312 lines
17 KiB
XML
312 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.56.37636" ID-GBIF-Dataset="2d79291b-713a-4311-ae6b-41a35889def7" ID-PMC="PMC6684523" ID-Pensoft-Pub="1314-4049-56-101" ID-Pensoft-UUID="21A48961443D5A14973AB868F57E980E" ID-PubMed="31402842" ModsDocID="1314-4049-56-101" checkinTime="1564475665910" checkinUser="pensoft" docAuthor="Tasanathai, Kanoksri, Noisripoom, Wasana, Chaitika, Thanyarat, Khonsanit, Artit, Hasin, Sasitorn & Luangsa-ard, Jennifer" docDate="2019" docId="80498DBCD394554E8FAFEF6692582427" docLanguage="en" docName="MycoKeys 56: 101-129" docOrigin="MycoKeys 56" docSource="http://dx.doi.org/10.3897/mycokeys.56.37636" docTitle="Ophiocordyceps termiticola Tasanathai, Noisripoom & Luangsa-ard, sp. nov." docType="treatment" docVersion="35" lastPageNumber="101" masterDocId="D900C81EFFFBFFDFA570845BFFA5FFD9" masterDocTitle="Phylogenetic and morphological classification of Ophiocordyceps species on termites from Thailand" masterLastPageNumber="129" masterPageNumber="101" pageNumber="101" updateTime="1668136276120" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Phylogenetic and morphological classification of Ophiocordyceps species on termites from Thailand</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tasanathai, Kanoksri</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Noisripoom, Wasana</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chaitika, Thanyarat</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Khonsanit, Artit</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hasin, Sasitorn</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Luangsa-ard, Jennifer</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>56</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>101</mods:start>
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<mods:end>129</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.56.37636</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.56.37636</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-56-101</mods:identifier>
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<mods:identifier type="Pensoft-UUID">21A48961443D5A14973AB868F57E980E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159292997" LSID="urn:lsid:plazi:treatment:80498DBCD394554E8FAFEF6692582427" httpUri="http://treatment.plazi.org/id/80498DBCD394554E8FAFEF6692582427" lastPageNumber="101" pageId="0" pageNumber="101">
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<subSubSection pageId="0" pageNumber="101" type="nomenclature">
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<paragraph pageId="0" pageNumber="101">
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<taxonomicName LSID="80498dbc-d394-554e-8faf-ef6692582427" authority="Tasanathai, Noisripoom & Luangsa-ard" class="Sordariomycetes" family="Ophiocordycipitaceae" genus="Ophiocordyceps" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ophiocordyceps termiticola" order="Hypocreales" pageId="0" pageNumber="101" phylum="Ascomycota" rank="species" species="termiticola">Ophiocordyceps termiticola Tasanathai, Noisripoom & Luangsa-ard</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="101">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Ophiocordyceps termiticola (BBH 5634, BCC 1920) A stroma of fungus emerging from termite B part of stroma showing perithecia C perithecia D ascus E ascospore F phialides with conidia on synnema G conidium H colony on PDA at 20 d obverse and reverse I phialides with conidia on PDA J conidium K-O scanning electron micrographs of phialides with conidia on PDA P colony on PSA at 20 d obverse and reverse Q phialides with conidia on PSA R colony on SDYA / 4 at 20 d obverse and reverse S phialides with conidia. Scale bars: 2 cm (A); 1 μm (B, K, O); 100 μm (C); 15 μm (D); 8 μm (E); 5 μm (F, G); 7 mm (H, P, R); 3 μm (I, J, L, M, N, Q, S)." figureDoi="10.3897/mycokeys.56.37636.figure8" httpUri="https://binary.pensoft.net/fig/321650" pageId="0" pageNumber="101">Figure 8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="typification">
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<paragraph pageId="0" pageNumber="101">Typification.</paragraph>
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<paragraph pageId="0" pageNumber="101">
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THAILAND. Kanchanaburi Province, Khao Laem National Park; 14°746'N, 98°625'E; on termite; 20 June 1995; N.L. Hywel-Jones, R. Nasit, S. Sivichai (holotype BBH5634 dried culture; ex-type living culture, BCC 1920). GenBank: ITS = MH754724, LSU = MH753678,
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<emphasis italics="true" pageId="0" pageNumber="101">TEF</emphasis>
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= MK284265,
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<emphasis italics="true" pageId="0" pageNumber="101">RPB1</emphasis>
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= MK214108,
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<emphasis italics="true" pageId="0" pageNumber="101">RPB2</emphasis>
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= MK214094
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</paragraph>
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<caption doi="10.3897/mycokeys.56.37636.figure8" httpUri="https://binary.pensoft.net/fig/321650" pageId="0" pageNumber="101" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="101">
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Figure 8.
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<taxonomicName class="Sordariomycetes" family="Ophiocordycipitaceae" genus="Ophiocordyceps" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ophiocordyceps termiticola" order="Hypocreales" pageId="0" pageNumber="101" phylum="Ascomycota" rank="species" species="termiticola">
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<emphasis italics="true" pageId="0" pageNumber="101">Ophiocordyceps termiticola</emphasis>
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</taxonomicName>
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(BBH5634, BCC 1920)
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<emphasis bold="true" pageId="0" pageNumber="101">A</emphasis>
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stroma of fungus emerging from termite
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<emphasis bold="true" pageId="0" pageNumber="101">B</emphasis>
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part of stroma showing perithecia
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<emphasis bold="true" pageId="0" pageNumber="101">C</emphasis>
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perithecia
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<emphasis bold="true" pageId="0" pageNumber="101">D</emphasis>
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ascus
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<emphasis bold="true" pageId="0" pageNumber="101">E</emphasis>
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ascospore
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<emphasis bold="true" pageId="0" pageNumber="101">F</emphasis>
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phialides with conidia on synnema
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<emphasis bold="true" pageId="0" pageNumber="101">G</emphasis>
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conidium
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<emphasis bold="true" pageId="0" pageNumber="101">H</emphasis>
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colony on PDA at 20 d obverse and reverse
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<emphasis bold="true" pageId="0" pageNumber="101">I</emphasis>
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phialides with conidia on PDA
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<emphasis bold="true" pageId="0" pageNumber="101">J</emphasis>
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conidium
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<emphasis bold="true" pageId="0" pageNumber="101">
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<normalizedToken originalValue="K–O">K-O</normalizedToken>
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</emphasis>
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scanning electron micrographs of phialides with conidia on PDA
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<emphasis bold="true" pageId="0" pageNumber="101">P</emphasis>
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colony on PSA at 20 d obverse and reverse
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<emphasis bold="true" pageId="0" pageNumber="101">Q</emphasis>
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phialides with conidia on PSA
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<emphasis bold="true" pageId="0" pageNumber="101">R</emphasis>
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colony on SDYA/4 at 20 d obverse and reverse
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<emphasis bold="true" pageId="0" pageNumber="101">S</emphasis>
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phialides with conidia. Scale bars: 2 cm (
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<emphasis bold="true" pageId="0" pageNumber="101">A</emphasis>
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); 1
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="101">B, K, O</emphasis>
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); 100
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="101">C</emphasis>
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); 15
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="101">D</emphasis>
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); 8
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="101">E</emphasis>
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); 5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="101">F, G</emphasis>
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); 7 mm (
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<emphasis bold="true" pageId="0" pageNumber="101">H, P, R</emphasis>
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); 3
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="101">I, J, L, M, N, Q, S</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="etymology">
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<paragraph pageId="0" pageNumber="101">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="101">
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'
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<taxonomicName class="Pilidiophora" family="Lineidae" genus="Acari" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acari termiticola" order="Heteronemertea" pageId="0" pageNumber="101" phylum="Nemertea" rank="species" species="termiticola">
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<emphasis italics="true" pageId="0" pageNumber="101">termiticola</emphasis>
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</taxonomicName>
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' referring to the host family,
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<taxonomicName family="Termitidae" lsidName="" pageId="0" pageNumber="101" rank="family">Termitidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="description">
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<paragraph pageId="0" pageNumber="101">Description.</paragraph>
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<paragraph pageId="0" pageNumber="101">
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Stroma solitary, simple, filiform, up to 14 cm long, 1 mm wide, yellow-brown, ca. 6 cm emerging above the leaf litter, ca. 8 cm buried in the soil. Asexual state (
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<taxonomicName class="Sordariomycetes" family="Ophiocordycipitaceae" genus="Hymenostilbe" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenostilbe" order="Hypocreales" pageId="0" pageNumber="101" phylum="Ascomycota" rank="genus">Hymenostilbe</taxonomicName>
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-like) produced ca. 1 cm at the terminal part of the stroma, grey.
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<emphasis italics="true" pageId="0" pageNumber="101">Perithecia</emphasis>
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pseudo-immersed, globose to subglobose, produced on one-third of the terminal part of the stroma ending near the apex, (200
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<normalizedToken originalValue="–)225–261(–">-)225-261(-</normalizedToken>
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280)
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<normalizedToken originalValue="×">x</normalizedToken>
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(150
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<normalizedToken originalValue="–)178–229(–">-)178-229(-</normalizedToken>
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250)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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<emphasis italics="true" pageId="0" pageNumber="101">Asci</emphasis>
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8-spored, filiform, (62.5
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<normalizedToken originalValue="–)76.5–100.5(–">-)76.5-100.5(-</normalizedToken>
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110)
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<normalizedToken originalValue="×">x</normalizedToken>
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(4
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<normalizedToken originalValue="–)4.5–5.5(–">-)4.5-5.5(-</normalizedToken>
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6)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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<emphasis italics="true" pageId="0" pageNumber="101">Ascospores</emphasis>
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whole, filiform, 85
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<normalizedToken originalValue="×">x</normalizedToken>
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2
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, Asexual state
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<taxonomicName class="Sordariomycetes" family="Ophiocordycipitaceae" genus="Hymenostilbe" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hymenostilbe" order="Hypocreales" pageId="0" pageNumber="101" phylum="Ascomycota" rank="genus">Hymenostilbe</taxonomicName>
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-like, conidiogenous cells formed a compact hymenium-like layer and had from two to four denticles at their apices, cylindrical to clavate, (10
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<normalizedToken originalValue="–)11.5–16(–">-)11.5-16(-</normalizedToken>
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17)
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<normalizedToken originalValue="×">x</normalizedToken>
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3
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<normalizedToken originalValue="–5(–">-5(-</normalizedToken>
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6)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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. Conidia, hyaline, fusiform 7
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<normalizedToken originalValue="×">x</normalizedToken>
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3
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="description">
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<paragraph pageId="0" pageNumber="101">Culture characteristics.</paragraph>
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<paragraph pageId="0" pageNumber="101">
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Colonies on PDA, attaining a diam. of 28 mm within 20 d at 25 °C, grey (oac781) to pale grey (oac851).
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<emphasis italics="true" pageId="0" pageNumber="101">Conidiogenous cells</emphasis>
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monophialidic to polyphialidic, arising from hyphae laterally, with an inflated base (7
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<normalizedToken originalValue="–)7.5–10(–">-)7.5-10(-</normalizedToken>
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11)
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<normalizedToken originalValue="×">x</normalizedToken>
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(2.5-) 3
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<normalizedToken originalValue="–3.5(–">-3.5(-</normalizedToken>
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4)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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<emphasis italics="true" pageId="0" pageNumber="101">Conidia</emphasis>
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hyaline, globose, 2.5-3 (-3.5)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, one-celled with warty surface.
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</paragraph>
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<paragraph pageId="0" pageNumber="101">
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Colonies on PSA, attaining a diam. of 22 mm within 20 d at 25 °C, white to pale grey, cotton-like.
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<emphasis italics="true" pageId="0" pageNumber="101">Conidiogenous cells</emphasis>
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monophialidic to polyphialidic, hyaline, smooth, with an inflated base (7
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<normalizedToken originalValue="–)8–10.5(–">-)8-10.5(-</normalizedToken>
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13)
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<normalizedToken originalValue="×">x</normalizedToken>
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3-4 (-5)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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<emphasis italics="true" pageId="0" pageNumber="101">Conidia</emphasis>
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hyaline, globose, (2
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<normalizedToken originalValue="–)2.7–3.4(–">-)2.7-3.4(-</normalizedToken>
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4)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, one celled with warty surface.
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</paragraph>
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<paragraph pageId="0" pageNumber="101">
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Colonies on SDYA/4, attaining a diam. of 29 mm within 20 d at 25 °C, grey to pale grey (oac851).
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<emphasis italics="true" pageId="0" pageNumber="101">Conidiogenous cells</emphasis>
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monophialidic to polyphialidic, hyaline, smooth, with an inflated base (7
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<normalizedToken originalValue="–)8–10.5(–">-)8-10.5(-</normalizedToken>
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13)
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<normalizedToken originalValue="×">x</normalizedToken>
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3-4
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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<emphasis italics="true" pageId="0" pageNumber="101">Conidia</emphasis>
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hyaline, globose, 3
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<normalizedToken originalValue="–3.5(–">-3.5(-</normalizedToken>
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4)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, one-celled with warty surface.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="distribution">
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<paragraph pageId="0" pageNumber="101">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="101">Thailand.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="ecology">
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<paragraph pageId="0" pageNumber="101">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="101">
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Parasitic on a pair of termites from a reproductive caste (Order
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<taxonomicName lsidName="Acari" order="Isoptera" pageId="0" pageNumber="101" rank="order">Isoptera</taxonomicName>
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: Family
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<taxonomicName family="Termitidae" lsidName="" pageId="0" pageNumber="101" rank="family">Termitidae</taxonomicName>
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, Subfamily
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<taxonomicName lsidName="" pageId="0" pageNumber="101" rank="subfamily" subfamily="Macrotermitinae">Macrotermitinae</taxonomicName>
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) and these specimens were buried in the soil. The fungus emerged from the segment between the prothorax and mesothorax of one of the termite pairs.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="materials_examined">
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<paragraph pageId="0" pageNumber="101">Additional specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="101">THAILAND. Chanthaburi Province, Khao Soi Dao Wildlife Reserve; 13°136'N, 102°218'E; on termite; 20 June 1996; R. Nasit, S. Sivichai, K. Tasanathai (BBH5179, BCC1770).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="notes">
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<paragraph pageId="0" pageNumber="101">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="101">
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Both
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<taxonomicName lsidName="O. termiticola" pageId="0" pageNumber="101" rank="species" species="termiticola">
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<emphasis italics="true" pageId="0" pageNumber="101">O. termiticola</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="O. khokpasiensis" pageId="0" pageNumber="101" rank="species" species="khokpasiensis">
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<emphasis italics="true" pageId="0" pageNumber="101">O. khokpasiensis</emphasis>
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</taxonomicName>
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produce pseudo-immersed reddish perithecia on a stroma. In
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<taxonomicName lsidName="O. termiticola" pageId="0" pageNumber="101" rank="species" species="termiticola">
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<emphasis italics="true" pageId="0" pageNumber="101">O. termiticola</emphasis>
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</taxonomicName>
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, the perithecia are tightly packed, while in
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<taxonomicName lsidName="O. khokpasiensis" pageId="0" pageNumber="101" rank="species" species="khokpasiensis">
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<emphasis italics="true" pageId="0" pageNumber="101">O. khokpasiensis</emphasis>
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</taxonomicName>
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, they are loosely aggregated and the length of the anamorphic layer at the end of the fertile part is longer in the latter.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |