treatments-xml/data/03/E1/87/03E187FCFFCD696FFDC40FEFFCBDFC0C.xml
2024-06-21 12:22:17 +02:00

452 lines
66 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="3C516C2E5931A480B04C68B60D6B0389" ID-CLB-Dataset="284910" ID-DOI="10.5852/ejt.2023.908.2337" ID-GBIF-Dataset="6156c195-5953-4af2-9999-a1a03fd0f87e" ID-ISSN="2118-9773" ID-Zenodo-Dep="10149298" ID-ZooBank="1728B5B0-A130-4370-9610-9DA9CEDF4717" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700226465688" checkinUser="plazi" docAuthor="Antić, Dragan, Šević, Mirko, Barjadze, Shalva &amp; Makarov, Slobodan" docDate="2023" docId="03E187FCFFCD696FFDC40FEFFCBDFC0C" docLanguage="en" docName="EJT.2023.908.39-76.pdf" docOrigin="European Journal of Taxonomy 908" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2337/10171" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Acanthophorella aurita Antic 2023, sp. nov." docType="treatment" docUuid="ADCB7F33-A824-42E3-B7F5-D2EA41BC93DA" docUuidSource="ZooBank" docVersion="6" lastPageNumber="49" masterDocId="FFD8FF84FFC96965FFF20A17FFD4FFE7" masterDocTitle="Review of the genus Acanthophorella Antić &amp; Makarov, 2016 (Diplopoda, Chordeumatida, Anthroleucosomatidae), with descriptions of three new species from the Caucasus" masterLastPageNumber="76" masterPageNumber="39" pageNumber="43" updateTime="1705069979865" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1D8C6A5E3D506258A86C77FDF97116D5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="051A513B2E22DC0530354AFBE969C399">
<mods:title id="DE513CA8F9AE73EDC215163F0DE47C46">Review of the genus Acanthophorella Antić &amp; Makarov, 2016 (Diplopoda, Chordeumatida, Anthroleucosomatidae), with descriptions of three new species from the Caucasus</mods:title>
</mods:titleInfo>
<mods:name id="D20161BF6E1580C506A2A088CB7AB56C" type="personal">
<mods:role id="4015E141C376E09282D4444E1F90CAD9">
<mods:roleTerm id="EF8F4F28DFD252586D3DC0FBDB211125">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1F352050848CCA5E67AAF167DB464D64">Antić, Dragan</mods:namePart>
<mods:nameIdentifier id="5979A24E075CBA4556E8E452DD8AA5A9" type="LSID">B5FACDD3-ACCA-4050-8118-80EBD2BB942F</mods:nameIdentifier>
<mods:affiliation id="EE73BCFBD2F1737B569AE2FA8C46DEA8">University of Belgrade Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. &amp; Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
<mods:nameIdentifier id="BF00803B8D865FACD34FA8FFE58263A1" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
</mods:name>
<mods:name id="94536C0A6E26C841295970352D2B2757" type="personal">
<mods:role id="978F73E6FB6E4B15C2FF7A233190E5CC">
<mods:roleTerm id="9D3DFA0128FCE7E795127039C777AFF3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E0A361EB8AE1C742050A4B0B093CF3A5">Šević, Mirko</mods:namePart>
<mods:nameIdentifier id="7B6665A420C578C5B821701412C9637C" type="LSID">AE92D931-6893-4884-8173-31C0B5D327FD</mods:nameIdentifier>
<mods:affiliation id="5DEE0B290C966C2187215CD7AA693B60">University of Belgrade Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. &amp; Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
<mods:nameIdentifier id="CD77F40162E6D52BD6D5855C84AD5FF8" type="email">mirko.sevic@bio.bg.ac.rs</mods:nameIdentifier>
</mods:name>
<mods:name id="74C9C360255C00CFE142035897E9C038" type="personal">
<mods:role id="E50082D861D65DD1103B528D421A31CA">
<mods:roleTerm id="FBE140BA632221A865DA9E23DA1DBA0E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05A59E8CDA737ED213EB1213A390D6DF">Barjadze, Shalva</mods:namePart>
<mods:nameIdentifier id="982EA952EAE6D778CF763AD902CFE368" type="LSID">AB36BEF1-C006-41A3-861E-E0B79EE35FBF</mods:nameIdentifier>
<mods:affiliation id="161A20790ACC344321D1BD0E3CDA26B1">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia.</mods:affiliation>
<mods:nameIdentifier id="802943A5C0898CB3F4F5959F052172B3" type="email">shalva.barjadze@iliauni.edu.ge</mods:nameIdentifier>
</mods:name>
<mods:name id="BBDB5AC25B3D1B5ADD06A2BF99737C16" type="personal">
<mods:role id="BE4C6EFDE9850DCB2D3C6CCC32A33BE7">
<mods:roleTerm id="1AAA169BDB025D28BAA605B91E8FBDF0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4BCEC8CDEB8981924B715CCDEE97B77A">Makarov, Slobodan</mods:namePart>
<mods:nameIdentifier id="B0D2E9F83119FFED15AC93B5C3CC23A5" type="LSID">7F5C25A5-615E-427B-B4FC-DB1A0EE1B534</mods:nameIdentifier>
<mods:affiliation id="9CADA40567F786B85CB8B93BE848EBA3">University of Belgrade Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. &amp; Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
<mods:nameIdentifier id="0930A9BFE29C6BF630D2CD2A93825272" type="email">slobodan@bio.bg.ac.rs</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="75F13753D54AE35590741826ED9574F8">text</mods:typeOfResource>
<mods:relatedItem id="3BF9FD650F74311CAF71B18B23BCB08E" type="host">
<mods:titleInfo id="273E27629759738DF40C8490A6942B5D">
<mods:title id="774C58FE066E59C74E6E454378CC0B25">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="809E5201F7834446EB398D22CB612045">
<mods:date id="0884E2CB51BCD897E622C6600290EC24">2023</mods:date>
<mods:detail id="1D9EFF49BDC206EE8DCBA415A510F34A" type="pubDate">
<mods:number id="9F4E092F1CD5DD91FE49A4F5A54F3CD7">2023-11-17</mods:number>
</mods:detail>
<mods:detail id="DE9EDDB16B984B6054058545EAB9BA17" type="volume">
<mods:number id="C383CE2114BB1FEB7CF9942AFF421264">908</mods:number>
</mods:detail>
<mods:extent id="A9746E1759EB2CD0BA6997874BBBF8A7" unit="page">
<mods:start id="B6C80D1223A269633997490B5BBCC10E">39</mods:start>
<mods:end id="CA49C4BE3ABD5ED1C6197E26381066D9">76</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3AF0403FA8FEF0367CD2BBFC04FB9354">
<mods:url id="8DBFC43CBFCEC77292AF4F9AB7CE8055">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2337/10171</mods:url>
</mods:location>
<mods:classification id="F6228A7F0EF54C9CB6A501F3E595C1E2">journal article</mods:classification>
<mods:identifier id="ED583766671B136952A2267862D11BC8" type="CLB-Dataset">284910</mods:identifier>
<mods:identifier id="6DC30A84DE3416E7F3591252AA958956" type="DOI">10.5852/ejt.2023.908.2337</mods:identifier>
<mods:identifier id="67266F23AE49F1586B2B0186C97A8674" type="GBIF-Dataset">6156c195-5953-4af2-9999-a1a03fd0f87e</mods:identifier>
<mods:identifier id="D129BC8AB1267B27D88D0E88D72E7AD8" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="59B7493B2DEDE11F94AD2818DFA3E10E" type="Zenodo-Dep">10149298</mods:identifier>
<mods:identifier id="0D056F83A7934F733340E05A7B1E6487" type="ZooBank">1728B5B0-A130-4370-9610-9DA9CEDF4717</mods:identifier>
</mods:mods>
<treatment id="03E187FCFFCD696FFDC40FEFFCBDFC0C" ID-DOI="http://doi.org/10.5281/zenodo.10168138" ID-Zenodo-Dep="10168138" LSID="urn:lsid:zoobank.org:act:ADCB7F33-A824-42E3-B7F5-D2EA41BC93DA" httpUri="http://treatment.plazi.org/id/03E187FCFFCD696FFDC40FEFFCBDFC0C" lastPageId="10" lastPageNumber="49" pageId="4" pageNumber="43">
<subSubSection id="C3526561FFCD6961FDC40FEFFC29F9F4" box="[566,1021,1528,1555]" pageId="4" pageNumber="43" type="nomenclature">
<paragraph id="8BF736EAFFCD6961FDC40FEFFC29F9F4" blockId="4.[354,1235,1528,1626]" box="[566,1021,1528,1555]" pageId="4" pageNumber="43">
<heading id="D0BF8186FFCD6961FDC40FEFFC29F9F4" box="[566,1021,1528,1555]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="43" reason="2">
<taxonomicName id="4C484D69FFCD6961FDC40FEFFC4FF9F4" ID-CoL="BW7HV" authority="Antic" authorityName="Antic" authorityYear="2023" box="[566,923,1528,1555]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aurita" status="sp. nov.">
<emphasis id="B93CEAF8FFCD6961FDC40FEFFC87F9F4" bold="true" box="[566,851,1528,1555]" italics="true" pageId="4" pageNumber="43">Acanthophorella aurita</emphasis>
Antić
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFCD6961FC510FEEFC29F9F4" box="[931,1021,1529,1555]" pageId="4" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFCD6961FE900C0BFC57F9BE" pageId="4" pageNumber="43" type="description">
<paragraph id="8BF736EAFFCD6961FE900C0BFB07F9D0" blockId="4.[354,1235,1528,1626]" box="[354,1235,1564,1591]" pageId="4" pageNumber="43">
<uri id="FFD93AE8FFCD6961FE900C0BFB07F9D0" box="[354,1235,1564,1591]" pageId="4" pageNumber="43">
urn:lsid:zoobank.org:act:
<uuid id="FFEE0C3FFFCD6961FD710C0BFB07F9D0" box="[643,1235,1564,1591]" pageId="4" pageNumber="43">ADCB7F33-A824-42E3-B7F5-D2EA41BC93DA</uuid>
</uri>
</paragraph>
<paragraph id="8BF736EAFFCD6961FD420C28FC57F9BE" blockId="4.[354,1235,1528,1626]" box="[688,899,1599,1626]" pageId="4" pageNumber="43">
<figureCitation id="13732A6FFFCD6961FD420C28FCCDF9BD" box="[688,793,1599,1626]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="6.[189,232,1892,1918]" captionStartId-1="7.[189,232,1822,1848]" captionStartId-2="8.[189,232,1892,1918]" captionStartId-3="9.[189,232,1822,1848]" captionTargetBox-0="[189,1398,274,1838]" captionTargetBox-1="[189,1398,269,1764]" captionTargetBox-2="[189,1398,829,1844]" captionTargetBox-3="[199,1388,793,1738]" captionTargetId-0="figure-16@6.[189,1398,274,1838]" captionTargetId-1="figure-12@7.[189,1398,269,1764]" captionTargetId-2="figure-162@8.[189,1398,829,1844]" captionTargetId-3="figure-147@9.[189,1398,773,1781]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="Fig. 2. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Habitus, lateral view. B. Right field with ommatidia, lateral view. C. Head, anterior view, white arrows indicate rectangular lobes. D. Midbody rings, lateral view. E. Midbody rings, dorsal view. Scale bars: A, DE = 1 mm; BC = 0.5 mm." captionText-1="Fig. 3. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Leg-pair 2, posterior view. B. Left leg 3, anterior view, white arrow indicates the distoventral femoral pad. C. Right leg 3 with a distoventral femoral pad, anterior view. D. Leg-pair 7, anterior view. E. Leg-pair 10, anterior view. Scale bars = 0.2 mm." captionText-2="Fig. 4. AB. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). CD. Acanthophorella cf. A. aurita from Kveda Tlughi. A, C. Anterior gonopods, anterior and posterior views, respectively. B, D. Posterior gonopods, anterior views. Scale bars = 0.2 mm; CD not to scale." captionText-3="Fig. 5. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363), gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, anterior view. E. Right posterior gonopod, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; cv = coxal vesicle; ll = lateral lamella; mp = medial part; s = sternum; sl = sternal lamella; t = telopodital pigment remnants; tf = angiocoxal tuft. Drawings by DA. Scale bars = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10149304" figureDoi-1="http://doi.org/10.5281/zenodo.10149306" figureDoi-2="http://doi.org/10.5281/zenodo.10149308" figureDoi-3="http://doi.org/10.5281/zenodo.10149310" httpUri-0="https://zenodo.org/record/10149304/files/figure.png" httpUri-1="https://zenodo.org/record/10149306/files/figure.png" httpUri-2="https://zenodo.org/record/10149308/files/figure.png" httpUri-3="https://zenodo.org/record/10149310/files/figure.png" pageId="4" pageNumber="43">Figs 25</figureCitation>
,
<figureCitation id="13732A6FFFCD6961FCD70C28FC8EF9BE" box="[805,858,1599,1625]" captionStart="Fig" captionStartId="33.[189,232,1822,1848]" captionTargetBox="[202,1378,705,1770]" captionTargetId="figure-170@33.[189,1398,686,1780]" captionTargetPageId="33" captionText="Fig. 21. Anterior gonopods in the genera Acanthophorella Antić &amp; Makarov, 2016 and PseudoflagellophorellaAntić &amp; Makarov, 2016, anterior views. A. A. auritaAntić sp. nov. B. A. barjadzei Antić &amp; Makarov, 2016. C. A. chegemi Antić &amp; Makarov, 2016. D. A. devi Antić sp. nov. E. A. irystoni Antić &amp; Makarov, 2016. F. A. valerii Antić sp. nov. G. P. eskovi Antić &amp; Makarov, 2016. H. P. mirabilis Antić &amp; Makarov, 2016. I. P. papilioformis Antić &amp; Makarov, 2016. BC, E, GI after Antić &amp; Makarov (2016). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149345" httpUri="https://zenodo.org/record/10149345/files/figure.png" pageId="4" pageNumber="43">21A</figureCitation>
,
<figureCitation id="13732A6FFFCD6961FC940C28FC57F9BE" box="[870,899,1599,1625]" captionStart="Fig" captionStartId="35.[189,232,1998,2024]" captionTargetBox="[189,1398,1153,1959]" captionTargetId="figure-355@35.[189,1398,1150,1959]" captionTargetPageId="35" captionText="Fig. 22. Distribution of the genus Acanthophorella Antić &amp; Makarov, 2016." figureDoi="http://doi.org/10.5281/zenodo.10149347" httpUri="https://zenodo.org/record/10149347/files/figure.png" pageId="4" pageNumber="43">22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFCD6961FF4F0C97FD08F8D4" pageId="4" pageNumber="43" type="diagnosis">
<paragraph id="8BF736EAFFCD6961FF4F0C97FEE2F97D" blockId="4.[189,310,1664,1690]" box="[189,310,1664,1690]" pageId="4" pageNumber="43">
<heading id="D0BF8186FFCD6961FF4F0C97FEE2F97D" bold="true" box="[189,310,1664,1690]" fontSize="11" level="3" pageId="4" pageNumber="43" reason="3">
<emphasis id="B93CEAF8FFCD6961FF4F0C97FEE2F97D" bold="true" box="[189,310,1664,1690]" pageId="4" pageNumber="43">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFCD6961FF4F0CB8FD08F8D4" blockId="4.[189,1399,1711,1843]" pageId="4" pageNumber="43">
Distinguished from congeners and members of the genus
