257 lines
36 KiB
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257 lines
36 KiB
XML
<document id="274118B59BA04D7E570F6D0267DDE02D" ID-DOI="10.11646/zootaxa.4995.3.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5057385" ID-ZooBank="AF31D6D4-AFD4-46A2-A400-29B0281810E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625172752147" checkinUser="plazi" docAuthor="Hara, Hideho, Smith, David R. & Shinohara, Akihiko" docDate="2021" docId="03F1576FFFCF694AFBB5FB93FA0CFDCF" docLanguage="en" docName="zootaxa.4995.3.4.pdf" docOrigin="Zootaxa 4995 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gilpinia hakonensis" docType="key" docVersion="3" lastPageNumber="473" masterDocId="FFC82F17FFCD6948FB22FFACFF90FA5B" masterDocTitle="Gilpinia hakonensis and similar species in Japan and ovipositors of five European Gilpinia species (Hymenoptera, Diprionidae)" masterLastPageNumber="491" masterPageNumber="471" pageNumber="473" updateTime="1698982284166" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1C141F0D87C11238271420E8865B79F2">Gilpinia hakonensis and similar species in Japan and ovipositors of five European Gilpinia species (Hymenoptera, Diprionidae)</mods:title>
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<mods:name id="C88506F0385B94387049B18C5A5575C4" type="personal">
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<mods:roleTerm id="2AFD653BF12CBD2D786DD582EF1EA61B">Author</mods:roleTerm>
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<mods:namePart id="44BB058A4CC8C39357079FEFFE0EFCCD">Hara, Hideho</mods:namePart>
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<mods:roleTerm id="1ACC8285E8A703D77184389BA6FF27E1">Author</mods:roleTerm>
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<mods:namePart id="C62EA612B8BE224412BD1061AA1B4B9D">Smith, David R.</mods:namePart>
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<mods:affiliation id="FFD2200B0C0AD72F551A73CCF5AF181E">Systematic Entomology Laboratory, c / o National Museum of Natural History, Smithsonian Institution, P. O. Box 37012, MRC 168, Washington, DC 20013 - 7012, USA.</mods:affiliation>
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<mods:roleTerm id="E26F6E68246E0AD77485508D688E0FD2">Author</mods:roleTerm>
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<mods:namePart id="F2DE88B4FB45A7A731C6644AD4C11628">Shinohara, Akihiko</mods:namePart>
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<mods:affiliation id="17F49742B6E046B28500CD91BD12F21C">National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
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<mods:typeOfResource id="319C2BF65EE6B76100046D02F7855AB9">text</mods:typeOfResource>
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<mods:title id="BBC501A134C72B9C8B27CED9D920D51F">Zootaxa</mods:title>
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<mods:part id="2B3A6DB36442A9C4ECF8B81EBF7A8F4E">
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<mods:date id="BF385BC35ACE85EADB261F7F22EBFEC8">2021</mods:date>
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<mods:number id="5EEE22786D054F69F4D6C148DFBB27F6">2021-07-01</mods:number>
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<mods:number id="C570AE0C57C1398A1203E6FD5FF3D45F">4995</mods:number>
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<mods:identifier id="FD4FECA2FECB134801FAD49407CA7C12" type="DOI">10.11646/zootaxa.4995.3.4</mods:identifier>
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<paragraph id="8BE7E679FFCF694AFBB5FB93FCDCFE02" blockId="2.[151,844,1086,1113]" box="[151,844,1086,1113]" pageId="2" pageNumber="473">
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<heading id="D0AF5115FFCF694AFBB5FB93FCDCFE02" bold="true" box="[151,844,1086,1113]" fontSize="11" level="1" pageId="2" pageNumber="473" reason="1">
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<emphasis id="B92C3A6BFFCF694AFBB5FB93FCDCFE02" bold="true" box="[151,844,1086,1113]" pageId="2" pageNumber="473">
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Key to
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<taxonomicName id="4C589DFAFFCF694AFBD3FB92FE75FE02" baseAuthorityName="Matsumura" baseAuthorityYear="1912" box="[241,485,1086,1113]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="hakonensis">
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<emphasis id="B92C3A6BFFCF694AFBD3FB92FE75FE02" bold="true" box="[241,485,1086,1113]" italics="true" pageId="2" pageNumber="473">Gilpinia hakonensis</emphasis>
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</taxonomicName>
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and similar species in
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<collectingCountry id="F34FA6E9FFCF694AF9DDFB93FCDCFE02" box="[767,844,1087,1113]" name="Japan" pageId="2" pageNumber="473">Japan</collectingCountry>
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</emphasis>
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<paragraph id="8BE7E679FFCF694AFBB5FB28FA0CFEC2" blockId="2.[151,1437,1156,1940]" box="[151,1436,1156,1178]" pageId="2" pageNumber="473">
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<keyLead id="30A9454CFFCF694AFBB5FB28FA0CFEC2" box="[151,1436,1156,1178]" pageId="2" pageNumber="473">
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<specimenCount id="9D5E2DF0FFCF694AFBB5FB28FEBCFEC1" box="[151,300,1156,1178]" pageId="2" pageNumber="473" type="female">1 Female</specimenCount>
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............................................................................................. 2
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</keyLead>
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||
</paragraph>
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||
<paragraph id="8BE7E679FFCF694AFBB5FB0DFA0CFEEC" blockId="2.[151,1437,1156,1940]" box="[151,1436,1185,1207]" pageId="2" pageNumber="473">
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<keyLead id="30A9454CFFCF694AFBB5FB0DFA0CFEEC" box="[151,1436,1185,1207]" pageId="2" pageNumber="473">
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- Male (unknown in
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<taxonomicName id="4C589DFAFFCF694AFABBFB0DFD97FEED" authorityName="Hara & Smith & Shinohara" authorityYear="2021" box="[409,519,1185,1206]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="okinawa">
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<emphasis id="B92C3A6BFFCF694AFABBFB0DFD97FEED" box="[409,519,1185,1206]" italics="true" pageId="2" pageNumber="473">G. okinawa</emphasis>
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</taxonomicName>
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).......................................................................... 6
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30ACFEDCFFCF694AFBB5FB12FA0CFEAA" pageId="2" pageNumber="473">
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<paragraph id="8BE7E679FFCF694AFBB5FB12FA0CFE88" blockId="2.[151,1437,1156,1940]" box="[151,1436,1214,1236]" pageId="2" pageNumber="473">
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<keyLead id="30A9454CFFCF694AFBB5FB12FA0CFE88" box="[151,1436,1214,1236]" pageId="2" pageNumber="473">
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2(1) Hind tibia with posterior spur slender, rod- or thornlike........................................
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<taxonomicName id="4C589DFAFFCF694AFFC6FB13FAA5FE8F" baseAuthorityName="Klug" baseAuthorityYear="1812" box="[1252,1333,1215,1236]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="socia">
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<emphasis id="B92C3A6BFFCF694AFFC6FB13FAA5FE8F" box="[1252,1333,1215,1236]" italics="true" pageId="2" pageNumber="473">G. socia</emphasis>
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</taxonomicName>
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group ... 3
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</keyLead>
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</paragraph>
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<paragraph id="8BE7E679FFCF694AFBB5FB70FA0CFEAA" blockId="2.[151,1437,1156,1940]" box="[151,1436,1244,1266]" pageId="2" pageNumber="473">
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<keyLead id="30A9454CFFCF694AFBB5FB70FA0CFEAA" box="[151,1436,1244,1266]" pageId="2" pageNumber="473">
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- Hind tibia with posterior spur expanded, scalelike.............................................
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<taxonomicName id="4C589DFAFFCF694AFFCAFB70FACEFEAA" baseAuthorityName="Hartig" baseAuthorityYear="1834" box="[1256,1374,1244,1265]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="polytoma">
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<emphasis id="B92C3A6BFFCF694AFFCAFB70FACEFEAA" box="[1256,1374,1244,1265]" italics="true" pageId="2" pageNumber="473">G. polytoma</emphasis>
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</taxonomicName>
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group
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30ACFEDCFFCF694AFBB5FB55FA1BFF12" pageId="2" pageNumber="473">
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<paragraph id="8BE7E679FFCF694AFBB5FB55FA0CFF55" blockId="2.[151,1437,1156,1940]" box="[151,1436,1273,1295]" pageId="2" pageNumber="473">
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<keyLead id="30A9454CFFCF694AFBB5FB55FA0CFF55" box="[151,1436,1273,1295]" pageId="2" pageNumber="473">
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3(2) Valvula 3 with scopa located at apex of apico-lateral projection (
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<figureCitation id="1363FAFCFFCF694AF86FFB55FC26FF54" box="[845,950,1273,1295]" captionStart="FIGURES 5" captionStartId="11.[151,264,1868,1892]" captionTargetBox="[204,1384,181,1835]" captionTargetId="figure-19@11.[204,1384,181,1843]" captionTargetPageId="11" captionText="FIGURES 5A–U.A–P, Valvula 3, dorsal and posterior views; Q–U, ovipositor or lance, dorsal view (Q, S, T) or lateral view (R, U). A–F, Q, R, Gilpinia hakonensis: A–B, lectotype of Lophyrus hakonensis; C, D, holotype of G. hakonensis var. laticincta; E, F, P, paratype of Diprion fukudai; Q, R, Honshu. G–J, S, Gilpinia amamiana: G, H, holotype; I, J, Tokunoshima Isl.; S, Amami Oshima Isl. K, L, T, U, Gilpinia okinawa, holotype. M, N, Gilpinia frutetorum, Sweden. O, P, Gilpinia laricis, Germany. R, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057397" httpUri="https://zenodo.org/record/5057397/files/figure.png" pageId="2" pageNumber="473">Fig. 5A–L</figureCitation>
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).........
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<taxonomicName id="4C589DFAFFCF694AFF0CFB55FB26FF55" baseAuthorityName="Matsumura" baseAuthorityYear="1912" box="[1070,1206,1273,1294]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="hakonensis">
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<emphasis id="B92C3A6BFFCF694AFF0CFB55FB26FF55" box="[1070,1206,1273,1294]" italics="true" pageId="2" pageNumber="473">G. hakonensis</emphasis>
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</taxonomicName>
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and similar species ... 4
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</keyLead>
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</paragraph>
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<paragraph id="8BE7E679FFCF694AFBB5FABBFA1BFF12" blockId="2.[151,1437,1156,1940]" pageId="2" pageNumber="473">
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<keyLead id="30A9454CFFCF694AFBB5FABBFA1BFF12" pageId="2" pageNumber="473">
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- Valvula 3 with scopa located from apex to inner side of apico-lateral projection..................
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<taxonomicName id="4C589DFAFFCF694AFF98FABBFA08FF70" authority="Takeuchi, 1940" authorityName="Takeuchi" authorityYear="1940" box="[1210,1432,1302,1324]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="tohi">
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<emphasis id="B92C3A6BFFCF694AFF98FABBFB6CFF70" box="[1210,1276,1302,1324]" italics="true" pageId="2" pageNumber="473">G. tohi</emphasis>
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<bibRefCitation id="EFC99B88FFCF694AFE21FABAFA08FF70" author="Takeuchi, K." box="[1283,1432,1302,1324]" pageId="2" pageNumber="473" pagination="187 - 199" refId="ref15041" refString="Takeuchi, K. (1940) A systematic study on the suborder Symphyta (Hymenoptera) of the Japanese Empire (III). Tenthredo, 3, 187 - 199." type="journal article" year="1940">Takeuchi, 1940</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C589DFAFFCF694AFBC0FA98FDF0FF12" authority="Hara & Nakamura, 2015" authorityName="Hara & Nakamura" authorityYear="2015" box="[226,608,1331,1354]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="albiclavata">
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<emphasis id="B92C3A6BFFCF694AFBC0FA98FEFBFF12" box="[226,363,1332,1353]" italics="true" pageId="2" pageNumber="473">G. albiclavata</emphasis>
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<bibRefCitation id="EFC99B88FFCF694AFA53FA98FDF0FF12" author="Hara, H. & Nakamura, H." box="[369,608,1331,1354]" pageId="2" pageNumber="473" pagination="31 - 40" refId="ref13271" refString="Hara, H. & Nakamura, H. (2015) A pine sawfly, Gilpinia albiclavata sp. nov. (Hymenoptera: Diprionidae), infesting Pinus pumila in the Japanese Alps. Entomological Science, 18, 31 - 40. https: // doi. org / 10.1111 / ens. 12088" type="journal article" year="2015">Hara & Nakamura, 2015</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C589DFAFFCF694AF948FA98FB92FF12" authority="Hara & Shinohara, 2015" authorityName="Hara & Shinohara" authorityYear="2015" box="[618,1026,1331,1354]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="hokkaidoensis">
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<emphasis id="B92C3A6BFFCF694AF948FA98FC80FF12" box="[618,784,1332,1353]" italics="true" pageId="2" pageNumber="473">G. hokkaidoensis</emphasis>
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<bibRefCitation id="EFC99B88FFCF694AF834FA98FB92FF12" author="Hara, H. & Shinohara, A." box="[790,1026,1331,1354]" pageId="2" pageNumber="473" pagination="21 - 41" refId="ref13328" refString="Hara, H. & Shinohara, A. (2015) The Gilpinia abieticola Species Group (Hymenoptera, Diprionidae). Bulletin of the National Museum of Nature and Science, Series A, 41, 21 - 41." type="journal article" year="2015">Hara & Shinohara, 2015</bibRefCitation>
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||
</taxonomicName>
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, and
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||
<taxonomicName id="4C589DFAFFCF694AFF17FA98FA1BFF12" authority="Hara & Shinohara, 2015" authorityName="Hara & Shinohara" authorityYear="2015" box="[1077,1419,1331,1354]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="kojimai">
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<emphasis id="B92C3A6BFFCF694AFF17FA98FB0AFF12" box="[1077,1178,1332,1353]" italics="true" pageId="2" pageNumber="473">G. kojimai</emphasis>
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<bibRefCitation id="EFC99B88FFCF694AFF82FA98FA1BFF12" author="Hara, H. & Shinohara, A." box="[1184,1419,1331,1354]" pageId="2" pageNumber="473" pagination="21 - 41" refId="ref13328" refString="Hara, H. & Shinohara, A. (2015) The Gilpinia abieticola Species Group (Hymenoptera, Diprionidae). Bulletin of the National Museum of Nature and Science, Series A, 41, 21 - 41." type="journal article" year="2015">Hara & Shinohara, 2015</bibRefCitation>
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</taxonomicName>
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||
</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep id="30ACFEDCFFCF694AFBB5FAFDFA0CFFA2" pageId="2" pageNumber="473">
|
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<paragraph id="8BE7E679FFCF694AFBB5FAFDFA0DFFFA" blockId="2.[151,1437,1156,1940]" pageId="2" pageNumber="473">
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||
<keyLead id="30A9454CFFCF694AFBB5FAFDFA0DFFFA" pageId="2" pageNumber="473">
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||
4(3) Thorax with median mesoscutal lobe black, sometimes narrowly yellowish posteriorly and mesepisternum black, often with yellow spot posteriorly (
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<figureCitation id="1363FAFCFFCF694AFAEBFAC2FDA7FFDF" box="[457,567,1390,1412]" captionStart="FIGURES 1" captionStartId="3.[151,264,1869,1893]" captionTargetBox="[151,1435,185,1845]" captionTargetId="figure-19@3.[151,1435,181,1845]" captionTargetPageId="3" captionText="FIGURES 1A–N. Gilpinia hakonensis: A, B, K, lectotype of Lophyrus hakonensis, female; C, D, L, holotype of G. hakonensis var. laticincta, female; E, female, Honshu; F, female, Honshu; G, H, M, paratype of Diprion fukudai, female; I, J, N, male, Hon- shu. A, C, E, F, G, I, Dorsal or dorsolateral view; B, D, H, J, ventrolateral or lateral view; K–N, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057387" httpUri="https://zenodo.org/record/5057387/files/figure.png" pageId="2" pageNumber="473">Figs 1A–H</figureCitation>
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). Supraclypeal area black (
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<figureCitation id="1363FAFCFFCF694AF814FAC2FC37FFDF" box="[822,935,1390,1412]" captionStart="FIGURES 1" captionStartId="3.[151,264,1869,1893]" captionTargetBox="[151,1435,185,1845]" captionTargetId="figure-19@3.[151,1435,181,1845]" captionTargetPageId="3" captionText="FIGURES 1A–N. Gilpinia hakonensis: A, B, K, lectotype of Lophyrus hakonensis, female; C, D, L, holotype of G. hakonensis var. laticincta, female; E, female, Honshu; F, female, Honshu; G, H, M, paratype of Diprion fukudai, female; I, J, N, male, Hon- shu. A, C, E, F, G, I, Dorsal or dorsolateral view; B, D, H, J, ventrolateral or lateral view; K–N, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057387" httpUri="https://zenodo.org/record/5057387/files/figure.png" pageId="2" pageNumber="473">Figs 1K–M</figureCitation>
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). In lancet, serrula of second annulus with postcalcar widely separated from precalcar (
|
||
<figureCitation id="1363FAFCFFCF694AF901FA20FD00FFF9" box="[547,656,1420,1442]" captionStart="FIGURES 6" captionStartId="12.[151,264,1869,1893]" captionTargetBox="[154,1435,181,1833]" captionTargetId="figure-19@12.[151,1435,181,1845]" captionTargetPageId="12" captionText="FIGURES 6A–J. Lancet. A–F, Gilpinia hakonensis: A, lectotype of Lophyrus hakonensis; B, holotype of G. hakonensis var. laticincta; C, Kyushu, D, Honshu, E, paratype of Diprion fukudai. F–H, Gilpinia amamiana: F, holotype; G, Amami Oshima Isl.; H, Tokunoshima Isl. I, J, Gilpinia okinawa: I, holotype; J, Okinawa Isl. 1, 2, 5, first (most basal), second and fifth annuli. D–F, H, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057401" httpUri="https://zenodo.org/record/5057401/files/figure.png" pageId="2" pageNumber="473">Figs 6A–E</figureCitation>
|
||
). [Distribution: Honshu and Kyushu,
|
||
<collectingCountry id="F34FA6E9FFCF694AF8DAFA20FBA2FFFA" box="[1016,1074,1420,1441]" name="Japan" pageId="2" pageNumber="473">Japan</collectingCountry>
|
||
;
|
||
<collectingCountry id="F34FA6E9FFCF694AFF1EFA27FBECFFFA" box="[1084,1148,1419,1441]" name="South Korea" pageId="2" pageNumber="473">Korea</collectingCountry>
|
||
.]..........
|
||
<taxonomicName id="4C589DFAFFCF694AFE36FA20FA0DFFFA" baseAuthorityName="Matsumura" baseAuthorityYear="1912" box="[1300,1437,1420,1441]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="hakonensis">
|
||
<emphasis id="B92C3A6BFFCF694AFE36FA20FA0DFFFA" box="[1300,1437,1420,1441]" italics="true" pageId="2" pageNumber="473">G. hakonensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE7E679FFCF694AFBB5FA05FA0CFFA2" blockId="2.[151,1437,1156,1940]" pageId="2" pageNumber="473">
|
||
<keyLead id="30A9454CFFCF694AFBB5FA05FA0CFFA2" pageId="2" pageNumber="473">
|
||
- Thorax with median mesoscutal lobe widely or entirely yellow or red yellow and mesepisternum yellow on dorsal half or entirely (
|
||
<figureCitation id="1363FAFCFFCF694AFA1BFA6AFE76FF87" box="[313,486,1478,1500]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2A–H. A–F, I–K, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; I–L, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="2" pageNumber="473">Figs 2A–D, G, H</figureCitation>
|
||
). Supraclypeal area yellow (
|
||
<figureCitation id="1363FAFCFFCF694AF9D9FA6AFCE7FF87" box="[763,887,1478,1500]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2A–H. A–F, I–K, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; I–L, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="2" pageNumber="473">Figs 2I, J, L</figureCitation>
|
||
). In lancet, serrula of second annulus with postcalcar (if distinct) relatively narrowly separated from precalcar (
|
||
<figureCitation id="1363FAFCFFCF694AF9CCFA48FCF9FFA2" box="[750,873,1508,1530]" captionStart="FIGURES 6" captionStartId="12.[151,264,1869,1893]" captionTargetBox="[154,1435,181,1833]" captionTargetId="figure-19@12.[151,1435,181,1845]" captionTargetPageId="12" captionText="FIGURES 6A–J. Lancet. A–F, Gilpinia hakonensis: A, lectotype of Lophyrus hakonensis; B, holotype of G. hakonensis var. laticincta; C, Kyushu, D, Honshu, E, paratype of Diprion fukudai. F–H, Gilpinia amamiana: F, holotype; G, Amami Oshima Isl.; H, Tokunoshima Isl. I, J, Gilpinia okinawa: I, holotype; J, Okinawa Isl. 1, 2, 5, first (most basal), second and fifth annuli. D–F, H, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057401" httpUri="https://zenodo.org/record/5057401/files/figure.png" pageId="2" pageNumber="473">Figs 6F, G, I</figureCitation>
|
||
)............................................. 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30ACFEDCFFCF694AFBB5F9ADFA0CFCD0" pageId="2" pageNumber="473">
|
||
<paragraph id="8BE7E679FFCF694AFBB5F9ADFA0CFC0A" blockId="2.[151,1437,1156,1940]" pageId="2" pageNumber="473">
|
||
<keyLead id="30A9454CFFCF694AFBB5F9ADFA0CFC0A" pageId="2" pageNumber="473">
|
||
5(4) Median mesoscutal lobe anteromedially black (
|
||
<figureCitation id="1363FAFCFFCF694AF98BF9ADFC86FC4C" box="[681,790,1537,1559]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2A–H. A–F, I–K, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; I–L, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="2" pageNumber="473">Figs 2A, C</figureCitation>
|
||
). Mesepisternum ventrally black (
|
||
<figureCitation id="1363FAFCFFCF694AFF7CF9ADFB5BFC4D" box="[1118,1227,1537,1559]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2A–H. A–F, I–K, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; I–L, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="2" pageNumber="473">Figs 2B, D</figureCitation>
|
||
). Metascutellum with posterior margin convex in dorsal view (Fig. 4I). [Distribution: Amami Oshima Island and Tokunoshima Island,
|
||
<collectingCountry id="F34FA6E9FFCF694AFE38F9B2FAC6FC6F" box="[1306,1366,1566,1588]" name="Japan" pageId="2" pageNumber="473">Japan</collectingCountry>
|
||
.].............................................................................................
|
||
<taxonomicName id="4C589DFAFFCF694AFE38F990FA0CFC0A" authorityName="Okutani" authorityYear="1965" box="[1306,1436,1596,1617]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="amamiana">
|
||
<emphasis id="B92C3A6BFFCF694AFE38F990FA0CFC0A" box="[1306,1436,1596,1617]" italics="true" pageId="2" pageNumber="473">G. amamiana</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE7E679FFCF694AFBB5F9F5FA0CFCD0" blockId="2.[151,1437,1156,1940]" pageId="2" pageNumber="473">
|
||
<keyLead id="30A9454CFFCF694AFBB5F9F5FA0CFCD0" pageId="2" pageNumber="473">
|
||
- Median mesoscutal lobe entirely red yellow (
|
||
<figureCitation id="1363FAFCFFCF694AF9B6F9F5FD72FC34" box="[660,738,1625,1647]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2A–H. A–F, I–K, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; I–L, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="2" pageNumber="473">Fig. 2G</figureCitation>
|
||
). Mesepisternum entirely red yellow (
|
||
<figureCitation id="1363FAFCFFCF694AFF6DF9F5FB0DFC34" box="[1103,1181,1625,1647]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2A–H. A–F, I–K, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; I–L, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="2" pageNumber="473">Fig. 2H</figureCitation>
|
||
). Metascutellum with pos- terior margin barely convex in dorsal view (Fig. 4M). [Distribution:
|
||
<collectingRegion id="499C289BFFCF694AF851F9DAFC5AFCD7" box="[883,970,1654,1676]" country="Japan" name="Okinawa" pageId="2" pageNumber="473">Okinawa</collectingRegion>
|
||
Island,
|
||
<collectingCountry id="F34FA6E9FFCF694AFF34F9DAFBC2FCD7" box="[1046,1106,1654,1676]" name="Japan" pageId="2" pageNumber="473">Japan</collectingCountry>
|
||
.]................
|
||
<taxonomicName id="4C589DFAFFCF694AFE0CF9DBFA0CFCD0" authorityName="Hara & Smith & Shinohara" authorityYear="2021" box="[1326,1436,1654,1676]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="okinawa">
|
||
<emphasis id="B92C3A6BFFCF694AFE0CF9DBFA0CFCD0" box="[1326,1436,1654,1676]" italics="true" pageId="2" pageNumber="473">G. okinawa</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30ACFEDCFFCF694AFBB5F938FA0CFCBF" pageId="2" pageNumber="473">
|
||
<paragraph id="8BE7E679FFCF694AFBB5F938FA0CFC9D" blockId="2.[151,1437,1156,1940]" pageId="2" pageNumber="473">
|
||
<keyLead id="30A9454CFFCF694AFBB5F938FA0CFC9D" pageId="2" pageNumber="473">
|
||
6(1) Genitalia in ventral view with inner margin of basiparamere horizontal and lateral margin of parapenis short (
|
||
<figureCitation id="1363FAFCFFCF694AFFD2F938FAC9FCF2" box="[1264,1369,1684,1706]" captionStart="FIGURES 7" captionStartId="13.[151,264,1915,1939]" captionTargetBox="[151,1423,960,1891]" captionTargetId="figure-347@13.[151,1435,957,1891]" captionTargetPageId="13" captionText="FIGURES 7A–F. A, D, Male genitalia, dorsal view; B, E, male genitalia (penis valve removed), ventral view; C, F, penis valve, lateral view (left dorsal). A–C, Gilpinia hakonensis, Honshu. D–F, Gilpinia amamiana, Amami Oshima Isl." figureDoi="http://doi.org/10.5281/zenodo.5057405" httpUri="https://zenodo.org/record/5057405/files/figure.png" pageId="2" pageNumber="473">Figs 7B, E</figureCitation>
|
||
); penis valve with very long ergot and spinose swelling near apex (
|
||
<figureCitation id="1363FAFCFFCF694AF83EF91DFC15FC9D" box="[796,901,1713,1735]" captionStart="FIGURES 7" captionStartId="13.[151,264,1915,1939]" captionTargetBox="[151,1423,960,1891]" captionTargetId="figure-347@13.[151,1435,957,1891]" captionTargetPageId="13" captionText="FIGURES 7A–F. A, D, Male genitalia, dorsal view; B, E, male genitalia (penis valve removed), ventral view; C, F, penis valve, lateral view (left dorsal). A–C, Gilpinia hakonensis, Honshu. D–F, Gilpinia amamiana, Amami Oshima Isl." figureDoi="http://doi.org/10.5281/zenodo.5057405" httpUri="https://zenodo.org/record/5057405/files/figure.png" pageId="2" pageNumber="473">Figs 7C, F</figureCitation>
|
||
)..............
|
||
<taxonomicName id="4C589DFAFFCF694AFF0CF91DFB26FC9D" baseAuthorityName="Matsumura" baseAuthorityYear="1912" box="[1070,1206,1713,1734]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="hakonensis">
|
||
<emphasis id="B92C3A6BFFCF694AFF0CF91DFB26FC9D" box="[1070,1206,1713,1734]" italics="true" pageId="2" pageNumber="473">G. hakonensis</emphasis>
|
||
</taxonomicName>
|
||
and similar species ... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE7E679FFCF694AFBB5F963FA0CFCBF" blockId="2.[151,1437,1156,1940]" box="[151,1436,1742,1764]" pageId="2" pageNumber="473">
|
||
<keyLead id="30A9454CFFCF694AFBB5F963FA0CFCBF" box="[151,1436,1742,1764]" pageId="2" pageNumber="473">- Genitalia not as above....................................................................... Other species</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30ACFEDCFFCF694AFBB5F940FA0CFDCF" pageId="2" pageNumber="473">
|
||
<paragraph id="8BE7E679FFCF694AFBB5F940FA0DFD60" blockId="2.[151,1437,1156,1940]" pageId="2" pageNumber="473">
|
||
<keyLead id="30A9454CFFCF694AFBB5F940FA0DFD60" pageId="2" pageNumber="473">
|
||
7(6) Parapenis with lateral margin rather long (
|
||
<figureCitation id="1363FAFCFFCF694AF95DF940FD5CFD5A" box="[639,716,1772,1794]" captionStart="FIGURES 7" captionStartId="13.[151,264,1915,1939]" captionTargetBox="[151,1423,960,1891]" captionTargetId="figure-347@13.[151,1435,957,1891]" captionTargetPageId="13" captionText="FIGURES 7A–F. A, D, Male genitalia, dorsal view; B, E, male genitalia (penis valve removed), ventral view; C, F, penis valve, lateral view (left dorsal). A–C, Gilpinia hakonensis, Honshu. D–F, Gilpinia amamiana, Amami Oshima Isl." figureDoi="http://doi.org/10.5281/zenodo.5057405" httpUri="https://zenodo.org/record/5057405/files/figure.png" pageId="2" pageNumber="473">Fig. 7B</figureCitation>
|
||
). Penis valve with valviceps slightly convex ventrally at apical third (
|
||
<figureCitation id="1363FAFCFFCF694AFE57F940FE91FD44" captionStart="FIGURES 7" captionStartId="13.[151,264,1915,1939]" captionTargetBox="[151,1423,960,1891]" captionTargetId="figure-347@13.[151,1435,957,1891]" captionTargetPageId="13" captionText="FIGURES 7A–F. A, D, Male genitalia, dorsal view; B, E, male genitalia (penis valve removed), ventral view; C, F, penis valve, lateral view (left dorsal). A–C, Gilpinia hakonensis, Honshu. D–F, Gilpinia amamiana, Amami Oshima Isl." figureDoi="http://doi.org/10.5281/zenodo.5057405" httpUri="https://zenodo.org/record/5057405/files/figure.png" pageId="2" pageNumber="473">Fig. 7C</figureCitation>
|
||
). Pronotum entirely black (
|
||
<figureCitation id="1363FAFCFFCF694AF92FF8A5FDFCFD45" box="[525,620,1801,1823]" captionStart="FIGURES 1" captionStartId="3.[151,264,1869,1893]" captionTargetBox="[151,1435,185,1845]" captionTargetId="figure-19@3.[151,1435,181,1845]" captionTargetPageId="3" captionText="FIGURES 1A–N. Gilpinia hakonensis: A, B, K, lectotype of Lophyrus hakonensis, female; C, D, L, holotype of G. hakonensis var. laticincta, female; E, female, Honshu; F, female, Honshu; G, H, M, paratype of Diprion fukudai, female; I, J, N, male, Hon- shu. A, C, E, F, G, I, Dorsal or dorsolateral view; B, D, H, J, ventrolateral or lateral view; K–N, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057387" httpUri="https://zenodo.org/record/5057387/files/figure.png" pageId="2" pageNumber="473">Figs 1I, J</figureCitation>
|
||
). Whole trochanters and wide basal parts of femora black. Abdomen ventrally dark brown to black............................................................................
