1238 lines
94 KiB
XML
1238 lines
94 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.96.50821" ID-GBIF-Dataset="fb568d02-c3d7-48b9-9729-2f85ea601db7" ID-GBIF-Taxon="164363161" ID-Pensoft-Pub="1860-0743-1-217" ID-Pensoft-UUID="930A6280AF085CA9B6F8AC6CE0E1C120" ID-ZooBank="0B80FF1B5ED7430BA471A5DE02E8E6D3" LSID="urn:lsid:zoobank.org:act:48DA4500-2806-491E-8AD8-D2C830AF39F4" ModsDocID="1860-0743-1-217" checkinTime="1590646559532" checkinUser="pensoft" docAuthor="Gasiorek, Piotr, Voncina, Katarzyna, Degma, Peter & Michalczyk, Lukasz" docDate="2020" docId="048ADD2264FA50DB8E10D4274F347B4C" docLanguage="en" docName="ZoosystEvol 96(1): 217-236" docOrigin="Zoosystematics and Evolution 96 (1)" docSource="http://dx.doi.org/10.3897/zse.96.50821" docTitle="Bryodelphax nigripunctatus Degma, Gasiorek, Voncina & Michalczyk 2020, sp. nov." docType="treatment" docUuid="48DA4500-2806-491E-8AD8-D2C830AF39F4" docUuidSource="ZooBank" docVersion="4" id="930A6280AF085CA9B6F8AC6CE0E1C120" lastPageNumber="217" masterDocId="930A6280AF085CA9B6F8AC6CE0E1C120" masterDocTitle="Small is beautiful: the first phylogenetic analysis of Bryodelphax Thulin, 1928 (Heterotardigrada, Echiniscidae)" masterLastPageNumber="236" masterPageNumber="217" pageId="0" pageNumber="217" updateTime="1634708808286" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Small is beautiful: the first phylogenetic analysis of Bryodelphax Thulin, 1928 (Heterotardigrada, Echiniscidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Gasiorek, Piotr</mods:namePart>
|
||
<mods:affiliation>Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
|
||
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2814-8117</mods:nameIdentifier>
|
||
<mods:nameIdentifier type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Voncina, Katarzyna</mods:namePart>
|
||
<mods:affiliation>Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Degma, Peter</mods:namePart>
|
||
<mods:affiliation>Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina, Ilkovicova 6 / B 1, 84215, Bratislava, Slovakia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Michalczyk, Lukasz</mods:namePart>
|
||
<mods:affiliation>Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
|
||
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2912-4870</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zoosystematics and Evolution</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2020</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>96</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>217</mods:start>
|
||
<mods:end>236</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zse.96.50821</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.50821</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1860-0743-1-217</mods:identifier>
|
||
<mods:identifier type="ZooBank">0B80FF1B5ED7430BA471A5DE02E8E6D3</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">930A6280AF085CA9B6F8AC6CE0E1C120</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="164363161" LSID="urn:lsid:zoobank.org:act:48DA4500-2806-491E-8AD8-D2C830AF39F4" httpUri="http://treatment.plazi.org/id/048ADD2264FA50DB8E10D4274F347B4C" lastPageNumber="217" pageId="0" pageNumber="217">
|
||
<subSubSection pageId="0" pageNumber="217" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<taxonomicName LSID="048ADD22-64FA-50DB-8E10-D4274F347B4C" authority="Degma, Gąsiorek, Vončina & Michalczyk" authorityName="Degma, Gasiorek, Voncina & Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax nigripunctatus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="nigripunctatus" status="sp. nov.">
|
||
Bryodelphax nigripunctatus Degma,
|
||
<normalizedToken originalValue="Gąsiorek">Gasiorek</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="Vončina">Voncina</normalizedToken>
|
||
& Michalczyk
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="217">sp. nov.</taxonomicNameLabel>
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">Figures 7</figureCitation>
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of adult male of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - dorsal view; B - lateral view. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217">, 8</figureCitation>
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Detailed morphology of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - frontal part of the body; B - central part of the body; C - caudal part of the body. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure9" httpUri="https://binary.pensoft.net/fig/413956" pageId="0" pageNumber="217">, 9</figureCitation>
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Details of cuticular sculpturing of Bryodelphax nigripunctatus sp. nov.: A - microscope focused on the epicuticular granules (allotype, NCM); B - microscope focused on the intracuticular pillars (allotype, NCM); C - frontal part of the body (allotype, PCM); D - frontal part of the body (holotype, PCM). Scale bars in µm." figureDoi="10.3897/zse.96.50821.figure10" httpUri="https://binary.pensoft.net/fig/413957" pageId="0" pageNumber="217">, 10</figureCitation>
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Variability of the dentate collar IV in Bryodelphax nigripunctatus sp. nov. (paratypes, PCM). Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure11" httpUri="https://binary.pensoft.net/fig/413958" pageId="0" pageNumber="217">, 11</figureCitation>
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Schematic arrangement of the ventral plates in all members of the weglarskae group (species are arranged in order of the increasing reduction of the ventral armature). Known cases of ontogenetic variability and sexual dimorphism are depicted. Following species taken from Kaczmarek et al. (2012): B. aaseae, B. iohannis, B. parvuspolaris, B. sinensis and B. weglarskae; Gasiorek et al. (2017 a): B. maculatus; Gasiorek & Degma (2018): B. instabilis." figureDoi="10.3897/zse.96.50821.figure12" httpUri="https://binary.pensoft.net/fig/413959" pageId="0" pageNumber="217">, 12</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurements [in µm] of selected morphological structures of mature females of Bryodelphax nigripunctatus sp. nov. mounted in Hoyer's medium (N - number of specimens / structures measured, Range refers to the smallest and the largest structure amongst all measured specimens; SD - standard deviation; sp - the ratio of the length of a given structure to the length of the scapular plate expressed as a percentage)." pageId="0" pageNumber="217">Tables 5</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Measurements [in µm] of selected morphological structures of mature males of Bryodelphax nigripunctatus sp. nov. mounted in Hoyer's medium (N - number of specimens / structures measured, Range refers to the smallest and the largest structure amongst all measured specimens; SD - standard deviation; sp - the ratio of the length of a given structure to the length of the scapular plate expressed as a percentage)." pageId="0" pageNumber="217">6</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="217" type="type material">
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<taxonomicName genus="Locus" lsidName="Locus typicus" pageId="0" pageNumber="217" rank="species" species="typicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Locus typicus</emphasis>
|
||
</taxonomicName>
|
||
and type material.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<geoCoordinate degrees="39" direction="north" minutes="57" orientation="latitude" precision="15" seconds="00" value="39.95">39°57'00"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="3" direction="east" minutes="10" orientation="longitude" precision="15" seconds="50" value="3.1805556">3°10'50"E</geoCoordinate>
|
||
, 160 m a.s.l.; near the road above Cala Figuera beach, Cap de Formentor, NE Mallorca, the Balearic Islands, Spain. Holotype (adult female; together with one male paratype in slide 716/45), allotype (adult male; slide 716/9) and 30 paratypes (9 females, 14 males, 3 specimens of unknown sex, 2 juveniles and 2 larvae; slides 716/1-5, 10, 13, 17 -20, 25, 27-29, 32-34, 41, 45, 47-48, 50-51) deposited in the Department of Zoology, Comenius University in Bratislava; 13 paratypes (6 females, 6 males and one specimen of unknown sex; slides 716/26, 38, 40, 42-44, 46, 49) deposited in the Institute of Zoology and Biomedical Research, Jagiellonian University; 9 paratypes (4 females and 5 males; slides 716/6-8, 11-12, 22-24, 35) deposited in the Natural History Museum of Denmark, University of Copenhagen; 9 paratypes (4 females and 5 males; slides 716/14-16, 21, 31, 36-37) deposited in the Department of Animal Biology, University of Catania. Single paratype (male) mounted on a SEM stub (14.19) deposited in the Institute of Zoology and Biomedical Research, Jagiellonian University. Eight specimens used for DNA extraction.
|
||
<taxonomicName authorityName="Degma, Gasiorek, Voncina & Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax nigripunctatus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="nigripunctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax nigripunctatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. was not accompanied by other species in the sample.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="C8DD6F01157095B3023157F71BF200AE" httpUri="http://table.plazi.org/id/C8DD6F01157095B3023157F71BF200AE" pageId="0" pageNumber="217" start="Table 5" startId="T5">
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<emphasis bold="true" pageId="0" pageNumber="217">Table 5.</emphasis>
|
||
Measurements [in
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
] of selected morphological structures of mature females of
|
||
<taxonomicName authorityName="Degma, Gasiorek, Voncina & Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax nigripunctatus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="nigripunctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax nigripunctatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. mounted in
|
||
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
|
||
medium (
|
||
<emphasis bold="true" pageId="0" pageNumber="217">N</emphasis>
|
||
- number of specimens/structures measured,
|
||
<emphasis bold="true" pageId="0" pageNumber="217">Range</emphasis>
|
||
refers to the smallest and the largest structure amongst all measured specimens;
|
||
<emphasis bold="true" pageId="0" pageNumber="217">SD</emphasis>
|
||
- standard deviation;
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
- the ratio of the length of a given structure to the length of the scapular plate expressed as a percentage).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<table pageId="0" pageNumber="217">
|
||
<tr pageId="0" pageNumber="217">
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="2">Character</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="2">N</th>
|
||
<th colspan="6" pageId="0" pageNumber="217" rowspan="1">Range</th>
|
||
<th colspan="2" pageId="0" pageNumber="217" rowspan="1">Mean</th>
|
||
<th colspan="2" pageId="0" pageNumber="217" rowspan="1">SD</th>
|
||
<th colspan="2" pageId="0" pageNumber="217" rowspan="1">Holotype</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<th colspan="3" pageId="0" pageNumber="217" rowspan="1">
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
</th>
|
||
<th colspan="3" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Body length</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">114</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">144</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">562</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">689</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">126</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">645</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">38</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">131</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">657</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Scapular plate length</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">18.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">21.7</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">19.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">19.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="14" pageId="0" pageNumber="217" rowspan="1">Head appendages lengths</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
Cirrus
|
||
<emphasis italics="true" pageId="0" pageNumber="217">internus</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">11</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">6.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">32.9</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">42.8</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">37.4</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">3.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">37.4</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Cephalic papilla</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">14</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">12.6</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">19.7</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">15.9</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">1.9</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">16.5</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
Cirrus
|
||
<emphasis italics="true" pageId="0" pageNumber="217">externus</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">12</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">10.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">12.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">54.1</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">66.9</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">11.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">60.1</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">4.4</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">12.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">61.4</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Clava</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">10</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">12.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">16.9</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">14.8</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">1.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">15.0</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
Cirrus
|
||
<emphasis italics="true" pageId="0" pageNumber="217">A</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">24.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">31.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">120.7</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">163.7</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">28.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">143.6</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">11.0</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">26.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">132.4</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
Cirrus
|
||
<emphasis italics="true" pageId="0" pageNumber="217">A</emphasis>
|
||
/Body length ratio
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">20%</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">25%</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">22%</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2%</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">20%</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="14" pageId="0" pageNumber="217" rowspan="1">Body appendages lengths</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Papilla on leg IV length</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">7.2</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">12.5</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">9.5</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">2.0</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">7.6</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Number of teeth on the collar</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">10</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="14" pageId="0" pageNumber="217" rowspan="1">Claw heights</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw I</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">12</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">35.1</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">44.4</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">39.8</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">3.1</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">36.0</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw II</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">36.1</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">44.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">39.6</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">2.8</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">38.8</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw III</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">13</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">6.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">36.1</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">46.2</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">39.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">3.1</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">38.1</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw IV</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">38.1</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">48.0</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">43.0</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">3.5</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">?</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">?</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption ID-Table-UUID="619F38CCA267D24DDDA634C56F651D24" httpUri="http://table.plazi.org/id/619F38CCA267D24DDDA634C56F651D24" pageId="0" pageNumber="217" start="Table 6" startId="T6">
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<emphasis bold="true" pageId="0" pageNumber="217">Table 6.</emphasis>
|
||
Measurements [in
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
] of selected morphological structures of mature males of
|
||
<taxonomicName authorityName="Degma, Gasiorek, Voncina & Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax nigripunctatus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="nigripunctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax nigripunctatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. mounted in
|
||
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
|
||
medium (
|
||
<emphasis bold="true" pageId="0" pageNumber="217">N</emphasis>
|
||
- number of specimens/structures measured,
|
||
<emphasis bold="true" pageId="0" pageNumber="217">Range</emphasis>
|
||
refers to the smallest and the largest structure amongst all measured specimens;
|
||
<emphasis bold="true" pageId="0" pageNumber="217">SD</emphasis>
|
||
- standard deviation;
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
- the ratio of the length of a given structure to the length of the scapular plate expressed as a percentage).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<table pageId="0" pageNumber="217">
|
||
<tr pageId="0" pageNumber="217">
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="2">Character</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="2">N</th>
|
||
<th colspan="6" pageId="0" pageNumber="217" rowspan="1">Range</th>
|
||
<th colspan="2" pageId="0" pageNumber="217" rowspan="1">Mean</th>
|
||
<th colspan="2" pageId="0" pageNumber="217" rowspan="1">SD</th>
|
||
<th colspan="2" pageId="0" pageNumber="217" rowspan="1">Allotype</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<th colspan="3" pageId="0" pageNumber="217" rowspan="1">
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
</th>
|
||
<th colspan="3" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Body length</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">106</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">134</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">618</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">725</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">123</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">685</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">30</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">130</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">704</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Scapular plate length</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">16.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">19.5</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">17.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">18.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="14" pageId="0" pageNumber="217" rowspan="1">Head appendages lengths</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
Cirrus
|
||
<emphasis italics="true" pageId="0" pageNumber="217">internus</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">13</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">6.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">9.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">35.8</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">51.7</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">42.8</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">4.6</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">46.0</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Cephalic papilla</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">14</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">19.0</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">32.0</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">4.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">25.5</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">3.9</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">28.1</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
Cirrus
|
||
<emphasis italics="true" pageId="0" pageNumber="217">externus</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">13</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">12.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">70.6</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">85.8</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">14.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">78.6</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">4.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">13.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">75.4</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Clava</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">11</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">4.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">17.6</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">24.2</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">21.2</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">1.9</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">4.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">21.9</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
Cirrus
|
||
<emphasis italics="true" pageId="0" pageNumber="217">A</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">26.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">32.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">141.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">187.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">29.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">163.8</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">11.4</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">30.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">163.4</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
Cirrus
|
||
<emphasis italics="true" pageId="0" pageNumber="217">A</emphasis>
|
||
/Body length ratio
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">21%</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">26%</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">24%</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2%</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">23%</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="14" pageId="0" pageNumber="217" rowspan="1">Body appendages lengths</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Papilla on leg IV length</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">9.7</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">12.8</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">11.4</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">1.0</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">10.9</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Number of teeth on the collar</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">12</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="14" pageId="0" pageNumber="217" rowspan="1">Claw heights</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw I</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">11</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">37.6</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">50.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">45.1</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">3.4</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">44.8</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw II</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">10</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">41.2</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">48.5</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">44.7</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">2.5</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">47.9</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw III</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">14</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">9.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">38.5</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">49.5</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">44.7</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">3.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">9.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">48.6</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="217">
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw IV</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">9.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">43.6</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">52.6</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">48.3</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">2.8</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">?</td>
|
||
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">?</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="217" type="etymology">
|
||
<paragraph pageId="0" pageNumber="217">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
From Latin
|
||
<emphasis italics="true" pageId="0" pageNumber="217">niger</emphasis>
|
||
= black +
|
||
<emphasis italics="true" pageId="0" pageNumber="217">punctum</emphasis>
|
||
= dot, spot. The name underlines evident epicuticular granules appearing dark in PCM and contrasting with other elements of dorsal sculpture. Adjective in the nominative singular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="217" type="adults">
|
||
<paragraph pageId="0" pageNumber="217">Adults.</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
Body translucent without distinct colour and usually stout in females and more slender in males (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of adult male of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - dorsal view; B - lateral view. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217">8</figureCitation>
|
||
),
|
||
<emphasis italics="true" pageId="0" pageNumber="217">bs</emphasis>
|
||
= 43.0-49.2% (x̅ = 45.6%, N = 8) in females and 37.5-44.7% (x̅ = 39.8%, N = 10) in males. Eyes absent or not visible after mounting in
|
||
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
|
||
medium. Cephalic papillae and clavae elliptical with rounded apex. Cephalic papilla is relatively broader in males than in females. Both cephalic papillae and primary clavae relatively longer in males than in females (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Details of cuticular sculpturing of Bryodelphax nigripunctatus sp. nov.: A - microscope focused on the epicuticular granules (allotype, NCM); B - microscope focused on the intracuticular pillars (allotype, NCM); C - frontal part of the body (allotype, PCM); D - frontal part of the body (holotype, PCM). Scale bars in µm." figureDoi="10.3897/zse.96.50821.figure10" httpUri="https://binary.pensoft.net/fig/413957" pageId="0" pageNumber="217">10C, D</figureCitation>
|
||
; compare the
|
||
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
|
||
of the cephalic papilla and clava in Tables
|
||
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurements [in µm] of selected morphological structures of mature females of Bryodelphax nigripunctatus sp. nov. mounted in Hoyer's medium (N - number of specimens / structures measured, Range refers to the smallest and the largest structure amongst all measured specimens; SD - standard deviation; sp - the ratio of the length of a given structure to the length of the scapular plate expressed as a percentage)." httpUri="http://table.plazi.org/id/C8DD6F01157095B3023157F71BF200AE" pageId="0" pageNumber="217" tableUuid="C8DD6F01157095B3023157F71BF200AE">5</tableCitation>
|
||
,
|
||
<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Measurements [in µm] of selected morphological structures of mature males of Bryodelphax nigripunctatus sp. nov. mounted in Hoyer's medium (N - number of specimens / structures measured, Range refers to the smallest and the largest structure amongst all measured specimens; SD - standard deviation; sp - the ratio of the length of a given structure to the length of the scapular plate expressed as a percentage)." httpUri="http://table.plazi.org/id/619F38CCA267D24DDDA634C56F651D24" pageId="0" pageNumber="217" tableUuid="619F38CCA267D24DDDA634C56F651D24">6</tableCitation>
|
||
). Cirri
|
||
<emphasis italics="true" pageId="0" pageNumber="217">interni</emphasis>
|
||
always shorter than cirri
|
||
<emphasis italics="true" pageId="0" pageNumber="217">externi</emphasis>
|
||
and both with poorly-developed cirrophores. Cirri
|
||
<emphasis italics="true" pageId="0" pageNumber="217">A</emphasis>
|
||
reach around 1/5-1/4 of the total body length (Tables
|
||
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurements [in µm] of selected morphological structures of mature females of Bryodelphax nigripunctatus sp. nov. mounted in Hoyer's medium (N - number of specimens / structures measured, Range refers to the smallest and the largest structure amongst all measured specimens; SD - standard deviation; sp - the ratio of the length of a given structure to the length of the scapular plate expressed as a percentage)." httpUri="http://table.plazi.org/id/C8DD6F01157095B3023157F71BF200AE" pageId="0" pageNumber="217" tableUuid="C8DD6F01157095B3023157F71BF200AE">5</tableCitation>
|
||
,
|
||
<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Measurements [in µm] of selected morphological structures of mature males of Bryodelphax nigripunctatus sp. nov. mounted in Hoyer's medium (N - number of specimens / structures measured, Range refers to the smallest and the largest structure amongst all measured specimens; SD - standard deviation; sp - the ratio of the length of a given structure to the length of the scapular plate expressed as a percentage)." httpUri="http://table.plazi.org/id/619F38CCA267D24DDDA634C56F651D24" pageId="0" pageNumber="217" tableUuid="619F38CCA267D24DDDA634C56F651D24">6</tableCitation>
|
||
). Unappendaged. Cuticular sculpture consists of large epicuticular granules, true pores and intra-cuticular pillars (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Details of cuticular sculpturing of Bryodelphax nigripunctatus sp. nov.: A - microscope focused on the epicuticular granules (allotype, NCM); B - microscope focused on the intracuticular pillars (allotype, NCM); C - frontal part of the body (allotype, PCM); D - frontal part of the body (holotype, PCM). Scale bars in µm." figureDoi="10.3897/zse.96.50821.figure10" httpUri="https://binary.pensoft.net/fig/413957" pageId="0" pageNumber="217">10</figureCitation>
|
||
). In PCM, these structures appear, respectively, as conspicuous large dark spots, smaller bright spots and fine dark and dense punctuation (pseudogranulation). Epicuticular granules of irregular shape and size (up to ca. 1.6
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
) are merged together (as visible in SEM, Figs
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of adult male of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - dorsal view; B - lateral view. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217">8</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Details of cuticular sculpturing of Bryodelphax nigripunctatus sp. nov.: A - microscope focused on the epicuticular granules (allotype, NCM); B - microscope focused on the intracuticular pillars (allotype, NCM); C - frontal part of the body (allotype, PCM); D - frontal part of the body (holotype, PCM). Scale bars in µm." figureDoi="10.3897/zse.96.50821.figure10" httpUri="https://binary.pensoft.net/fig/413957" pageId="0" pageNumber="217">10</figureCitation>
|
||
) and arranged in rows along the margins of all plates, although they are least visible or absent in posterior median plate 1 and posterior median plate 2. Moreover, in the cervical (neck) plate, the row or a double row of granules is also present along its transverse axis. Granules in rows appear as dark spots in PCM (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7A-C</figureCitation>
|
||
). Similar rows of granules cover also folds which create the faceting of the scapular and caudal plate: median and two lateral longitudinal folds (at the level of cirrophores
|
||
<emphasis italics="true" pageId="0" pageNumber="217">A</emphasis>
|
||
) together with 3-4 posterior transverse folds on the scapular plate and two longitudinal folds dividing the caudal plate into three parallel portions (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of adult male of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - dorsal view; B - lateral view. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217">8</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Detailed morphology of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - frontal part of the body; B - central part of the body; C - caudal part of the body. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure9" httpUri="https://binary.pensoft.net/fig/413956" pageId="0" pageNumber="217">9A, C</figureCitation>
|
||
). Finally, short longitudinal rows of granules divide both paired segmental plates and posterior portions of anterior m1-2 plates into left and right portions (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of adult male of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - dorsal view; B - lateral view. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217">8</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Detailed morphology of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - frontal part of the body; B - central part of the body; C - caudal part of the body. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure9" httpUri="https://binary.pensoft.net/fig/413956" pageId="0" pageNumber="217">9B</figureCitation>
|
||
). Scattered granules also irregularly cover the surface of the cephalic, paired, caudal, anterior m1-2 and anterior parts of scapular plates (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of adult male of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - dorsal view; B - lateral view. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217">8</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Detailed morphology of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - frontal part of the body; B - central part of the body; C - caudal part of the body. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure9" httpUri="https://binary.pensoft.net/fig/413956" pageId="0" pageNumber="217">9</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Details of cuticular sculpturing of Bryodelphax nigripunctatus sp. nov.: A - microscope focused on the epicuticular granules (allotype, NCM); B - microscope focused on the intracuticular pillars (allotype, NCM); C - frontal part of the body (allotype, PCM); D - frontal part of the body (holotype, PCM). Scale bars in µm." figureDoi="10.3897/zse.96.50821.figure10" httpUri="https://binary.pensoft.net/fig/413957" pageId="0" pageNumber="217">10A</figureCitation>
|
||
). Round, focusable pores (0.3-0.4
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
in diameter) are unequally distributed on dorsal plates in spaces between the scattered granules, on pedal plates IV and between transverse rows of granules in the scapular plate, but they are absent on the rows of granules (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of adult male of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - dorsal view; B - lateral view. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217">8</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Detailed morphology of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - frontal part of the body; B - central part of the body; C - caudal part of the body. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure9" httpUri="https://binary.pensoft.net/fig/413956" pageId="0" pageNumber="217">9A-C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Details of cuticular sculpturing of Bryodelphax nigripunctatus sp. nov.: A - microscope focused on the epicuticular granules (allotype, NCM); B - microscope focused on the intracuticular pillars (allotype, NCM); C - frontal part of the body (allotype, PCM); D - frontal part of the body (holotype, PCM). Scale bars in µm." figureDoi="10.3897/zse.96.50821.figure10" httpUri="https://binary.pensoft.net/fig/413957" pageId="0" pageNumber="217">10B</figureCitation>
|
||
). The density of pores varies between the sexes and elements of armour, with the largest pores present on the segmental plate II, anterior m2 and the scapular plate (21-40 pores/100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2, x̅ = 29, 14-28 pores/100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2, x̅ = 22 and 18-31pores/100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2, x̅ = 26, respectively, N = 15 in females and 7-35 pores/100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2, x̅ = 29, 0-34 pores/100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2, x̅ = 25 and 11-33 pores/100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2, x̅ = 25, respectively; N = 15 in males) and lowest density on caudal plate (16-27 pores/100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2, x̅ = 22 in females and 7-28 pores/100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2, x̅ = 20 in males; N = 15). On the scapular plate (in the area delimited with lateral longitudinal rows of granules), lines of pores tend to alternate with transverse rows of granules (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7A-C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of adult male of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - dorsal view; B - lateral view. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217">8A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Detailed morphology of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - frontal part of the body; B - central part of the body; C - caudal part of the body. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure9" httpUri="https://binary.pensoft.net/fig/413956" pageId="0" pageNumber="217">9A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Details of cuticular sculpturing of Bryodelphax nigripunctatus sp. nov.: A - microscope focused on the epicuticular granules (allotype, NCM); B - microscope focused on the intracuticular pillars (allotype, NCM); C - frontal part of the body (allotype, PCM); D - frontal part of the body (holotype, PCM). Scale bars in µm." figureDoi="10.3897/zse.96.50821.figure10" httpUri="https://binary.pensoft.net/fig/413957" pageId="0" pageNumber="217">10A, B</figureCitation>
|
||
). Regularly distributed round intra-cuticular pillars (0.1-0.2
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
in diameter) reinforce the entire cuticle (also under the granules), but they are well-visible only in the cephalic, scapular, both paired segmental, caudal, anterior median 1 and anterior median 2 plates (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Details of cuticular sculpturing of Bryodelphax nigripunctatus sp. nov.: A - microscope focused on the epicuticular granules (allotype, NCM); B - microscope focused on the intracuticular pillars (allotype, NCM); C - frontal part of the body (allotype, PCM); D - frontal part of the body (holotype, PCM). Scale bars in µm." figureDoi="10.3897/zse.96.50821.figure10" httpUri="https://binary.pensoft.net/fig/413957" pageId="0" pageNumber="217">10B</figureCitation>
|
||
), as well as on pedal plates IV. On the remaining cuticle, they are weakly (venter) or scarcely (legs) detectable.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<emphasis bold="true" pageId="0" pageNumber="217">Figure 7.</emphasis>
|
||
Habitus of adults of
|
||
<taxonomicName authorityName="Degma, Gasiorek, Voncina & Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax nigripunctatus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="nigripunctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax nigripunctatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (PCM):
|
||
<emphasis bold="true" pageId="0" pageNumber="217">A</emphasis>
|
||
- female (holotype, dorsolateral view);
|
||
<emphasis bold="true" pageId="0" pageNumber="217">B</emphasis>
|
||
- male (allotype, dorsal view);
|
||
<emphasis bold="true" pageId="0" pageNumber="217">C</emphasis>
|
||
- female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs);
|
||
<emphasis bold="true" pageId="0" pageNumber="217">D</emphasis>
|
||
- male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<emphasis bold="true" pageId="0" pageNumber="217">Figure 8.</emphasis>
|
||
Habitus of adult male of
|
||
<taxonomicName authorityName="Degma, Gasiorek, Voncina & Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax nigripunctatus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="nigripunctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax nigripunctatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (paratype, SEM):
|
||
<emphasis bold="true" pageId="0" pageNumber="217">A</emphasis>
|
||
- dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="217">B</emphasis>
|
||
- lateral view. Scale bars: 50
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zse.96.50821.figure9" httpUri="https://binary.pensoft.net/fig/413956" pageId="0" pageNumber="217" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<emphasis bold="true" pageId="0" pageNumber="217">Figure 9.</emphasis>
|
||
Detailed morphology of
|
||
<taxonomicName authorityName="Degma, Gasiorek, Voncina & Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax nigripunctatus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="nigripunctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax nigripunctatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (paratype, SEM):
|
||
<emphasis bold="true" pageId="0" pageNumber="217">A</emphasis>
|
||
- frontal part of the body;
|
||
<emphasis bold="true" pageId="0" pageNumber="217">B</emphasis>
|
||
- central part of the body;
|
||
<emphasis bold="true" pageId="0" pageNumber="217">C</emphasis>
|
||
- caudal part of the body. Scale bars: 10
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zse.96.50821.figure10" httpUri="https://binary.pensoft.net/fig/413957" pageId="0" pageNumber="217" start="Figure 10" startId="F10">
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<emphasis bold="true" pageId="0" pageNumber="217">Figure 10.</emphasis>
|
||
Details of cuticular sculpturing of
|
||
<taxonomicName authorityName="Degma, Gasiorek, Voncina & Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax nigripunctatus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="nigripunctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax nigripunctatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.:
|
||
<emphasis bold="true" pageId="0" pageNumber="217">A</emphasis>
|
||
- microscope focused on the epicuticular granules (allotype, NCM);
|
||
<emphasis bold="true" pageId="0" pageNumber="217">B</emphasis>
|
||
- microscope focused on the intracuticular pillars (allotype, NCM);
|
||
<emphasis bold="true" pageId="0" pageNumber="217">C</emphasis>
|
||
- frontal part of the body (allotype, PCM);
|
||
<emphasis bold="true" pageId="0" pageNumber="217">D</emphasis>
|
||
- frontal part of the body (holotype, PCM). Scale bars in
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
Cephalic plate with an anterior chalice-like incision. Each segmental and median plate consists of the anterior and the posterior portion separated each from other with a transverse bright poreless stripe in PCM. Therefore, paired segmental plates are subdivided into the narrower anterior (ca. 1/3-2/5 of the plate length) and the wider posterior portions, trapezoidal anterior median plate 1 is subdivided just behind its anterior margin, pentagonal anterior m2 (the largest amongst the median plates) is divided at approximately equally-long portions, triangular anterior portion of median plate 3 is ca. two times as long (along median body axis) as the posterior one with rounded posterior margin (dividing transverse line of anterior median plates 2-3 correspond with posterior corners of paired segmental plates). Pentagonal posterior median plates 1-2 subdivided at portions of about same lengths. On each body side, the first two pairs of supplementary lateral platelets are connected with anterior and posterior median plates 1-2 and the third pair is connected with the posterior portion of m3 and with the anterior edge of caudal plate. Anterior platelets of each pair have very distinctly-thickened lateral (lower) margins (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of adult male of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - dorsal view; B - lateral view. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217">8</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Detailed morphology of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - frontal part of the body; B - central part of the body; C - caudal part of the body. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure9" httpUri="https://binary.pensoft.net/fig/413956" pageId="0" pageNumber="217">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
Venter with transverse rows of weakly-developed plates unevenly sculptured with epicuticular granules similar to those on the dorsal plates, but a bit smaller. There are three rows of ventral plates in females (the plate formula III:2-2-1) and two rows in males (II:2-2) (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7D</figureCitation>
|
||
). The outer surface of legs with a narrow well-visible proximal pulvinus and a wide weak distal pedal plate placed in the central part of the leg. A single row of small epicuticular granules (similar to those on ventral plates) on the distal edge of pulvini in legs I-III (rarely, the second row also on their proximal edge). Pedal plates I-III sculptured usually with three (sometimes with more) transverse rows of epicuticular granules, which can be either shortened or connected at their ends (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7A-C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of adult male of Bryodelphax nigripunctatus sp. nov. (paratype, SEM): A - dorsal view; B - lateral view. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure8" httpUri="https://binary.pensoft.net/fig/413955" pageId="0" pageNumber="217">8B</figureCitation>
|
||
). The pedal plate IV sculptured with distinct intra-cuticular pillars and scattered pores and distally hemmed with dentate collar. The collar with sharp teeth, always longer than the width of their bases and with the distance between teeth similar to their basal widths, although some pairs of teeth can be merged (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Variability of the dentate collar IV in Bryodelphax nigripunctatus sp. nov. (paratypes, PCM). Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure11" httpUri="https://binary.pensoft.net/fig/413958" pageId="0" pageNumber="217">11</figureCitation>
|
||
). Papilla or spine on legs I-III absent, papilla on legs IV well developed. Claws slender, claws IV always slightly longer than claws I-III. External claws smooth, internal ones with a small spur pointing downwards and placed very close to the claw bases (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7A-C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of adults of Bryodelphax nigripunctatus sp. nov. (PCM): A - female (holotype, dorsolateral view); B - male (allotype, dorsal view); C - female (paratype, lateral view; Roman numerals signify epicuticular belts of granules on legs); D - male (allotype, ventral view, Roman numerals point out reduced ventral armature; insert with the claws II, arrowhead indicates spur on internal claw). Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure7" httpUri="https://binary.pensoft.net/fig/413954" pageId="0" pageNumber="217">7D</figureCitation>
|
||
, insert).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.96.50821.figure11" httpUri="https://binary.pensoft.net/fig/413958" pageId="0" pageNumber="217" start="Figure 11" startId="F11">
|
||
<paragraph pageId="0" pageNumber="217">
|
||
<emphasis bold="true" pageId="0" pageNumber="217">Figure 11.</emphasis>
|
||
Variability of the dentate collar IV in
|
||
<taxonomicName authorityName="Degma, Gasiorek, Voncina & Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax nigripunctatus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="nigripunctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax nigripunctatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (paratypes, PCM). Scale bars: 10
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="217" type="juveniles">
|
||
<paragraph pageId="0" pageNumber="217">Juveniles.</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
In appearance as adults, but smaller (111-112
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
) and with ventral plates just marked with rows of granules. Selected measurements of a shorter specimen: cephalic papilla 3.1
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
, scapular plate 14.9
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
long, claws I-III 5.1-5.6 and claws IV 6.5
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
long.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="217" type="larvae">
|
||
<paragraph pageId="0" pageNumber="217">Larvae.</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
83-85
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
long. Dorsal plates (especially median ones) mostly with poorly-delineated margins, supplementary lateral platelets absent. Epicuticular granules less numerous than in adults, concentrated mainly on posterior margins of the cephalic, scapular, both paired and caudal plates. Cuticular pores less numerous than in adults, but intracuticular pillars, stripes of granules on the outer surface of legs, papilla on legs IV and dentate collar IV well developed. Ventral plates not visible in laterally orientated larvae. Claws with spurs formed as in adults. Some measurements of shorter specimen: cephalic papilla 2.5
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
, claws II-III 4.6- 5.3 and claws IV 6.4
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
long.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="217" type="eggs">
|
||
<paragraph pageId="0" pageNumber="217">Eggs.</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">Not found.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="217" type="dna sequences">
|
||
<paragraph pageId="0" pageNumber="217">DNA sequences.</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">Two 18S rRNA haplotypes (MT333472-3), single 28S rRNA haplotype (MT333465).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="217" type="phenotypic differential diagnosis">
|
||
<paragraph pageId="0" pageNumber="217">Phenotypic differential diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
Having ventral plates,
|
||
<taxonomicName authorityName="Degma, Gasiorek, Voncina & Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax nigripunctatus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="nigripunctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax nigripunctatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. belongs to the
|
||
<taxonomicName baseAuthorityName="Pilato" baseAuthorityYear="1972" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="weglarskae" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="weglarskae">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">weglarskae</emphasis>
|
||
</taxonomicName>
|
||
group. Within the group, only
|
||
<taxonomicName genus="B." lsidName="B. amphoterus" pageId="0" pageNumber="217" rank="species" species="amphoterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">B. amphoterus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="B." lsidName="B. maculatus" pageId="0" pageNumber="217" rank="species" species="maculatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">B. maculatus</emphasis>
|
||
</taxonomicName>
|
||
have a reduced number of ventral plate rows to two or three, as in the new species. Consecutively,
|
||
<taxonomicName genus="B." lsidName="B. nigripunctatus" pageId="0" pageNumber="217" rank="species" species="nigripunctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">B. nigripunctatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from:
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">B. amphoterus, known from Croatia (Istria) and Greece (Crete) (McInnes 1994), by: the mode of reproduction (dioecy in the new species vs. parthenogenesis in B. amphoterus), the presence of lateral supplementary platelets (absent in B. amphoterus), the presence of epicuticular granules on dorsal and ventral plates (absent in B. amphoterus), a different number of ventral plate rows in females (ventral plate formula III:2-2-1 in females of the new species vs. II:2-2 in B. amphoterus) and by the lack of spurs on external claws (extremely small spur very difficult to observe just near the base in B. amphoterus);</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
B. maculatus, known from Tunisia and Greece, by: the mode of reproduction (dioecy in the new species vs. parthenogenesis in B. maculatus), large contrasting granules on dorsal plates (granules not contrasting and clearly visible only in SEM in B. maculatus), the absence of patches or transverse stripes of epicuticular granules on ventral cuticle between legs (present in B. maculatus), a smaller maximal pore density (21-40 pores per 100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2 in segmental II plate in the new species females vs. 48-61 pores per 100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2 in the same plate in B. maculatus), a relatively longer internal peribuccal cirrus (sp is 33-43% in females of the new species vs. 21-30% in B. maculatus) and by relatively longer claws II-IV (sp for claws II is 36-44%, for claws III 36-46%, for claws IV 38-48% in females of the new species vs. 29-36%, 30-34% and 32-38%, respectively in B. maculatus).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">
|
||
Genotypic differential diagnosis:
|
||
<emphasis italics="true" pageId="0" pageNumber="217">p</emphasis>
|
||
-distances between the new species and the remaining
|
||
<taxonomicName authorityName="Thulin" authorityYear="1928" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax</emphasis>
|
||
</taxonomicName>
|
||
spp., for which DNA sequences are available, were as follows:
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">18S rRNA: from 0.4% (B. maculatus, KY609137 and MT333471) to 2.9% (B. australasiaticus sp. nov., MT333468);</paragraph>
|
||
<paragraph pageId="0" pageNumber="217">28S rRNA from 4.2% (B. instabilis, MH414965) to 8.1% (B. decoratus sp. nov., MT333462, MT333463).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |