treatments-xml/data/B3/5D/66/B35D667A002DB3039DD2F93EFAD6DFCF.xml

<document id="8C27BE57AA3EB784D84AE880E3059F62" ID-DOI="10.5281/zenodo.176562" ID-GBIF-Dataset="04dc18fd-f14c-487b-b693-c9bd0bffd3c8" ID-ISSN="1175-5326" ID-Zenodo-Dep="176562" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459835818516" checkinUser="plazi" docAuthor="Shirokaya, Alena" docDate="2007" docId="B35D667A002DB3039DD2F93EFAD6DFCF" docLanguage="en" docName="zt01466p067.pdf" docOrigin="Zootaxa 1466" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Gerstfeldtiancylus ushunensis Shirokaya, 2007, sp. nov." docType="treatment" docVersion="10" lastPageNumber="64" masterDocId="4F641E02002CB30A9D45FFE1FFABDE00" masterDocTitle="A new species of Gerstfeldtiancylus Starobogatov, 1989 (Pulmonata: Basommatophora: Acroloxidae) from Lake Baikal" masterLastPageNumber="67" masterPageNumber="55" pageNumber="56" updateTime="1698223613284" updateUser="plazi">
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<mods:title id="EEC8562DA9EE015FEFC0956249EB92F5">A new species of Gerstfeldtiancylus Starobogatov, 1989 (Pulmonata: Basommatophora: Acroloxidae) from Lake Baikal</mods:title>
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<mods:namePart id="D20CFF3CAB033952F5EC6F054924298D">Shirokaya, Alena</mods:namePart>
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<mods:date id="7194D7C2DF004A2D1D5361FF4482583E">2007</mods:date>
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<treatment id="B35D667A002DB3039DD2F93EFAD6DFCF" ID-DOI="http://doi.org/10.5281/zenodo.5618836" ID-GBIF-Taxon="119358876" ID-Zenodo-Dep="5618836" LSID="urn:lsid:plazi:treatment:B35D667A002DB3039DD2F93EFAD6DFCF" httpUri="http://treatment.plazi.org/id/B35D667A002DB3039DD2F93EFAD6DFCF" lastPageId="9" lastPageNumber="64" pageId="1" pageNumber="56">
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<paragraph id="3B4BD76C002DB30B9DD2F93EFDCCD8F9" blockId="1.[151,615,1759,1820]" box="[151,615,1759,1785]" pageId="1" pageNumber="56">
<heading id="60036000002DB30B9DD2F93EFDCCD8F9" bold="true" box="[151,615,1759,1785]" fontSize="11" level="1" pageId="1" pageNumber="56" reason="1">
<emphasis id="09800B7E002DB30B9DD2F93EFDCCD8F9" bold="true" box="[151,615,1759,1785]" pageId="1" pageNumber="56">
<taxonomicName id="FCF4ACEF002DB30B9DD2F93EFDA9D8F9" ID-CoL="6KFSZ" box="[151,514,1759,1785]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="1" pageNumber="56" phylum="Mollusca" rank="species" species="ushunensis" status="sp. nov.">
<emphasis id="09800B7E002DB30B9DD2F93EFDA9D8F9" bold="true" box="[151,514,1759,1785]" italics="true" pageId="1" pageNumber="56">Gerstfeldtiancylus ushunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="12B3B605002DB30B9F4CF93EFDCCD8F9" box="[521,615,1759,1785]" pageId="1" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3B4BD76C002DB30B9DD2F8E3FEBBD91C" blockId="1.[151,615,1759,1820]" box="[151,272,1794,1820]" pageId="1" pageNumber="56">
(
<figureCitation id="A3CFCBE9002DB30B9DDAF8E3FEA3D91C" box="[159,264,1794,1820]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5. A – B" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStartId-0="5.[151,255,1488,1512]" captionStartId-1="6.[151,255,1716,1740]" captionStartId-2="7.[151,255,1718,1742]" captionStartId-3="8.[151,255,1659,1683]" captionStartId-4="9.[151,255,1900,1924]" captionTargetBox-0="[264,1322,194,1463]" captionTargetBox-1="[264,1322,1146,1692]" captionTargetBox-2="[274,1303,187,1701]" captionTargetBox-3="[290,1306,186,1659]" captionTargetBox-4="[288,1303,533,1860]" captionTargetId-0="figure@5.[264,1323,194,1464]" captionTargetId-1="figure@6.[264,1323,1146,1692]" captionTargetId-2="figure@7.[264,1323,170,1720]" captionTargetId-3="figure@8.[275,1316,172,1659]" captionTargetId-4="figure@9.[264,1323,500,1876]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="FIGURE 3. Shells of Gerstfeldtiancylus ushunensis, sp. nov. (SEM). A – D. Teleoconch (ZIN RAS 1 / 521 – 2006): A. Top view. B. Right side view. C. Left side view. D. Rear view. E – G. Protoconch (LIN SB RAS 905): E. Top view. F. Right side view. G. Left side view. H. Initial protoconch plate. I – K. Protoconch sculpture (LIN SB RAS 904). Scale bars: A – D = 2.0 mm; E = 1.0 mm; F – G = 0.25 mm; H, J = 0.1 mm; I = 0.2 mm; K = 0.02 mm." captionText-1="FIGURE 4. Radula of Gerstfeldtiancylus ushunensis, sp. nov. (SEM). A – C. ZIN RAS 2 / 521 – 2006: A. Central tooth. B. Three transverse rows of teeth. C. Lateral teeth. Scale bars: A = 0.01 mm; B – C = 0.1 mm." captionText-2="FIGURE 5. A – B. Anatomy of Gerstfeldtiancylus ushunensis, sp. nov.. А. General topography of main organs of the alimentary system, dorsal view. Digestive gland removed. B. Dorsal view of specimen without shell. C – D. Odontophore of G. ushunensis, sp. nov.. E – L. Jaws of different Gerstfeldtiancylus species. E – H. G. ushunensis, sp. nov. (A, C – E. ZIN RAS 1 / 521 – 2006. B, F – H. ZIN RAS 2 / 521 – 2006). I. G. renardii (Dybowski, 1884). J. G. roepstorfi Shirokaya et al., 2003. K. G. benedictiae Starobogatov, 1989 (whole mounts in Faure-Berlezet liquid). L. G. kotyensis Starobogatov, 1989 (frontal histological section). I – L. From Shirokaya &amp; Röpstorf (2004). aal, left anterior adductor; aar, right anterior adductor; ap, anal pore; bm, buccal mass; cae, caecum; dgp, digestive gland pores; int 1, first intestinal loop; int 2, second intestinal loop; mb, mantle border; oa, original or posterior adductor; oe, oesophagus; rs, radular sac; sg, salivary glands; st, stomach. Scale bars: A – D = 1 mm; E = 0.8 mm; F – H, J = 0.5 mm; I, K – L = 0.3 mm." captionText-3="FIGURE 6. Longitudinally sectioned male copulatory organ of Gerstfeldtiancylus species (A – D, G. diagrammic). A. G. benedictiae Starobogatov, 1989. B, E. G. roepstorfi Shirokaya et al., 2003. C. G. kotyensis Starobogatov, 1989. D. G. renardii (Dybowski, 1884). A – E. From Kruglov &amp; Starobogatov (1991 a), Shirokaya (2005). F – G. G. ushunensis, sp. nov. (ZIN RAS 2 / 521 – 2006). fl, flagellum; mp, male genital pore; p, penis; ps, penis sheath; psr, penis sheath retractor; pr, preputium; prc, preputium chamber; prp, preputium pilasters; sar, sarcobellum; vd, vas deferens; ve, velum. Scale bars: A = 0.5 mm; B – G = 1 mm." captionText-4="FIGURE 7. Egg masses of Gerstfeldtiancylus ushunensis, sp. nov.. A – C. Lateral and dorsal view (going from top to bottom). cam, capsular albuminous matrix; cm, capsular membrane; emb, embryo; et, extrasyncapsular tunic; sm, syncapsular matrix; stu, syncapsular tunic; tt, terminal tail. All scale bars: 2 mm." httpUri-0="https://zenodo.org/record/176565/files/figure.png" httpUri-1="https://zenodo.org/record/176566/files/figure.png" httpUri-2="https://zenodo.org/record/176567/files/figure.png" httpUri-3="https://zenodo.org/record/176568/files/figure.png" httpUri-4="https://zenodo.org/record/176569/files/figure.png" pageId="1" pageNumber="56">Figs 3–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="73EE84E7002DB30B9DD2F8B3FE19D9E4" pageId="1" pageNumber="56" type="materials_examined">
<paragraph id="3B4BD76C002DB30B9DD2F8B3FE19D9E4" blockId="1.[151,1436,1874,2020]" pageId="1" pageNumber="56">
<emphasis id="09800B7E002DB30B9DD2F8B3FEE5D96C" bold="true" box="[151,334,1874,1900]" pageId="1" pageNumber="56">
<typeStatus id="E44F69CE002DB30B9DD2F8B3FF79D96C" box="[151,210,1874,1900]" pageId="1" pageNumber="56">Type</typeStatus>
material:
</emphasis>
Sixteen specimens from the 
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locality. 
<typeStatus id="E44F69CE002DB30B9E0DF8B3FC11D96C" box="[840,954,1874,1900]" pageId="1" pageNumber="56" type="holotype">Holotype</typeStatus>
: 
<collectingCountry id="43E397FC002DB30B9E80F8B3FBBDD96C" box="[965,1046,1874,1900]" name="Russia" pageId="1" pageNumber="56">Russia</collectingCountry>
, Lake Baikal, Malye Olkhonskie Vorota, Ushun Bay, 
<geoCoordinate id="5EC0B1AB002DB30B9CD4F89BFD9AD994" box="[401,561,1914,1940]" direction="north" orientation="latitude" pageId="1" pageNumber="56" precision="15" value="53.015">53°00’54’’ N</geoCoordinate>
<geoCoordinate id="5EC0B1AB002DB30B9F78F89BFD41D994" box="[573,746,1914,1940]" direction="east" orientation="longitude" pageId="1" pageNumber="56" precision="15" value="106.96">106°56’96’’ E</geoCoordinate>
, 
<quantity id="FC0C7A89002DB30B9FB9F89BFCC3D994" box="[764,872,1914,1940]" metricMagnitude="1" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="1" pageNumber="56" unit="m" value="21.0" valueMax="22.0" valueMin="20.0">20–22 m</quantity>
depth, on stones, coll. I.V. Khanaev and V.F. Skudenko 0 7.07.2003 (
<collectionCode id="5DE54FA9002DB30B9CE3F843FE72D9BC" box="[422,473,1954,1980]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="56">ZIN</collectionCode>
<collectionCode id="5DE54FA9002DB30B9C9AF843FDB3D9BC" box="[479,536,1954,1980]" pageId="1" pageNumber="56">RAS</collectionCode>
1/521–2006). 
<typeStatus id="E44F69CE002DB30B9F81F843FC91D9BC" box="[708,826,1954,1980]" pageId="1" pageNumber="56" type="paratype">Paratypes</typeStatus>
: 
<quantity id="FC0C7A89002DB30B9E01F843FC36D9BC" box="[836,925,1954,1980]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="2.2" metricValueMin="0.4" pageId="1" pageNumber="56" unit="m" value="13.0" valueMax="22.0" valueMin="4.0">4–22 m</quantity>
depth (
<collectionCode id="5DE54FA9002DB30B9EB1F843FB8CD9BC" box="[1012,1063,1954,1980]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="56">ZIN</collectionCode>
<collectionCode id="5DE54FA9002DB30B9968F843FBCDD9BC" box="[1069,1126,1954,1980]" pageId="1" pageNumber="56">RAS</collectionCode>
2/521–2006, 3/521–2006; LIN SB 
<collectionCode id="5DE54FA9002DB30B9DBFF82BFE98D9E4" box="[250,307,1994,2020]" pageId="1" pageNumber="56">RAS</collectionCode>
904, 905).
</paragraph>
</subSubSection>
<caption id="6F8B87E4002EB3089DD2F926FF76D9D1" httpUri="https://zenodo.org/record/176563/files/figure.png" pageId="2" pageNumber="57" targetBox="[270,1317,210,1692]" targetPageId="2">
<paragraph id="3B4BD76C002EB3089DD2F926FF76D9D1" blockId="2.[151,1436,1735,2001]" pageId="2" pageNumber="57">
<emphasis id="09800B7E002EB3089DD2F926FEB1D8DF" bold="true" box="[151,282,1735,1759]" pageId="2" pageNumber="57">FIGURE 1.</emphasis>
Schematic maps of Lake Baikal. A. Geographical regions of four shallow-water (0–100 m) provinces. The boundaries of the provinces and regions are indicated according to Starobogatov (1970), modified by Sitnikova (2006). Regions: South-Baikal (SB), Anginsky (ANG), Malomorsky (М), East-Olkhon (EOL), Turkinsky (TUR), Barguzin- Chivyrkuysky (BCH), North-Baikal, eastern coast (NBE), North-Baikal, western coast (NBW), Ushkansky (USH). B. Localities of abundant species of 
<taxonomicName id="FCF4ACEF002EB3089CBAF8B3FC30D969" authority="Starobogatov, 1989" authorityName="Starobogatov" authorityYear="1989" box="[511,923,1874,1897]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="2" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="09800B7E002EB3089CBAF8B3FD68D969" box="[511,707,1874,1897]" italics="true" pageId="2" pageNumber="57">Gerstfeldtiancylus</emphasis>
Starobogatov, 1989
</taxonomicName>
. Symbols same as in C. C. Localities of species occupying disjunctive and local areas. 1. 
<taxonomicName id="FCF4ACEF002EB3089F0AF894FC6AD98B" authority="Starobogatov, 1989" authorityName="Starobogatov" authorityYear="1989" box="[591,961,1908,1932]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="2" pageNumber="57" phylum="Mollusca" rank="species" species="benedictiae">
<emphasis id="09800B7E002EB3089F0AF894FD43D98C" box="[591,744,1909,1932]" italics="true" pageId="2" pageNumber="57">G. benedictiae</emphasis>
Starobogatov, 1989
</taxonomicName>
. 2. 
<emphasis id="09800B7E002EB3089EA0F894FBFED98C" box="[997,1109,1909,1932]" italics="true" pageId="2" pageNumber="57">G. re n a rd i i</emphasis>
(Dybowski, 1884). 3. 
<emphasis id="09800B7E002EB308980CF894FF67D9AF" italics="true" pageId="2" pageNumber="57">G. k o t y - ensis</emphasis>
Starobogatov, 1989. 4. 
<taxonomicName id="FCF4ACEF002EB3089C8CF879FDF0D9AF" box="[457,603,1944,1967]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="2" pageNumber="57" phylum="Mollusca" rank="species" species="ushunensis" status="sp. nov.">
<emphasis id="09800B7E002EB3089C8CF879FDF0D9AF" box="[457,603,1944,1967]" italics="true" pageId="2" pageNumber="57">G. ushunensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="12B3B605002EB3089F2DF876FD16D9AF" box="[616,701,1943,1967]" pageId="2" pageNumber="57" rank="species">
<emphasis id="09800B7E002EB3089F2DF876FD16D9AF" bold="true" box="[616,701,1943,1967]" pageId="2" pageNumber="57">sp. nov.</emphasis>
</taxonomicNameLabel>
. 5. 
<taxonomicName id="FCF4ACEF002EB3089FA1F879FBF6D9AE" authority="Shirokaya et al., 2003" authorityName="Shirokaya et al." authorityYear="2003" box="[740,1117,1943,1967]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="2" pageNumber="57" phylum="Mollusca" rank="species" species="roepstorfi">
<emphasis id="09800B7E002EB3089FA1F879FCC1D9AF" box="[740,874,1944,1967]" italics="true" pageId="2" pageNumber="57">G. roepstorfi</emphasis>
Shirokaya et al., 2003
</taxonomicName>
. B–C. From Shirokaya (2004, 2005).
</paragraph>
</caption>
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<paragraph id="3B4BD76C002FB3099DD2FB7DFD54DB1B" blockId="3.[151,1436,1180,1307]" pageId="3" pageNumber="58">
<emphasis id="09800B7E002FB3099DD2FB7DFEB2DAB4" bold="true" box="[151,281,1180,1204]" pageId="3" pageNumber="58">FIGURE 2.</emphasis>
Measurements of acroloxid shells (after Kruglov &amp; Starobogatov (1991b), modified). L, length of aperture; La, distance from apex to frontal apertural edge (projected on longitudinal axis of aperture); W, width of aperture; wL, distance from apex to left apertural edge; a, chord of anterior (maximal) slope; H, height of teleoconch; l, length of protoconch; w, width of protoconch; h, height of protoconch.
</paragraph>
</caption>
<subSubSection id="73EE84E7002FB3099D82FAAEFBA8DB69" box="[199,1027,1359,1385]" pageId="3" pageNumber="58" type="etymology">
<paragraph id="3B4BD76C002FB3099D82FAAEFBA8DB69" blockId="3.[151,1437,1359,2025]" box="[199,1027,1359,1385]" pageId="3" pageNumber="58">
<emphasis id="09800B7E002FB3099D82FAAEFEFFDB69" bold="true" box="[199,340,1359,1385]" pageId="3" pageNumber="58">Etymology.</emphasis>
The species is named after its 
<typeStatus id="E44F69CE002FB3099FF9FAAEFD45DB69" box="[700,750,1359,1385]" pageId="3" pageNumber="58">type</typeStatus>
locality — Ushun Bay.
</paragraph>
</subSubSection>
<subSubSection id="73EE84E7002FB3099D82FA96FC59D831" pageId="3" pageNumber="58" type="materials_examined">
<paragraph id="3B4BD76C002FB3099D82FA96FC59D831" blockId="3.[151,1437,1359,2025]" pageId="3" pageNumber="58">
<emphasis id="09800B7E002FB3099D82FA96FE17DB91" bold="true" box="[199,444,1399,1425]" pageId="3" pageNumber="58">Material examined.</emphasis>
Teleoconch: 
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(
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<collectionCode id="5DE54FA9002FB3099E54FA96FCE1DB91" box="[785,842,1399,1425]" pageId="3" pageNumber="58">RAS</collectionCode>
1/521–2006) and 15 
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(
<collectionCode id="5DE54FA9002FB3099989FA96FB54DB91" box="[1228,1279,1399,1425]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="58">ZIN</collectionCode>
<collectionCode id="5DE54FA9002FB309984DFA96FAEADB91" box="[1288,1345,1399,1425]" pageId="3" pageNumber="58">RAS</collectionCode>
2/521– 2006, 3/521–2006, LIN SB 
<collectionCode id="5DE54FA9002FB3099CB0FA7EFD85DBB9" box="[501,558,1439,1465]" pageId="3" pageNumber="58">RAS</collectionCode>
904, 905); alimentary system, odontophore and jaw: 
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and ten 
<typeStatus id="E44F69CE002FB3099DD2FA26FEACDBE1" box="[151,263,1479,1505]" pageId="3" pageNumber="58" type="paratype">paratypes</typeStatus>
(
<collectionCode id="5DE54FA9002FB3099C52FA26FEE1DBE1" box="[279,330,1479,1505]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="58">ZIN</collectionCode>
<collectionCode id="5DE54FA9002FB3099C16FA26FE27DBE1" box="[339,396,1479,1505]" pageId="3" pageNumber="58">RAS</collectionCode>
2/521–2006, 3/521–2006); protoconch: four 
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(LIN SB 
<collectionCode id="5DE54FA9002FB30999CBFA26FB6CDBE1" box="[1166,1223,1479,1505]" pageId="3" pageNumber="58">RAS</collectionCode>
904, 905); radula and male copulatory organ: five 
<typeStatus id="E44F69CE002FB3099F50FA0EFD2ED809" box="[533,645,1519,1545]" pageId="3" pageNumber="58" type="paratype">paratypes</typeStatus>
(
<collectionCode id="5DE54FA9002FB3099FD0FA0EFD63D809" box="[661,712,1519,1545]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="58">ZIN</collectionCode>
<collectionCode id="5DE54FA9002FB3099F8AFA0EFCA3D809" box="[719,776,1519,1545]" pageId="3" pageNumber="58">RAS</collectionCode>
2/521–2006); mantle and shell adductor muscles: seven 
<typeStatus id="E44F69CE002FB3099DD2F9F6FEACD831" box="[151,263,1559,1585]" pageId="3" pageNumber="58" type="paratype">paratypes</typeStatus>
(
<collectionCode id="5DE54FA9002FB3099C53F9F6FEE2D831" box="[278,329,1559,1585]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="58">ZIN</collectionCode>
<collectionCode id="5DE54FA9002FB3099C15F9F6FE22D831" box="[336,393,1559,1585]" pageId="3" pageNumber="58">RAS</collectionCode>
2/521–2006). Fourteen syncapsulae were measured.
</paragraph>
</subSubSection>
<subSubSection id="73EE84E7002FB3039D82F9DEFD8EDF52" lastPageId="9" lastPageNumber="64" pageId="3" pageNumber="58" type="description">
<paragraph id="3B4BD76C002FB30E9D82F9DEFCE8DF02" blockId="3.[151,1437,1359,2025]" lastBlockId="4.[151,1436,151,258]" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58">
<emphasis id="09800B7E002FB3099D82F9DEFEF4D859" bold="true" box="[199,351,1599,1625]" pageId="3" pageNumber="58">Description.</emphasis>
Shell (
<figureCitation id="A3CFCBE9002FB3099CF6F9DEFE5CD859" box="[435,503,1599,1625]" captionStart="FIGURE 3" captionStartId="5.[151,255,1488,1512]" captionTargetBox="[264,1322,194,1463]" captionTargetId="figure@5.[264,1323,194,1464]" captionTargetPageId="5" captionText="FIGURE 3. Shells of Gerstfeldtiancylus ushunensis, sp. nov. (SEM). A – D. Teleoconch (ZIN RAS 1 / 521 – 2006): A. Top view. B. Right side view. C. Left side view. D. Rear view. E – G. Protoconch (LIN SB RAS 905): E. Top view. F. Right side view. G. Left side view. H. Initial protoconch plate. I – K. Protoconch sculpture (LIN SB RAS 904). Scale bars: A – D = 2.0 mm; E = 1.0 mm; F – G = 0.25 mm; H, J = 0.1 mm; I = 0.2 mm; K = 0.02 mm." httpUri="https://zenodo.org/record/176565/files/figure.png" pageId="3" pageNumber="58">Fig. 3</figureCitation>
A–D): Large (aperture up to 
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long), relatively high (in mature specimens height and aperture length ratio is 0.34–0.43). Apex slightly displaced to the left (distance from apex to the left aperture edge is 0.76–0.99 of the distance to the right one) and distant from the anterior edge of aperture by 0.58–0.68 of its length (in projection to longitudinal axis of aperture). Distance from apex to the frontal aperture edge along a chord of the anterior (maximal) slope is 0.69–0.77. All four shell slopes slightly convex. Aperture broad oval, its width and length ratio is 0.79–0.84. Protoconch cap-shaped, relatively low (H/h = 8.5); its apical part bears two tubercles, the slopes underneath are slightly convex (
<figureCitation id="A3CFCBE9002FB309993BF8CEFB6FD949" box="[1150,1220,1839,1865]" captionStart="FIGURE 3" captionStartId="5.[151,255,1488,1512]" captionTargetBox="[264,1322,194,1463]" captionTargetId="figure@5.[264,1323,194,1464]" captionTargetPageId="5" captionText="FIGURE 3. Shells of Gerstfeldtiancylus ushunensis, sp. nov. (SEM). A – D. Teleoconch (ZIN RAS 1 / 521 – 2006): A. Top view. B. Right side view. C. Left side view. D. Rear view. E – G. Protoconch (LIN SB RAS 905): E. Top view. F. Right side view. G. Left side view. H. Initial protoconch plate. I – K. Protoconch sculpture (LIN SB RAS 904). Scale bars: A – D = 2.0 mm; E = 1.0 mm; F – G = 0.25 mm; H, J = 0.1 mm; I = 0.2 mm; K = 0.02 mm." httpUri="https://zenodo.org/record/176565/files/figure.png" pageId="3" pageNumber="58">Fig. 3</figureCitation>
E–G). Protoconch surface has a distinct reticular microsculpture formed by transecting radial costae and concentric cords and distinct rectangular grooves 
<quantity id="FC0C7A89002FB3099CADF89EFD1AD999" box="[488,689,1919,1945]" metricMagnitude="-5" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.7" metricValueMin="2.0" pageId="3" pageNumber="58" unit="mm" value="0.0235" valueMax="0.027" valueMin="0.02">0.020–0.027 mm</quantity>
long and 
<quantity id="FC0C7A89002FB3099E6FF89EFC58D999" box="[810,1011,1919,1945]" metricMagnitude="-5" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="3" pageNumber="58" unit="mm" value="0.017" valueMax="0.02" valueMin="0.014">0.014–0.020 mm</quantity>
wide (
<figureCitation id="A3CFCBE9002FB3099903F89EFB26D999" box="[1094,1165,1919,1945]" captionStart="FIGURE 3" captionStartId="5.[151,255,1488,1512]" captionTargetBox="[264,1322,194,1463]" captionTargetId="figure@5.[264,1323,194,1464]" captionTargetPageId="5" captionText="FIGURE 3. Shells of Gerstfeldtiancylus ushunensis, sp. nov. (SEM). A – D. Teleoconch (ZIN RAS 1 / 521 – 2006): A. Top view. B. Right side view. C. Left side view. D. Rear view. E – G. Protoconch (LIN SB RAS 905): E. Top view. F. Right side view. G. Left side view. H. Initial protoconch plate. I – K. Protoconch sculpture (LIN SB RAS 904). Scale bars: A – D = 2.0 mm; E = 1.0 mm; F – G = 0.25 mm; H, J = 0.1 mm; I = 0.2 mm; K = 0.02 mm." httpUri="https://zenodo.org/record/176565/files/figure.png" pageId="3" pageNumber="58">Fig. 3</figureCitation>
K). Reticular sculpture gradually turns in a concentric one in the direction from the apical part to the protoconch edge (
<figureCitation id="A3CFCBE9002FB30999B4F846FA9ED9C1" box="[1265,1333,1959,1985]" captionStart="FIGURE 3" captionStartId="5.[151,255,1488,1512]" captionTargetBox="[264,1322,194,1463]" captionTargetId="figure@5.[264,1323,194,1464]" captionTargetPageId="5" captionText="FIGURE 3. Shells of Gerstfeldtiancylus ushunensis, sp. nov. (SEM). A – D. Teleoconch (ZIN RAS 1 / 521 – 2006): A. Top view. B. Right side view. C. Left side view. D. Rear view. E – G. Protoconch (LIN SB RAS 905): E. Top view. F. Right side view. G. Left side view. H. Initial protoconch plate. I – K. Protoconch sculpture (LIN SB RAS 904). Scale bars: A – D = 2.0 mm; E = 1.0 mm; F – G = 0.25 mm; H, J = 0.1 mm; I = 0.2 mm; K = 0.02 mm." httpUri="https://zenodo.org/record/176565/files/figure.png" pageId="3" pageNumber="58">
Fig. 
<date id="4F4AF1AC002FB3099862F846FA9ED9C1" box="[1319,1333,1959,1985]" pageId="3" pageNumber="58">3</date>
</figureCitation>
I–J). Initial plate lacks any sculpture (
<figureCitation id="A3CFCBE9002FB3099CB1F82EFD92D9E9" box="[500,569,1999,2025]" captionStart="FIGURE 3" captionStartId="5.[151,255,1488,1512]" captionTargetBox="[264,1322,194,1463]" captionTargetId="figure@5.[264,1323,194,1464]" captionTargetPageId="5" captionText="FIGURE 3. Shells of Gerstfeldtiancylus ushunensis, sp. nov. (SEM). A – D. Teleoconch (ZIN RAS 1 / 521 – 2006): A. Top view. B. Right side view. C. Left side view. D. Rear view. E – G. Protoconch (LIN SB RAS 905): E. Top view. F. Right side view. G. Left side view. H. Initial protoconch plate. I – K. Protoconch sculpture (LIN SB RAS 904). Scale bars: A – D = 2.0 mm; E = 1.0 mm; F – G = 0.25 mm; H, J = 0.1 mm; I = 0.2 mm; K = 0.02 mm." httpUri="https://zenodo.org/record/176565/files/figure.png" pageId="3" pageNumber="58">Fig. 3</figureCitation>
H), its diameter being about 
<quantity id="FC0C7A89002FB3099ECDF82EFC5AD9E9" box="[904,1009,1999,2025]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="58" unit="mm" value="0.12">0.12 mm</quantity>
and forms an angle of 13–14° to the protoconch aperture plane. Protoconch forms an angle of 13–14° to the plane of teleoconch aperture. Dimensions of 
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protoconch: length (l) = 
<quantity id="FC0C7A890028B30E9FC3FF21FD5ADEDA" box="[646,753,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="2.44" pageId="4" pageNumber="59" unit="mm" value="2.44">2.44 mm</quantity>
, width (w) = 
<quantity id="FC0C7A890028B30E9ECAFF21FC50DEDA" box="[911,1019,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="59" unit="mm" value="1.7">1.70 mm</quantity>
, height (h) = 
<quantity id="FC0C7A890028B30E99DDFF21FAAEDEDA" box="[1176,1285,192,218]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="59" unit="mm" value="0.35">0.35 mm</quantity>
. Dimensions of 
<typeStatus id="E44F69CE0028B30E9DF3FF09FEB6DF02" box="[182,285,232,258]" pageId="4" pageNumber="59" type="holotype">holotype</typeStatus>
and 
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teleoconchs are given in 
<tableCitation id="7676E2D70028B30E9FA0FF09FC96DF02" box="[741,829,232,258]" captionStart="TABLE 1" captionStartId="4.[151,239,311,335]" captionTargetBox="[159,1334,402,1151]" captionTargetPageId="4" captionText="TABLE 1. Shell dimensions of Gerstfeldtiancylus ushunensis, sp. nov. holotype (ZIN RAS 1 / 521 – 2006) and paratypes (in mm). For abbreviations of dimensions see Fig. 2." httpUri="http://table.plazi.org/id/6F8B87E40028B30E9DD2FED6FD6CDF71" pageId="4" pageNumber="59" tableUuid="6F8B87E40028B30E9DD2FED6FD6CDF71">Table 1</tableCitation>
.
</paragraph>
<caption id="6F8B87E40028B30E9DD2FED6FD6CDF71" ID-Table-UUID="6F8B87E40028B30E9DD2FED6FD6CDF71" httpUri="http://table.plazi.org/id/6F8B87E40028B30E9DD2FED6FD6CDF71" pageId="4" pageNumber="59" targetBox="[159,1334,402,1151]" targetIsTable="true" targetPageId="4">
<paragraph id="3B4BD76C0028B30E9DD2FED6FD6CDF71" blockId="4.[151,1436,311,369]" pageId="4" pageNumber="59">
<emphasis id="09800B7E0028B30E9DD2FED6FEA2DF4F" bold="true" box="[151,265,311,335]" pageId="4" pageNumber="59">TABLE 1.</emphasis>
Shell dimensions of 
<taxonomicName id="FCF4ACEF0028B30E9CAAFED9FC84DF4F" box="[495,815,312,335]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="4" pageNumber="59" phylum="Mollusca" rank="species" species="ushunensis" status="sp. nov.">
<emphasis id="09800B7E0028B30E9CAAFED9FC84DF4F" box="[495,815,312,335]" italics="true" pageId="4" pageNumber="59">Gerstfeldtiancylus ushunensis</emphasis>
</taxonomicName>
, 
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<emphasis id="09800B7E0028B30E9E78FED6FC3FDF4F" bold="true" box="[829,916,311,335]" pageId="4" pageNumber="59">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype (ZIN RAS 1/521–2006) and paratypes (in mm). For abbreviations of dimensions see Fig. 2.
</paragraph>
</caption>
<paragraph id="3B4BD76C0028B30E9DDAFE73FA9DDA7F" pageId="4" pageNumber="59">
<table id="49F425CC00284CF59DDAFE73FA9DDA7F" box="[159,1334,402,1151]" gridcols="7" gridrows="17" pageId="4" pageNumber="59">
<tr id="85C4D52E00284CF59DDAFE73FA9DDFA9" box="[159,1334,402,425]" gridrow="0" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFE73FE27DFA9" box="[159,396,402,425]" gridcol="0" gridrow="0" pageId="4" pageNumber="59">Specimen number</th>
<th id="C615BC5200284CF59CBCFE73FD83DFA9" box="[505,552,402,425]" gridcol="1" gridrow="0" pageId="4" pageNumber="59">L</th>
<th id="C615BC5200284CF59FD0FE73FD6FDFA9" box="[661,708,402,425]" gridcol="2" gridrow="0" pageId="4" pageNumber="59">La</th>
<th id="C615BC5200284CF59E77FE73FCCADFA9" box="[818,865,402,425]" gridcol="3" gridrow="0" pageId="4" pageNumber="59">W</th>
<th id="C615BC5200284CF59E8BFE73FC56DFA9" box="[974,1021,402,425]" gridcol="4" gridrow="0" pageId="4" pageNumber="59">wL</th>
<th id="C615BC5200284CF5992FFE73FB32DFA9" box="[1130,1177,402,425]" gridcol="5" gridrow="0" pageId="4" pageNumber="59">a</th>
<th id="C615BC5200284CF59842FE73FA9DDFA9" box="[1287,1334,402,425]" gridcol="6" gridrow="0" pageId="4" pageNumber="59">H</th>
</tr>
<tr id="85C4D52E00284CF59DDAFE21FA9DDFD7" box="[159,1334,448,471]" gridrow="1" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFE21FE27DFD7" box="[159,396,448,471]" gridcol="0" gridrow="1" pageId="4" pageNumber="59">ZIN RAS 1/521–2006</th>
<td id="C615BC5200284CF59CBCFE21FD83DFD7" box="[505,552,448,471]" gridcol="1" gridrow="1" pageId="4" pageNumber="59">7.20</td>
<td id="C615BC5200284CF59FD0FE21FD6FDFD7" box="[661,708,448,471]" gridcol="2" gridrow="1" pageId="4" pageNumber="59">4.20</td>
<td id="C615BC5200284CF59E77FE21FCCADFD7" box="[818,865,448,471]" gridcol="3" gridrow="1" pageId="4" pageNumber="59">5.80</td>
<td id="C615BC5200284CF59E8BFE21FC56DFD7" box="[974,1021,448,471]" gridcol="4" gridrow="1" pageId="4" pageNumber="59">2.50</td>
<td id="C615BC5200284CF5992FFE21FB32DFD7" box="[1130,1177,448,471]" gridcol="5" gridrow="1" pageId="4" pageNumber="59">5.50</td>
<td id="C615BC5200284CF59842FE21FA9DDFD7" box="[1287,1334,448,471]" gridcol="6" gridrow="1" pageId="4" pageNumber="59">3.00</td>
</tr>
<tr id="85C4D52E00284CF59DDAFE0CFA9DDC04" box="[159,1334,493,516]" gridrow="2" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFE0CFE27DC04" box="[159,396,493,516]" gridcol="0" gridrow="2" pageId="4" pageNumber="59">ZIN RAS 2/521–2006</th>
<td id="C615BC5200284CF59CBCFE0CFD83DC04" box="[505,552,493,516]" gridcol="1" gridrow="2" pageId="4" pageNumber="59">7.40</td>
<td id="C615BC5200284CF59FD0FE0CFD6FDC04" box="[661,708,493,516]" gridcol="2" gridrow="2" pageId="4" pageNumber="59">5.00</td>
<td id="C615BC5200284CF59E77FE0CFCCADC04" box="[818,865,493,516]" gridcol="3" gridrow="2" pageId="4" pageNumber="59">5.90</td>
<td id="C615BC5200284CF59E8BFE0CFC56DC04" box="[974,1021,493,516]" gridcol="4" gridrow="2" pageId="4" pageNumber="59">2.90</td>
<td id="C615BC5200284CF5992FFE0CFB32DC04" box="[1130,1177,493,516]" gridcol="5" gridrow="2" pageId="4" pageNumber="59">5.70</td>
<td id="C615BC5200284CF59842FE0CFA9DDC04" box="[1287,1334,493,516]" gridcol="6" gridrow="2" pageId="4" pageNumber="59">3.15</td>
</tr>
<tr id="85C4D52E00284CF59DDAFDFBFA9DDC31" box="[159,1334,538,561]" gridrow="3" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFDFBFE27DC31" box="[159,396,538,561]" gridcol="0" gridrow="3" pageId="4" pageNumber="59">ZIN RAS 2/521–2006</th>
<td id="C615BC5200284CF59CBCFDFBFD83DC31" box="[505,552,538,561]" gridcol="1" gridrow="3" pageId="4" pageNumber="59">6.95</td>
<td id="C615BC5200284CF59FD0FDFBFD6FDC31" box="[661,708,538,561]" gridcol="2" gridrow="3" pageId="4" pageNumber="59">4.55</td>
<td id="C615BC5200284CF59E77FDFBFCCADC31" box="[818,865,538,561]" gridcol="3" gridrow="3" pageId="4" pageNumber="59">5.55</td>
<td id="C615BC5200284CF59E8BFDFBFC56DC31" box="[974,1021,538,561]" gridcol="4" gridrow="3" pageId="4" pageNumber="59">2.50</td>
<td id="C615BC5200284CF5992FFDFBFB32DC31" box="[1130,1177,538,561]" gridcol="5" gridrow="3" pageId="4" pageNumber="59">4.95</td>
<td id="C615BC5200284CF59842FDFBFA9DDC31" box="[1287,1334,538,561]" gridcol="6" gridrow="3" pageId="4" pageNumber="59">2.85</td>
</tr>
<tr id="85C4D52E00284CF59DDAFDA9FA9DDC5F" box="[159,1334,584,607]" gridrow="4" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFDA9FE27DC5F" box="[159,396,584,607]" gridcol="0" gridrow="4" pageId="4" pageNumber="59">ZIN RAS 2/521–2006</th>
<td id="C615BC5200284CF59CBCFDA9FD83DC5F" box="[505,552,584,607]" gridcol="1" gridrow="4" pageId="4" pageNumber="59">7.20</td>
<td id="C615BC5200284CF59FD0FDA9FD6FDC5F" box="[661,708,584,607]" gridcol="2" gridrow="4" pageId="4" pageNumber="59">4.60</td>
<td id="C615BC5200284CF59E77FDA9FCCADC5F" box="[818,865,584,607]" gridcol="3" gridrow="4" pageId="4" pageNumber="59">5.70</td>
<td id="C615BC5200284CF59E8BFDA9FC56DC5F" box="[974,1021,584,607]" gridcol="4" gridrow="4" pageId="4" pageNumber="59">2.70</td>
<td id="C615BC5200284CF5992FFDA9FB32DC5F" box="[1130,1177,584,607]" gridcol="5" gridrow="4" pageId="4" pageNumber="59">5.30</td>
<td id="C615BC5200284CF59842FDA9FA9DDC5F" box="[1287,1334,584,607]" gridcol="6" gridrow="4" pageId="4" pageNumber="59">2.65</td>
</tr>
<tr id="85C4D52E00284CF59DDAFD94FA9DDC8C" box="[159,1334,629,652]" gridrow="5" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFD94FE27DC8C" box="[159,396,629,652]" gridcol="0" gridrow="5" pageId="4" pageNumber="59">ZIN RAS 2/521–2006</th>
<td id="C615BC5200284CF59CBCFD94FD83DC8C" box="[505,552,629,652]" gridcol="1" gridrow="5" pageId="4" pageNumber="59">6.90</td>
<td id="C615BC5200284CF59FD0FD94FD6FDC8C" box="[661,708,629,652]" gridcol="2" gridrow="5" pageId="4" pageNumber="59">4.35</td>
<td id="C615BC5200284CF59E77FD94FCCADC8C" box="[818,865,629,652]" gridcol="3" gridrow="5" pageId="4" pageNumber="59">5.60</td>
<td id="C615BC5200284CF59E8BFD94FC56DC8C" box="[974,1021,629,652]" gridcol="4" gridrow="5" pageId="4" pageNumber="59">2.70</td>
<td id="C615BC5200284CF5992FFD94FB32DC8C" box="[1130,1177,629,652]" gridcol="5" gridrow="5" pageId="4" pageNumber="59">4.75</td>
<td id="C615BC5200284CF59842FD94FA9DDC8C" box="[1287,1334,629,652]" gridcol="6" gridrow="5" pageId="4" pageNumber="59">2.55</td>
</tr>
<tr id="85C4D52E00284CF59DDAFD43FA9DDCB9" box="[159,1334,674,697]" gridrow="6" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFD43FE27DCB9" box="[159,396,674,697]" gridcol="0" gridrow="6" pageId="4" pageNumber="59">ZIN RAS 2/521–2006</th>
<td id="C615BC5200284CF59CBCFD43FD83DCB9" box="[505,552,674,697]" gridcol="1" gridrow="6" pageId="4" pageNumber="59">6.40</td>
<td id="C615BC5200284CF59FD0FD43FD6FDCB9" box="[661,708,674,697]" gridcol="2" gridrow="6" pageId="4" pageNumber="59">3.85</td>
<td id="C615BC5200284CF59E77FD43FCCADCB9" box="[818,865,674,697]" gridcol="3" gridrow="6" pageId="4" pageNumber="59">5.10</td>
<td id="C615BC5200284CF59E8BFD43FC56DCB9" box="[974,1021,674,697]" gridcol="4" gridrow="6" pageId="4" pageNumber="59">2.50</td>
<td id="C615BC5200284CF5992FFD43FB32DCB9" box="[1130,1177,674,697]" gridcol="5" gridrow="6" pageId="4" pageNumber="59">4.40</td>
<td id="C615BC5200284CF59842FD43FA9DDCB9" box="[1287,1334,674,697]" gridcol="6" gridrow="6" pageId="4" pageNumber="59">2.20</td>
</tr>
<tr id="85C4D52E00284CF59DDAFD31FA9DDCE7" box="[159,1334,720,743]" gridrow="7" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFD31FE27DCE7" box="[159,396,720,743]" gridcol="0" gridrow="7" pageId="4" pageNumber="59">ZIN RAS 2/521–2006</th>
<td id="C615BC5200284CF59CBCFD31FD83DCE7" box="[505,552,720,743]" gridcol="1" gridrow="7" pageId="4" pageNumber="59">6.05</td>
<td id="C615BC5200284CF59FD0FD31FD6FDCE7" box="[661,708,720,743]" gridcol="2" gridrow="7" pageId="4" pageNumber="59">3.60</td>
<td id="C615BC5200284CF59E77FD31FCCADCE7" box="[818,865,720,743]" gridcol="3" gridrow="7" pageId="4" pageNumber="59">5.10</td>
<td id="C615BC5200284CF59E8BFD31FC56DCE7" box="[974,1021,720,743]" gridcol="4" gridrow="7" pageId="4" pageNumber="59">2.50</td>
<td id="C615BC5200284CF5992FFD31FB32DCE7" box="[1130,1177,720,743]" gridcol="5" gridrow="7" pageId="4" pageNumber="59">4.40</td>
<td id="C615BC5200284CF59842FD31FA9DDCE7" box="[1287,1334,720,743]" gridcol="6" gridrow="7" pageId="4" pageNumber="59">2.10</td>
</tr>
<tr id="85C4D52E00284CF59DDAFD1CFA9DDD14" box="[159,1334,765,788]" gridrow="8" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFD1CFE27DD14" box="[159,396,765,788]" gridcol="0" gridrow="8" pageId="4" pageNumber="59">ZIN RAS 3/521–2006</th>
<td id="C615BC5200284CF59CBCFD1CFD83DD14" box="[505,552,765,788]" gridcol="1" gridrow="8" pageId="4" pageNumber="59">4.00</td>
<td id="C615BC5200284CF59FD0FD1CFD6FDD14" box="[661,708,765,788]" gridcol="2" gridrow="8" pageId="4" pageNumber="59">2.45</td>
<td id="C615BC5200284CF59E77FD1CFCCADD14" box="[818,865,765,788]" gridcol="3" gridrow="8" pageId="4" pageNumber="59">3.30</td>
<td id="C615BC5200284CF59E8BFD1CFC56DD14" box="[974,1021,765,788]" gridcol="4" gridrow="8" pageId="4" pageNumber="59">1.65</td>
<td id="C615BC5200284CF5992FFD1CFB32DD14" box="[1130,1177,765,788]" gridcol="5" gridrow="8" pageId="4" pageNumber="59">2.90</td>
<td id="C615BC5200284CF59842FD1CFA9DDD14" box="[1287,1334,765,788]" gridcol="6" gridrow="8" pageId="4" pageNumber="59">1.20</td>
</tr>
<tr id="85C4D52E00284CF59DDAFCCBFA9DDD41" box="[159,1334,810,833]" gridrow="9" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFCCBFE27DD41" box="[159,396,810,833]" gridcol="0" gridrow="9" pageId="4" pageNumber="59">ZIN RAS 2/521–2006</th>
<td id="C615BC5200284CF59CBCFCCBFD83DD41" box="[505,552,810,833]" gridcol="1" gridrow="9" pageId="4" pageNumber="59">4.70</td>
<td id="C615BC5200284CF59FD0FCCBFD6FDD41" box="[661,708,810,833]" gridcol="2" gridrow="9" pageId="4" pageNumber="59">3.05</td>
<td id="C615BC5200284CF59E77FCCBFCCADD41" box="[818,865,810,833]" gridcol="3" gridrow="9" pageId="4" pageNumber="59">4.10</td>
<td id="C615BC5200284CF59E8BFCCBFC56DD41" box="[974,1021,810,833]" gridcol="4" gridrow="9" pageId="4" pageNumber="59">1.90</td>
<td id="C615BC5200284CF5992FFCCBFB32DD41" box="[1130,1177,810,833]" gridcol="5" gridrow="9" pageId="4" pageNumber="59">3.50</td>
<td id="C615BC5200284CF59842FCCBFA9DDD41" box="[1287,1334,810,833]" gridcol="6" gridrow="9" pageId="4" pageNumber="59">1.80</td>
</tr>
<tr id="85C4D52E00284CF59DDAFCB9FA9DDD6F" box="[159,1334,856,879]" gridrow="10" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFCB9FE27DD6F" box="[159,396,856,879]" gridcol="0" gridrow="10" pageId="4" pageNumber="59">ZIN RAS 2/521–2006</th>
<td id="C615BC5200284CF59CBCFCB9FD83DD6F" box="[505,552,856,879]" gridcol="1" gridrow="10" pageId="4" pageNumber="59">4.50</td>
<td id="C615BC5200284CF59FD0FCB9FD6FDD6F" box="[661,708,856,879]" gridcol="2" gridrow="10" pageId="4" pageNumber="59">2.85</td>
<td id="C615BC5200284CF59E77FCB9FCCADD6F" box="[818,865,856,879]" gridcol="3" gridrow="10" pageId="4" pageNumber="59">3.90</td>
<td id="C615BC5200284CF59E8BFCB9FC56DD6F" box="[974,1021,856,879]" gridcol="4" gridrow="10" pageId="4" pageNumber="59">1.70</td>
<td id="C615BC5200284CF5992FFCB9FB32DD6F" box="[1130,1177,856,879]" gridcol="5" gridrow="10" pageId="4" pageNumber="59">3.40</td>
<td id="C615BC5200284CF59842FCB9FA9DDD6F" box="[1287,1334,856,879]" gridcol="6" gridrow="10" pageId="4" pageNumber="59">1.85</td>
</tr>
<tr id="85C4D52E00284CF59DDAFC64FA9DDD9C" box="[159,1334,901,924]" gridrow="11" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFC64FE27DD9C" box="[159,396,901,924]" gridcol="0" gridrow="11" pageId="4" pageNumber="59">ZIN RAS 2/521–2006</th>
<td id="C615BC5200284CF59CBCFC64FD83DD9C" box="[505,552,901,924]" gridcol="1" gridrow="11" pageId="4" pageNumber="59">4.50</td>
<td id="C615BC5200284CF59FD0FC64FD6FDD9C" box="[661,708,901,924]" gridcol="2" gridrow="11" pageId="4" pageNumber="59">2.90</td>
<td id="C615BC5200284CF59E77FC64FCCADD9C" box="[818,865,901,924]" gridcol="3" gridrow="11" pageId="4" pageNumber="59">3.80</td>
<td id="C615BC5200284CF59E8BFC64FC56DD9C" box="[974,1021,901,924]" gridcol="4" gridrow="11" pageId="4" pageNumber="59">1.80</td>
<td id="C615BC5200284CF5992FFC64FB32DD9C" box="[1130,1177,901,924]" gridcol="5" gridrow="11" pageId="4" pageNumber="59">3.30</td>
<td id="C615BC5200284CF59842FC64FA9DDD9C" box="[1287,1334,901,924]" gridcol="6" gridrow="11" pageId="4" pageNumber="59">1.50</td>
</tr>
<tr id="85C4D52E00284CF59DDAFC53FA9DDDC9" box="[159,1334,946,969]" gridrow="12" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFC53FE27DDC9" box="[159,396,946,969]" gridcol="0" gridrow="12" pageId="4" pageNumber="59">LIN SB RAS 904</th>
<td id="C615BC5200284CF59CBCFC53FD83DDC9" box="[505,552,946,969]" gridcol="1" gridrow="12" pageId="4" pageNumber="59">2.70</td>
<td id="C615BC5200284CF59FD0FC53FD6FDDC9" box="[661,708,946,969]" gridcol="2" gridrow="12" pageId="4" pageNumber="59">1.75</td>
<td id="C615BC5200284CF59E77FC53FCCADDC9" box="[818,865,946,969]" gridcol="3" gridrow="12" pageId="4" pageNumber="59">2.15</td>
<td id="C615BC5200284CF59E8BFC53FC56DDC9" box="[974,1021,946,969]" gridcol="4" gridrow="12" pageId="4" pageNumber="59">0.90</td>
<td id="C615BC5200284CF5992FFC53FB32DDC9" box="[1130,1177,946,969]" gridcol="5" gridrow="12" pageId="4" pageNumber="59">1.85</td>
<td id="C615BC5200284CF59842FC53FA9DDDC9" box="[1287,1334,946,969]" gridcol="6" gridrow="12" pageId="4" pageNumber="59">0.75</td>
</tr>
<tr id="85C4D52E00284CF59DDAFC01FA9DDDF7" box="[159,1334,992,1015]" gridrow="13" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFC01FE27DDF7" box="[159,396,992,1015]" gridcol="0" gridrow="13" pageId="4" pageNumber="59">LIN SB RAS 904</th>
<td id="C615BC5200284CF59CBCFC01FD83DDF7" box="[505,552,992,1015]" gridcol="1" gridrow="13" pageId="4" pageNumber="59">2.40</td>
<td id="C615BC5200284CF59FD0FC01FD6FDDF7" box="[661,708,992,1015]" gridcol="2" gridrow="13" pageId="4" pageNumber="59">1.65</td>
<td id="C615BC5200284CF59E77FC01FCCADDF7" box="[818,865,992,1015]" gridcol="3" gridrow="13" pageId="4" pageNumber="59">2.05</td>
<td id="C615BC5200284CF59E8BFC01FC56DDF7" box="[974,1021,992,1015]" gridcol="4" gridrow="13" pageId="4" pageNumber="59">0.90</td>
<td id="C615BC5200284CF5992FFC01FB32DDF7" box="[1130,1177,992,1015]" gridcol="5" gridrow="13" pageId="4" pageNumber="59">1.75</td>
<td id="C615BC5200284CF59842FC01FA9DDDF7" box="[1287,1334,992,1015]" gridcol="6" gridrow="13" pageId="4" pageNumber="59">0.80</td>
</tr>
<tr id="85C4D52E00284CF59DDAFBECFA9DDA24" box="[159,1334,1037,1060]" gridrow="14" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFBECFE27DA24" box="[159,396,1037,1060]" gridcol="0" gridrow="14" pageId="4" pageNumber="59">LIN SB RAS 905</th>
<td id="C615BC5200284CF59CBCFBECFD83DA24" box="[505,552,1037,1060]" gridcol="1" gridrow="14" pageId="4" pageNumber="59">2.95</td>
<td id="C615BC5200284CF59FD0FBECFD6FDA24" box="[661,708,1037,1060]" gridcol="2" gridrow="14" pageId="4" pageNumber="59">1.85</td>
<td id="C615BC5200284CF59E77FBECFCCADA24" box="[818,865,1037,1060]" gridcol="3" gridrow="14" pageId="4" pageNumber="59">2.35</td>
<td id="C615BC5200284CF59E8BFBECFC56DA24" box="[974,1021,1037,1060]" gridcol="4" gridrow="14" pageId="4" pageNumber="59">1.10</td>
<td id="C615BC5200284CF5992FFBECFB32DA24" box="[1130,1177,1037,1060]" gridcol="5" gridrow="14" pageId="4" pageNumber="59">1.90</td>
<td id="C615BC5200284CF59842FBECFA9DDA24" box="[1287,1334,1037,1060]" gridcol="6" gridrow="14" pageId="4" pageNumber="59">0.75</td>
</tr>
<tr id="85C4D52E00284CF59DDAFBDBFA9DDA51" box="[159,1334,1082,1105]" gridrow="15" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFBDBFE27DA51" box="[159,396,1082,1105]" gridcol="0" gridrow="15" pageId="4" pageNumber="59">LIN SB RAS 905</th>
<td id="C615BC5200284CF59CBCFBDBFD83DA51" box="[505,552,1082,1105]" gridcol="1" gridrow="15" pageId="4" pageNumber="59">3.25</td>
<td id="C615BC5200284CF59FD0FBDBFD6FDA51" box="[661,708,1082,1105]" gridcol="2" gridrow="15" pageId="4" pageNumber="59">2.00</td>
<td id="C615BC5200284CF59E77FBDBFCCADA51" box="[818,865,1082,1105]" gridcol="3" gridrow="15" pageId="4" pageNumber="59">2.50</td>
<td id="C615BC5200284CF59E8BFBDBFC56DA51" box="[974,1021,1082,1105]" gridcol="4" gridrow="15" pageId="4" pageNumber="59">1.00</td>
<td id="C615BC5200284CF5992FFBDBFB32DA51" box="[1130,1177,1082,1105]" gridcol="5" gridrow="15" pageId="4" pageNumber="59">2.05</td>
<td id="C615BC5200284CF59842FBDBFA9DDA51" box="[1287,1334,1082,1105]" gridcol="6" gridrow="15" pageId="4" pageNumber="59">0.80</td>
</tr>
<tr id="85C4D52E00284CF59DDAFB89FA9DDA7F" box="[159,1334,1128,1151]" gridrow="16" pageId="4" pageNumber="59">
<th id="C615BC5200284CF59DDAFB89FE27DA7F" box="[159,396,1128,1151]" gridcol="0" gridrow="16" pageId="4" pageNumber="59">LIN SB RAS 905</th>
<td id="C615BC5200284CF59CBCFB89FD83DA7F" box="[505,552,1128,1151]" gridcol="1" gridrow="16" pageId="4" pageNumber="59">2.85</td>
<td id="C615BC5200284CF59FD0FB89FD6FDA7F" box="[661,708,1128,1151]" gridcol="2" gridrow="16" pageId="4" pageNumber="59">1.65</td>
<td id="C615BC5200284CF59E77FB89FCCADA7F" box="[818,865,1128,1151]" gridcol="3" gridrow="16" pageId="4" pageNumber="59">2.30</td>
<td id="C615BC5200284CF59E8BFB89FC56DA7F" box="[974,1021,1128,1151]" gridcol="4" gridrow="16" pageId="4" pageNumber="59">0.95</td>
<td id="C615BC5200284CF5992FFB89FB32DA7F" box="[1130,1177,1128,1151]" gridcol="5" gridrow="16" pageId="4" pageNumber="59">2.00</td>
<td id="C615BC5200284CF59842FB89FA9DDA7F" box="[1287,1334,1128,1151]" gridcol="6" gridrow="16" pageId="4" pageNumber="59">0.60</td>
</tr>
</table>
</paragraph>
<paragraph id="3B4BD76C0028B30E9D82FB33FD5FDB8C" blockId="4.[151,1437,1234,2020]" pageId="4" pageNumber="59">
Radula (
<figureCitation id="A3CFCBE90028B30E9C6EFB33FED9DAEC" box="[299,370,1234,1260]" captionStart="FIGURE 4" captionStartId="6.[151,255,1716,1740]" captionTargetBox="[264,1322,1146,1692]" captionTargetId="figure@6.[264,1323,1146,1692]" captionTargetPageId="6" captionText="FIGURE 4. Radula of Gerstfeldtiancylus ushunensis, sp. nov. (SEM). A – C. ZIN RAS 2 / 521 – 2006: A. Central tooth. B. Three transverse rows of teeth. C. Lateral teeth. Scale bars: A = 0.01 mm; B – C = 0.1 mm." httpUri="https://zenodo.org/record/176566/files/figure.png" pageId="4" pageNumber="59">Fig. 4</figureCitation>
) of mature specimens: 74–90 transverse teeth rows. Each row has 57–65 teeth: 16–23 lateral and 7–15 marginal teeth on each side of the central tooth. Central tooth has a narrow base and a short cutting edge with one median cusp and 3–4 wrinkles on both sides. Lateral teeth have a narrow long cutting edge with two terminal cusps. Inner marginal teeth have a broad base and short cutting edge (cusps not developed). The remaining marginal teeth have no cutting edge.
</paragraph>
<paragraph id="3B4BD76C0028B30E9D82FA7BFD03D8CC" blockId="4.[151,1437,1234,2020]" pageId="4" pageNumber="59">
Alimentary system (
<figureCitation id="A3CFCBE90028B30E9CF1FA7BFDBADBB4" box="[436,529,1434,1460]" captionStart="FIGURE 5. A – B" captionStartId="7.[151,255,1718,1742]" captionTargetBox="[274,1303,187,1701]" captionTargetId="figure@7.[264,1323,170,1720]" captionTargetPageId="7" captionText="FIGURE 5. A – B. Anatomy of Gerstfeldtiancylus ushunensis, sp. nov.. А. General topography of main organs of the alimentary system, dorsal view. Digestive gland removed. B. Dorsal view of specimen without shell. C – D. Odontophore of G. ushunensis, sp. nov.. E – L. Jaws of different Gerstfeldtiancylus species. E – H. G. ushunensis, sp. nov. (A, C – E. ZIN RAS 1 / 521 – 2006. B, F – H. ZIN RAS 2 / 521 – 2006). I. G. renardii (Dybowski, 1884). J. G. roepstorfi Shirokaya et al., 2003. K. G. benedictiae Starobogatov, 1989 (whole mounts in Faure-Berlezet liquid). L. G. kotyensis Starobogatov, 1989 (frontal histological section). I – L. From Shirokaya &amp; Röpstorf (2004). aal, left anterior adductor; aar, right anterior adductor; ap, anal pore; bm, buccal mass; cae, caecum; dgp, digestive gland pores; int 1, first intestinal loop; int 2, second intestinal loop; mb, mantle border; oa, original or posterior adductor; oe, oesophagus; rs, radular sac; sg, salivary glands; st, stomach. Scale bars: A – D = 1 mm; E = 0.8 mm; F – H, J = 0.5 mm; I, K – L = 0.3 mm." httpUri="https://zenodo.org/record/176567/files/figure.png" pageId="4" pageNumber="59">Fig. 5A</figureCitation>
, C–H): Jaw horseshoe-shaped, consists of 76–101 lanceolate scales. Each lateral part of the jaw has up to 32–43 scales, dorsal — 12–15. Buccal bulb is rounded, its length making about 1/3 of the body length. Odontophore of living specimens is red. Salivary glands are short, slender (about 
<quantity id="FC0C7A890028B30E983DFA0BFF6ED82C" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="59" unit="mm" value="0.1">0.1 mm</quantity>
in diameter), proximal section expanded to 
<quantity id="FC0C7A890028B30E9F94F9F3FCC5D82C" box="[721,878,1554,1580]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="4" pageNumber="59" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.25–0.3 mm</quantity>
in diameter. Radular sac is short and its length does not exceed 1/6 of the body length. Stomach caecum is pyriform, branching dorso-laterally from the stomach and directed to the left. Dorsal digestive gland pore larger than ventral one. First intestinal loop is positioned on the right side of the body, the second one in the left posterior quarter of it. Digestive gland is large and occupies almost 2/3 of the visceral mass.
</paragraph>
<paragraph id="3B4BD76C0028B30E9D82F93BFD42D9E4" blockId="4.[151,1437,1234,2020]" pageId="4" pageNumber="59">
Shell adductor muscles (
<figureCitation id="A3CFCBE90028B30E9CA3F93BFDE9D8F4" box="[486,578,1754,1780]" captionStart="FIGURE 5. A – B" captionStartId="7.[151,255,1718,1742]" captionTargetBox="[274,1303,187,1701]" captionTargetId="figure@7.[264,1323,170,1720]" captionTargetPageId="7" captionText="FIGURE 5. A – B. Anatomy of Gerstfeldtiancylus ushunensis, sp. nov.. А. General topography of main organs of the alimentary system, dorsal view. Digestive gland removed. B. Dorsal view of specimen without shell. C – D. Odontophore of G. ushunensis, sp. nov.. E – L. Jaws of different Gerstfeldtiancylus species. E – H. G. ushunensis, sp. nov. (A, C – E. ZIN RAS 1 / 521 – 2006. B, F – H. ZIN RAS 2 / 521 – 2006). I. G. renardii (Dybowski, 1884). J. G. roepstorfi Shirokaya et al., 2003. K. G. benedictiae Starobogatov, 1989 (whole mounts in Faure-Berlezet liquid). L. G. kotyensis Starobogatov, 1989 (frontal histological section). I – L. From Shirokaya &amp; Röpstorf (2004). aal, left anterior adductor; aar, right anterior adductor; ap, anal pore; bm, buccal mass; cae, caecum; dgp, digestive gland pores; int 1, first intestinal loop; int 2, second intestinal loop; mb, mantle border; oa, original or posterior adductor; oe, oesophagus; rs, radular sac; sg, salivary glands; st, stomach. Scale bars: A – D = 1 mm; E = 0.8 mm; F – H, J = 0.5 mm; I, K – L = 0.3 mm." httpUri="https://zenodo.org/record/176567/files/figure.png" pageId="4" pageNumber="59">Fig. 5B</figureCitation>
): Left anterior adductor is slender with an elongated surface directed perpendicular to the longitudinal axis of the body or curved like an arch. Right anterior adductor consists of two parts. One is located closer to the anterior body end; it has an arch-shaped surface and is positioned on one line with the left anterior adductor. Other one with a broad oval surface is parallel to the longitudinal axis of the body. Left and right adductors are almost equal in height. Right half of posterior (original) adductor is substantially elongated and anteriorly directed. Its surface is arch-shaped. Surface area of posterior adductor is 2– 2.5 times as large as that of each anterior adductor.
</paragraph>
<caption id="6F8B87E40029B30F9DD2FA31FBB9D84F" httpUri="https://zenodo.org/record/176565/files/figure.png" pageId="5" pageNumber="60" targetBox="[264,1322,194,1463]" targetPageId="5">
<paragraph id="3B4BD76C0029B30F9DD2FA31FBB9D84F" blockId="5.[151,1437,1488,1615]" pageId="5" pageNumber="60">
<emphasis id="09800B7E0029B30F9DD2FA31FEB2DBE8" bold="true" box="[151,281,1488,1512]" pageId="5" pageNumber="60">FIGURE 3.</emphasis>
Shells of 
<taxonomicName id="FCF4ACEF0029B30F9CC1FA31FD6FDBE7" box="[388,708,1488,1511]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="5" pageNumber="60" phylum="Mollusca" rank="species" species="ushunensis" status="sp. nov.">
<emphasis id="09800B7E0029B30F9CC1FA31FD6FDBE7" box="[388,708,1488,1511]" italics="true" pageId="5" pageNumber="60">Gerstfeldtiancylus ushunensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="12B3B6050029B30F9F94FA31FC8CDBE8" box="[721,807,1488,1512]" pageId="5" pageNumber="60" rank="species">
<emphasis id="09800B7E0029B30F9F94FA31FC8CDBE8" bold="true" box="[721,807,1488,1512]" pageId="5" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
(SEM). A–D. Teleoconch (ZIN RAS 1/521–2006): A. Top view. B. Right side view. C. Left side view. D. Rear view. E–G. Protoconch (LIN SB RAS 905): E. Top view. F. Right side view. G. Left side view. H. Initial protoconch plate. I–K. Protoconch sculpture (LIN SB RAS 904). Scale bars: A–D = 2.0 mm; E = 1.0 mm; F–G = 0.25 mm; H, J = 0.1 mm; I = 0.2 mm; K = 0.02 mm.
</paragraph>
</caption>
<paragraph id="3B4BD76C0029B30F9D82F962FC60D98D" blockId="5.[151,1437,1667,2013]" pageId="5" pageNumber="60">
Male copulatory organ (
<figureCitation id="A3CFCBE90029B30F9CABF962FD93D89D" box="[494,568,1667,1693]" captionStart="FIGURE 6" captionStartId="8.[151,255,1659,1683]" captionTargetBox="[290,1306,186,1659]" captionTargetId="figure@8.[275,1316,172,1659]" captionTargetPageId="8" captionText="FIGURE 6. Longitudinally sectioned male copulatory organ of Gerstfeldtiancylus species (A – D, G. diagrammic). A. G. benedictiae Starobogatov, 1989. B, E. G. roepstorfi Shirokaya et al., 2003. C. G. kotyensis Starobogatov, 1989. D. G. renardii (Dybowski, 1884). A – E. From Kruglov &amp; Starobogatov (1991 a), Shirokaya (2005). F – G. G. ushunensis, sp. nov. (ZIN RAS 2 / 521 – 2006). fl, flagellum; mp, male genital pore; p, penis; ps, penis sheath; psr, penis sheath retractor; pr, preputium; prc, preputium chamber; prp, preputium pilasters; sar, sarcobellum; vd, vas deferens; ve, velum. Scale bars: A = 0.5 mm; B – G = 1 mm." httpUri="https://zenodo.org/record/176568/files/figure.png" pageId="5" pageNumber="60">Fig. 6</figureCitation>
F–G): Flagellum nearly as long as the total length of penis sheath and preputium. In cross-section flagellum is even all the way through and the distal end is slightly inflated. Penis sheath is long, cylindrical, evenly wide. Penis clavate, having a terminally located deferent duct opening. Penis length is 3/5 of the sheath length. Proximal part of penis is twice as wide as the distal one. Preputium is cylindrical and it forms a globose chamber with muscular walls in the proximal part. The ratio of the chamber width to the preputium width is 1.2–1.3. Sarcobellum and velum are well developed. Preputium is as long as the penis sheath. Pilasters are pronounced along the whole preputium.
</paragraph>
<paragraph id="3B4BD76C0029B30C9D82F87AFEB4DF2A" blockId="5.[151,1437,1667,2013]" lastBlockId="6.[151,1436,152,298]" lastPageId="6" lastPageNumber="61" pageId="5" pageNumber="60">
Egg masses (
<figureCitation id="A3CFCBE90029B30F9C22F87AFE04D9B5" box="[359,431,1947,1973]" captionStart="FIGURE 7" captionStartId="9.[151,255,1900,1924]" captionTargetBox="[288,1303,533,1860]" captionTargetId="figure@9.[264,1323,500,1876]" captionTargetPageId="9" captionText="FIGURE 7. Egg masses of Gerstfeldtiancylus ushunensis, sp. nov.. A – C. Lateral and dorsal view (going from top to bottom). cam, capsular albuminous matrix; cm, capsular membrane; emb, embryo; et, extrasyncapsular tunic; sm, syncapsular matrix; stu, syncapsular tunic; tt, terminal tail. All scale bars: 2 mm." httpUri="https://zenodo.org/record/176569/files/figure.png" pageId="5" pageNumber="60">Fig. 7</figureCitation>
A–C, 
<tableCitation id="7676E2D70029B30F9CBCF87AFDFCD9B5" box="[505,599,1947,1973]" captionStart="TABLE 2" captionStartId="6.[151,239,351,375]" captionTargetBox="[159,1422,408,1063]" captionTargetPageId="6" captionText="TABLE 2. Dimensional characteristics of Gerstfeldtiancylus ushunensis, sp. nov. egg masses (in mm)." httpUri="http://table.plazi.org/id/6F8B87E4002AB30C9DD2FEBEFB4ADF77" pageId="5" pageNumber="60" tableUuid="6F8B87E4002AB30C9DD2FEBEFB4ADF77">Table 2</tableCitation>
): New syncapsulae are bluish-translucent with convex upper surface, large (l = 4.8–7.0 mm, w = 4.0– 
<quantity id="FC0C7A890029B30F9F54F822FDC6D9DD" box="[529,621,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="60" unit="mm" value="6.5">6.5 mm</quantity>
, h = 
<quantity id="FC0C7A890029B30F9FEDF822FC90D9DD" box="[680,827,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.5" metricValueMin="1.3" pageId="5" pageNumber="60" unit="mm" value="1.9" valueMax="2.5" valueMin="1.3">1.3–2.5 mm</quantity>
). The clutch is oval or ellipsoidal. Extrasyncapsular tunic is very broad, with uneven fimbriated margins. Terminal part of the clutch protrudes like a spine. One syncapsula contains two to five egg capsulae. Egg capsulae are very large (up to 
<quantity id="FC0C7A89002AB30C992CFF21FB6EDEDA" box="[1129,1221,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="61" unit="mm" value="2.8">2.8 mm</quantity>
long and 
<quantity id="FC0C7A89002AB30C9805FF21FA37DEDA" box="[1344,1436,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="61" unit="mm" value="2.3">2.3 mm</quantity>
wide), oval with centrally or peripherally positioned eggs. Unrounded stones free of plant coatings serve as substratum.
</paragraph>
<caption id="6F8B87E4002AB30C9DD2FEBEFB4ADF77" ID-Table-UUID="6F8B87E4002AB30C9DD2FEBEFB4ADF77" box="[151,1249,351,375]" httpUri="http://table.plazi.org/id/6F8B87E4002AB30C9DD2FEBEFB4ADF77" pageId="6" pageNumber="61" targetBox="[159,1422,408,1063]" targetIsTable="true" targetPageId="6">
<paragraph id="3B4BD76C002AB30C9DD2FEBEFB4ADF77" blockId="6.[151,1249,351,375]" box="[151,1249,351,375]" pageId="6" pageNumber="61">
<emphasis id="09800B7E002AB30C9DD2FEBEFEA3DF77" bold="true" box="[151,264,351,375]" pageId="6" pageNumber="61">TABLE 2.</emphasis>
Dimensional characteristics of 
<taxonomicName id="FCF4ACEF002AB30C9F1EFE81FC31DF77" box="[603,922,352,375]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="ushunensis" status="sp. nov.">
<emphasis id="09800B7E002AB30C9F1EFE81FC31DF77" box="[603,922,352,375]" italics="true" pageId="6" pageNumber="61">Gerstfeldtiancylus ushunensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="12B3B605002AB30C9EEDFEBEFC55DF77" box="[936,1022,351,375]" pageId="6" pageNumber="61" rank="species">
<emphasis id="09800B7E002AB30C9EEDFEBEFC55DF77" bold="true" box="[936,1022,351,375]" pageId="6" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
egg masses (in mm).
</paragraph>
</caption>
<paragraph id="3B4BD76C002AB30C9DDAFE79FA27DA27" pageId="6" pageNumber="61">
<table id="49F425CC002A4CF59DDAFE79FA25DA27" box="[159,1422,408,1063]" gridcols="7" gridrows="11" pageId="6" pageNumber="61">
<tr id="85C4D52E002A4CF59DDAFE79FA25DFD1" box="[159,1422,408,465]" gridrow="0" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFE79FF6CDFD1" box="[159,199,408,465]" gridcol="0" gridrow="0" pageId="6" pageNumber="61">No.</th>
<th id="C615BC52002A4CF59DA7FE79FEFCDFD1" box="[226,343,408,465]" gridcol="1" gridrow="0" pageId="6" pageNumber="61">Length of syncapsule</th>
<th id="C615BC52002A4CF59CCDFE79FDB0DFD1" box="[392,539,408,465]" gridcol="2" gridrow="0" pageId="6" pageNumber="61">Width of syn- capsule</th>
<th id="C615BC52002A4CF59F76FE79FD14DFD1" box="[563,703,408,465]" gridcol="3" gridrow="0" pageId="6" pageNumber="61">Convexity of syncapsule</th>
<th id="C615BC52002A4CF59F9AFE79FCC3DFD1" box="[735,872,408,465]" gridcol="4" gridrow="0" pageId="6" pageNumber="61">Number of egg capsulae</th>
<th id="C615BC52002A4CF59EC2FE79FBD4DFD1" box="[903,1151,408,465]" gridcol="5" gridrow="0" pageId="6" pageNumber="61">Length of egg capsulae</th>
<th id="C615BC52002A4CF599DAFE79FA25DFD1" box="[1183,1422,408,465]" gridcol="6" gridrow="0" pageId="6" pageNumber="61">Width of egg capsulae</th>
</tr>
<tr id="85C4D52E002A4CF59DDAFE04FA25DFFC" box="[159,1422,485,508]" gridrow="1" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFE04FF6CDFFC" box="[159,199,485,508]" gridcol="0" gridrow="1" pageId="6" pageNumber="61">1</th>
<td id="C615BC52002A4CF59DA7FE04FEFCDFFC" box="[226,343,485,508]" gridcol="1" gridrow="1" pageId="6" pageNumber="61">5.12</td>
<td id="C615BC52002A4CF59CCDFE04FDB0DFFC" box="[392,539,485,508]" gridcol="2" gridrow="1" pageId="6" pageNumber="61">4.60</td>
<td id="C615BC52002A4CF59F76FE04FD14DFFC" box="[563,703,485,508]" gridcol="3" gridrow="1" pageId="6" pageNumber="61">1.38</td>
<td id="C615BC52002A4CF59F9AFE04FCC3DFFC" box="[735,872,485,508]" gridcol="4" gridrow="1" pageId="6" pageNumber="61">2</td>
<td id="C615BC52002A4CF59EC2FE04FBD4DFFC" box="[903,1151,485,508]" gridcol="5" gridrow="1" pageId="6" pageNumber="61">1.75–2.79</td>
<td id="C615BC52002A4CF599DAFE04FA25DFFC" box="[1183,1422,485,508]" gridcol="6" gridrow="1" pageId="6" pageNumber="61">1.25–2.27</td>
</tr>
<tr id="85C4D52E002A4CF59DDAFDF1FA25DC51" box="[159,1422,528,593]" gridrow="2" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFDF1FF6CDC51" box="[159,199,528,593]" gridcol="0" gridrow="2" pageId="6" pageNumber="61">2 3</th>
<td id="C615BC52002A4CF59DA7FDF1FEFCDC51" box="[226,343,528,593]" gridcol="1" gridrow="2" pageId="6" pageNumber="61">5.26 6.59</td>
<td id="C615BC52002A4CF59CCDFDF1FDB0DC51" box="[392,539,528,593]" gridcol="2" gridrow="2" pageId="6" pageNumber="61">4.52 4.83</td>
<td id="C615BC52002A4CF59F76FDF1FD14DC51" box="[563,703,528,593]" gridcol="3" gridrow="2" pageId="6" pageNumber="61">1.63 1.86</td>
<td id="C615BC52002A4CF59F9AFDF1FCC3DC51" box="[735,872,528,593]" gridcol="4" gridrow="2" pageId="6" pageNumber="61">4 3</td>
<td id="C615BC52002A4CF59EC2FDF1FBD4DC51" box="[903,1151,528,593]" gridcol="5" gridrow="2" pageId="6" pageNumber="61">1.54±0.59 (1.06–2.37) 1.96±0.33 (1.59–2.22)</td>
<td id="C615BC52002A4CF599DAFDF1FA25DC51" box="[1183,1422,528,593]" gridcol="6" gridrow="2" pageId="6" pageNumber="61">1.25±0.41 (0.88–1.84) 1.54±0.27 (1.23–1.73)</td>
</tr>
<tr id="85C4D52E002A4CF59DDAFD84FA25DC7C" box="[159,1422,613,636]" gridrow="3" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFD84FF6CDC7C" box="[159,199,613,636]" gridcol="0" gridrow="3" pageId="6" pageNumber="61">4</th>
<td id="C615BC52002A4CF59DA7FD84FEFCDC7C" box="[226,343,613,636]" gridcol="1" gridrow="3" pageId="6" pageNumber="61">5.30</td>
<td id="C615BC52002A4CF59CCDFD84FDB0DC7C" box="[392,539,613,636]" gridcol="2" gridrow="3" pageId="6" pageNumber="61">4.00</td>
<td id="C615BC52002A4CF59F76FD84FD14DC7C" box="[563,703,613,636]" gridcol="3" gridrow="3" pageId="6" pageNumber="61">1.60</td>
<td id="C615BC52002A4CF59F9AFD84FCC3DC7C" box="[735,872,613,636]" gridcol="4" gridrow="3" pageId="6" pageNumber="61">5</td>
<td id="C615BC52002A4CF59EC2FD84FBD4DC7C" box="[903,1151,613,636]" gridcol="5" gridrow="3" pageId="6" pageNumber="61">2.25±0.42 (1.57–2.64)</td>
<td id="C615BC52002A4CF599DAFD84FA25DC7C" box="[1183,1422,613,636]" gridcol="6" gridrow="3" pageId="6" pageNumber="61">1.64±0.19 (1.51–1.97)</td>
</tr>
<tr id="85C4D52E002A4CF59DDAFD71FA25DCD1" box="[159,1422,656,721]" gridrow="4" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFD71FF6CDCD1" box="[159,199,656,721]" gridcol="0" gridrow="4" pageId="6" pageNumber="61">5 6</th>
<td id="C615BC52002A4CF59DA7FD71FEFCDCD1" box="[226,343,656,721]" gridcol="1" gridrow="4" pageId="6" pageNumber="61">6.90 7.00</td>
<td id="C615BC52002A4CF59CCDFD71FDB0DCD1" box="[392,539,656,721]" gridcol="2" gridrow="4" pageId="6" pageNumber="61">5.20 6.50</td>
<td id="C615BC52002A4CF59F76FD71FD14DCD1" box="[563,703,656,721]" gridcol="3" gridrow="4" pageId="6" pageNumber="61">2.30 2.80</td>
<td id="C615BC52002A4CF59F9AFD71FCC3DCD1" box="[735,872,656,721]" gridcol="4" gridrow="4" pageId="6" pageNumber="61">5 5</td>
<td id="C615BC52002A4CF59EC2FD71FBD4DCD1" box="[903,1151,656,721]" gridcol="5" gridrow="4" pageId="6" pageNumber="61">2.67±0.35 (2.17–2.99) 2.80±0.56 (2.11–3.26)</td>
<td id="C615BC52002A4CF599DAFD71FA25DCD1" box="[1183,1422,656,721]" gridcol="6" gridrow="4" pageId="6" pageNumber="61">1.97±0.16 (1.84–2.18) 2.28±0.48 (1.63–2.79)</td>
</tr>
<tr id="85C4D52E002A4CF59DDAFD04FA25DCFC" box="[159,1422,741,764]" gridrow="5" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFD04FF6CDCFC" box="[159,199,741,764]" gridcol="0" gridrow="5" pageId="6" pageNumber="61">7</th>
<td id="C615BC52002A4CF59DA7FD04FEFCDCFC" box="[226,343,741,764]" gridcol="1" gridrow="5" pageId="6" pageNumber="61">6.00</td>
<td id="C615BC52002A4CF59CCDFD04FDB0DCFC" box="[392,539,741,764]" gridcol="2" gridrow="5" pageId="6" pageNumber="61">5.30</td>
<td id="C615BC52002A4CF59F76FD04FD14DCFC" box="[563,703,741,764]" gridcol="3" gridrow="5" pageId="6" pageNumber="61">2.94</td>
<td id="C615BC52002A4CF59F9AFD04FCC3DCFC" box="[735,872,741,764]" gridcol="4" gridrow="5" pageId="6" pageNumber="61">4</td>
<td id="C615BC52002A4CF59EC2FD04FBD4DCFC" box="[903,1151,741,764]" gridcol="5" gridrow="5" pageId="6" pageNumber="61">2.45±0.47 (1.88–2.92)</td>
<td id="C615BC52002A4CF599DAFD04FA25DCFC" box="[1183,1422,741,764]" gridcol="6" gridrow="5" pageId="6" pageNumber="61">1.96±0.59 (1.35–2.77)</td>
</tr>
<tr id="85C4D52E002A4CF59DDAFCF1FA25DD51" box="[159,1422,784,849]" gridrow="6" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFCF1FF6CDD51" box="[159,199,784,849]" gridcol="0" gridrow="6" pageId="6" pageNumber="61">8 9</th>
<td id="C615BC52002A4CF59DA7FCF1FEFCDD51" box="[226,343,784,849]" gridcol="1" gridrow="6" pageId="6" pageNumber="61">5.90 5.30</td>
<td id="C615BC52002A4CF59CCDFCF1FDB0DD51" box="[392,539,784,849]" gridcol="2" gridrow="6" pageId="6" pageNumber="61">4.20 5.00</td>
<td id="C615BC52002A4CF59F76FCF1FD14DD51" box="[563,703,784,849]" gridcol="3" gridrow="6" pageId="6" pageNumber="61">2.00 1.90</td>
<td id="C615BC52002A4CF59F9AFCF1FCC3DD51" box="[735,872,784,849]" gridcol="4" gridrow="6" pageId="6" pageNumber="61">3 2</td>
<td id="C615BC52002A4CF59EC2FCF1FBD4DD51" box="[903,1151,784,849]" gridcol="5" gridrow="6" pageId="6" pageNumber="61">3.01±0.1 (2.95–3.13) 2.34–3.27</td>
<td id="C615BC52002A4CF599DAFCF1FA25DD51" box="[1183,1422,784,849]" gridcol="6" gridrow="6" pageId="6" pageNumber="61">2.11±0.29 (1.81–2.39) 1.65–2.95</td>
</tr>
<tr id="85C4D52E002A4CF59DDAFC84FA25DD7C" box="[159,1422,869,892]" gridrow="7" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFC84FF6CDD7C" box="[159,199,869,892]" gridcol="0" gridrow="7" pageId="6" pageNumber="61">10</th>
<td id="C615BC52002A4CF59DA7FC84FEFCDD7C" box="[226,343,869,892]" gridcol="1" gridrow="7" pageId="6" pageNumber="61">5.70</td>
<td id="C615BC52002A4CF59CCDFC84FDB0DD7C" box="[392,539,869,892]" gridcol="2" gridrow="7" pageId="6" pageNumber="61">5.50</td>
<td id="C615BC52002A4CF59F76FC84FD14DD7C" box="[563,703,869,892]" gridcol="3" gridrow="7" pageId="6" pageNumber="61">1.50</td>
<td id="C615BC52002A4CF59F9AFC84FCC3DD7C" box="[735,872,869,892]" gridcol="4" gridrow="7" pageId="6" pageNumber="61">2</td>
<td id="C615BC52002A4CF59EC2FC84FBD4DD7C" box="[903,1151,869,892]" gridcol="5" gridrow="7" pageId="6" pageNumber="61">2.68–3.49</td>
<td id="C615BC52002A4CF599DAFC84FA25DD7C" box="[1183,1422,869,892]" gridcol="6" gridrow="7" pageId="6" pageNumber="61">2.25–2.30</td>
</tr>
<tr id="85C4D52E002A4CF59DDAFC71FA25DDD1" box="[159,1422,912,977]" gridrow="8" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFC71FF6CDDD1" box="[159,199,912,977]" gridcol="0" gridrow="8" pageId="6" pageNumber="61">11 12</th>
<td id="C615BC52002A4CF59DA7FC71FEFCDDD1" box="[226,343,912,977]" gridcol="1" gridrow="8" pageId="6" pageNumber="61">5.70 4.80</td>
<td id="C615BC52002A4CF59CCDFC71FDB0DDD1" box="[392,539,912,977]" gridcol="2" gridrow="8" pageId="6" pageNumber="61">5.55 4.35</td>
<td id="C615BC52002A4CF59F76FC71FD14DDD1" box="[563,703,912,977]" gridcol="3" gridrow="8" pageId="6" pageNumber="61">2.30 1.40</td>
<td id="C615BC52002A4CF59F9AFC71FCC3DDD1" box="[735,872,912,977]" gridcol="4" gridrow="8" pageId="6" pageNumber="61">3 2</td>
<td id="C615BC52002A4CF59EC2FC71FBD4DDD1" box="[903,1151,912,977]" gridcol="5" gridrow="8" pageId="6" pageNumber="61">2.74±1.20 (1.97–4.12) 2.60–2.66</td>
<td id="C615BC52002A4CF599DAFC71FA25DDD1" box="[1183,1422,912,977]" gridcol="6" gridrow="8" pageId="6" pageNumber="61">2.08±0.59 (1.65–2.75) 1.74–2.14</td>
</tr>
<tr id="85C4D52E002A4CF59DDAFC04FA25DDFC" box="[159,1422,997,1020]" gridrow="9" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFC04FF6CDDFC" box="[159,199,997,1020]" gridcol="0" gridrow="9" pageId="6" pageNumber="61">13</th>
<td id="C615BC52002A4CF59DA7FC04FEFCDDFC" box="[226,343,997,1020]" gridcol="1" gridrow="9" pageId="6" pageNumber="61">5.50</td>
<td id="C615BC52002A4CF59CCDFC04FDB0DDFC" box="[392,539,997,1020]" gridcol="2" gridrow="9" pageId="6" pageNumber="61">4.00</td>
<td id="C615BC52002A4CF59F76FC04FD14DDFC" box="[563,703,997,1020]" gridcol="3" gridrow="9" pageId="6" pageNumber="61">2.10</td>
<td id="C615BC52002A4CF59F9AFC04FCC3DDFC" box="[735,872,997,1020]" gridcol="4" gridrow="9" pageId="6" pageNumber="61">3</td>
<td id="C615BC52002A4CF59EC2FC04FBD4DDFC" box="[903,1151,997,1020]" gridcol="5" gridrow="9" pageId="6" pageNumber="61">3.15±0.22 (2.92–3.36)</td>
<td id="C615BC52002A4CF599DAFC04FA25DDFC" box="[1183,1422,997,1020]" gridcol="6" gridrow="9" pageId="6" pageNumber="61">2.11±0.09 (2.05–2.21)</td>
</tr>
<tr id="85C4D52E002A4CF59DDAFBF1FA25DA27" box="[159,1422,1040,1063]" gridrow="10" pageId="6" pageNumber="61">
<th id="C615BC52002A4CF59DDAFBF1FF6CDA27" box="[159,199,1040,1063]" gridcol="0" gridrow="10" pageId="6" pageNumber="61">14</th>
<td id="C615BC52002A4CF59DA7FBF1FEFCDA27" box="[226,343,1040,1063]" gridcol="1" gridrow="10" pageId="6" pageNumber="61">5.80</td>
<td id="C615BC52002A4CF59CCDFBF1FDB0DA27" box="[392,539,1040,1063]" gridcol="2" gridrow="10" pageId="6" pageNumber="61">5.50</td>
<td id="C615BC52002A4CF59F76FBF1FD14DA27" box="[563,703,1040,1063]" gridcol="3" gridrow="10" pageId="6" pageNumber="61">2.30</td>
<td id="C615BC52002A4CF59F9AFBF1FCC3DA27" box="[735,872,1040,1063]" gridcol="4" gridrow="10" pageId="6" pageNumber="61">3</td>
<td id="C615BC52002A4CF59EC2FBF1FBD4DA27" box="[903,1151,1040,1063]" gridcol="5" gridrow="10" pageId="6" pageNumber="61">3.06±0.40 (2.61–3.37)</td>
<td id="C615BC52002A4CF599DAFBF1FA25DA27" box="[1183,1422,1040,1063]" gridcol="6" gridrow="10" pageId="6" pageNumber="61">2.27±0.16 (2.11–2.42)</td>
</tr>
</table>
</paragraph>
<caption id="6F8B87E4002AB30C9DD2F955FBFCD8EE" httpUri="https://zenodo.org/record/176566/files/figure.png" pageId="6" pageNumber="61" targetBox="[264,1322,1146,1692]" targetPageId="6">
<paragraph id="3B4BD76C002AB30C9DD2F955FBFCD8EE" blockId="6.[151,1436,1716,1774]" pageId="6" pageNumber="61">
<emphasis id="09800B7E002AB30C9DD2F955FEB2D8CC" bold="true" box="[151,281,1716,1740]" pageId="6" pageNumber="61">FIGURE 4.</emphasis>
Radula of 
<taxonomicName id="FCF4ACEF002AB30C9CCAF955FD64D8CB" box="[399,719,1716,1739]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="ushunensis" status="sp. nov.">
<emphasis id="09800B7E002AB30C9CCAF955FD64D8CB" box="[399,719,1716,1739]" italics="true" pageId="6" pageNumber="61">Gerstfeldtiancylus ushunensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="12B3B605002AB30C9F99F955FC99D8CC" box="[732,818,1716,1740]" pageId="6" pageNumber="61" rank="species">
<emphasis id="09800B7E002AB30C9F99F955FC99D8CC" bold="true" box="[732,818,1716,1740]" pageId="6" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
(SEM). A–C. ZIN RAS 2/521–2006: A. Central tooth. B. Three transverse rows of teeth. C. Lateral teeth. Scale bars: A = 0.01 mm; B–C = 0.1 mm.
</paragraph>
</caption>
<paragraph id="3B4BD76C002AB3029D82F8C0FA37D9DE" blockId="6.[151,1436,1825,2012]" lastBlockId="8.[151,1436,1908,2014]" lastPageId="8" lastPageNumber="63" pageId="6" pageNumber="61">
<emphasis id="09800B7E002AB30C9D82F8C0FE76D93B" bold="true" box="[199,477,1825,1851]" pageId="6" pageNumber="61">Differential diagnosis.</emphasis>
By shell morphology 
<taxonomicName id="FCF4ACEF002AB30C9FAAF8C3FBFAD93B" box="[751,1105,1826,1851]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="ushunensis">
<emphasis id="09800B7E002AB30C9FAAF8C3FBFAD93B" box="[751,1105,1826,1851]" italics="true" pageId="6" pageNumber="61">Gerstfeldtiancylus ushunensis</emphasis>
</taxonomicName>
is most similar to 
<emphasis id="09800B7E002AB30C987FF8C3FF7ED963" italics="true" pageId="6" pageNumber="61">G. ro e p - storfi</emphasis>
. The teleoconchs of both species are relatively low (H/L = 0.30–0.47) and the apex is distant from the anterior edge of aperture by little over 1/2 of its length. 
<emphasis id="09800B7E002AB30C9E68F893FC15D98B" box="[813,958,1906,1931]" italics="true" pageId="6" pageNumber="61">G. ro e p s t o r f i</emphasis>
differs by an apex location precisely on the longitudinal axis of the shell. The shells of 
<taxonomicName id="FCF4ACEF002AB30C9F81F87BFCFBD9B3" box="[708,848,1946,1971]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="kotyensis">
<emphasis id="09800B7E002AB30C9F81F87BFCFBD9B3" box="[708,848,1946,1971]" italics="true" pageId="6" pageNumber="61">G. kotyensis</emphasis>
</taxonomicName>
and 
<emphasis id="09800B7E002AB30C9ECEF87BFBACD9B3" box="[907,1031,1946,1971]" italics="true" pageId="6" pageNumber="61">G. re n a rd i i</emphasis>
are higher (H/L = 0.45–0.60) and the apex is more distant from the anterior edge of the aperture (3/4 of its length). The apex of 
<taxonomicName id="FCF4ACEF002AB30C99B3F823FA2AD9DB" box="[1270,1409,1986,2011]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="6" pageNumber="61" phylum="Mollusca" rank="species" species="kotyensis">
<emphasis id="09800B7E002AB30C99B3F823FA2AD9DB" box="[1270,1409,1986,2011]" italics="true" pageId="6" pageNumber="61">G. kotyensis</emphasis>
</taxonomicName>
is located on the longitudinal shell axis, that of 
<emphasis id="09800B7E0024B3029FF2F895FC9FD98D" box="[695,820,1908,1933]" italics="true" pageId="8" pageNumber="63">G. re n a rd i i</emphasis>
is considerably shifted to the left. In contrast to 
<taxonomicName id="FCF4ACEF0024B30298C6F895FEB3D9B5" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="8" pageNumber="63" phylum="Mollusca" rank="species" species="ushunensis">
<emphasis id="09800B7E0024B30298C6F895FEB3D9B5" italics="true" pageId="8" pageNumber="63">G. ushunensis</emphasis>
</taxonomicName>
, all remaining species of the subgenus have a concave or straight posterior teleoconch slope and a relatively high protoconch (H/h = 4–5). Radular morphology of 
<taxonomicName id="FCF4ACEF0024B3029EC9F825FB86D9DD" box="[908,1069,1988,2013]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="8" pageNumber="63" phylum="Mollusca" rank="species" species="ushunensis">
<emphasis id="09800B7E0024B3029EC9F825FB86D9DD" box="[908,1069,1988,2013]" italics="true" pageId="8" pageNumber="63">G. ushunensis</emphasis>
</taxonomicName>
and 
<emphasis id="09800B7E0024B3029922F825FB49D9DD" box="[1127,1250,1988,2013]" italics="true" pageId="8" pageNumber="63">G. re n a rd i i</emphasis>
is very similar.
</paragraph>
<caption id="6F8B87E4002BB30D9DD2F957FB95D9E2" httpUri="https://zenodo.org/record/176567/files/figure.png" pageId="7" pageNumber="62" targetBox="[274,1303,187,1701]" targetPageId="7">
<paragraph id="3B4BD76C002BB30D9DD2F957FB95D9E2" blockId="7.[151,1436,1718,2018]" pageId="7" pageNumber="62">
<emphasis id="09800B7E002BB30D9DD2F957FEB2D8CE" bold="true" box="[151,281,1718,1742]" pageId="7" pageNumber="62">FIGURE 5.</emphasis>
A–B. Anatomy of 
<taxonomicName id="FCF4ACEF002BB30D9CA0F957FC8FD8CD" box="[485,804,1718,1741]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="7" pageNumber="62" phylum="Mollusca" rank="species" species="ushunensis" status="sp. nov.">
<emphasis id="09800B7E002BB30D9CA0F957FC8FD8CD" box="[485,804,1718,1741]" italics="true" pageId="7" pageNumber="62">Gerstfeldtiancylus ushunensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="12B3B605002BB30D9E77F957FC2CD8CE" box="[818,903,1718,1742]" pageId="7" pageNumber="62" rank="species">
<emphasis id="09800B7E002BB30D9E77F957FC2CD8CE" bold="true" box="[818,903,1718,1742]" pageId="7" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
. А. General topography of main organs of the alimentary system, dorsal view. Digestive gland removed. B. Dorsal view of specimen without shell. C–D. Odontophore of 
<taxonomicName id="FCF4ACEF002BB30D9DD2F91AFE80D912" box="[151,299,1787,1810]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="7" pageNumber="62" phylum="Mollusca" rank="species" species="ushunensis" status="sp. nov.">
<emphasis id="09800B7E002BB30D9DD2F91AFE80D912" box="[151,299,1787,1810]" italics="true" pageId="7" pageNumber="62">G. ushunensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="12B3B605002BB30D9C7FF91AFE3AD913" box="[314,401,1787,1811]" pageId="7" pageNumber="62" rank="species">
<emphasis id="09800B7E002BB30D9C7FF91AFE3AD913" bold="true" box="[314,401,1787,1811]" pageId="7" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
. E–L. Jaws of different 
<taxonomicName id="FCF4ACEF002BB30D9FD9F91AFCF4D912" box="[668,863,1787,1810]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="7" pageNumber="62" phylum="Mollusca" rank="genus">
<emphasis id="09800B7E002BB30D9FD9F91AFCF4D912" box="[668,863,1787,1810]" italics="true" pageId="7" pageNumber="62">Gerstfeldtiancylus</emphasis>
</taxonomicName>
species. E–H. 
<taxonomicName id="FCF4ACEF002BB30D9946F91AFB3CD912" box="[1027,1175,1787,1810]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="7" pageNumber="62" phylum="Mollusca" rank="species" species="ushunensis" status="sp. nov.">
<emphasis id="09800B7E002BB30D9946F91AFB3CD912" box="[1027,1175,1787,1810]" italics="true" pageId="7" pageNumber="62">G. ushunensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="12B3B605002BB30D99E3F91AFB55D913" box="[1190,1278,1787,1811]" pageId="7" pageNumber="62" rank="species">
<emphasis id="09800B7E002BB30D99E3F91AFB55D913" bold="true" box="[1190,1278,1787,1811]" pageId="7" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
(A, C–E. ZIN RAS 1/521–2006. B, F–H. ZIN RAS 2/521–2006). I. 
<taxonomicName id="FCF4ACEF002BB30D9FACF8FFFB9BD935" authority="Dybowski, 1884" authorityName="Dybowski" authorityYear="1884" box="[745,1072,1822,1845]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="7" pageNumber="62" phylum="Mollusca" rank="species" species="renardii">
<emphasis id="09800B7E002BB30D9FACF8FFFCF5D935" box="[745,862,1822,1845]" italics="true" pageId="7" pageNumber="62">G. renardii</emphasis>
(Dybowski, 1884)
</taxonomicName>
. J. 
<taxonomicName id="FCF4ACEF002BB30D9912F8FFFF7BD958" authority="Shirokaya et al., 2003" authorityName="Shirokaya et al." authorityYear="2003" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="7" pageNumber="62" phylum="Mollusca" rank="species" species="roepstorfi">
<emphasis id="09800B7E002BB30D9912F8FFFB74D935" box="[1111,1247,1822,1845]" italics="true" pageId="7" pageNumber="62">G. roepstorfi</emphasis>
Shirokaya et al., 2003
</taxonomicName>
. K. 
<taxonomicName id="FCF4ACEF002BB30D9DBFF8A0FDCCD958" authority="Starobogatov, 1989" authorityName="Starobogatov" authorityYear="1989" box="[250,615,1857,1880]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="7" pageNumber="62" phylum="Mollusca" rank="species" species="benedictiae">
<emphasis id="09800B7E002BB30D9DBFF8A0FE39D958" box="[250,402,1857,1880]" italics="true" pageId="7" pageNumber="62">G. benedictiae</emphasis>
Starobogatov, 1989
</taxonomicName>
(whole mounts in Faure-Berlezet liquid). L. 
<taxonomicName id="FCF4ACEF002BB30D9902F8A0FA37D958" authority="Starobogatov, 1989" authorityName="Starobogatov" authorityYear="1989" box="[1095,1436,1857,1880]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="7" pageNumber="62" phylum="Mollusca" rank="species" species="kotyensis">
<emphasis id="09800B7E002BB30D9902F8A0FB6CD958" box="[1095,1223,1857,1880]" italics="true" pageId="7" pageNumber="62">G. kotyensis</emphasis>
Starobogatov, 1989
</taxonomicName>
(frontal histological section). I–L. From Shirokaya &amp; Röpstorf (2004). aal, left anterior adductor; aar, right anterior adductor; ap, anal pore; bm, buccal mass; cae, caecum; dgp, digestive gland pores; int1, first intestinal loop; int2, second intestinal loop; mb, mantle border; oa, original or posterior adductor; oe, oesophagus; rs, radular sac; sg, salivary glands; st, stomach. Scale bars: A–D = 1 mm; E = 0.8 mm; F–H, J = 0.5 mm; I, K–L = 0.3 mm.
</paragraph>
</caption>
<caption id="6F8B87E40024B3029DD2F99AFE11D940" httpUri="https://zenodo.org/record/176568/files/figure.png" pageId="8" pageNumber="63" targetBox="[290,1306,186,1659]" targetPageId="8">
<paragraph id="3B4BD76C0024B3029DD2F99AFE11D940" blockId="8.[151,1437,1659,1856]" pageId="8" pageNumber="63">
<emphasis id="09800B7E0024B3029DD2F99AFEB2D893" bold="true" box="[151,281,1659,1683]" pageId="8" pageNumber="63">FIGURE 6.</emphasis>
Longitudinally sectioned male copulatory organ of 
<taxonomicName id="FCF4ACEF0024B3029E02F99DFBA1D893" box="[839,1034,1660,1683]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="8" pageNumber="63" phylum="Mollusca" rank="genus">
<emphasis id="09800B7E0024B3029E02F99DFBA1D893" box="[839,1034,1660,1683]" italics="true" pageId="8" pageNumber="63">Gerstfeldtiancylus</emphasis>
</taxonomicName>
species (A–D, G. diagrammic). A. 
<taxonomicName id="FCF4ACEF0024B30298C0F99DFE44D8B6" authority="Starobogatov, 1989" authorityName="Starobogatov" authorityYear="1989" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="8" pageNumber="63" phylum="Mollusca" rank="species" species="benedictiae">
<emphasis id="09800B7E0024B30298C0F99DFEB9D8B6" italics="true" pageId="8" pageNumber="63">G. benedictiae</emphasis>
Starobogatov, 1989
</taxonomicName>
. B, E. 
<taxonomicName id="FCF4ACEF0024B3029F7AF97EFC6CD8B6" authority="Shirokaya et al., 2003" authorityName="Shirokaya et al." authorityYear="2003" box="[575,967,1695,1718]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="8" pageNumber="63" phylum="Mollusca" rank="species" species="roepstorfi">
<emphasis id="09800B7E0024B3029F7AF97EFD6CD8B6" box="[575,711,1695,1718]" italics="true" pageId="8" pageNumber="63">G. roepstorfi</emphasis>
Shirokaya et al., 2003
</taxonomicName>
. C. 
<taxonomicName id="FCF4ACEF0024B3029EB0F97EFAFED8B6" authority="Starobogatov, 1989" authorityName="Starobogatov" authorityYear="1989" box="[1013,1365,1695,1718]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="8" pageNumber="63" phylum="Mollusca" rank="species" species="kotyensis">
<emphasis id="09800B7E0024B3029EB0F97EFBDCD8B6" box="[1013,1143,1695,1718]" italics="true" pageId="8" pageNumber="63">G. kotyensis</emphasis>
Starobogatov, 1989
</taxonomicName>
. D. 
<taxonomicName id="FCF4ACEF0024B30298C0F97EFE1CD8D8" authority="Dybowski, 1884" authorityName="Dybowski" authorityYear="1884" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="8" pageNumber="63" phylum="Mollusca" rank="species" species="renardii">
<emphasis id="09800B7E0024B30298C0F97EFF47D8D9" italics="true" pageId="8" pageNumber="63">G. renardii</emphasis>
(Dybowski, 1884)
</taxonomicName>
. A–E. From Kruglov &amp; Starobogatov (1991a), Shirokaya (2005). F–G. 
<taxonomicName id="FCF4ACEF0024B30299ECF923FA91D8D9" box="[1193,1338,1730,1753]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="8" pageNumber="63" phylum="Mollusca" rank="species" species="ushunensis" status="sp. nov.">
<emphasis id="09800B7E0024B30299ECF923FA91D8D9" box="[1193,1338,1730,1753]" italics="true" pageId="8" pageNumber="63">G. ushunensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="12B3B6050024B3029802F920FA37D8D9" box="[1351,1436,1729,1753]" pageId="8" pageNumber="63" rank="species">
<emphasis id="09800B7E0024B3029802F920FA37D8D9" bold="true" box="[1351,1436,1729,1753]" pageId="8" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
(ZIN RAS 2/521–2006). fl, flagellum; mp, male genital pore; p, penis; ps, penis sheath; psr, penis sheath retractor; pr, preputium; prc, preputium chamber; prp, preputium pilasters; sar, sarcobellum; vd, vas deferens; ve, velum. Scale bars: A = 0.5 mm; B–G = 1 mm.
</paragraph>
</caption>
<paragraph id="3B4BD76C0025B3039DD2FF79FD8EDF52" blockId="9.[151,1437,152,463]" pageId="9" pageNumber="64">
The latter differs by having a lower number of teeth in one transverse row. All other representatives of the genus differ from 
<taxonomicName id="FCF4ACEF0025B3039C31FF21FDB2DED9" box="[372,537,192,217]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="9" pageNumber="64" phylum="Mollusca" rank="species" species="ushunensis">
<emphasis id="09800B7E0025B3039C31FF21FDB2DED9" box="[372,537,192,217]" italics="true" pageId="9" pageNumber="64">G. ushunensis</emphasis>
</taxonomicName>
by having long salivary glands, a rounded or oval surface of left anterior adductor, a bilaterally symmetrical position of the posterior adductor and by the absence of a velum in the male copulatory organ (
<figureCitation id="A3CFCBE90025B3039CE9FEF1FE5BDF2A" box="[428,496,272,298]" captionStart="FIGURE 6" captionStartId="8.[151,255,1659,1683]" captionTargetBox="[290,1306,186,1659]" captionTargetId="figure@8.[275,1316,172,1659]" captionTargetPageId="8" captionText="FIGURE 6. Longitudinally sectioned male copulatory organ of Gerstfeldtiancylus species (A – D, G. diagrammic). A. G. benedictiae Starobogatov, 1989. B, E. G. roepstorfi Shirokaya et al., 2003. C. G. kotyensis Starobogatov, 1989. D. G. renardii (Dybowski, 1884). A – E. From Kruglov &amp; Starobogatov (1991 a), Shirokaya (2005). F – G. G. ushunensis, sp. nov. (ZIN RAS 2 / 521 – 2006). fl, flagellum; mp, male genital pore; p, penis; ps, penis sheath; psr, penis sheath retractor; pr, preputium; prc, preputium chamber; prp, preputium pilasters; sar, sarcobellum; vd, vas deferens; ve, velum. Scale bars: A = 0.5 mm; B – G = 1 mm." httpUri="https://zenodo.org/record/176568/files/figure.png" pageId="9" pageNumber="64">Fig. 6</figureCitation>
A–E). 
<taxonomicName id="FCF4ACEF0025B3039F7AFEF1FD74DF29" box="[575,735,272,297]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="9" pageNumber="64" phylum="Mollusca" rank="species" species="ushunensis">
<emphasis id="09800B7E0025B3039F7AFEF1FD74DF29" box="[575,735,272,297]" italics="true" pageId="9" pageNumber="64">G. ushunensis</emphasis>
</taxonomicName>
is also the only species among acroloxids from Lake Baikal with scales on the dorsal jaw part.
</paragraph>
</subSubSection>
<subSubSection id="73EE84E70025B3039D82FEBEFAD6DFCF" pageId="9" pageNumber="64" type="distribution">
<paragraph id="3B4BD76C0025B3039D82FEBEFAD6DFCF" blockId="9.[151,1437,152,463]" pageId="9" pageNumber="64">
<emphasis id="09800B7E0025B3039D82FEBEFDA3DF79" bold="true" box="[199,520,351,377]" pageId="9" pageNumber="64">Distribution and habitat.</emphasis>
<taxonomicName id="FCF4ACEF0025B3039F51FE81FCD3DF79" box="[532,888,352,377]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="9" pageNumber="64" phylum="Mollusca" rank="species" species="ushunensis">
<emphasis id="09800B7E0025B3039F51FE81FCD3DF79" box="[532,888,352,377]" italics="true" pageId="9" pageNumber="64">Gerstfeldtiancylus ushunensis</emphasis>
</taxonomicName>
is only known from the 
<typeStatus id="E44F69CE0025B30399F6FE81FB4EDF7A" box="[1203,1253,352,378]" pageId="9" pageNumber="64">type</typeStatus>
locality and it coexists with 
<taxonomicName id="FCF4ACEF0025B3039C7DFE6AFE74DFA4" box="[312,479,395,420]" class="Gastropoda" family="Acroloxidae" genus="Gerstfeldtiancylus" kingdom="Animalia" order="Hygrophila" pageId="9" pageNumber="64" phylum="Mollusca" rank="species" species="benedictiae">
<emphasis id="09800B7E0025B3039C7DFE6AFE74DFA4" box="[312,479,395,420]" italics="true" pageId="9" pageNumber="64">G. benedictiae</emphasis>
</taxonomicName>
and 
<emphasis id="09800B7E0025B3039F5DFE6AFD0ADFA4" box="[536,673,395,420]" italics="true" pageId="9" pageNumber="64">G. k o t y e n s i s</emphasis>
. The new species is most abundant (up to 
<specimenCount id="2DF21CE50025B30399D3FE6BFAEADFA4" box="[1174,1345,394,420]" pageId="9" pageNumber="64" type="generic">114 specimens</specimenCount>
/m2 and 3.81 biomass g/m2, respectively) at the depth of 
<quantity id="FC0C7A890025B3039F8EFE54FC98DFCF" box="[715,819,437,463]" metricMagnitude="1" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="9" pageNumber="64" unit="m" value="21.0" valueMax="22.0" valueMin="20.0">20–22 m</quantity>
on a bottom composed of multi-layered boulders.
</paragraph>
</subSubSection>
</treatment>
</document>