|
||
<taxonomicName id="4C589DFAFFCF694AFE36F88BFA0DFD60" baseAuthorityName="Matsumura" baseAuthorityYear="1912" box="[1300,1437,1830,1852]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="hakonensis">
|
||
<emphasis id="B92C3A6BFFCF694AFE36F88BFA0DFD60" box="[1300,1437,1830,1852]" italics="true" pageId="2" pageNumber="473">G. hakonensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE7E679FFCF694AFBB5F8E8FA0CFDCF" blockId="2.[151,1437,1156,1940]" pageId="2" pageNumber="473">
|
||
<keyLead id="30A9454CFFCF694AFBB5F8E8FA0CFDCF" pageId="2" pageNumber="473">
|
||
- Parapenis with lateral margin very short (
|
||
<figureCitation id="1363FAFCFFCF694AF949F8E8FD25FD02" box="[619,693,1860,1882]" captionStart="FIGURES 7" captionStartId="13.[151,264,1915,1939]" captionTargetBox="[151,1423,960,1891]" captionTargetId="figure-347@13.[151,1435,957,1891]" captionTargetPageId="13" captionText="FIGURES 7A–F. A, D, Male genitalia, dorsal view; B, E, male genitalia (penis valve removed), ventral view; C, F, penis valve, lateral view (left dorsal). A–C, Gilpinia hakonensis, Honshu. D–F, Gilpinia amamiana, Amami Oshima Isl." figureDoi="http://doi.org/10.5281/zenodo.5057405" httpUri="https://zenodo.org/record/5057405/files/figure.png" pageId="2" pageNumber="473">Fig. 7E</figureCitation>
|
||
). Penis valve with valviceps exceedingly convex ventrally at apical third (
|
||
<figureCitation id="1363FAFCFFCF694AFE57F8E8FF6FFD2D" captionStart="FIGURES 7" captionStartId="13.[151,264,1915,1939]" captionTargetBox="[151,1423,960,1891]" captionTargetId="figure-347@13.[151,1435,957,1891]" captionTargetPageId="13" captionText="FIGURES 7A–F. A, D, Male genitalia, dorsal view; B, E, male genitalia (penis valve removed), ventral view; C, F, penis valve, lateral view (left dorsal). A–C, Gilpinia hakonensis, Honshu. D–F, Gilpinia amamiana, Amami Oshima Isl." figureDoi="http://doi.org/10.5281/zenodo.5057405" httpUri="https://zenodo.org/record/5057405/files/figure.png" pageId="2" pageNumber="473">Fig. 7F</figureCitation>
|
||
). Pronotum mostly yellow (
|
||
<figureCitation id="1363FAFCFFCF694AF929F8CDFD1FFD2D" box="[523,655,1889,1911]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2A–H. A–F, I–K, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; I–L, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="2" pageNumber="473">Figs 2E, F, K</figureCitation>
|
||
). Trochanters widely and femora entirely yellow. Abdomen ventrally mostly yellow......................................................................................
|
||
<taxonomicName id="4C589DFAFFCF694AFE38F8D3FA0CFDCF" authorityName="Okutani" authorityYear="1965" box="[1306,1436,1919,1940]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="amamiana">
|
||
<emphasis id="B92C3A6BFFCF694AFE38F8D3FA0CFDCF" box="[1306,1436,1919,1940]" italics="true" pageId="2" pageNumber="473">G. amamiana</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |