treatments-xml/data/11/7F/22/117F220BFF8B751111D9114995E4FE23.xml

<document id="DC2D39A938E50D100C558078D5D88CF1" ID-DOI="10.11646/zootaxa.3964.4.5" ID-GBIF-Dataset="c86698a4-d6e3-454e-b175-3ab9beb9a5e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="238043" ID-ZooBank="8A90D889-4C33-401A-A1A8-AB31663402BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461077564734" checkinUser="plazi" docAuthor="Huynh, Cuong &amp; Veenstra, Anneke A." docDate="2015" docId="117F220BFF8B751111D9114995E4FE23" docLanguage="en" docName="zt03964p474.pdf" docOrigin="Zootaxa 3964 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monographis dongnaiensis Huynh &amp; Veenstra, 2015, sp. n." docType="treatment" docVersion="8" lastPageNumber="470" masterDocId="ED465A73FF88751A114E15599738FFF3" masterDocTitle="Description of a new species of Penicillate Millipede from the genus Monographis (Diplopoda: Polyxenidae) found in Vietnam" masterLastPageNumber="474" masterPageNumber="460" pageNumber="463" updateTime="1698586781965" updateUser="plazi">
<mods:mods id="0C1128DAC2740DFFC8C4AE96059BED42" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="282A1E19D86EA65519C9FBCAD86AF7B5">
<mods:title id="DAFA3AFFBF1E42D9C819CC866B55EBD1">Description of a new species of Penicillate Millipede from the genus Monographis (Diplopoda: Polyxenidae) found in Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="EDDE674D559CD1B8C8986B2AC167BAD3" type="personal">
<mods:role id="6FA87163A28FF8FAF8ED7DB5DD3A5D3C">
<mods:roleTerm id="8595281829FD50C7F6E5413D02591D35">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="414FC450DFEAE3AC2DFBA5B133B5C938">Huynh, Cuong</mods:namePart>
</mods:name>
<mods:name id="C1F2A30288B78B59957272A995DB1A6F" type="personal">
<mods:role id="F9230270BD815C8BA0F53448D4C0BABD">
<mods:roleTerm id="A8C42FDB97F93905949ACD640E303E63">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="90CE54F03E6ECAE4C4B9328F0B4F15AD">Veenstra, Anneke A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5054DEAB2601A0A24904FFA5C4B6F32A">text</mods:typeOfResource>
<mods:relatedItem id="2FD04982E641DD9651BC500046096341" type="host">
<mods:titleInfo id="E61FF905E6ADF6CD9D057F79072B5A42">
<mods:title id="6CDEA34603C3407981A640C8BA554E34">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="240EC2C286B0A46CA3BBC4005FC409FD">
<mods:date id="D82C73926FC8B643AF287E2C28BEC7B7">2015</mods:date>
<mods:detail id="B91D1B1D214A7E3F62D68149551D4836" type="volume">
<mods:number id="F3AE3BA1ED1FBFFBCEFF3A21BFAC8E27">3964</mods:number>
</mods:detail>
<mods:detail id="6A73DF35E5A1366CE81BD7D23EF89B47" type="issue">
<mods:number id="FE4F182B968B3AE7773196F7596C0E8A">4</mods:number>
</mods:detail>
<mods:extent id="F1F52EF2F8A59113FF99A8B50CD1CA4B" unit="page">
<mods:start id="371B37D707D49E04697A743A928FD9CB">460</mods:start>
<mods:end id="DF47F54CE08E24E81F8C7F691B47A59B">474</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7D581D962EF2BFFDD5EC61A9759F26BD">journal article</mods:classification>
<mods:identifier id="004454FB06EEB0C95F01C237755B3092" type="DOI">10.11646/zootaxa.3964.4.5</mods:identifier>
<mods:identifier id="729D6FE6F2434EACCACB8957B92319E3" type="GBIF-Dataset">c86698a4-d6e3-454e-b175-3ab9beb9a5e1</mods:identifier>
<mods:identifier id="15E9DA871E4D4F648C1F1A491E7F0598" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="83D355F218D861158692A1EECCFCCBF7" type="Zenodo-Dep">238043</mods:identifier>
<mods:identifier id="DAB794BC786C11D7B7304462427A0D6C" type="ZooBank">8A90D889-4C33-401A-A1A8-AB31663402BC</mods:identifier>
</mods:mods>
<treatment id="117F220BFF8B751111D9114995E4FE23" ID-DOI="http://doi.org/10.5281/zenodo.6108090" ID-GBIF-Taxon="119617995" ID-Zenodo-Dep="6108090" LSID="urn:lsid:plazi:treatment:117F220BFF8B751111D9114995E4FE23" httpUri="http://treatment.plazi.org/id/117F220BFF8B751111D9114995E4FE23" lastPageId="11" lastPageNumber="470" pageId="3" pageNumber="463">
<subSubSection id="D1CCC096FF8B751911D911499630FBB8" pageId="3" pageNumber="463" type="nomenclature">
<paragraph id="9969931DFF8B751911D911499512FBD9" blockId="3.[151,554,1040,1100]" box="[151,554,1040,1066]" pageId="3" pageNumber="463">
<heading id="C2212471FF8B751911D911499512FBD9" bold="true" box="[151,554,1040,1066]" fontSize="11" level="1" pageId="3" pageNumber="463" reason="1">
<emphasis id="ABA24F0FFF8B751911D911499512FBD9" bold="true" box="[151,554,1040,1066]" pageId="3" pageNumber="463">
<taxonomicName id="5ED6E89EFF8B751911D9114996D8FBD9" box="[151,480,1040,1066]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="3" pageNumber="463" phylum="Arthropoda" rank="species" species="dongnaiensis" status="sp. nov.">
<emphasis id="ABA24F0FFF8B751911D9114996D8FBD9" bold="true" box="[151,480,1040,1066]" italics="true" pageId="3" pageNumber="463">Monographis dongnaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B091F274FF8B751910A611499512FBD9" box="[488,554,1040,1066]" pageId="3" pageNumber="463" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9969931DFF8B751911D9116A9630FBB8" blockId="3.[151,554,1040,1100]" box="[151,264,1075,1100]" pageId="3" pageNumber="463">
<emphasis id="ABA24F0FFF8B751911D9116A9798FBBF" bold="true" box="[151,160,1075,1100]" pageId="3" pageNumber="463">(</emphasis>
<figureCitation id="01ED8F98FF8B751911EE116A97C7FBB8" box="[160,255,1075,1100]" captionStart="FIGURE 2. A" captionStartId="4.[151,250,1111,1133]" captionTargetBox="[272,1315,193,1089]" captionTargetId="figure@4.[272,1315,193,1090]" captionTargetPageId="4" captionText="FIGURE 2. A live specimen of Monographis dongnaiensis from Dong Nai province, southern Viet Nam." httpUri="https://zenodo.org/record/238045/files/figure.png" pageId="3" pageNumber="463">Figure 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D1CCC096FF8B751911D9112296A0FA62" pageId="3" pageNumber="463" type="materials_examined">
<paragraph id="9969931DFF8B751911D911229369FB2E" blockId="3.[151,1437,1147,2004]" pageId="3" pageNumber="463">
<emphasis id="ABA24F0FFF8B751911D911229633FB67" bold="true" box="[151,267,1147,1172]" pageId="3" pageNumber="463">
<typeStatus id="466D2DBFFF8B751911D91122963EFB67" box="[151,262,1147,1172]" pageId="3" pageNumber="463" type="holotype">Holotype</typeStatus>
.
</emphasis>
An adult male, stadium VIII with 13 pairs of legs, was collected by the author, Cuong Huynh, from Trang Bom district, (10°54′59″N, 106°59′31″E northeast Ho Chi Minh City); in Dong Nai province, southern 
<collectingCountry id="E1C1D38DFF8B751911D9119D97C1FB2E" box="[151,249,1220,1245]" name="Vietnam" pageId="3" pageNumber="463">Vietnam</collectingCountry>
, 
<date id="ED68B5DDFF8B7519104A119D96CBFB2E" box="[260,499,1220,1245]" pageId="3" pageNumber="463" value="2012-12-20">20th December 2012</date>
. Queensland Museum accession number QMS 98960.
</paragraph>
<paragraph id="9969931DFF8B7519118911B196A0FA62" blockId="3.[151,1437,1147,2004]" pageId="3" pageNumber="463">
<emphasis id="ABA24F0FFF8B7519118911B1967EFAF2" bold="true" box="[199,326,1256,1281]" pageId="3" pageNumber="463">
<typeStatus id="466D2DBFFF8B7519118911B19679FAF2" box="[199,321,1256,1281]" pageId="3" pageNumber="463" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Two adult males (QMS 98961, 98962), seven adult females (QMS 98963–98969) and sub-adult female (stadium 
<date id="ED68B5DDFF8B7519101510549681FAD6" box="[347,441,1293,1317]" pageId="3" pageNumber="463" value="2012-07">VII—12</date>
pairs of legs) (QMS 98970) are from the 
<typeStatus id="466D2DBFFF8B751912E1105494E7FAD6" box="[943,991,1293,1317]" pageId="3" pageNumber="463">type</typeStatus>
locality (location 1). An adult female (QMS 98971) and a sub-adult male (stadium 
<date id="ED68B5DDFF8B751913C1106895D6FABA" box="[655,750,1329,1353]" pageId="3" pageNumber="463" value="2012-07">VII—12</date>
pairs of legs) (QMS 98972) are from location 2. All mounted slides and specimens preserved in 80% ethanol, have been lodged in the Queensland Museum, Brisbane, Queensland, 
<collectingCountry id="E1C1D38DFF8B75191067102196ABFA62" box="[297,403,1400,1425]" name="Australia" pageId="3" pageNumber="463">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D1CCC096FF8B7519118910C59429FA2A" pageId="3" pageNumber="463" type="etymology">
<paragraph id="9969931DFF8B7519118910C59429FA2A" blockId="3.[151,1437,1147,2004]" pageId="3" pageNumber="463">
<emphasis id="ABA24F0FFF8B7519118910C59676FA46" bold="true" box="[199,334,1436,1461]" pageId="3" pageNumber="463">Etymology.</emphasis>
The specific name of 
<taxonomicName id="5ED6E89EFF8B7519130410C59444FA46" box="[586,892,1436,1461]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="3" pageNumber="463" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF8B7519130410C59444FA46" box="[586,892,1436,1461]" italics="true" pageId="3" pageNumber="463">Monographis dongnaiensis</emphasis>
</taxonomicName>
refers to the name of the province of Dong Nai, southern 
<collectingCountry id="E1C1D38DFF8B751911B010999658FA2A" box="[254,352,1472,1497]" name="Vietnam" pageId="3" pageNumber="463">Vietnam</collectingCountry>
, where this species was first collected.
</paragraph>
</subSubSection>
<subSubSection id="D1CCC096FF8B7519118910BD9531F963" pageId="3" pageNumber="463" type="diagnosis">
<paragraph id="9969931DFF8B7519118910BD9531F963" blockId="3.[151,1437,1147,2004]" pageId="3" pageNumber="463">
<emphasis id="ABA24F0FFF8B7519118910BD9679FA0E" bold="true" box="[199,321,1508,1533]" pageId="3" pageNumber="463">Diagnosis.</emphasis>
<taxonomicName id="5ED6E89EFF8B7519100210BD95BAFA0E" box="[332,642,1508,1533]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="3" pageNumber="463" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF8B7519100210BD95BAFA0E" box="[332,642,1508,1533]" italics="true" pageId="3" pageNumber="463">Monographis dongnaiensis</emphasis>
</taxonomicName>
differs from other species of genus 
<taxonomicName id="5ED6E89EFF8B7519157A10BD93F3FA0E" box="[1076,1227,1508,1533]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="3" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF8B7519157A10BD93F3FA0E" box="[1076,1227,1508,1533]" italics="true" pageId="3" pageNumber="463">Monographis</emphasis>
</taxonomicName>
by having 19–22 sensilla, with 3 anterior thick sensilla and 5 internal sensilla, which form a triangular shape on the 6th antennal article. The surface of the labrum is smooth and asetose. Telotarsus bears a posterior process with a curved and enlarged base and is half the length of the claw. A small lateral process is present and the anterior setiform process is equal to the length of the claw.
</paragraph>
</subSubSection>
<subSubSection id="D1CCC096FF8B7513118913C294EEFBEF" lastPageId="9" lastPageNumber="469" pageId="3" pageNumber="463" type="description">
<paragraph id="9969931DFF8B7519118913C2941FF908" blockId="3.[151,1437,1147,2004]" pageId="3" pageNumber="463">
<emphasis id="ABA24F0FFF8B7519118913C29661F947" bold="true" box="[199,345,1691,1716]" pageId="3" pageNumber="463">Description.</emphasis>
<emphasis id="ABA24F0FFF8B7519102F13C5953BF940" box="[353,515,1692,1715]" italics="true" pageId="3" pageNumber="463">Measurements</emphasis>
: 
<typeStatus id="466D2DBFFF8B7519135D13C29543F947" box="[531,635,1691,1716]" pageId="3" pageNumber="463" type="holotype">Holotype</typeStatus>
male body length was 
<quantity id="5E2E3EF8FF8B751912C813C294D8F940" box="[902,992,1691,1715]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="463" unit="mm" value="3.2">3.2 mm</quantity>
, male (
<typeStatus id="466D2DBFFF8B7519157D13C2939AF940" box="[1075,1186,1691,1715]" pageId="3" pageNumber="463" type="paratype">paratypes</typeStatus>
) range in length from 
<quantity id="5E2E3EF8FF8B751911D913E6961DF924" box="[151,293,1727,1751]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.2" metricValueMin="2.4" pageId="3" pageNumber="463" unit="mm" value="2.8" valueMax="3.2" valueMin="2.4">2.4–3.2 mm</quantity>
(
<emphasis id="ABA24F0FFF8B7519107313999673F924" box="[317,331,1728,1751]" italics="true" pageId="3" pageNumber="463">n</emphasis>
= 10) and females (
<emphasis id="ABA24F0FFF8B7519131C13999558F924" box="[594,608,1728,1751]" italics="true" pageId="3" pageNumber="463">n</emphasis>
= 10) range from 3.0– 
<quantity id="5E2E3EF8FF8B751912CA13E694DCF924" box="[900,996,1727,1751]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="463" unit="mm" value="3.4">3.4 mm</quantity>
. The caudal bundles of females are approximately 
<quantity id="5E2E3EF8FF8B7519107113BA96A0F908" box="[319,408,1763,1787]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="463" unit="mm" value="0.9">0.9 mm</quantity>
, longer than in the males (
<quantity id="5E2E3EF8FF8B751913F113BA9423F908" box="[703,795,1763,1787]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="463" unit="mm" value="0.7">0.7 mm</quantity>
).
</paragraph>
<paragraph id="9969931DFF8B75191189125E9342F827" blockId="3.[151,1437,1147,2004]" pageId="3" pageNumber="463">
<emphasis id="ABA24F0FFF8B75191189125E9668F8D3" box="[199,336,1799,1824]" italics="true" pageId="3" pageNumber="463">Colouration</emphasis>
: The body is yellowish brown in colour and darker on the latero-posterior rosette trichomes, which form 2 dark lateral bands contrasting with a light yellow medial band. Eyes are bright red with two black horizontal bands apparent in the vertex area, which distinguishes the head from the rest of the animal. The last tergite is darkest in colour and extended into the telson region. Here, the caudal bundle trichomes form light silvery bands near the base. The colour of these bands gradually fades to white and reflects light to produce a black-silver colouration towards the end of the telson. The ventral side of the body is white in colour.
</paragraph>
<caption id="CDA9C395FF8C751E11D9110E93FEFB9E" box="[151,1222,1111,1133]" httpUri="https://zenodo.org/record/238045/files/figure.png" pageId="4" pageNumber="464" targetBox="[272,1315,193,1089]" targetPageId="4">
<paragraph id="9969931DFF8C751E11D9110E93FEFB9E" blockId="4.[151,1222,1111,1133]" box="[151,1222,1111,1133]" pageId="4" pageNumber="464">
<emphasis id="ABA24F0FFF8C751E11D9110E9635FB9E" bold="true" box="[151,269,1111,1133]" pageId="4" pageNumber="464">FIGURE 2</emphasis>
. A live specimen of 
<taxonomicName id="5ED6E89EFF8C751E1091110E95CCFB9E" box="[479,756,1111,1133]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="464" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF8C751E1091110E95CCFB9E" box="[479,756,1111,1133]" italics="true" pageId="4" pageNumber="464">Monographis dongnaiensis</emphasis>
</taxonomicName>
from Dong Nai province, southern Viet Nam.
</paragraph>
</caption>
<paragraph id="9969931DFF8C751E118911CD9781F9E6" blockId="4.[151,1437,1172,2037]" pageId="4" pageNumber="464">
<emphasis id="ABA24F0FFF8C751E118911CD963CFB5E" box="[199,260,1172,1197]" italics="true" pageId="4" pageNumber="464">Head</emphasis>
: There are 8 ommatidia on each side: 4 located dorsally and 
<quantity id="5E2E3EF8FF8C751E128C11CC94D7FB5E" box="[962,1007,1172,1197]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="464" unit="in" value="4.0">4 in</quantity>
lateral positions (1 anterior, 2 medial and 1 posterior). The vertex has 2 posterior trichome groups, a large gap is present between the 2 groups and each group is with 2 rows. The anterior, oblique row has a variable number of large trichome sockets in the centre. These trichome sockets reduce in size toward both ends. The posterior rows have fewer small trichome sockets. There is a narrow space between the anterior and posterior rows. The 
<typeStatus id="466D2DBFFF8C751E121A107D948EFACE" box="[852,950,1316,1341]" pageId="4" pageNumber="464" type="holotype">holotype</typeStatus>
male had 13 + 13 anterior row trichome sockets and 3 + 3 posterior row trichome sockets. The 
<typeStatus id="466D2DBFFF8C751E124A10109457FA93" box="[772,879,1353,1376]" pageId="4" pageNumber="464" type="paratype">paratypes</typeStatus>
showed that variation is common in this species, regardless of sex (QMS 98962 with 11(L: left) + 12 (R: right) and 4 (L) + 3(R), QMS 98964 12 (L) + 14 (R) and 3 (L) + 7 (R)). Trichobothria (trichobothrium 
<emphasis id="ABA24F0FFF8C751E13E910C9958DFA5A" bold="true" box="[679,693,1424,1449]" pageId="4" pageNumber="464">a</emphasis>
, internal position, trichobothrium 
<emphasis id="ABA24F0FFF8C751E150110C99366FA5A" bold="true" box="[1103,1118,1424,1449]" pageId="4" pageNumber="464">b</emphasis>
, intermediate position and trichobothrium 
<emphasis id="ABA24F0FFF8C751E100910ED966BFA3E" bold="true" box="[327,339,1460,1485]" pageId="4" pageNumber="464">c</emphasis>
, external position) are typically thin, funicular and cylindrical structures. All these trichobothria (a, b and c) are equal in size and form an isosceles triangle with equal distance between ab and bc (
<figureCitation id="01ED8F98FF8C751E15B110819250FA02" box="[1279,1384,1496,1521]" captionStart="FIGURE 3" captionStartId="5.[151,250,1331,1353]" captionTargetBox="[155,1429,381,1299]" captionTargetId="figure@5.[151,1435,373,1309]" captionTargetPageId="5" captionText="FIGURE 3. Holotype male (QMS 98960), A. Head capsule of Monographis dongnaiensis showing ommatidia pattern, 2 posterior vertex trichomes and 3 trichobothria: trichobothrium a, b and c. B. Collum and lateral protuberances with pattern of trichome sockets (all trichomes were removed). C. Trichome socket arrangements from tergite 2. D. Tergite 10 with densely packed smaller sockets. (A, B, C, D drawn to the same scale). E. Structure of trichobothria: Trichobothrium a (internal position), Trichobothrium b (intermediate position), Trichobothrium c (external position), sockets and their funicular structures. F. Structure of clypeo-labrum with 2 lateral lamellae, 2 lamellae and a median cleft. G. Structures of 2 lateral palps (a conical sensillum was missed on each lateral palp) and medial palps from gnathochilaria (Lp: lateral palp, Mp: medial palp)." httpUri="https://zenodo.org/record/238046/files/figure.png" pageId="4" pageNumber="464">Figures 3</figureCitation>
A &amp; E).
</paragraph>
<paragraph id="9969931DFF8C751E118913789475F85F" blockId="4.[151,1437,1172,2037]" pageId="4" pageNumber="464">
<emphasis id="ABA24F0FFF8C751E118913789609F9CB" box="[199,305,1569,1592]" italics="true" pageId="4" pageNumber="464">Antennae</emphasis>
: These typically consist of 8 articles and four sensory cones, which is characteristic of 
<taxonomicName id="5ED6E89EFF8C751E1444137992A0F9CA" box="[1290,1432,1568,1593]" class="Diplopoda" family="Polyxenidae" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="464" phylum="Arthropoda" rank="family">Polyxenidae</taxonomicName>
. The 6th antennal article (Figures 4C &amp; D) has 20 bacilliform sensilla in males and 
<quantity id="5E2E3EF8FF8C751E1578131193A2F993" box="[1078,1178,1607,1632]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.588" metricValueMin="4.826" pageId="4" pageNumber="464" unit="in" value="20.5" valueMax="22.0" valueMin="19.0">19–22 in</quantity>
females. The 
<typeStatus id="466D2DBFFF8C751E1474131E92A4F993" box="[1338,1436,1607,1632]" pageId="4" pageNumber="464" type="holotype">holotype</typeStatus>
has 20 sensilla, including 3 thick bacilliform sensilla and 17 thin bacilliform sensilla. Two adult female 
<typeStatus id="466D2DBFFF8C751E1464133592A0F977" box="[1322,1432,1644,1668]" pageId="4" pageNumber="464" type="paratype">paratypes</typeStatus>
, QMS 98969 from Giang Dien, QMS 98971 from Dinh Quan, both have a greater number of sensilla (22) on the 6th antennal article. The typical 3 thick bacilliform sensilla: anterior 
<emphasis id="ABA24F0FFF8C751E123213EF94B2F93C" bold="true" box="[892,906,1718,1743]" pageId="4" pageNumber="464">a</emphasis>
(Tbs 
<emphasis id="ABA24F0FFF8C751E128013EF94E4F93C" bold="true" box="[974,988,1718,1743]" pageId="4" pageNumber="464">a</emphasis>
), intermediate 
<emphasis id="ABA24F0FFF8C751E15C413EF93AAF93C" bold="true" box="[1162,1170,1718,1743]" pageId="4" pageNumber="464">i</emphasis>
(Tbs 
<emphasis id="ABA24F0FFF8C751E159813EF93DFF93C" bold="true" box="[1238,1255,1718,1743]" pageId="4" pageNumber="464">i)</emphasis>
and posterior 
<emphasis id="ABA24F0FFF8C751E14C213EF92A3F93C" bold="true" box="[1420,1435,1718,1743]" pageId="4" pageNumber="464">p</emphasis>
(Tbs 
<emphasis id="ABA24F0FFF8C751E119D138397DAF900" bold="true" box="[211,226,1754,1779]" pageId="4" pageNumber="464">p</emphasis>
) are located in the anterior portion. These thick bacilliform sensilla combine with 11–14 thin bacilliform sensilla (tbs) forming a triangular shape, surrounding an inner group of 5 thin bacilliform sensilla (Figure 4B). A conical sensillum is absent. The 7th antennal article has 2 thick bacilliform sensilla (Tbs), the anterior one being slightly longer than the one located posteriorly, with one setiform sensillum (s) between them and a conical sensillum (c) locates in the posterior position (Figure 4A). This pattern of sensilla on the 7th article is commonly seen in all species of 
<taxonomicName id="5ED6E89EFF8C751E10CB12CA9470F85F" authority="Huynh &amp; Veenstra 2013" authorityName="Huynh &amp; Veenstra" authorityYear="2013" box="[389,840,1939,1964]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF8C751E10CB12CA9524F85F" box="[389,540,1939,1964]" italics="true" pageId="4" pageNumber="464">Monographis</emphasis>
(
<bibRefCitation id="FD47EEECFF8C751E136412CA9479F85F" author="Huynh" box="[554,833,1939,1964]" pageId="4" pageNumber="464" refString="Huynh, C. &amp; Veenstra, A. (2013) Taxonomy and biology of a new species of pincushion Millipede of the genus Monographis (Diplopoda: Polyxenidae) from Australia. Zootaxa, 3721 (6), 573 - 588. http: // dx. doi. org / 10.11646 / zootaxa. 3721.6.4" type="journal article" year="2013">Huynh &amp; Veenstra 2013</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="9969931DFF8C751F118912EE94D7FF27" blockId="4.[151,1437,1172,2037]" lastBlockId="5.[151,1437,151,320]" lastPageId="5" lastPageNumber="465" pageId="4" pageNumber="464">
<emphasis id="ABA24F0FFF8C751E118912EE9649F823" box="[199,369,1975,2000]" italics="true" pageId="4" pageNumber="464">Clypeo-labrum</emphasis>
: 9 setae as long as the width of the labrum are present. Setae on the 
<typeStatus id="466D2DBFFF8C751E153E12E193E3F823" box="[1136,1243,1976,2000]" pageId="4" pageNumber="464" type="paratype">paratypes</typeStatus>
range from 
<quantity id="5E2E3EF8FF8C751E142B12E19795F806" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="4" pageNumber="464" unit="in" value="9.5" valueMax="11.0" valueMin="8.0">8–11 in</quantity>
both sexes. The labrum surface on the 
<typeStatus id="466D2DBFFF8C751E132C128595FCF806" box="[610,708,2012,2037]" pageId="4" pageNumber="464" type="holotype">holotype</typeStatus>
is smooth, with a line of minute setae along the posterior portion and the anterior margin asetose. Some 
<typeStatus id="466D2DBFFF8D751F132E15C195F3FF43" box="[608,715,152,176]" pageId="5" pageNumber="465" type="paratype">paratypes</typeStatus>
have a few setae on the labrum surface. A 1 + 1 lamellae is associated with lateral lamellae on each side of the median cleft (
<figureCitation id="01ED8F98FF8D751F122115E594F7FF27" box="[879,975,188,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1331,1353]" captionTargetBox="[155,1429,381,1299]" captionTargetId="figure@5.[151,1435,373,1309]" captionTargetPageId="5" captionText="FIGURE 3. Holotype male (QMS 98960), A. Head capsule of Monographis dongnaiensis showing ommatidia pattern, 2 posterior vertex trichomes and 3 trichobothria: trichobothrium a, b and c. B. Collum and lateral protuberances with pattern of trichome sockets (all trichomes were removed). C. Trichome socket arrangements from tergite 2. D. Tergite 10 with densely packed smaller sockets. (A, B, C, D drawn to the same scale). E. Structure of trichobothria: Trichobothrium a (internal position), Trichobothrium b (intermediate position), Trichobothrium c (external position), sockets and their funicular structures. F. Structure of clypeo-labrum with 2 lateral lamellae, 2 lamellae and a median cleft. G. Structures of 2 lateral palps (a conical sensillum was missed on each lateral palp) and medial palps from gnathochilaria (Lp: lateral palp, Mp: medial palp)." httpUri="https://zenodo.org/record/238046/files/figure.png" pageId="5" pageNumber="465">Figure 3</figureCitation>
F).
</paragraph>
<paragraph id="9969931DFF8D751F1189158697F1FEB3" blockId="5.[151,1437,151,320]" pageId="5" pageNumber="465">
<emphasis id="ABA24F0FFF8D751F118915869509FF0B" box="[199,561,223,248]" italics="true" pageId="5" pageNumber="465">Lateral palp of gnathochilarium</emphasis>
: This is 1.25 times the length of the medial palp. There are 11 conical sensilla on the lateral palp and medial palp with 22 slender conical sensilla, found in all 
<typeStatus id="466D2DBFFF8D751F155F145C9344FEEE" box="[1041,1148,261,285]" pageId="5" pageNumber="465" type="paratype">paratypes</typeStatus>
and the 
<typeStatus id="466D2DBFFF8D751F1594145D9278FEEE" box="[1242,1344,260,285]" pageId="5" pageNumber="465" type="holotype">holotype</typeStatus>
. (
<figureCitation id="01ED8F98FF8D751F141F145D979DFEB3" captionStart="FIGURE 3" captionStartId="5.[151,250,1331,1353]" captionTargetBox="[155,1429,381,1299]" captionTargetId="figure@5.[151,1435,373,1309]" captionTargetPageId="5" captionText="FIGURE 3. Holotype male (QMS 98960), A. Head capsule of Monographis dongnaiensis showing ommatidia pattern, 2 posterior vertex trichomes and 3 trichobothria: trichobothrium a, b and c. B. Collum and lateral protuberances with pattern of trichome sockets (all trichomes were removed). C. Trichome socket arrangements from tergite 2. D. Tergite 10 with densely packed smaller sockets. (A, B, C, D drawn to the same scale). E. Structure of trichobothria: Trichobothrium a (internal position), Trichobothrium b (intermediate position), Trichobothrium c (external position), sockets and their funicular structures. F. Structure of clypeo-labrum with 2 lateral lamellae, 2 lamellae and a median cleft. G. Structures of 2 lateral palps (a conical sensillum was missed on each lateral palp) and medial palps from gnathochilaria (Lp: lateral palp, Mp: medial palp)." httpUri="https://zenodo.org/record/238046/files/figure.png" pageId="5" pageNumber="465">Figure 3</figureCitation>
G).
</paragraph>
<caption id="CDA9C395FF8D751F11D9106A92AFF9F2" httpUri="https://zenodo.org/record/238046/files/figure.png" pageId="5" pageNumber="465" targetBox="[155,1429,381,1299]" targetPageId="5">
<paragraph id="9969931DFF8D751F11D9106A92AFF9F2" blockId="5.[151,1436,1331,1537]" pageId="5" pageNumber="465">
<emphasis id="ABA24F0FFF8D751F11D9106A9628FABA" bold="true" box="[151,272,1331,1353]" pageId="5" pageNumber="465">FIGURE 3</emphasis>
. Holotype male (QMS 98960), 
<emphasis id="ABA24F0FFF8D751F1329106A9541FABA" bold="true" box="[615,633,1331,1353]" pageId="5" pageNumber="465">A</emphasis>
. Head capsule of 
<taxonomicName id="5ED6E89EFF8D751F1271106A9360FABA" box="[831,1112,1331,1353]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="5" pageNumber="465" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF8D751F1271106A9360FABA" box="[831,1112,1331,1353]" italics="true" pageId="5" pageNumber="465">Monographis dongnaiensis</emphasis>
</taxonomicName>
showing ommatidia pattern, 2 posterior vertex trichomes and 3 trichobothria: trichobothrium 
<emphasis id="ABA24F0FFF8D751F12571008941EFA94" bold="true" box="[793,806,1361,1383]" pageId="5" pageNumber="465">a</emphasis>
, 
<emphasis id="ABA24F0FFF8D751F127D10089479FA94" bold="true" box="[819,833,1361,1383]" pageId="5" pageNumber="465">b</emphasis>
and 
<emphasis id="ABA24F0FFF8D751F123A10089447FA94" bold="true" box="[884,895,1361,1383]" pageId="5" pageNumber="465">c</emphasis>
. 
<emphasis id="ABA24F0FFF8D751F12C2100894A5FA94" bold="true" box="[908,925,1361,1383]" pageId="5" pageNumber="465">B</emphasis>
. Collum and lateral protuberances with pattern of trichome sockets (all trichomes were removed). 
<emphasis id="ABA24F0FFF8D751F13C7102995A3FA75" bold="true" box="[649,667,1392,1414]" pageId="5" pageNumber="465">C</emphasis>
. Trichome socket arrangements from tergite 2. 
<emphasis id="ABA24F0FFF8D751F15C7102993A3FA75" bold="true" box="[1161,1179,1392,1414]" pageId="5" pageNumber="465">D</emphasis>
. Tergite 10 with densely packed smaller sockets. (
<emphasis id="ABA24F0FFF8D751F10ED10D69526FA56" bold="true" box="[419,542,1423,1445]" pageId="5" pageNumber="465">A, B, C, D</emphasis>
drawn to the same scale). 
<emphasis id="ABA24F0FFF8D751F120810D6946FFA56" bold="true" box="[838,855,1423,1445]" pageId="5" pageNumber="465">E</emphasis>
. Structure of trichobothria: Trichobothrium 
<emphasis id="ABA24F0FFF8D751F146010D69202FA56" bold="true" box="[1326,1338,1423,1445]" pageId="5" pageNumber="465">a</emphasis>
(internal position), Trichobothrium 
<emphasis id="ABA24F0FFF8D751F10FA10F496FAFA30" bold="true" box="[436,450,1453,1475]" pageId="5" pageNumber="465">b</emphasis>
(intermediate position), Trichobothrium 
<emphasis id="ABA24F0FFF8D751F12CA10F494B7FA30" bold="true" box="[900,911,1453,1475]" pageId="5" pageNumber="465">c</emphasis>
(external position), sockets and their funicular structures. 
<emphasis id="ABA24F0FFF8D751F10481095962BFA11" bold="true" box="[262,275,1484,1506]" pageId="5" pageNumber="465">F</emphasis>
. Structure of clypeo-labrum with 2 lateral lamellae, 2 lamellae and a median cleft. 
<emphasis id="ABA24F0FFF8D751F152B1095934EFA11" bold="true" box="[1125,1142,1484,1506]" pageId="5" pageNumber="465">G</emphasis>
. Structures of 2 lateral palps (a conical sensillum was missed on each lateral palp) and medial palps from gnathochilaria (Lp: lateral palp, Mp: medial palp).
</paragraph>
</caption>
<paragraph id="9969931DFF8D751F11891371954AF85A" blockId="5.[151,1437,1576,2033]" pageId="5" pageNumber="465">
<emphasis id="ABA24F0FFF8D751F118913719630F9B2" box="[199,264,1576,1601]" italics="true" pageId="5" pageNumber="465">Trunk</emphasis>
: This consists of 10 segments, 9 pleural projections, excluding the telson and caudal bundle; 13 pairs of legs. The collum is the only tergite with lateral protuberances bearing 3–6 trichomes on each side. All other tergites have a pair of pleural projections located antero-laterally. The tergal trichome socket arrangements typically have 2 broad oval shapes, slightly enlarged laterally, and a posterior row slightly upwardly curved toward the centre with a large gap between these trichome socket rows. The collum is considered to be tergite 1 (smallest tergite) with 57 (Left: L), 57 (Right: R) trichome sockets and 6(L) / 6(R) in lateral protuberance on 
<typeStatus id="466D2DBFFF8D751F1523138593F7F906" box="[1133,1231,1756,1781]" pageId="5" pageNumber="465" type="holotype">holotype</typeStatus>
(
<figureCitation id="01ED8F98FF8D751F15AE1385927BF906" box="[1248,1347,1756,1781]" captionStart="FIGURE 3" captionStartId="5.[151,250,1331,1353]" captionTargetBox="[155,1429,381,1299]" captionTargetId="figure@5.[151,1435,373,1309]" captionTargetPageId="5" captionText="FIGURE 3. Holotype male (QMS 98960), A. Head capsule of Monographis dongnaiensis showing ommatidia pattern, 2 posterior vertex trichomes and 3 trichobothria: trichobothrium a, b and c. B. Collum and lateral protuberances with pattern of trichome sockets (all trichomes were removed). C. Trichome socket arrangements from tergite 2. D. Tergite 10 with densely packed smaller sockets. (A, B, C, D drawn to the same scale). E. Structure of trichobothria: Trichobothrium a (internal position), Trichobothrium b (intermediate position), Trichobothrium c (external position), sockets and their funicular structures. F. Structure of clypeo-labrum with 2 lateral lamellae, 2 lamellae and a median cleft. G. Structures of 2 lateral palps (a conical sensillum was missed on each lateral palp) and medial palps from gnathochilaria (Lp: lateral palp, Mp: medial palp)." httpUri="https://zenodo.org/record/238046/files/figure.png" pageId="5" pageNumber="465">Figure 3</figureCitation>
B). The numbers varied in 
<typeStatus id="466D2DBFFF8D751F1029125896FFF8EA" box="[359,455,1793,1817]" pageId="5" pageNumber="465" type="paratype">paratype</typeStatus>
males, range 43–56; and 
<typeStatus id="466D2DBFFF8D751F13A81258947EF8EA" box="[742,838,1793,1817]" pageId="5" pageNumber="465" type="paratype">paratype</typeStatus>
females, range 51–66 trichome sockets. Tergite 2 has the same structural pattern with the posterior row slightly longer and with 58(L) / 57(R) trichome sockets (
<figureCitation id="01ED8F98FF8D751F141F127D979DF892" captionStart="FIGURE 3" captionStartId="5.[151,250,1331,1353]" captionTargetBox="[155,1429,381,1299]" captionTargetId="figure@5.[151,1435,373,1309]" captionTargetPageId="5" captionText="FIGURE 3. Holotype male (QMS 98960), A. Head capsule of Monographis dongnaiensis showing ommatidia pattern, 2 posterior vertex trichomes and 3 trichobothria: trichobothrium a, b and c. B. Collum and lateral protuberances with pattern of trichome sockets (all trichomes were removed). C. Trichome socket arrangements from tergite 2. D. Tergite 10 with densely packed smaller sockets. (A, B, C, D drawn to the same scale). E. Structure of trichobothria: Trichobothrium a (internal position), Trichobothrium b (intermediate position), Trichobothrium c (external position), sockets and their funicular structures. F. Structure of clypeo-labrum with 2 lateral lamellae, 2 lamellae and a median cleft. G. Structures of 2 lateral palps (a conical sensillum was missed on each lateral palp) and medial palps from gnathochilaria (Lp: lateral palp, Mp: medial palp)." httpUri="https://zenodo.org/record/238046/files/figure.png" pageId="5" pageNumber="465">Figure 3</figureCitation>
C). For Tergites 3–9, the patterns are very similar with the characteristic large gaps. Tergite 10 is an exception where the trichome sockets are smaller and denser. There is almost no space between the lateral rosette trichome sockets and the posterior row (
<figureCitation id="01ED8F98FF8D751F10A312C99575F85A" box="[493,589,1936,1961]" captionStart="FIGURE 3" captionStartId="5.[151,250,1331,1353]" captionTargetBox="[155,1429,381,1299]" captionTargetId="figure@5.[151,1435,373,1309]" captionTargetPageId="5" captionText="FIGURE 3. Holotype male (QMS 98960), A. Head capsule of Monographis dongnaiensis showing ommatidia pattern, 2 posterior vertex trichomes and 3 trichobothria: trichobothrium a, b and c. B. Collum and lateral protuberances with pattern of trichome sockets (all trichomes were removed). C. Trichome socket arrangements from tergite 2. D. Tergite 10 with densely packed smaller sockets. (A, B, C, D drawn to the same scale). E. Structure of trichobothria: Trichobothrium a (internal position), Trichobothrium b (intermediate position), Trichobothrium c (external position), sockets and their funicular structures. F. Structure of clypeo-labrum with 2 lateral lamellae, 2 lamellae and a median cleft. G. Structures of 2 lateral palps (a conical sensillum was missed on each lateral palp) and medial palps from gnathochilaria (Lp: lateral palp, Mp: medial palp)." httpUri="https://zenodo.org/record/238046/files/figure.png" pageId="5" pageNumber="465">Figure 3</figureCitation>
D).
</paragraph>
<paragraph id="9969931DFF8D751C118912EC94B4FE5E" blockId="5.[151,1437,1576,2033]" lastBlockId="6.[151,1436,151,429]" lastPageId="6" lastPageNumber="466" pageId="5" pageNumber="465">
<emphasis id="ABA24F0FFF8D751F118912EC97C4F83F" box="[199,252,1973,1996]" italics="true" pageId="5" pageNumber="465">Legs</emphasis>
: Leg segments are named following 
<bibRefCitation id="FD47EEECFF8D751F13D212EC9470F83E" author="Manton" box="[668,840,1972,1997]" pageId="5" pageNumber="465" refString="Manton, S. M. (1956) The Evolution of Arthropodan Locomotory Mechanisms-Part 5: The Structure, Habits and Evolution of the Pselaphognatha (Diplopoda). Journal of the Linnean Society London, 43, 153 - 187. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1957. tb 02516. x" type="journal article" year="1956">Manton (1956)</bibRefCitation>
. Legs 1 and 2 are without trochanter, leg 1 also lacks tarsus 1. Chaetotaxy as follows: coxa 1: 2 sensilla, coxa 2: 2 sensilla, coxae 3–13: 1–4 sensilla; all pre-femurs, postfemurs, tibia with 1 sensillum, except femurs with 2–4 sensilla, tarsus 1 and tarsus 2 with a spine (
<figureCitation id="01ED8F98FF8E751C145B15CE92A8FF43" box="[1301,1424,151,176]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="6" pageNumber="466">Figure 5A</figureCitation>
). Coxa, pre-femur and femur have a ridged biarticulate, funicular cylindrical sensillum (
<figureCitation id="01ED8F98FF8E751C152015E593E8FF26" box="[1134,1232,188,213]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="6" pageNumber="466">Figure 5</figureCitation>
B). There are also 2–3 smaller biarticulate, funicular cylindrical sensillum present on the posterior end of the femur and some on the penis (
<figureCitation id="01ED8F98FF8E751C11AE145D96A1FEEE" box="[224,409,260,285]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="6" pageNumber="466">Figure 5A and 5</figureCitation>
C). At their distal edge, post-femur and tibia are with a setiform sensillum (
<figureCitation id="01ED8F98FF8E751C15AB145D927DFEEE" box="[1253,1349,260,285]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="6" pageNumber="466">Figure 5</figureCitation>
D). The spine on tarsus 2 is much larger, and sharply pointed (
<figureCitation id="01ED8F98FF8E751C13BC147E9469FEB3" box="[754,849,295,320]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="6" pageNumber="466">Figure 5</figureCitation>
F) compared to a spine in the antero-sternal position of tarsus 1 (
<figureCitation id="01ED8F98FF8E751C106C141596BCFE96" box="[290,388,332,357]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="6" pageNumber="466">Figure 5</figureCitation>
E). Two to four ridged biarticulate, funicular cylindrical sensilla are found on the posterior edge of the last sternite. These sensilla are similar to those present on the coxa. The number of sensilla varies; 4 present on the 
<typeStatus id="466D2DBFFF8E751C107514CD96A5FE5E" box="[315,413,404,429]" pageId="6" pageNumber="466" type="holotype">holotype</typeStatus>
male and 2–4 on the 
<typeStatus id="466D2DBFFF8E751C13C014CC95C2FE5E" box="[654,762,405,429]" pageId="6" pageNumber="466" type="paratype">paratypes</typeStatus>
(
<figureCitation id="01ED8F98FF8E751C124614CD9450FE5E" box="[776,872,404,429]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="6" pageNumber="466">Figure 5</figureCitation>
G).
</paragraph>
<paragraph id="9969931DFF8E751C11D910E4948BF9A1" blockId="6.[151,1436,1466,1618]" pageId="6" pageNumber="466">
<emphasis id="ABA24F0FFF8E751C11D910E49635FA20" bold="true" box="[151,269,1469,1491]" pageId="6" pageNumber="466">FIGURE 4</emphasis>
. 
<emphasis id="ABA24F0FFF8E751C105410E49614FA20" bold="true" box="[282,300,1469,1491]" pageId="6" pageNumber="466">A</emphasis>
. Arrangement of sensilla on 7th antennal article, a conical sensillum (
<emphasis id="ABA24F0FFF8E751C12A210E494CFFA20" bold="true" box="[1004,1015,1469,1491]" pageId="6" pageNumber="466">c</emphasis>
), thick bacilliform sensillum (
<emphasis id="ABA24F0FFF8E751C146310E4926EFA20" bold="true" box="[1325,1366,1469,1491]" pageId="6" pageNumber="466">Tbs</emphasis>
) and a setiform sensillum (
<emphasis id="ABA24F0FFF8E751C102C10879654FA07" bold="true" box="[354,364,1502,1524]" pageId="6" pageNumber="466">s</emphasis>
). 
<emphasis id="ABA24F0FFF8E751C10CF108796AAFA07" bold="true" box="[385,402,1502,1524]" pageId="6" pageNumber="466">B</emphasis>
. Arrangement of sensilla of 6th article, showing position of 3 thick bacilliform sensilla (
<emphasis id="ABA24F0FFF8E751C14401087920EFA07" bold="true" box="[1294,1334,1502,1524]" pageId="6" pageNumber="466">Tbs</emphasis>
): anterior 
<emphasis id="ABA24F0FFF8E751C11D910A4979CF9E0" bold="true" box="[151,164,1533,1555]" pageId="6" pageNumber="466">a</emphasis>
, intermediate 
<emphasis id="ABA24F0FFF8E751C107810A49605F9E0" bold="true" box="[310,317,1533,1555]" pageId="6" pageNumber="466">i</emphasis>
and posterior 
<emphasis id="ABA24F0FFF8E751C109F10A496E7F9E0" bold="true" box="[465,479,1533,1555]" pageId="6" pageNumber="466">p</emphasis>
; 17 thin bacilliform sensilla (
<emphasis id="ABA24F0FFF8E751C124010A49416F9E0" bold="true" box="[782,814,1533,1555]" pageId="6" pageNumber="466">tbs</emphasis>
) included 5 inner sensilla (arrows). 
<emphasis id="ABA24F0FFF8E751C15EF10A4938BF9E0" bold="true" box="[1185,1203,1533,1555]" pageId="6" pageNumber="466">C</emphasis>
. Left side antenna and its article. 
<emphasis id="ABA24F0FFF8E751C104A1347962EF9C7" bold="true" box="[260,278,1566,1588]" pageId="6" pageNumber="466">D</emphasis>
. Scanning Electron Microscopic (SEM) image showed articles 6th, 7th and 8th with 4 sensory cones (A, B, and C were from the 
<typeStatus id="466D2DBFFF8E751C1065136596BBF9A1" box="[299,387,1596,1618]" pageId="6" pageNumber="466" type="holotype">holotype</typeStatus>
male (QMS 98960), D was an antenna from a female).
</paragraph>
<paragraph id="9969931DFF8E751C1189132693D2F913" blockId="6.[151,1436,1663,2018]" pageId="6" pageNumber="466">
<emphasis id="ABA24F0FFF8E751C118913269647F96B" box="[199,383,1663,1688]" italics="true" pageId="6" pageNumber="466">Claw -Telotarsus</emphasis>
: This is a slender structure comprised of a posterior process that is curved with an enlarged base, half as long as the claw. An anterior process is absent. A tiny lateral process is present. The anterior setiform process is almost equal in length to the claw and a common lamella process is present (
<figureCitation id="01ED8F98FF8E751C1528139E93FEF913" box="[1126,1222,1735,1760]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="6" pageNumber="466">Figure 5</figureCitation>
H).
</paragraph>
<paragraph id="9969931DFF8E751C118913B69643F8DD" blockId="6.[151,1436,1663,2018]" pageId="6" pageNumber="466">
<emphasis id="ABA24F0FFF8E751C118913B6969BF8F4" box="[199,419,1774,1799]" italics="true" pageId="6" pageNumber="466">Sex organs in male</emphasis>
: A pair of penes is present on the 2nd coxa and 2 pairs of coxal glands located on coxal plates of 8th–9th legs.
</paragraph>
<paragraph id="9969931DFF8E751C1189126093C9F811" blockId="6.[151,1436,1663,2018]" pageId="6" pageNumber="466">
<emphasis id="ABA24F0FFF8E751C118912609635F8A1" box="[199,269,1849,1874]" italics="true" pageId="6" pageNumber="466">Telson</emphasis>
: The dorsal ornamental trichome sockets are arranged almost symmetrically on each side, with 16 trichome sockets 
<emphasis id="ABA24F0FFF8E751C102F12079657F886" bold="true" box="[353,367,1886,1909]" italics="true" pageId="6" pageNumber="466">a</emphasis>
in the 
<typeStatus id="466D2DBFFF8E751C108B1204951FF885" box="[453,551,1885,1910]" pageId="6" pageNumber="466" type="holotype">holotype</typeStatus>
male. The 
<typeStatus id="466D2DBFFF8E751C13E012079422F885" box="[686,794,1886,1910]" pageId="6" pageNumber="466" type="paratype">paratypes</typeStatus>
have 8–23 trichome sockets 
<emphasis id="ABA24F0FFF8E751C152012079344F886" bold="true" box="[1134,1148,1886,1909]" italics="true" pageId="6" pageNumber="466">a</emphasis>
in both sexes. Trichome sockets 
<emphasis id="ABA24F0FFF8E751C11BC12D89638F86B" bold="true" box="[242,256,1921,1944]" italics="true" pageId="6" pageNumber="466">a</emphasis>
form 2 rows; top row with small sockets and bottom row with larger trichome sockets. The two largest holders are situated near trichome 
<emphasis id="ABA24F0FFF8E751C137A12FD957AF84E" bold="true" box="[564,578,1956,1981]" italics="true" pageId="6" pageNumber="466">b</emphasis>
. Three large protruding base trichome sockets 
<emphasis id="ABA24F0FFF8E751C153B12FF93B9F84E" bold="true" box="[1141,1153,1958,1981]" italics="true" pageId="6" pageNumber="466">c</emphasis>
: 
<emphasis id="ABA24F0FFF8E751C15D812FC93E6F84E" bold="true" box="[1174,1246,1957,1981]" italics="true" pageId="6" pageNumber="466">c1, c2</emphasis>
and 
<emphasis id="ABA24F0FFF8E751C145112FC9201F84E" bold="true" box="[1311,1337,1957,1981]" italics="true" pageId="6" pageNumber="466">c3</emphasis>
, form a triangular shape. Circular indentation 
<emphasis id="ABA24F0FFF8E751C130E12919576F812" bold="true" box="[576,590,1992,2017]" italics="true" pageId="6" pageNumber="466">d</emphasis>
is apparent near the exterior side of trichomes 
<emphasis id="ABA24F0FFF8E751C151212909350F813" bold="true" box="[1116,1128,1993,2016]" italics="true" pageId="6" pageNumber="466">c</emphasis>
(
<figureCitation id="01ED8F98FF8E751C1539129093E0F811" box="[1143,1240,1993,2018]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="6" pageNumber="466">
Figure 
<date id="ED68B5DDFF8E751C1587129093E0F811" box="[1225,1240,1993,2018]" pageId="6" pageNumber="466">5</date>
</figureCitation>
I).
</paragraph>
<caption id="CDA9C395FF8F751D11D913AB9203F82C" httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="7" pageNumber="467" targetBox="[200,1380,211,1747]" targetPageId="7">
<paragraph id="9969931DFF8F751D11D913AB9203F82C" blockId="7.[151,1437,1778,2015]" pageId="7" pageNumber="467">
<emphasis id="ABA24F0FFF8F751D11D913AB9637F8FB" bold="true" box="[151,271,1778,1800]" pageId="7" pageNumber="467">FIGURE 5</emphasis>
. 
<emphasis id="ABA24F0FFF8F751D105013AB9608F8FB" bold="true" box="[286,304,1778,1800]" pageId="7" pageNumber="467">A</emphasis>
. Leg segments and chaetotaxy of the second leg and penis (
<emphasis id="ABA24F0FFF8F751D12E213AB9482F8FB" bold="true" box="[940,954,1778,1800]" pageId="7" pageNumber="467">p</emphasis>
: penis, 
<emphasis id="ABA24F0FFF8F751D154013AB931EF8FB" bold="true" box="[1038,1062,1778,1800]" pageId="7" pageNumber="467">co</emphasis>
: coxa, 
<emphasis id="ABA24F0FFF8F751D153B13AB938BF8FB" bold="true" box="[1141,1203,1778,1800]" pageId="7" pageNumber="467">pre.fe</emphasis>
: pre-femur, 
<emphasis id="ABA24F0FFF8F751D147813AB9272F8FB" bold="true" box="[1334,1354,1778,1800]" pageId="7" pageNumber="467">fe</emphasis>
: femur, 
<emphasis id="ABA24F0FFF8F751D11D9124897E6F8D4" bold="true" box="[151,222,1809,1831]" pageId="7" pageNumber="467">post.fe</emphasis>
: post-femur, 
<emphasis id="ABA24F0FFF8F751D103F124896B6F8D4" bold="true" box="[369,398,1809,1831]" pageId="7" pageNumber="467">tib</emphasis>
: tibia, 
<emphasis id="ABA24F0FFF8F751D10AE124896C5F8D4" bold="true" box="[480,509,1809,1831]" pageId="7" pageNumber="467">T1</emphasis>
: tarsus 1, 
<emphasis id="ABA24F0FFF8F751D1338124895ACF8D4" bold="true" box="[630,660,1809,1831]" pageId="7" pageNumber="467">T2</emphasis>
: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (
<emphasis id="ABA24F0FFF8F751D102612769641F8B6" bold="true" box="[360,377,1839,1861]" pageId="7" pageNumber="467">B</emphasis>
) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (
<emphasis id="ABA24F0FFF8F751D15B412769234F8B6" bold="true" box="[1274,1292,1839,1861]" pageId="7" pageNumber="467">C</emphasis>
) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (
<emphasis id="ABA24F0FFF8F751D15CE121793AAF897" bold="true" box="[1152,1170,1870,1892]" pageId="7" pageNumber="467">D</emphasis>
). 
<emphasis id="ABA24F0FFF8F751D15E612179381F897" bold="true" box="[1192,1209,1870,1892]" pageId="7" pageNumber="467">E</emphasis>
&amp; 
<emphasis id="ABA24F0FFF8F751D1595121793D0F897" bold="true" box="[1243,1256,1870,1892]" pageId="7" pageNumber="467">F</emphasis>
. Spines found on Tarsus 1 and 2. 
<emphasis id="ABA24F0FFF8F751D107912359670F871" bold="true" box="[311,328,1900,1922]" pageId="7" pageNumber="467">G</emphasis>
. Last sternite, with 3 sensilla and a socket without sensillum. 
<emphasis id="ABA24F0FFF8F751D12F0123594EAF871" bold="true" box="[958,978,1900,1922]" pageId="7" pageNumber="467">H</emphasis>
. Telotarsus structure with processes: 
<emphasis id="ABA24F0FFF8F751D140012359261F871" bold="true" box="[1358,1369,1900,1922]" pageId="7" pageNumber="467">c</emphasis>
: claw, 
<emphasis id="ABA24F0FFF8F751D11D912D297A6F852" bold="true" box="[151,158,1931,1953]" pageId="7" pageNumber="467">l</emphasis>
: lamella process, 
<emphasis id="ABA24F0FFF8F751D101812D29653F852" bold="true" box="[342,363,1931,1953]" pageId="7" pageNumber="467">lp</emphasis>
: lateral process, 
<emphasis id="ABA24F0FFF8F751D135612D2951EF852" bold="true" box="[536,550,1931,1953]" pageId="7" pageNumber="467">p</emphasis>
: posterior process, 
<emphasis id="ABA24F0FFF8F751D13A012D295C0F852" bold="true" box="[750,760,1931,1953]" pageId="7" pageNumber="467">s</emphasis>
: anterior setiform process. 
<emphasis id="ABA24F0FFF8F751D154112D29321F852" bold="true" box="[1039,1049,1931,1953]" pageId="7" pageNumber="467">I</emphasis>
. Dorsal ornamental trichome sockets: trichome sockets 
<emphasis id="ABA24F0FFF8F751D100912F2966BF833" bold="true" box="[327,339,1963,1984]" italics="true" pageId="7" pageNumber="467">a</emphasis>
: 
<emphasis id="ABA24F0FFF8F751D102F12F29655F833" bold="true" box="[353,365,1963,1984]" italics="true" pageId="7" pageNumber="467">a</emphasis>
1 to 
<emphasis id="ABA24F0FFF8F751D10D712F2969EF833" bold="true" box="[409,422,1963,1984]" italics="true" pageId="7" pageNumber="467">a</emphasis>
16; trichome socket 
<emphasis id="ABA24F0FFF8F751D133D12F395B8F833" bold="true" box="[627,640,1962,1984]" italics="true" pageId="7" pageNumber="467">b</emphasis>
; trichome sockets 
<emphasis id="ABA24F0FFF8F751D127212F2947FF833" bold="true" box="[828,839,1963,1984]" italics="true" pageId="7" pageNumber="467">c</emphasis>
: 
<emphasis id="ABA24F0FFF8F751D121A12F29467F833" bold="true" box="[852,863,1963,1984]" italics="true" pageId="7" pageNumber="467">c</emphasis>
1, 
<emphasis id="ABA24F0FFF8F751D123712F294BCF833" bold="true" box="[889,900,1963,1984]" italics="true" pageId="7" pageNumber="467">c</emphasis>
2 and 
<emphasis id="ABA24F0FFF8F751D128F12F294F4F833" bold="true" box="[961,972,1963,1984]" italics="true" pageId="7" pageNumber="467">c</emphasis>
3. Circular indentation 
<emphasis id="ABA24F0FFF8F751D15F812F393FAF833" bold="true" box="[1206,1218,1962,1984]" italics="true" pageId="7" pageNumber="467">d</emphasis>
. 
<emphasis id="ABA24F0FFF8F751D158012F393E3F833" bold="true" box="[1230,1243,1962,1984]" pageId="7" pageNumber="467">J</emphasis>
. Nest trichome. 
<emphasis id="ABA24F0FFF8F751D14CC12F392AEF833" bold="true" box="[1410,1430,1962,1984]" pageId="7" pageNumber="467">K</emphasis>
. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female).
</paragraph>
</caption>
<paragraph id="9969931DFF80751211D9112E967CFB1A" blockId="8.[151,1436,1140,1257]" pageId="8" pageNumber="468">
<emphasis id="ABA24F0FFF80751211D9112E9637FB7E" bold="true" box="[151,271,1143,1165]" pageId="8" pageNumber="468">FIGURE 6</emphasis>
. Genus 
<taxonomicName id="5ED6E89EFF8075121026112E96C9FB7E" box="[360,497,1143,1165]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF8075121026112E96C9FB7E" box="[360,497,1143,1165]" italics="true" pageId="8" pageNumber="468">Monographis</emphasis>
</taxonomicName>
divided into 2 groups of species based on the arrangement of sensilla on the 6th antennal article (
<bibRefCitation id="FD47EEECFF80751211AB11CC96DCFB58" author="Huynh" box="[229,484,1173,1195]" pageId="8" pageNumber="468" refString="Huynh, C. &amp; Veenstra, A. (2013) Taxonomy and biology of a new species of pincushion Millipede of the genus Monographis (Diplopoda: Polyxenidae) from Australia. Zootaxa, 3721 (6), 573 - 588. http: // dx. doi. org / 10.11646 / zootaxa. 3721.6.4" type="journal article" year="2013">Huynh &amp; Veenstra 2013</bibRefCitation>
); 
<taxonomicName id="5ED6E89EFF80751210B811CC9434FB58" box="[502,780,1173,1195]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF80751210B811CC9434FB58" box="[502,780,1173,1195]" italics="true" pageId="8" pageNumber="468">Monographis dongnaiensis</emphasis>
</taxonomicName>
was placed in Group 2, which has sensilla forming a triangular shape. The drawings of sensilla observed on 
<taxonomicName id="5ED6E89EFF807512132D11ED9461FB39" box="[611,857,1204,1226]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="kraepelini">
<emphasis id="ABA24F0FFF807512132D11ED9461FB39" box="[611,857,1204,1226]" italics="true" pageId="8" pageNumber="468">Monographis kraepelini</emphasis>
</taxonomicName>
and 
<taxonomicName id="5ED6E89EFF80751212C211EC9314FB39" box="[908,1068,1205,1226]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="tamoyoensis">
<emphasis id="ABA24F0FFF80751212C211EC9314FB39" box="[908,1068,1205,1226]" italics="true" pageId="8" pageNumber="468">M. tamoyoensis</emphasis>
</taxonomicName>
were from 
<bibRefCitation id="FD47EEECFF80751215EA11EC9678FB1A" pageId="8" pageNumber="468" refString="Nguyen Duy-Jacquemin, M. &amp; Conde, B. (1967) Morphologie et Geonemie du genre Monographis. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 64, 43 - 81." type="journal article">Nguyen Duy-Jacquemin &amp; Condé (1967)</bibRefCitation>
.
</paragraph>
<paragraph id="9969931DFF8075121189104C920BF92D" blockId="8.[151,1436,1301,2016]" pageId="8" pageNumber="468">
<emphasis id="ABA24F0FFF8075121189104C9641FADD" box="[199,377,1301,1326]" italics="true" pageId="8" pageNumber="468">Caudal bundles</emphasis>
: In both males and females of 
<taxonomicName id="5ED6E89EFF8075121392104C9328FADD" box="[732,1040,1301,1326]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF8075121392104C9328FADD" box="[732,1040,1301,1326]" italics="true" pageId="8" pageNumber="468">Monographis dongnaiensis</emphasis>
</taxonomicName>
, these are similar in appearance to those of 
<taxonomicName id="5ED6E89EFF80751211B7106295C3FAA1" authority="Huynh &amp; Veenstra 2013" authorityName="Huynh &amp; Veenstra" authorityYear="2013" box="[249,763,1337,1362]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF80751211B7106296F4FAA1" box="[249,460,1337,1362]" italics="true" pageId="8" pageNumber="468">M. queenslandicus</emphasis>
(
<bibRefCitation id="FD47EEECFF8075121095106095CCFAA1" author="Huynh" box="[475,756,1337,1362]" pageId="8" pageNumber="468" refString="Huynh, C. &amp; Veenstra, A. (2013) Taxonomy and biology of a new species of pincushion Millipede of the genus Monographis (Diplopoda: Polyxenidae) from Australia. Zootaxa, 3721 (6), 573 - 588. http: // dx. doi. org / 10.11646 / zootaxa. 3721.6.4" type="journal article" year="2013">Huynh &amp; Veenstra 2013</bibRefCitation>
)
</taxonomicName>
. In males, the caudal bundle is uniform in structure and the caudal trichome sockets are in similar sizes. The caudal trichomes have 2–4 hooks commonly with 2–3 hooks facing ventrally. The hooks are connected with distal facing, small barbates in the middle section and join with a larger barbate stem, forming 3–4 rings of 3 spines, distally facing, at the base of this trichome stem (
<figureCitation id="01ED8F98FF807512145810FC924FFA4D" box="[1302,1399,1445,1470]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="8" pageNumber="468">Figure 5</figureCitation>
K). Few larger barbate trichome sockets are present; commonly 2–3 rows, and form slightly uneven lateral rows that extend toward the centre of the telson. The barbate trichomes (which are used to prevent hooks on trichomes becoming tangled) are large with their length exceeding the longest caudal trichomes in the bundle. In females, the caudal bundle has 2 distinguishing structures: the caudal trichomes, which are the main dorsal structures and are similar to the male caudal bundle, and 2 latero-sternal bundles of nest trichomes (
<figureCitation id="01ED8F98FF8075121562130393B5F980" box="[1068,1165,1626,1651]" captionStart="FIGURE 5. A" captionStartId="7.[151,250,1778,1800]" captionTargetBox="[200,1380,211,1747]" captionTargetId="figure@7.[197,1389,193,1757]" captionTargetPageId="7" captionText="FIGURE 5. A. Leg segments and chaetotaxy of the second leg and penis (p: penis, co: coxa, pre. fe: pre-femur, fe: femur, post. fe: post-femur, tib: tibia, T 1: tarsus 1, T 2: tarsus 2). Three types of seta present: 1) Ridged biarticulate funicular cylindrical sensilla (B) were found in coxa, pre-femur, femur and the last sternite. 2) A similar small form (C) found in the posterior position on femur, sex organs—penis or vulva. 3) Setiform seta in post-femur and tibia (D). E &amp; F. Spines found on Tarsus 1 and 2. G. Last sternite, with 3 sensilla and a socket without sensillum. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. I. Dorsal ornamental trichome sockets: trichome sockets a: a 1 to a 16; trichome socket b; trichome sockets c: c 1, c 2 and c 3. Circular indentation d. J. Nest trichome. K. Caudal trichome. (A, B, C, D, E, F, G, H &amp; I were from the holotype male (QMS 98960), J &amp; K were from a female)." httpUri="https://zenodo.org/record/238047/files/figure.png" pageId="8" pageNumber="468">Figure 5</figureCitation>
J). These nest trichomes have small, fine trichome sockets, with trichomes half the length of those on the main caudal trichomes. They differ from caudal trichomes in having smaller, slender stems with double facing spines connected by 3–5 hooks. These nest trichomes are used for nest building to provide protection for egg clusters (
<bibRefCitation id="FD47EEECFF807512155F139C921FF92D" author="Huynh" box="[1041,1319,1733,1758]" pageId="8" pageNumber="468" refString="Huynh, C. &amp; Veenstra, A. (2014) Reproduction, egg morphology and development observed in Monographis queenslandicus (Diplopoda: Polyxenidae). Invertebrate Zoology, 11, 335 - 345." type="journal article" year="2014">Huynh &amp; Veenstra 2014</bibRefCitation>
).
</paragraph>
<paragraph id="9969931DFF807512118913B394C0F813" blockId="8.[151,1436,1301,2016]" pageId="8" pageNumber="468">
<emphasis id="ABA24F0FFF807512118913B39364F8F1" bold="true" box="[199,1116,1769,1795]" pageId="8" pageNumber="468">
Comparison of 
<taxonomicName id="5ED6E89EFF80751210C813B095FDF8F1" box="[390,709,1769,1794]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF80751210C813B095FDF8F1" bold="true" box="[390,709,1769,1794]" italics="true" pageId="8" pageNumber="468">Monographis dongnaiensis</emphasis>
</taxonomicName>
to other species in 
<emphasis id="ABA24F0FFF80751212F713B09364F8F1" bold="true" box="[953,1116,1769,1794]" italics="true" pageId="8" pageNumber="468">
<taxonomicName id="5ED6E89EFF80751212F713B09360F8F1" box="[953,1112,1769,1794]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="genus">Monographis</taxonomicName>
.
</emphasis>
</emphasis>
In addition to the common characteristics of the genus 
<taxonomicName id="5ED6E89EFF807512109E1254955FF8D5" box="[464,615,1805,1830]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF807512109E1254955FF8D5" box="[464,615,1805,1830]" italics="true" pageId="8" pageNumber="468">Monographis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5ED6E89EFF807512133B12569417F8D5" box="[629,815,1805,1830]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF807512133B12569417F8D5" box="[629,815,1805,1830]" italics="true" pageId="8" pageNumber="468">M. dongnaiensis</emphasis>
</taxonomicName>
has morphological characteristics similar to a group of 
<taxonomicName id="5ED6E89EFF80751211D9126D9616F8BE" box="[151,302,1844,1869]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF80751211D9126D9616F8BE" box="[151,302,1844,1869]" italics="true" pageId="8" pageNumber="468">Monographis</emphasis>
</taxonomicName>
species, which have the sensilla forming a triangular shape on the 6th antennal article (
<bibRefCitation id="FD47EEECFF8075121463126D9678F887" author="Huynh" pageId="8" pageNumber="468" refString="Huynh, C. &amp; Veenstra, A. (2013) Taxonomy and biology of a new species of pincushion Millipede of the genus Monographis (Diplopoda: Polyxenidae) from Australia. Zootaxa, 3721 (6), 573 - 588. http: // dx. doi. org / 10.11646 / zootaxa. 3721.6.4" type="journal article" year="2013">Huynh &amp; Veenstra 2013</bibRefCitation>
) (Figure 6). When comparing the sensilla structures on the 6th antennal article between 
<taxonomicName id="5ED6E89EFF80751214331204961AF864" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="ABA24F0FFF80751214331204961AF864" italics="true" pageId="8" pageNumber="468">M. yunnanensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="5ED6E89EFF807512101812D99537F86B" box="[342,527,1919,1944]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF807512101812D99537F86B" box="[342,527,1919,1944]" italics="true" pageId="8" pageNumber="468">M. dongnaiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5ED6E89EFF807512135212D995F3F864" box="[540,715,1920,1943]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="ABA24F0FFF807512135212D995F3F864" box="[540,715,1920,1943]" italics="true" pageId="8" pageNumber="468">M. yunnanensis</emphasis>
</taxonomicName>
has 17 sensilla with 3 inner sensilla, a conical sensillum (
<bibRefCitation id="FD47EEECFF807512140212269638F84F" author="Ishii" pageId="8" pageNumber="468" refString="Ishii, K. &amp; Yin, W. - Y. (2000) New species of Penicillata (Diplopoda) from Yunnan, Southwest China. In: Aoki, J., Yin, W. Y. &amp; Imadate, G. (Eds.), Taxonomical studies on the soil fauna of Yunnan Province in Southwest China. Tokai University Press, Tokyo, pp. 91 - 116." type="book chapter" year="2000">Ishii &amp; Yin 2000</bibRefCitation>
) and 
<taxonomicName id="5ED6E89EFF807512107312FC96CEF84F" box="[317,502,1955,1980]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF807512107312FC96CEF84F" box="[317,502,1955,1980]" italics="true" pageId="8" pageNumber="468">M. dongnaiensis</emphasis>
</taxonomicName>
has 20 sensilla with no conical sensillum but with 5 inner sensilla (Figure 7). These are obviously different species of 
<taxonomicName id="5ED6E89EFF807512135D129E9592F813" box="[531,682,1991,2016]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF807512135D129E9592F813" box="[531,682,1991,2016]" italics="true" pageId="8" pageNumber="468">Monographis</emphasis>
</taxonomicName>
occurring in the same region.
</paragraph>
<paragraph id="9969931DFF81751311D9169094EEFBEF" blockId="9.[151,1436,966,1052]" pageId="9" pageNumber="469">
<emphasis id="ABA24F0FFF81751311D916909634FC2C" bold="true" box="[151,268,969,991]" pageId="9" pageNumber="469">FIGURE 7</emphasis>
. Comparison of the number of sensilla observed on the 6th antennal article of 
<taxonomicName id="5ED6E89EFF817513155516939381FC2C" box="[1051,1209,970,991]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="ABA24F0FFF817513155516939381FC2C" box="[1051,1209,970,991]" italics="true" pageId="9" pageNumber="469">M. yunnanensis</emphasis>
</taxonomicName>
(
<emphasis id="ABA24F0FFF8175131586169093E2FC2C" bold="true" box="[1224,1242,969,991]" pageId="9" pageNumber="469">A</emphasis>
) and 
<taxonomicName id="5ED6E89EFF817513145A16909625FC0D" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF817513145A16909625FC0D" italics="true" pageId="9" pageNumber="469">Monographis dongnaiensis</emphasis>
</taxonomicName>
(
<emphasis id="ABA24F0FFF817513106516B19604FC0D" bold="true" box="[299,316,1000,1022]" pageId="9" pageNumber="469">B</emphasis>
). The pattern of sensilla of 
<taxonomicName id="5ED6E89EFF817513131C16B095C8FC0D" box="[594,752,1001,1022]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="ABA24F0FFF817513131C16B095C8FC0D" box="[594,752,1001,1022]" italics="true" pageId="9" pageNumber="469">M. yunnanensis</emphasis>
</taxonomicName>
is referred from 
<bibRefCitation id="FD47EEECFF81751312D416B19360FC0D" author="Ishii" box="[922,1112,1000,1022]" pageId="9" pageNumber="469" refString="Ishii, K. &amp; Yin, W. - Y. (2000) New species of Penicillata (Diplopoda) from Yunnan, Southwest China. In: Aoki, J., Yin, W. Y. &amp; Imadate, G. (Eds.), Taxonomical studies on the soil fauna of Yunnan Province in Southwest China. Tokai University Press, Tokyo, pp. 91 - 116." type="book chapter" year="2000">Ishii &amp; Yin (2000)</bibRefCitation>
. Small arrows indicate the inner sensilla, a large arrow indicates the location of the conical sensillum. Not to scale.
</paragraph>
</subSubSection>
<caption id="CDA9C395FF81751311D9132D97E8F914" httpUri="https://zenodo.org/record/238048/files/figure.png" pageId="9" pageNumber="469" targetBox="[152,1433,1102,1631]" targetPageId="9">
<paragraph id="9969931DFF81751311D9132D97E8F914" blockId="9.[151,1437,1652,1767]" pageId="9" pageNumber="469">
<emphasis id="ABA24F0FFF81751311D9132D9628F979" bold="true" box="[151,272,1652,1674]" pageId="9" pageNumber="469">FIGURE 8</emphasis>
. Comparison of the number of sensilla between 
<taxonomicName id="5ED6E89EFF817513125D132D9319F978" box="[787,1057,1652,1675]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="tamoyoensis">
<emphasis id="ABA24F0FFF817513125D132D9319F978" box="[787,1057,1652,1675]" italics="true" pageId="9" pageNumber="469">Monographis tamoyoensis</emphasis>
</taxonomicName>
(
<emphasis id="ABA24F0FFF817513157D132D937DF979" bold="true" box="[1075,1093,1652,1674]" pageId="9" pageNumber="469">A</emphasis>
), 
<taxonomicName id="5ED6E89EFF8175131513132F9226F979" box="[1117,1310,1652,1675]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF8175131513132F9226F979" box="[1117,1310,1652,1675]" italics="true" pageId="9" pageNumber="469">M. queenslandicus</emphasis>
</taxonomicName>
(
<emphasis id="ABA24F0FFF817513147E132D9279F979" bold="true" box="[1328,1345,1652,1674]" pageId="9" pageNumber="469">B</emphasis>
) and 
<taxonomicName id="5ED6E89EFF81751314CE132F9625F95A" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF81751314CE132F9625F95A" italics="true" pageId="9" pageNumber="469">M. dongnaiensis</emphasis>
</taxonomicName>
(
<emphasis id="ABA24F0FFF817513106313CA9607F95A" bold="true" box="[301,319,1683,1705]" pageId="9" pageNumber="469">C</emphasis>
). Sensilla pattern of 
<taxonomicName id="5ED6E89EFF817513135713CD9581F95A" box="[537,697,1684,1705]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="tamoyoensis">
<emphasis id="ABA24F0FFF817513135713CD9581F95A" box="[537,697,1684,1705]" italics="true" pageId="9" pageNumber="469">M. tamoyoensis</emphasis>
</taxonomicName>
is referred from Nguyen Duy-Jacquemin &amp; Condé (1967), and sensilla pattern of 
<taxonomicName id="5ED6E89EFF817513104E13EA9686F93B" box="[256,446,1714,1736]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF817513104E13EA9686F93B" box="[256,446,1714,1736]" italics="true" pageId="9" pageNumber="469">M. queenslandicus</emphasis>
</taxonomicName>
is from Huynh &amp; Veenstra (2013). Arrows indicate the positions of the conical sensillum. Not to scale.
</paragraph>
</caption>
<subSubSection id="D1CCC096FF8175111189125695E4FE23" lastPageId="11" lastPageNumber="471" pageId="9" pageNumber="469" type="reference_group">
<paragraph id="9969931DFF81751311891256968DF828" blockId="9.[151,1436,1806,2011]" pageId="9" pageNumber="469">
<taxonomicName id="5ED6E89EFF817513118912569646F8D5" ID-CoL="4475H" box="[199,382,1807,1830]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="tamoyoensis">
<emphasis id="ABA24F0FFF817513118912569646F8D5" box="[199,382,1807,1830]" italics="true" pageId="9" pageNumber="469">M. tamoyoensis</emphasis>
</taxonomicName>
has 18 sensilla with 5 inner sensilla (
<bibRefCitation id="FD47EEECFF817513121812569209F8D4" author="Nguyen" box="[854,1329,1806,1831]" pageId="9" pageNumber="469" refString="Nguyen Duy-Jacquemin, M. &amp; Conde, B. (1967) Morphologie et Geonemie du genre Monographis. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 64, 43 - 81." type="journal article" year="1967">Nguyen Duy-Jacquemin &amp; Condé 1967</bibRefCitation>
) and 
<taxonomicName id="5ED6E89EFF81751314331256967CF8B8" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF81751314331256967CF8B8" italics="true" pageId="9" pageNumber="469">M. queenslandicus</emphasis>
</taxonomicName>
has the same arrangement, but with 19 sensilla (
<bibRefCitation id="FD47EEECFF8175131238126B93AAF8B9" author="Huynh" box="[886,1170,1842,1867]" pageId="9" pageNumber="469" refString="Huynh, C. &amp; Veenstra, A. (2013) Taxonomy and biology of a new species of pincushion Millipede of the genus Monographis (Diplopoda: Polyxenidae) from Australia. Zootaxa, 3721 (6), 573 - 588. http: // dx. doi. org / 10.11646 / zootaxa. 3721.6.4" type="journal article" year="2013">Huynh &amp; Veenstra 2013</bibRefCitation>
). 
<taxonomicName id="5ED6E89EFF81751315E8126A926FF8B9" ID-CoL="4475H" box="[1190,1367,1843,1866]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="tamoyoensis">
<emphasis id="ABA24F0FFF81751315E8126A926FF8B9" box="[1190,1367,1843,1866]" italics="true" pageId="9" pageNumber="469">M. tamoyoensis</emphasis>
</taxonomicName>
has a conical sensillum located in the inner group of sensilla; 
<taxonomicName id="5ED6E89EFF8175131242120E94E6F89C" box="[780,990,1878,1903]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF8175131242120E94E6F89C" box="[780,990,1878,1903]" italics="true" pageId="9" pageNumber="469">M. queenslandicus</emphasis>
</taxonomicName>
is observed to have a conical sensillum located in the outer group of sensilla, whereas 
<taxonomicName id="5ED6E89EFF81751313D112229460F860" box="[671,856,1914,1939]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF81751313D112229460F860" box="[671,856,1914,1939]" italics="true" pageId="9" pageNumber="469">M. dongnaiensis</emphasis>
</taxonomicName>
is observed to have 20 sensilla with 5 inner sensilla and the absence of a conical sensillum (
<figureCitation id="01ED8F98FF817513131B12C79587F844" box="[597,703,1950,1975]" captionStart="FIGURE 8" captionStartId="9.[151,250,1652,1674]" captionTargetBox="[152,1433,1102,1631]" captionTargetId="figure@9.[152,1434,1102,1631]" captionTargetPageId="9" captionText="FIGURE 8. Comparison of the number of sensilla between Monographis tamoyoensis (A), M. queenslandicus (B) and M. dongnaiensis (C). Sensilla pattern of M. tamoyoensis is referred from Nguyen Duy-Jacquemin &amp; Condé (1967), and sensilla pattern of M. queenslandicus is from Huynh &amp; Veenstra (2013). Arrows indicate the positions of the conical sensillum. Not to scale." httpUri="https://zenodo.org/record/238048/files/figure.png" pageId="9" pageNumber="469">Figures 8</figureCitation>
&amp; 
<figureCitation id="01ED8F98FF81751313AA12C695CCF844" box="[740,756,1951,1975]" captionStart="FIGURE 9" captionStartId="10.[151,250,722,744]" captionTargetBox="[151,1436,193,700]" captionTargetId="figure@10.[151,1436,193,701]" captionTargetPageId="10" captionText="FIGURE 9. The Scanning Electron Microscopic images showed the similarity of the arrangement of sensilla between M. queenslandicus (A) and Monographis dongnaiensis (B). But M. dongnaiensis is different in lacking a conical sensillum. Small arrows indicate the inner sensilla, a large arrow indicates the position of the conical sensillum. Not to scale." httpUri="https://zenodo.org/record/238049/files/figure.png" pageId="9" pageNumber="469">9</figureCitation>
). This suggests that 
<taxonomicName id="5ED6E89EFF817513129412C693ABF844" box="[986,1171,1950,1975]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="469" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF817513129412C693ABF844" box="[986,1171,1950,1975]" italics="true" pageId="9" pageNumber="469">M. dongnaiensis</emphasis>
</taxonomicName>
is a species that should be included in this group.
</paragraph>
<caption id="CDA9C395FF82751011D9178B93EFFCD6" httpUri="https://zenodo.org/record/238049/files/figure.png" pageId="10" pageNumber="470" targetBox="[151,1436,193,700]" targetPageId="10">
<paragraph id="9969931DFF82751011D9178B93EFFCD6" blockId="10.[151,1437,722,805]" pageId="10" pageNumber="470">
<emphasis id="ABA24F0FFF82751011D9178B962EFD1B" bold="true" box="[151,278,722,744]" pageId="10" pageNumber="470">FIGURE 9.</emphasis>
The Scanning Electron Microscopic images showed the similarity of the arrangement of sensilla between 
<taxonomicName id="5ED6E89EFF82751014CE178A960BFCF4" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF82751014CE178A960BFCF4" italics="true" pageId="10" pageNumber="470">M. queenslandicus</emphasis>
</taxonomicName>
(
<emphasis id="ABA24F0FFF827510100C17A8966CFCF4" bold="true" box="[322,340,753,775]" pageId="10" pageNumber="470">A</emphasis>
) and 
<taxonomicName id="5ED6E89EFF82751010C017A8959BFCF4" box="[398,675,753,775]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF82751010C017A8959BFCF4" box="[398,675,753,775]" italics="true" pageId="10" pageNumber="470">Monographis dongnaiensis</emphasis>
</taxonomicName>
(
<emphasis id="ABA24F0FFF82751013FC17A895FBFCF4" bold="true" box="[690,707,753,775]" pageId="10" pageNumber="470">B</emphasis>
). But 
<taxonomicName id="5ED6E89EFF827510124D17AB9493FCF4" box="[771,939,753,775]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF827510124D17AB9493FCF4" box="[771,939,753,775]" italics="true" pageId="10" pageNumber="470">M. dongnaiensis</emphasis>
</taxonomicName>
is different in lacking a conical sensillum. Small arrows indicate the inner sensilla, a large arrow indicates the position of the conical sensillum. Not to scale.
</paragraph>
</caption>
<caption id="CDA9C395FF82751011D910A895D7F972" httpUri="https://zenodo.org/record/238050/files/figure.png" pageId="10" pageNumber="470" targetBox="[160,1433,862,1490]" targetPageId="10">
<paragraph id="9969931DFF82751011D910A895D7F972" blockId="10.[151,1437,1521,1665]" pageId="10" pageNumber="470">
<emphasis id="ABA24F0FFF82751011D910A89618F9F4" bold="true" box="[151,288,1521,1543]" pageId="10" pageNumber="470">FIGURE 10</emphasis>
. Comparison between claw structures of 
<taxonomicName id="5ED6E89EFF82751013BC10AB94A0F9F4" ID-CoL="4475H" box="[754,920,1522,1543]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="tamoyoensis">
<emphasis id="ABA24F0FFF82751013BC10AB94A0F9F4" box="[754,920,1522,1543]" italics="true" pageId="10" pageNumber="470">M. tamoyoensis</emphasis>
</taxonomicName>
(
<emphasis id="ABA24F0FFF82751012E110A894F9F9F4" bold="true" box="[943,961,1521,1543]" pageId="10" pageNumber="470">A</emphasis>
), 
<taxonomicName id="5ED6E89EFF827510129010AB939BF9F4" box="[990,1187,1521,1543]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF827510129010AB939BF9F4" box="[990,1187,1521,1543]" italics="true" pageId="10" pageNumber="470">M. queenslandicus</emphasis>
</taxonomicName>
(
<emphasis id="ABA24F0FFF82751015F410A893F3F9F4" bold="true" box="[1210,1227,1521,1543]" pageId="10" pageNumber="470">B</emphasis>
) and 
<taxonomicName id="5ED6E89EFF827510145A10A89625F9D6" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF827510145A10A89625F9D6" italics="true" pageId="10" pageNumber="470">Monographis dongnaiensis</emphasis>
</taxonomicName>
(
<emphasis id="ABA24F0FFF827510106013569678F9D6" bold="true" box="[302,320,1551,1573]" pageId="10" pageNumber="470">C</emphasis>
). A claw of 
<taxonomicName id="5ED6E89EFF827510108213489556F9D5" ID-CoL="4475H" box="[460,622,1553,1574]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="tamoyoensis">
<emphasis id="ABA24F0FFF827510108213489556F9D5" box="[460,622,1553,1574]" italics="true" pageId="10" pageNumber="470">M. tamoyoensis</emphasis>
</taxonomicName>
is referred from Nguyen Duy-Jacquemin &amp; Condé (1967), and a claw of 
<taxonomicName id="5ED6E89EFF82751014CE1348960BF9B7" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF82751014CE1348960BF9B7" italics="true" pageId="10" pageNumber="470">M. queenslandicus</emphasis>
</taxonomicName>
is from Huynh &amp; Veenstra (2013). Arrows indicate the length of the anterior setiform processes. The dot lines compared the length of anterior setiform processes to the claws. 
<emphasis id="ABA24F0FFF827510127D13159478F991" bold="true" box="[819,832,1612,1634]" pageId="10" pageNumber="470">a</emphasis>
: anterior process, 
<emphasis id="ABA24F0FFF82751012B11315933EF991" bold="true" box="[1023,1030,1612,1634]" pageId="10" pageNumber="470">l</emphasis>
: lamella process, 
<emphasis id="ABA24F0FFF827510158C131593EFF991" bold="true" box="[1218,1239,1612,1634]" pageId="10" pageNumber="470">lp</emphasis>
: lateral process, 
<emphasis id="ABA24F0FFF82751014C9131592ADF991" bold="true" box="[1415,1429,1612,1634]" pageId="10" pageNumber="470">p</emphasis>
: posterior process, 
<emphasis id="ABA24F0FFF827510101E13329662F972" bold="true" box="[336,346,1643,1665]" pageId="10" pageNumber="470">s</emphasis>
: anterior setiform process. Not to scale.
</paragraph>
</caption>
<paragraph id="9969931DFF827510118913F6947FF8A4" blockId="10.[151,1436,1710,2023]" pageId="10" pageNumber="470">
In this study, other structures such as the claws were used to distinguish between species in this group. 
<taxonomicName id="5ED6E89EFF827510143313F69613F918" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF827510143313F69613F918" italics="true" pageId="10" pageNumber="470">M. dongnaiensis</emphasis>
</taxonomicName>
has slender claws as does 
<taxonomicName id="5ED6E89EFF8275101325138A9479F918" box="[619,833,1746,1771]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF8275101325138A9479F918" box="[619,833,1746,1771]" italics="true" pageId="10" pageNumber="470">M. queenslandicus</emphasis>
</taxonomicName>
and differs from 
<taxonomicName id="5ED6E89EFF827510155C138A93FEF919" ID-CoL="4475H" box="[1042,1222,1747,1770]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="tamoyoensis">
<emphasis id="ABA24F0FFF827510155C138A93FEF919" box="[1042,1222,1747,1770]" italics="true" pageId="10" pageNumber="470">M. tamoyoensis</emphasis>
</taxonomicName>
, which has robust claws and short anterior setiform process. 
<taxonomicName id="5ED6E89EFF827510133F13AE9412F8FC" box="[625,810,1782,1807]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF827510133F13AE9412F8FC" box="[625,810,1782,1807]" italics="true" pageId="10" pageNumber="470">M. dongnaiensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="5ED6E89EFF82751012F013AE93A9F8FC" box="[958,1169,1782,1807]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF82751012F013AE93A9F8FC" box="[958,1169,1782,1807]" italics="true" pageId="10" pageNumber="470">M. queenslandicus</emphasis>
</taxonomicName>
in that the length of the anterior setiform process is equal to the claw, the anterior process is absent; while the anterior setiform process is longer than the claw in 
<taxonomicName id="5ED6E89EFF82751010D012669549F8A4" box="[414,625,1854,1879]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF82751010D012669549F8A4" box="[414,625,1854,1879]" italics="true" pageId="10" pageNumber="470">M. queenslandicus</emphasis>
</taxonomicName>
(
<figureCitation id="01ED8F98FF8275101331126795C0F8A4" box="[639,760,1854,1879]" captionStart="FIGURE 10" captionStartId="10.[151,250,1521,1543]" captionTargetBox="[160,1433,862,1490]" captionTargetId="figure@10.[151,1435,855,1499]" captionTargetPageId="10" captionText="FIGURE 10. Comparison between claw structures of M. tamoyoensis (A), M. queenslandicus (B) and Monographis dongnaiensis (C). A claw of M. tamoyoensis is referred from Nguyen Duy-Jacquemin &amp; Condé (1967), and a claw of M. queenslandicus is from Huynh &amp; Veenstra (2013). Arrows indicate the length of the anterior setiform processes. The dot lines compared the length of anterior setiform processes to the claws. a: anterior process, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. Not to scale." httpUri="https://zenodo.org/record/238050/files/figure.png" pageId="10" pageNumber="470">Figures 10</figureCitation>
&amp; 
<figureCitation id="01ED8F98FF827510125212669401F8A4" box="[796,825,1855,1879]" captionStart="FIGURE 11" captionStartId="11.[151,250,977,999]" captionTargetBox="[151,1435,519,955]" captionTargetId="figure@11.[151,1435,519,955]" captionTargetPageId="11" captionText="FIGURE 11. Images of SEM showed the difference in arrangement of processes in the claw structure of M. queenslandicus (A) (latro-posterior view) and Monographis dongnaiensis (B) (dorsal view). a: anterior process, c: claw, l: lamella process, lp: lateral process, p: posterior process, s: anterior setiform process. Arrows indicate position of the processes. Not to scale." httpUri="https://zenodo.org/record/238051/files/figure.png" pageId="10" pageNumber="470">11</figureCitation>
).
</paragraph>
<paragraph id="9969931DFF8275111189123B95E4FE23" blockId="10.[151,1436,1710,2023]" lastBlockId="11.[151,1437,151,464]" lastPageId="11" lastPageNumber="471" pageId="10" pageNumber="470">
<emphasis id="ABA24F0FFF8275101189123B9649F888" bold="true" box="[199,369,1890,1915]" pageId="10" pageNumber="470">DNA analysis.</emphasis>
A partial sequence with 1277 base pairs linear DNA was obtained from the DNA of five adult specimens of 
<taxonomicName id="5ED6E89EFF827510107E12DE96D1F86C" box="[304,489,1926,1951]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="470" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF827510107E12DE96D1F86C" box="[304,489,1926,1951]" italics="true" pageId="10" pageNumber="470">M. dongnaiensis</emphasis>
</taxonomicName>
(Genbank accession number 
<accessionNumber id="86850EFEFF827510127D12DF9492F86C" box="[819,938,1926,1951]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KP255446" pageId="10" pageNumber="470">KP255446</accessionNumber>
(accepted 
<date id="ED68B5DDFF827510156D12DE93E7F86C" box="[1059,1247,1927,1951]" pageId="10" pageNumber="470" value="2014-01-12">12 January 2014</date>
)). This truncated sequence excluded the first 27 base pairs and the base pairs from 1332 onward. Based on the results of a BLAST search (http://www.ncbi.nlm.nih.gov), the partial sequences of 18S ribosomal RNA gene from six closely-related penicillate millipedes were chosen for comparison with the sequence of 
<taxonomicName id="5ED6E89EFF837511157D15C093CFFF43" box="[1075,1271,151,176]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF837511157D15C093CFFF43" box="[1075,1271,151,176]" italics="true" pageId="11" pageNumber="471">M. dongnaiensis</emphasis>
</taxonomicName>
(
<accessionNumber id="86850EFEFF837511145F15CE92A9FF43" box="[1297,1425,151,176]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KP255446" pageId="11" pageNumber="471">KP255446</accessionNumber>
): 
<taxonomicName id="5ED6E89EFF83751111D915E59616FF26" ID-CoL="7PPF8" box="[151,302,188,213]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF83751111D915E59616FF26" box="[151,302,188,213]" italics="true" pageId="11" pageNumber="471">Monographis</emphasis>
</taxonomicName>
sp. from 
<collectingCountry id="E1C1D38DFF83751110D715E596E4FF26" box="[409,476,188,213]" name="China" pageId="11" pageNumber="471">China</collectingCountry>
(AY596371.1), 
<taxonomicName id="5ED6E89EFF83751113DD15E4945DFF26" box="[659,869,188,213]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF83751113DD15E4945DFF26" box="[659,869,188,213]" italics="true" pageId="11" pageNumber="471">M. queenslandicus</emphasis>
</taxonomicName>
(KF147166.1), two species from genus 
<taxonomicName id="5ED6E89EFF837511146715E592A4FF26" ID-CoL="7PTN7" box="[1321,1436,188,213]" class="Diplopoda" family="Polyxenidae" genus="Polyxenus" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF837511146715E592A4FF26" box="[1321,1436,188,213]" italics="true" pageId="11" pageNumber="471">Polyxenus</emphasis>
</taxonomicName>
(
<taxonomicName id="5ED6E89EFF83751111EE15B8962DFF0B" ID-CoL="786GT" box="[160,277,223,248]" class="Diplopoda" family="Polyxenidae" genus="Polyxenus" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="species" species="lagurus">
<emphasis id="ABA24F0FFF83751111EE15B8962DFF0B" box="[160,277,223,248]" italics="true" pageId="11" pageNumber="471">P. lagurus</emphasis>
</taxonomicName>
(EU368619.1) and 
<taxonomicName id="5ED6E89EFF837511134315B8958EFF0B" ID-CoL="6W6H3" box="[525,694,223,248]" class="Diplopoda" family="Polyxenidae" genus="Polyxenus" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="species" species="fasciculatus">
<emphasis id="ABA24F0FFF837511134315B8958EFF0B" box="[525,694,223,248]" italics="true" pageId="11" pageNumber="471">P. fasciculatus</emphasis>
</taxonomicName>
(AF173235.1)) and two millipedes are used as the outgroup (
<taxonomicName id="5ED6E89EFF83751111EE145D968AFEEF" ID-CoL="5D7HL" box="[160,434,260,285]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="species" species="neptunus">
<emphasis id="ABA24F0FFF83751111EE145D968AFEEF" box="[160,434,260,285]" italics="true" pageId="11" pageNumber="471">Zoosphaerium neptunus</emphasis>
</taxonomicName>
(FJ409972.1) &amp; 
<taxonomicName id="5ED6E89EFF83751113CE145D945AFEEE" ID-CoL="DY9" box="[640,866,260,285]" class="Diplopoda" family="Paradoxosomatidae" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="family">
<emphasis id="ABA24F0FFF83751113CE145D945AFEEE" box="[640,866,260,285]" italics="true" pageId="11" pageNumber="471">Paradoxosomatidae</emphasis>
</taxonomicName>
sp. (DQ666179.1)) in the phylogenetic analysis. The rooted consensus tree of 
<taxonomicName id="5ED6E89EFF83751110BE14709594FEB3" box="[496,684,295,320]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF83751110BE14709594FEB3" box="[496,684,295,320]" italics="true" pageId="11" pageNumber="471">M. dongnaiensis</emphasis>
</taxonomicName>
was generated by the bootstrap test with 1000 repetitions which yielded a strongly supported phylogenetic tree. There were four clear clades: 
<taxonomicName id="5ED6E89EFF83751115201414920AFE96" box="[1134,1330,332,357]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="species" species="dongnaiensis">
<emphasis id="ABA24F0FFF83751115201414920AFE96" box="[1134,1330,332,357]" italics="true" pageId="11" pageNumber="471">M. dongnaiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="5ED6E89EFF83751114331414967CFE7B" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="ABA24F0FFF83751114331414967CFE7B" italics="true" pageId="11" pageNumber="471">M. queenslandicus</emphasis>
</taxonomicName>
were a sister group, forming an independent clade. This clade branched with 
<taxonomicName id="5ED6E89EFF837511159414369249FE7B" ID-CoL="7PPF8" box="[1242,1393,367,392]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF837511159414369249FE7B" box="[1242,1393,367,392]" italics="true" pageId="11" pageNumber="471">Monographis</emphasis>
</taxonomicName>
sp. (AY596371.1) forming a monophyletic group. This group of 
<taxonomicName id="5ED6E89EFF837511121014CD94CDFE5E" ID-CoL="7PPF8" box="[862,1013,404,429]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF837511121014CD94CDFE5E" box="[862,1013,404,429]" italics="true" pageId="11" pageNumber="471">Monographis</emphasis>
</taxonomicName>
species was separated from the two 
<taxonomicName id="5ED6E89EFF83751111D914EE9632FE23" ID-CoL="7PTN7" box="[151,266,439,464]" class="Diplopoda" family="Polyxenidae" genus="Polyxenus" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="ABA24F0FFF83751111D914EE9632FE23" box="[151,266,439,464]" italics="true" pageId="11" pageNumber="471">Polyxenus</emphasis>
</taxonomicName>
species, as was the outgroup (
<figureCitation id="01ED8F98FF837511131314EE95F6FE23" box="[605,718,439,464]" captionStart="FIGURE 12" captionStartId="12.[151,250,1872,1894]" captionTargetBox="[201,1385,210,1827]" captionTargetId="figure@12.[186,1395,195,1836]" captionTargetPageId="12" captionText="FIGURE 12. Rooted consensus tree of Monographis dongnaiensis. Bootstrap test yielded a strong support for the phylogenetic tree. The partial sequences 18 S RNA gene of Polyxenus lagurus (EU 368619.1), Polyxenus fasciculatus (AF 173235.1), Monographis dongnaiensis (KP 255446.1), Monographis queenslandicus (KF 147166.1), Monographis sp. from China (AY 596371.1), Zoosphaerium neptunus (FJ 409972.1) and Paradoxosomatidae sp. (DQ 666179.1) were used in the phylogenetic analysis. The accession numbers of the used species are from Genbank." httpUri="https://zenodo.org/record/288711/files/figure.png" pageId="11" pageNumber="471">Figure 12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>