 17 pp." type="book" year="1958">Brian (1958)</bibRefCitation>
, has the same appendix terminating in a subtriangular extension and an overlapping expansion.
</paragraph>
<paragraph id="3B70A72471491579FF04FC48FB0DFB8A" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="260">
The differences between these genera and
<taxonomicName id="FCCFDCA771491579FD67FC48FCC8FC3E" authorityName="Tabacaru" authorityYear="1993" box="[676,883,907,933]" class="Malacostraca" family="Trichoniscidae" genus="Strouhaloniscellus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FD67FC48FCC8FC3E" box="[676,883,907,933]" italics="true" pageId="3" pageNumber="260">Strouhaloniscellus</emphasis>
</taxonomicName>
are much more important, especially with regard to the structure of the first male pleopod, the tegumentary structures and the shape of the pereonites, amongst other morphological characters (see
<bibRefCitation id="5F5EDAD571491579FE2FFC10FD2FFC76" author="Tabacaru, I." box="[492,660,979,1005]" pageId="3" pageNumber="260" pagination="43 - 85" refId="ref5173" refString="Tabacaru, I. (1993) Sur la classification des Trichoniscidae et la position systematique de Thaumatoniscellus orghidani Tabacaru, 1973 (Crustacea, Isopoda, Oniscidea). Travaux de l'Institut de Speologie &quot; Emile Racovitza &quot;, 32, 43 - 85." type="journal article" year="1993">Tabacaru 1993</bibRefCitation>
;
<bibRefCitation id="5F5EDAD571491579FD63FC10FC3BFC76" author="Bedek, J. &amp; Taiti, S." box="[672,896,979,1005]" pageId="3" pageNumber="260" pagination="59 - 64" refId="ref3791" refString="Bedek, J. &amp; Taiti, S. (2009) A new species of Strouhaloniscellus Tabacaru, 1993 (Crustacea: Isopoda: Oniscidea) from a cave in Mt Biokovo, Croatia. Zootaxa, 2196 (1), 59 - 64. https: // doi. org / 10.11646 / zootaxa. 2196.1.5" type="journal article" year="2009">Bedek &amp; Taiti 2009</bibRefCitation>
). This genus contains two species:
<taxonomicName id="FCCFDCA771491579FAC5FC10FE38FB8A" authority="(Strouhal, 1939)" baseAuthorityName="Strouhal" baseAuthorityYear="1939" class="Malacostraca" family="Trichoniscidae" genus="Strouhaloniscellus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="species" species="anophthalmus">
<emphasis id="09BB7B3671491579FAC5FC10FF7FFB8B" italics="true" pageId="3" pageNumber="260">S. anophthalmus</emphasis>
(
<bibRefCitation id="5F5EDAD571491579FF10FC34FEC0FB8A" author="Strouhal, H." box="[211,379,1015,1041]" pageId="3" pageNumber="260" pagination="291 - 307" refId="ref5144" refString="Strouhal, H. (1939) Landasseln aus Balkanhohlen, gesammelt von Prof. Dr. Karl Absolon. Zoologischer Anzeiger, 128, 291 - 307." type="journal article" year="1939">Strouhal, 1939</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="FCCFDCA771491579FE7AFC34FCDDFB8A" authority="(Bedek &amp; Taiti, 2009)" baseAuthorityName="Bedek &amp; Taiti" baseAuthorityYear="2009" box="[441,870,1015,1041]" class="Malacostraca" family="Trichoniscidae" genus="Strouhaloniscellus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="species" species="biokovoensis">
<emphasis id="09BB7B3671491579FE7AFC34FDDDFB8A" box="[441,614,1015,1041]" italics="true" pageId="3" pageNumber="260">S. biokovoensis</emphasis>
(
<bibRefCitation id="5F5EDAD571491579FDB7FC34FCE5FB8A" author="Bedek, J. &amp; Taiti, S." box="[628,862,1015,1041]" pageId="3" pageNumber="260" pagination="59 - 64" refId="ref3791" refString="Bedek, J. &amp; Taiti, S. (2009) A new species of Strouhaloniscellus Tabacaru, 1993 (Crustacea: Isopoda: Oniscidea) from a cave in Mt Biokovo, Croatia. Zootaxa, 2196 (1), 59 - 64. https: // doi. org / 10.11646 / zootaxa. 2196.1.5" type="journal article" year="2009">Bedek &amp; Taiti, 2009</bibRefCitation>
)
</taxonomicName>
, both from the Balkan region.
</paragraph>
<paragraph id="3B70A72471491579FF04FBD8FA9CFBE6" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="260">
On the other hand a comparison with
<taxonomicName id="FCCFDCA771491579FDB3FBD8FD49FBAE" authorityName="Vandel" authorityYear="1952" box="[624,754,1051,1077]" class="Malacostraca" family="Trichoniscidae" genus="Iberoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FDB3FBD8FD49FBAE" box="[624,754,1051,1077]" italics="true" pageId="3" pageNumber="260">Iberoniscus</emphasis>
</taxonomicName>
is necessary, since this genus is also represented in the same area of the Iberian Peninsula and it has been collected in the same locality as
<taxonomicName id="FCCFDCA771491579FC2BFBFCFB3FFBC3" authority="Garcia &amp; Miralles-Núñez &amp; Pérez-Fernández, 2020" authorityName="Garcia &amp; Miralles-Núñez &amp; Pérez-Fernández" authorityYear="2020" box="[1000,1156,1087,1112]" class="Malacostraca" family="Trichoniscidae" genus="Baeticoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="09BB7B3671491579FC2BFBFCFB3FFBC3" box="[1000,1156,1087,1112]" italics="true" pageId="3" pageNumber="260">Baeticoniscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1288C64D71491579FB4AFBFCFB6FFBC2" box="[1161,1236,1087,1113]" pageId="3" pageNumber="260" rank="genus">n. gen.</taxonomicNameLabel>
<taxonomicName id="FCCFDCA771491579FB19FBFCFAE7FBC2" authorityName="Vandel" authorityYear="1952" box="[1242,1372,1087,1113]" class="Malacostraca" family="Trichoniscidae" genus="Iberoniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FB19FBFCFAE7FBC2" box="[1242,1372,1087,1113]" italics="true" pageId="3" pageNumber="260">Iberoniscus</emphasis>
</taxonomicName>
show the same
<typeStatus id="E474198671491579FEC2FBA0FE8AFBE6" box="[257,305,1123,1149]" pageId="3" pageNumber="260">type</typeStatus>
of male first pleopod as
<taxonomicName id="FCCFDCA771491579FD84FBA0FD58FBE7" authorityName="Garcia &amp; Miralles-Núñez &amp; Pérez-Fernández" authorityYear="2020" box="[583,739,1123,1148]" class="Malacostraca" family="Trichoniscidae" genus="Baeticoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="09BB7B3671491579FD84FBA0FD58FBE7" box="[583,739,1123,1148]" italics="true" pageId="3" pageNumber="260">Baeticoniscus</emphasis>
</taxonomicName>
but it has a very large tubercle in the third pleonite.
</paragraph>
<paragraph id="3B70A72471491579FF04FB44FAF4FB72" blockId="3.[151,1437,150,1257]" pageId="3" pageNumber="260">Therefore, given the complexity of this group and while it is not possible to undertake a complete revision of the subfamily through molecular analysis, it is considered convenient to establish a new genus for these specimens from the south of the Iberian Peninsula, based on their distinctive combination of morphological characters.</paragraph>
</subSubSection>
</treatment>
</document>