<taxonomicName id="4C484D69FFCD6961FC890CB8FB58F92E" authorityName="Antic &amp; Makarov" authorityYear="2016" box="[891,1164,1711,1737]" class="Diplopoda" family="Anthroleucosomatidae" genus="Pseudoflagellophorella" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAF8FFCD6961FC890CB8FB58F92E" box="[891,1164,1711,1737]" italics="true" pageId="4" pageNumber="43">Pseudoflagellophorella</emphasis>
</taxonomicName>
by the presence of very characteristic, wide, more or less ear-shaped angiocoxites of the posterior gonopods in anterior or posterior views (vs angiocoxites are more or less slender or small in all other members of the genera
<taxonomicName id="4C484D69FFCD6961FF4D0D0EFE57F8D4" authorityName="Antic &amp; Makarov" authorityYear="2016" box="[191,387,1817,1843]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAF8FFCD6961FF4D0D0EFE57F8D4" box="[191,387,1817,1843]" italics="true" pageId="4" pageNumber="43">Acanthophorella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D69FFCD6961FE480D0EFD1FF8D4" authorityName="Antic &amp; Makarov" authorityYear="2016" box="[442,715,1817,1843]" class="Diplopoda" family="Anthroleucosomatidae" genus="Pseudoflagellophorella" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAF8FFCD6961FE480D0EFD1FF8D4" box="[442,715,1817,1843]" italics="true" pageId="4" pageNumber="43">Pseudoflagellophorella</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFCD6961FF4F0D4DFAB0F821" pageId="4" pageNumber="43" type="etymology">
<paragraph id="8BF736EAFFCD6961FF4F0D4DFE90F893" blockId="4.[189,324,1882,1908]" box="[189,324,1882,1908]" pageId="4" pageNumber="43">
<heading id="D0BF8186FFCD6961FF4F0D4DFE90F893" bold="true" box="[189,324,1882,1908]" fontSize="11" level="3" pageId="4" pageNumber="43" reason="3">
<emphasis id="B93CEAF8FFCD6961FF4F0D4DFE90F893" bold="true" box="[189,324,1882,1908]" pageId="4" pageNumber="43">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFCD6961FF4F0D9EFAB0F821" blockId="4.[189,1398,1929,1990]" pageId="4" pageNumber="43">
From Latin
<emphasis id="B93CEAF8FFCD6961FEA90D9DFE43F844" box="[347,407,1930,1955]" italics="true" pageId="4" pageNumber="43">auris</emphasis>
(ear) + -
<emphasis id="B93CEAF8FFCD6961FDC30D9DFD8FF844" box="[561,603,1930,1955]" italics="true" pageId="4" pageNumber="43">ītus</emphasis>
;
<taxonomicName id="4C484D69FFCD6961FD8B0D9EFD1FF844" authorityName="Antic" authorityYear="2023" box="[633,715,1929,1955]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aurita">
<emphasis id="B93CEAF8FFCD6961FD710D9DFD1FF844" box="[643,715,1930,1955]" italics="true" pageId="4" pageNumber="43">aurita</emphasis>
</taxonomicName>
(= earlike; having ears or having the shape of a ear) because of the characteristic shape of the angiocoxites of the posterior gonopods. Feminine adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFCC6960FF4F0B1FFAFDFE70" pageId="5" pageNumber="44" type="materials_examined">
<paragraph id="8BF736EAFFCC6960FF4F0B1FFE7EFEC5" blockId="5.[189,426,264,290]" box="[189,426,264,290]" pageId="5" pageNumber="44">
<heading id="D0BF8186FFCC6960FF4F0B1FFE7EFEC5" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="5" pageNumber="44" reason="3">
<emphasis id="B93CEAF8FFCC6960FF4F0B1FFE7EFEC5" bold="true" box="[189,426,264,290]" pageId="5" pageNumber="44">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFCC6960FF180B20FAFDFE70" blockId="5.[189,1398,311,408]" pageId="5" pageNumber="44">
<materialsCitation id="3B203CB7FFCC6960FF180B20FAFDFE70" collectingDate="2021-10-21" collectionCode="NHMW" collectorName="J. Grego &amp; M. Szekeres" country="Georgia" county="Khobi Municipality" location="Jegeta" municipality="Urta Mts" pageId="5" pageNumber="44" specimenCode="MY10363" specimenCount="1" specimenCount-male="1" stateProvince="Samegrelo-Zemo Svaneti" typeStatus="holotype">
<emphasis id="B93CEAF8FFCC6960FF180B20FE88FEB6" bold="true" box="[234,348,311,337]" pageId="5" pageNumber="44">
<typeStatus id="54F38848FFCC6960FF180B20FE88FEB6" box="[234,348,311,337]" pageId="5" pageNumber="44" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F35F767AFFCC6960FF4F0B4DFEEBFE93" box="[189,319,346,372]" name="Georgia" pageId="5" pageNumber="44">GEORGIA</collectingCountry>
<specimenCount id="9D4EFD63FFCC6960FEAA0B4FFEBBFE93" box="[344,367,344,372]" count="1" pageId="5" pageNumber="44" type="male"></specimenCount>
;
<collectingRegion id="498CF808FFCC6960FE8F0B4DFD7FFE93" box="[381,683,346,372]" country="Georgia" name="Samegrelo and Zemo Svaneti" pageId="5" pageNumber="44">Samegrelo-Zemo Svaneti</collectingRegion>
,
<collectingCounty id="62964E66FFCC6960FD440B4DFC74FE93" box="[694,928,346,372]" pageId="5" pageNumber="44">Khobi Municipality</collectingCounty>
,
<collectingMunicipality id="6B93AC90FFCC6960FC590B4CFBC2FE92" box="[939,1046,347,373]" pageId="5" pageNumber="44">Urta Mts</collectingMunicipality>
, small cavity near the ruins of
<location id="8E976031FFCC6960FF4F0B69FED2FE7F" LSID="urn:lsid:plazi:treatment:03E187FCFFCD696FFDC40FEFFCBDFC0C:8E976031FFCC6960FF4F0B69FED2FE7F" box="[189,262,382,408]" country="Georgia" county="Khobi Municipality" municipality="Urta Mts" name="Jegeta" pageId="5" pageNumber="44" stateProvince="Samegrelo-Zemo Svaneti">Jegeta</location>
church (
<location id="8E976031FFCC6960FE9F0B69FDC2FE7F" LSID="urn:lsid:plazi:treatment:03E187FCFFCD696FFDC40FEFFCBDFC0C:8E976031FFCC6960FE9F0B69FDC2FE7F" box="[365,534,382,408]" country="Georgia" county="Khobi Municipality" municipality="Urta Mts" name="Jegeta 2 Cave" pageId="5" pageNumber="44" stateProvince="Samegrelo-Zemo Svaneti">Jegeta 2 Cave</location>
);
<date id="FFF6102AFFCC6960FDD90B69FD12FE7F" box="[555,710,382,408]" pageId="5" pageNumber="44" value="2021-10-21">
<collectingDate id="EFB2E9C2FFCC6960FDD90B69FD12FE7F" box="[555,710,382,408]" pageId="5" pageNumber="44" value="2021-10-21">21 Oct. 2021</collectingDate>
</date>
;
<collectorName id="26BD533CFFCC6960FD210B69FCE2FE7F" box="[723,822,382,408]" pageId="5" pageNumber="44">J. Grego</collectorName>
and
<collectorName id="26BD533CFFCC6960FC9D0B69FBD4FE70" box="[879,1024,381,408]" pageId="5" pageNumber="44">M. Szekeres</collectorName>
leg.;
<collectionCode id="ED59AE2FFFCC6960FBB00B69FB76FE7F" box="[1090,1186,382,408]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="44" type="Museum">NHMW</collectionCode>
<specimenCode id="DBEE9E91FFCC6960FB5B0B6AFAF1FE70" box="[1193,1317,381,407]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="44" type="Museum">MY10363</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFCC696DFF4F0BA8FD1FFD99" lastPageId="8" lastPageNumber="47" pageId="5" pageNumber="44" type="description">
<paragraph id="8BF736EAFFCC6960FF4F0BA8FE9AFE3E" blockId="5.[189,334,447,473]" box="[189,334,447,473]" pageId="5" pageNumber="44">
<heading id="D0BF8186FFCC6960FF4F0BA8FE9AFE3E" bold="true" box="[189,334,447,473]" fontSize="11" level="3" pageId="5" pageNumber="44" reason="3">
<emphasis id="B93CEAF8FFCC6960FF4F0BA8FE9AFE3E" bold="true" box="[189,334,447,473]" pageId="5" pageNumber="44">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFCC6960FF4F0BF9FE27FDCC" blockId="5.[187,1399,493,555]" pageId="5" pageNumber="44">
<smallCapsWord id="8D11A036FFCC6960FF4F0BF9FF39FDE1" baselines="513,513" box="[189,237,494,520]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Size" pageId="5" pageNumber="44">SIZE</smallCapsWord>
<smallCapsWord id="8D11A036FFCC6960FF010BE4FEF4FDE1" baselines="513" box="[243,288,499,518]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="5" pageNumber="44">AND</smallCapsWord>
<smallCapsWord id="8D11A036FFCC6960FED70BE4FEAAFDE1" baselines="513" box="[293,382,499,518]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="number" pageId="5" pageNumber="44">NUMBER</smallCapsWord>
<smallCapsWord id="8D11A036FFCC6960FE760BE4FE4AFDE1" baselines="513" box="[388,414,499,518]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="5" pageNumber="44">OF</smallCapsWord>
<smallCapsWord id="8D11A036FFCC6960FE510BE4FE0AFDE1" baselines="513" box="[419,478,499,518]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="body" pageId="5" pageNumber="44">BODY</smallCapsWord>
<smallCapsWord id="8D11A036FFCC6960FE160BE4FDF6FDE1" baselines="513" box="[484,546,499,518]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="rings" pageId="5" pageNumber="44">RINGS</smallCapsWord>
. Body with 31 rings (including telson), length:
<quantity id="4CB09B0FFFCC6960FBB20BFAFB7EFDEF" box="[1088,1194,493,520]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="44" unit="mm" value="10.5">10.5 mm</quantity>
, vertical diameter of the largest ring 1.0 mm.
</paragraph>
<paragraph id="8BF736EAFFCC6960FF4F0845FB6EFD8B" blockId="5.[189,1210,594,621]" box="[189,1210,594,621]" pageId="5" pageNumber="44">
<smallCapsWord id="8D11A036FFCC6960FF4F0845FE9CFD8C" baselines="613,614" box="[189,328,594,620]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="5" pageNumber="44">COLORATION</smallCapsWord>
(
<figureCitation id="13732A6FFFCC6960FEAB0845FE4BFD8B" box="[345,415,594,621]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1838]" captionTargetId="figure-16@6.[189,1398,274,1838]" captionTargetPageId="6" captionText="Fig. 2. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Habitus, lateral view. B. Right field with ommatidia, lateral view. C. Head, anterior view, white arrows indicate rectangular lobes. D. Midbody rings, lateral view. E. Midbody rings, dorsal view. Scale bars: A, DE = 1 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149304" httpUri="https://zenodo.org/record/10149304/files/figure.png" pageId="5" pageNumber="44">Fig. 2</figureCitation>
). With brownish, greyish and yellowish patterns. Ommatidia black.
</paragraph>
<paragraph id="8BF736EAFFCC6960FF4F0883FC8CFC0D" blockId="5.[186,1400,659,1003]" pageId="5" pageNumber="44">
<smallCapsWord id="8D11A036FFCC6960FF4F0883FF28FD4B" baselines="679,679" box="[189,252,660,686]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="5" pageNumber="44">HEAD</smallCapsWord>
(
<figureCitation id="13732A6FFFCC6960FEFE0884FE52FD49" box="[268,390,659,686]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1838]" captionTargetId="figure-16@6.[189,1398,274,1838]" captionTargetPageId="6" captionText="Fig. 2. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Habitus, lateral view. B. Right field with ommatidia, lateral view. C. Head, anterior view, white arrows indicate rectangular lobes. D. Midbody rings, lateral view. E. Midbody rings, dorsal view. Scale bars: A, DE = 1 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149304" httpUri="https://zenodo.org/record/10149304/files/figure.png" pageId="5" pageNumber="44">Fig. 2BC</figureCitation>
). Setose, frontal side with a slight medial concavity and with a rectangular lobe (
<figureCitation id="13732A6FFFCC6960FAEA0884FAA5FD49" box="[1304,1393,659,686]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1838]" captionTargetId="figure-16@6.[189,1398,274,1838]" captionTargetPageId="6" captionText="Fig. 2. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Habitus, lateral view. B. Right field with ommatidia, lateral view. C. Head, anterior view, white arrows indicate rectangular lobes. D. Midbody rings, lateral view. E. Midbody rings, dorsal view. Scale bars: A, DE = 1 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149304" httpUri="https://zenodo.org/record/10149304/files/figure.png" pageId="5" pageNumber="44">Fig. 2C</figureCitation>
, white arrows), below each antennal socket. Labrum with three medial teeth and 5+4 labral and 2+2 supralabral setae. Promentum triangular, without setae. Lamellae linguales with 4+4 setae. Stipites with ca 20 setae each. Antennae
<quantity id="4CB09B0FFFCC6960FDF208EAFD8FFCFF" box="[512,603,765,792]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="44" unit="mm" value="1.8">1.8 mm</quantity>
long. Length of antennomeres (in mm): I (0.08), II (0.2), III (0.53), IV (0.22), V (0.47), VI (0.13), VII (0.14) and VIII (0.03). Length/breadth ratios of antennomeres IVII: I (1), II (2), III (5), IV (2.2), V (3.6), VI (1) and VII (1.5). Antennomeres II, IV, V, VI and VII with one, three, one, four and one long sensillum trichoideum, respectively. Antennomere 7 with one rather bacilliform sensillum (sensillum basiconicum?) curved distad, located below sensillum trichodeum. Lateral to antennal sockets a group of papilliform outgrowths present. Number of ommatidia: 12+
<quantity id="4CB09B0FFFCC6960FACE09BAFAA3FC20" box="[1340,1399,941,967]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="5" pageNumber="44" unit="in" value="13.0">13 in</quantity>
3 or 4 rows, arranged in elongated triangles (
<figureCitation id="13732A6FFFCC6960FD3909C7FC9FFC0D" box="[715,843,976,1002]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1838]" captionTargetId="figure-16@6.[189,1398,274,1838]" captionTargetPageId="6" captionText="Fig. 2. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Habitus, lateral view. B. Right field with ommatidia, lateral view. C. Head, anterior view, white arrows indicate rectangular lobes. D. Midbody rings, lateral view. E. Midbody rings, dorsal view. Scale bars: A, DE = 1 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149304" httpUri="https://zenodo.org/record/10149304/files/figure.png" pageId="5" pageNumber="44">Fig. 2BC</figureCitation>
).
</paragraph>
<paragraph id="8BF736EAFFCC6960FF4F0E05FDEBFBA8" blockId="5.[189,1398,1042,1103]" pageId="5" pageNumber="44">
<smallCapsWord id="8D11A036FFCC6960FF4F0E05FECFFBCD" baselines="1061,1061" box="[189,283,1042,1068]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Collum" pageId="5" pageNumber="44">COLLUM</smallCapsWord>
. Narrower than head, with six macrochaetae as all body rings. Anterior edge semi-circular, posterior margin gently concave.
</paragraph>
<paragraph id="8BF736EAFFCC6960FF4F0E60FE30FB30" blockId="5.[189,1399,1142,1240]" pageId="5" pageNumber="44">
<smallCapsWord id="8D11A036FFCC6960FF4F0E60FF2AFB68" baselines="1162,1162" box="[189,254,1143,1169]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="5" pageNumber="44">BODY</smallCapsWord>
<smallCapsWord id="8D11A036FFCC6960FEF50E6BFE91FB68" baselines="1162" box="[263,325,1148,1167]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="rings" pageId="5" pageNumber="44">RINGS</smallCapsWord>
(
<figureCitation id="13732A6FFFCC6960FEAA0E61FE2FFB77" box="[344,507,1142,1169]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1838]" captionTargetId="figure-16@6.[189,1398,274,1838]" captionTargetPageId="6" captionText="Fig. 2. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Habitus, lateral view. B. Right field with ommatidia, lateral view. C. Head, anterior view, white arrows indicate rectangular lobes. D. Midbody rings, lateral view. E. Midbody rings, dorsal view. Scale bars: A, DE = 1 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149304" httpUri="https://zenodo.org/record/10149304/files/figure.png" pageId="5" pageNumber="44">Fig. 2A, DE</figureCitation>
). With well-developed lateral keels. Macrochaetae long and rather rodlike, with longitudinal ribs, ending with spikes. CIX (ring 15) = 0.5; MIX (ring 15) ~ 1.5; PIX (ring 15) = 0.7; MA (ring15) ~ 100°.
</paragraph>
<paragraph id="8BF736EAFFCC6960FF4F0EE8FC29FADB" blockId="5.[189,1398,1278,1340]" pageId="5" pageNumber="44">
<smallCapsWord id="8D11A036FFCC6960FF4F0EE8FEC6FAF0" baselines="1298,1298" box="[189,274,1279,1305]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Telson" pageId="5" pageNumber="44">TELSON</smallCapsWord>
. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 distal setae. Paraprocts with 3+3 marginal setae in distal part.
</paragraph>
<paragraph id="8BF736EAFFCC6960FF4F0F74FAF0FA9A" blockId="5.[189,1316,1379,1405]" box="[189,1316,1379,1405]" pageId="5" pageNumber="44">
<smallCapsWord id="8D11A036FFCC6960FF4F0F74FEFAFA9C" baselines="1398,1398,1398,1398" box="[189,302,1379,1405]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg-pairs" pageId="5" pageNumber="44">LEG-PAIRS</smallCapsWord>
1
<smallCapsWord id="8D11A036FFCC6960FEB90F7FFEACFA9C" baselines="1398" box="[331,376,1384,1403]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="5" pageNumber="44">AND</smallCapsWord>
2. With tarsal combs; femora, postfemora and tibiae with long and robust setae.
</paragraph>
<paragraph id="8BF736EAFFCC6960FF4F0FB2FAEAF91C" blockId="5.[187,1400,1444,1788]" pageId="5" pageNumber="44">
<smallCapsWord id="8D11A036FFCC6960FF4F0FB2FF2BFA5A" baselines="1464,1464" box="[189,255,1445,1471]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="5" pageNumber="44">MALE</smallCapsWord>
<smallCapsWord id="8D11A036FFCC6960FEF70FBDFE83FA5A" baselines="1464" box="[261,343,1450,1469]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="sexual" pageId="5" pageNumber="44">SEXUAL</smallCapsWord>
<smallCapsWord id="8D11A036FFCC6960FEAE0FBDFE31FA5A" baselines="1464" box="[348,485,1450,1469]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="characters" pageId="5" pageNumber="44">CHARACTERS</smallCapsWord>
(
<figureCitation id="13732A6FFFCC6960FE070FB3FD86FA59" box="[501,594,1444,1470]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1838]" captionTargetId="figure-16@6.[189,1398,274,1838]" captionTargetPageId="6" captionText="Fig. 2. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Habitus, lateral view. B. Right field with ommatidia, lateral view. C. Head, anterior view, white arrows indicate rectangular lobes. D. Midbody rings, lateral view. E. Midbody rings, dorsal view. Scale bars: A, DE = 1 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149304" httpUri="https://zenodo.org/record/10149304/files/figure.png" pageId="5" pageNumber="44">Figs 2A</figureCitation>
,
<figureCitation id="13732A6FFFCC6960FDAD0FB3FDA4FA59" box="[607,624,1444,1470]" captionStart="Fig" captionStartId="7.[189,232,1822,1848]" captionTargetBox="[189,1398,269,1764]" captionTargetId="figure-12@7.[189,1398,269,1764]" captionTargetPageId="7" captionText="Fig. 3. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Leg-pair 2, posterior view. B. Left leg 3, anterior view, white arrow indicates the distoventral femoral pad. C. Right leg 3 with a distoventral femoral pad, anterior view. D. Leg-pair 7, anterior view. E. Leg-pair 10, anterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149306" httpUri="https://zenodo.org/record/10149306/files/figure.png" pageId="5" pageNumber="44">3</figureCitation>
). Gonopores mesally on coxae 2 (
<figureCitation id="13732A6FFFCC6960FC060FB3FB84FA59" box="[1012,1104,1444,1470]" captionStart="Fig" captionStartId="7.[189,232,1822,1848]" captionTargetBox="[189,1398,269,1764]" captionTargetId="figure-12@7.[189,1398,269,1764]" captionTargetPageId="7" captionText="Fig. 3. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Leg-pair 2, posterior view. B. Left leg 3, anterior view, white arrow indicates the distoventral femoral pad. C. Right leg 3 with a distoventral femoral pad, anterior view. D. Leg-pair 7, anterior view. E. Leg-pair 10, anterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149306" httpUri="https://zenodo.org/record/10149306/files/figure.png" pageId="5" pageNumber="44">Fig. 3A</figureCitation>
). Leg-pairs 37 enlarged, especially leg-pairs 3, 4 and 7 (
<figureCitation id="13732A6FFFCC6960FDC90FD0FD49FA05" box="[571,669,1479,1506]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1838]" captionTargetId="figure-16@6.[189,1398,274,1838]" captionTargetPageId="6" captionText="Fig. 2. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Habitus, lateral view. B. Right field with ommatidia, lateral view. C. Head, anterior view, white arrows indicate rectangular lobes. D. Midbody rings, lateral view. E. Midbody rings, dorsal view. Scale bars: A, DE = 1 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149304" httpUri="https://zenodo.org/record/10149304/files/figure.png" pageId="5" pageNumber="44">Fig. 2A</figureCitation>
). Leg-pairs 3 and 4 very thick, each with a proximal lateral protrusion on prefemora; prefemora and femora strong, rectangular; tarsi shorter and thicker compared to other legs (
<figureCitation id="13732A6FFFCC6960FE940C19FE10F9CF" box="[358,452,1550,1576]" captionStart="Fig" captionStartId="7.[189,232,1822,1848]" captionTargetBox="[189,1398,269,1764]" captionTargetId="figure-12@7.[189,1398,269,1764]" captionTargetPageId="7" captionText="Fig. 3. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Leg-pair 2, posterior view. B. Left leg 3, anterior view, white arrow indicates the distoventral femoral pad. C. Right leg 3 with a distoventral femoral pad, anterior view. D. Leg-pair 7, anterior view. E. Leg-pair 10, anterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149306" httpUri="https://zenodo.org/record/10149306/files/figure.png" pageId="5" pageNumber="44">Fig. 3B</figureCitation>
). Leg-pair 3 with a distoventral femoral pad (
<figureCitation id="13732A6FFFCC6960FC020C19FBA6F9CF" box="[1008,1138,1550,1576]" captionStart="Fig" captionStartId="7.[189,232,1822,1848]" captionTargetBox="[189,1398,269,1764]" captionTargetId="figure-12@7.[189,1398,269,1764]" captionTargetPageId="7" captionText="Fig. 3. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Leg-pair 2, posterior view. B. Left leg 3, anterior view, white arrow indicates the distoventral femoral pad. C. Right leg 3 with a distoventral femoral pad, anterior view. D. Leg-pair 7, anterior view. E. Leg-pair 10, anterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149306" httpUri="https://zenodo.org/record/10149306/files/figure.png" pageId="5" pageNumber="44">Fig. 3BC</figureCitation>
); observed on neither other podomeres nor legs. Leg-pair 4 with a mesal concavity on prefemora and femora. Leg-pair 5 with a proximal, anterior, triangular coxal protrusion. Leg-pair 6 without peculiarities. Leg-pair 7 robust; coxae with well-developed, subrectangular, flattened posterior processes, covered with long setae anteriorly, each with a lateral tooth curved anteriad (
<figureCitation id="13732A6FFFCC6960FD420C8CFCC5F952" box="[688,785,1691,1717]" captionStart="Fig" captionStartId="7.[189,232,1822,1848]" captionTargetBox="[189,1398,269,1764]" captionTargetId="figure-12@7.[189,1398,269,1764]" captionTargetPageId="7" captionText="Fig. 3. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Leg-pair 2, posterior view. B. Left leg 3, anterior view, white arrow indicates the distoventral femoral pad. C. Right leg 3 with a distoventral femoral pad, anterior view. D. Leg-pair 7, anterior view. E. Leg-pair 10, anterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149306" httpUri="https://zenodo.org/record/10149306/files/figure.png" pageId="5" pageNumber="44">Fig. 3D</figureCitation>
). Leg-pair 10 with coxal glands and with a bilobed protrusion, triangular anteriorly and rounded posteriorly, both lobes covered with tubercules (
<figureCitation id="13732A6FFFCC6960FAEA0CA9FAA7F93F" box="[1304,1395,1726,1752]" captionStart="Fig" captionStartId="7.[189,232,1822,1848]" captionTargetBox="[189,1398,269,1764]" captionTargetId="figure-12@7.[189,1398,269,1764]" captionTargetPageId="7" captionText="Fig. 3. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). A. Leg-pair 2, posterior view. B. Left leg 3, anterior view, white arrow indicates the distoventral femoral pad. C. Right leg 3 with a distoventral femoral pad, anterior view. D. Leg-pair 7, anterior view. E. Leg-pair 10, anterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149306" httpUri="https://zenodo.org/record/10149306/files/figure.png" pageId="5" pageNumber="44">Fig. 3E</figureCitation>
, only anterior, triangular part of lobe visible). Leg-pair 11 with coxal glands, no other peculiarities.
</paragraph>
<paragraph id="8BF736EAFFCC696DFF4F0D34FC65FE57" blockId="5.[189,1399,1826,1994]" lastBlockId="8.[188,1399,265,432]" lastPageId="8" lastPageNumber="47" pageId="5" pageNumber="44">
<smallCapsWord id="8D11A036FFCC6960FF4F0D34FEF8F8DC" baselines="1846,1846" box="[189,300,1827,1853]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Anterior" pageId="5" pageNumber="44">ANTERIOR</smallCapsWord>
<smallCapsWord id="8D11A036FFCC6960FECA0D3FFE7DF8DC" baselines="1846" box="[312,425,1832,1851]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="gonopods" pageId="5" pageNumber="44">GONOPODS</smallCapsWord>
(
<figureCitation id="13732A6FFFCC6960FE4C0D35FD9EF8DA" box="[446,586,1826,1853]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,829,1844]" captionTargetId="figure-162@8.[189,1398,829,1844]" captionTargetPageId="8" captionText="Fig. 4. AB. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). CD. Acanthophorella cf. A. aurita from Kveda Tlughi. A, C. Anterior gonopods, anterior and posterior views, respectively. B, D. Posterior gonopods, anterior views. Scale bars = 0.2 mm; CD not to scale." figureDoi="http://doi.org/10.5281/zenodo.10149308" httpUri="https://zenodo.org/record/10149308/files/figure.png" pageId="5" pageNumber="44">Figs 4A, C</figureCitation>
,
<figureCitation id="13732A6FFFCC6960FDA80D35FD77F8DA" box="[602,675,1826,1853]" captionStart="Fig" captionStartId="9.[189,232,1822,1848]" captionTargetBox="[199,1388,793,1738]" captionTargetId="figure-147@9.[189,1398,773,1781]" captionTargetPageId="9" captionText="Fig. 5. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363), gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, anterior view. E. Right posterior gonopod, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; cv = coxal vesicle; ll = lateral lamella; mp = medial part; s = sternum; sl = sternal lamella; t = telopodital pigment remnants; tf = angiocoxal tuft. Drawings by DA. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149310" httpUri="https://zenodo.org/record/10149310/files/figure.png" pageId="5" pageNumber="44">5AC</figureCitation>
,
<figureCitation id="13732A6FFFCC6960FD460D34FD32F8DA" box="[692,742,1827,1853]" captionStart="Fig" captionStartId="33.[189,232,1822,1848]" captionTargetBox="[202,1378,705,1770]" captionTargetId="figure-170@33.[189,1398,686,1780]" captionTargetPageId="33" captionText="Fig. 21. Anterior gonopods in the genera Acanthophorella Antić &amp; Makarov, 2016 and PseudoflagellophorellaAntić &amp; Makarov, 2016, anterior views. A. A. auritaAntić sp. nov. B. A. barjadzei Antić &amp; Makarov, 2016. C. A. chegemi Antić &amp; Makarov, 2016. D. A. devi Antić sp. nov. E. A. irystoni Antić &amp; Makarov, 2016. F. A. valerii Antić sp. nov. G. P. eskovi Antić &amp; Makarov, 2016. H. P. mirabilis Antić &amp; Makarov, 2016. I. P. papilioformis Antić &amp; Makarov, 2016. BC, E, GI after Antić &amp; Makarov (2016). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149345" httpUri="https://zenodo.org/record/10149345/files/figure.png" pageId="5" pageNumber="44">21A</figureCitation>
). Gonopodal sternum (s) wide, medially with a well-developed, subtriangular, fimbriate lamella (sl) on anterior side. Angiocoxites (a) consisting of a medial part (mp), lateral lamellae (ll) and a synangiocoxal base with anterior processes (ap). Medial parts well-developed, divided, but appressed to each other, shield like, tapering distolaterally into hook-shaped processes in anterior and posterior views, curved sharply proximad and posteriad; distally these processes wide, rather spoon-shaped; distomesal margins folded posteriad; additional three processes can be seen posteriorly (best seen in lateral view); angiocoxites posteroproximally with a pair of tufts (tf) with hairlike outgrowths. Lateral lamellae high, curving posterolaterad into two strongly developed layers of fringed extension. Anterior processes (ap) tapering distad, blunt, short, high as half as height of medial angiocoxal part. Coxal vesicle (?) (cv) poorly developed.
</paragraph>
<caption id="DF376662FFCF6963FF4F0D73FEA7F80E" ID-DOI="http://doi.org/10.5281/zenodo.10149304" ID-Zenodo-Dep="10149304" httpUri="https://zenodo.org/record/10149304/files/figure.png" pageId="6" pageNumber="45" startId="6.[189,232,1892,1918]" targetBox="[189,1398,274,1838]" targetPageId="6">
<paragraph id="8BF736EAFFCF6963FF4F0D73FEA7F80E" blockId="6.[189,1399,1891,2025]" pageId="6" pageNumber="45">
<emphasis id="B93CEAF8FFCF6963FF4F0D73FEDEF898" bold="true" box="[189,266,1892,1919]" pageId="6" pageNumber="45">Fig. 2.</emphasis>
<taxonomicName id="4C484D69FFCF6963FEFD0D72FDB2F898" authority="Antic" authorityName="Antic" authorityYear="2023" box="[271,614,1893,1919]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="aurita" status="sp. nov.">
<emphasis id="B93CEAF8FFCF6963FEFD0D72FDF5F898" box="[271,545,1893,1919]" italics="true" pageId="6" pageNumber="45">Acanthophorella aurita</emphasis>
Antić
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFCF6963FD9E0D72FD13F898" box="[620,711,1893,1919]" pageId="6" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMW MY10363).
<emphasis id="B93CEAF8FFCF6963FBAD0D73FBA0F899" bold="true" box="[1119,1140,1892,1918]" pageId="6" pageNumber="45">A</emphasis>
. Habitus, lateral view.
<emphasis id="B93CEAF8FFCF6963FF4F0D9FFF05F845" bold="true" box="[189,209,1928,1954]" pageId="6" pageNumber="45">B</emphasis>
. Right field with ommatidia, lateral view.
<emphasis id="B93CEAF8FFCF6963FD3E0D9FFD35F845" bold="true" box="[716,737,1928,1954]" pageId="6" pageNumber="45">C</emphasis>
. Head, anterior view, white arrows indicate rectangular lobes.
<emphasis id="B93CEAF8FFCF6963FEFF0DBCFEF6F822" bold="true" box="[269,290,1963,1989]" pageId="6" pageNumber="45">D</emphasis>
. Midbody rings, lateral view.
<emphasis id="B93CEAF8FFCF6963FD660DBCFD7CF822" bold="true" box="[660,680,1963,1989]" pageId="6" pageNumber="45">E</emphasis>
. Midbody rings, dorsal view. Scale bars: A, DE = 1 mm; BC = 0.5 mm.
</paragraph>
</caption>
<caption id="DF376662FFCE6962FF4F0D09FE63F845" ID-DOI="http://doi.org/10.5281/zenodo.10149306" ID-Zenodo-Dep="10149306" httpUri="https://zenodo.org/record/10149306/files/figure.png" pageId="7" pageNumber="46" startId="7.[189,232,1822,1848]" targetBox="[189,1398,269,1764]" targetPageId="7">
<paragraph id="8BF736EAFFCE6962FF4F0D09FE63F845" blockId="7.[188,1399,1820,1954]" pageId="7" pageNumber="46">
<emphasis id="B93CEAF8FFCE6962FF4F0D09FEDEF8DF" bold="true" box="[189,266,1822,1848]" pageId="7" pageNumber="46">Fig. 3.</emphasis>
<taxonomicName id="4C484D69FFCE6962FEE20D09FDBCF8DF" authority="Antic" authorityName="Antic" authorityYear="2023" box="[272,616,1822,1848]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="aurita" status="sp. nov.">
<emphasis id="B93CEAF8FFCE6962FEE20D09FDF7F8DF" box="[272,547,1822,1848]" italics="true" pageId="7" pageNumber="46">Acanthophorella aurita</emphasis>
Antić
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFCE6962FD9C0D08FD1EF8DE" box="[622,714,1823,1849]" pageId="7" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMW MY10363).
<emphasis id="B93CEAF8FFCE6962FB960D09FBADF8DF" bold="true" box="[1124,1145,1822,1848]" pageId="7" pageNumber="46">A</emphasis>
. Leg-pair 2, posterior view.
<emphasis id="B93CEAF8FFCE6962FEF00D56FEC2F8BC" bold="true" box="[258,278,1857,1883]" pageId="7" pageNumber="46">B</emphasis>
. Left leg 3, anterior view, white arrow indicates the distoventral femoral pad.
<emphasis id="B93CEAF8FFCE6962FB6A0D56FB79F8BC" bold="true" box="[1176,1197,1857,1883]" pageId="7" pageNumber="46">C</emphasis>
. Right leg 3 with a distoventral femoral pad, anterior view.
<emphasis id="B93CEAF8FFCE6962FD470D72FD1CF898" bold="true" box="[693,712,1893,1919]" pageId="7" pageNumber="46">D</emphasis>
. Leg-pair 7, anterior view.
<emphasis id="B93CEAF8FFCE6962FBE70D72FBF3F898" bold="true" box="[1045,1063,1893,1919]" pageId="7" pageNumber="46">E</emphasis>
. Leg-pair 10, anterior view. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BF736EAFFC1696DFF4F0BCFFD1FFD99" blockId="8.[188,1400,471,639]" pageId="8" pageNumber="47">
<smallCapsWord id="8D11A036FFC1696DFF4F0BCFFEE6FE17" baselines="491,491" box="[189,306,472,498]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Posterior" pageId="8" pageNumber="47">POSTERIOR</smallCapsWord>
<smallCapsWord id="8D11A036FFC1696DFECE0BCAFE79FE17" baselines="491" box="[316,429,477,496]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="gonopods" pageId="8" pageNumber="47">GONOPODS</smallCapsWord>
(
<figureCitation id="13732A6FFFC1696DFE320BC0FD90FE15" box="[448,580,471,498]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,829,1844]" captionTargetId="figure-162@8.[189,1398,829,1844]" captionTargetPageId="8" captionText="Fig. 4. AB. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). CD. Acanthophorella cf. A. aurita from Kveda Tlughi. A, C. Anterior gonopods, anterior and posterior views, respectively. B, D. Posterior gonopods, anterior views. Scale bars = 0.2 mm; CD not to scale." figureDoi="http://doi.org/10.5281/zenodo.10149308" httpUri="https://zenodo.org/record/10149308/files/figure.png" pageId="8" pageNumber="47">Figs 4B, D</figureCitation>
,
<figureCitation id="13732A6FFFC1696DFDA60BC0FD4DFE15" box="[596,665,471,498]" captionStart="Fig" captionStartId="9.[189,232,1822,1848]" captionTargetBox="[199,1388,793,1738]" captionTargetId="figure-147@9.[189,1398,773,1781]" captionTargetPageId="9" captionText="Fig. 5. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363), gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, anterior view. E. Right posterior gonopod, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; cv = coxal vesicle; ll = lateral lamella; mp = medial part; s = sternum; sl = sternal lamella; t = telopodital pigment remnants; tf = angiocoxal tuft. Drawings by DA. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149310" httpUri="https://zenodo.org/record/10149310/files/figure.png" pageId="8" pageNumber="47">5DE</figureCitation>
). Gonopodal sternum (s) wide, well-developed. Angiocoxites (a) strongly developed, more or less ear-shaped in anterior and posterior views, with lateral and distal margins curved anteriad; proximally with a short anterior process (ap). Coxal vesicles (cv) poorly developed, visible on mesal sides of angiocoxites at their mid-height. Telopodites (t) reduced to pigment remnants (a claw is visible on the right side).
</paragraph>
</subSubSection>
<caption id="DF376662FFC1696DFF4F0D73FAB0F822" ID-DOI="http://doi.org/10.5281/zenodo.10149308" ID-Zenodo-Dep="10149308" httpUri="https://zenodo.org/record/10149308/files/figure.png" pageId="8" pageNumber="47" startId="8.[189,232,1892,1918]" targetBox="[189,1398,829,1844]" targetPageId="8">
<paragraph id="8BF736EAFFC1696DFF4F0D73FAB0F822" blockId="8.[188,1399,1891,1990]" pageId="8" pageNumber="47">
<emphasis id="B93CEAF8FFC1696DFF4F0D73FEBAF899" bold="true" box="[189,366,1892,1919]" pageId="8" pageNumber="47">Fig. 4. AB</emphasis>
.
<taxonomicName id="4C484D69FFC1696DFE7C0D72FCD8F898" authority="Antic" authorityName="Antic" authorityYear="2023" box="[398,780,1893,1919]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="aurita" status="sp. nov.">
<emphasis id="B93CEAF8FFC1696DFE7C0D72FD60F898" box="[398,692,1893,1919]" italics="true" pageId="8" pageNumber="47">Acanthophorella aurita</emphasis>
Antić
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC1696DFCD40D72FC41F898" box="[806,917,1893,1919]" pageId="8" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMW MY10363).
<emphasis id="B93CEAF8FFC1696DFF4F0D9FFF22F845" bold="true" box="[189,246,1928,1954]" pageId="8" pageNumber="47">CD</emphasis>
.
<taxonomicName id="4C484D69FFC1696DFEF60D9FFE1CF845" authorityName="Antic &amp; Makarov" authorityYear="2016" box="[260,456,1928,1954]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAF8FFC1696DFEF60D9FFE1CF845" box="[260,456,1928,1954]" italics="true" pageId="8" pageNumber="47">Acanthophorella</emphasis>
</taxonomicName>
cf.
<taxonomicName id="4C484D69FFC1696DFE000D9FFD8DF845" authorityName="Antic" authorityYear="2023" box="[498,601,1928,1954]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="aurita">
<emphasis id="B93CEAF8FFC1696DFE000D9FFD8DF845" box="[498,601,1928,1954]" italics="true" pageId="8" pageNumber="47">A. aurita</emphasis>
</taxonomicName>
from Kveda Tlughi.
<emphasis id="B93CEAF8FFC1696DFCBF0D9FFC51F845" bold="true" box="[845,901,1928,1954]" pageId="8" pageNumber="47">A, C</emphasis>
. Anterior gonopods, anterior and posterior views, respectively.
<emphasis id="B93CEAF8FFC1696DFE590DBCFE36F822" bold="true" box="[427,482,1963,1989]" pageId="8" pageNumber="47">B, D</emphasis>
. Posterior gonopods, anterior views. Scale bars = 0.2 mm; CD not to scale.
</paragraph>
</caption>
<subSubSection id="C3526561FFC0696FFF4F0B1EFAD7FE8E" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" type="discussion">
<paragraph id="8BF736EAFFC0696CFF4F0B1EFEFBFEC4" blockId="9.[189,303,265,291]" box="[189,303,265,291]" pageId="9" pageNumber="48">
<heading id="D0BF8186FFC0696CFF4F0B1EFEFBFEC4" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="9" pageNumber="48" reason="3">
<emphasis id="B93CEAF8FFC0696CFF4F0B1EFEFBFEC4" bold="true" box="[189,303,265,291]" pageId="9" pageNumber="48">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFC0696CFF4F0B20FC63FDA0" blockId="9.[188,1399,311,584]" pageId="9" pageNumber="48">
An apparently conspecific specimen was found in the forest, near the Cholaga spring, village Kveda Tlughi, in the Racha region. This locality is about 100 air-km from the
<typeStatus id="54F38848FFC0696CFC1D0B4CFBF6FE92" box="[1007,1058,347,373]" pageId="9" pageNumber="48">type</typeStatus>
and only locality of
<taxonomicName id="4C484D69FFC0696CFAFC0B4CFAA2FE93" authority="Antić &amp; Šević &amp; Barjadze &amp; Makarov, 2023" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[1294,1398,347,373]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="aurita" status="sp. nov.">
<emphasis id="B93CEAF8FFC0696CFAFC0B4CFAA2FE93" box="[1294,1398,347,373]" italics="true" pageId="9" pageNumber="48">A. aurita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC0696CFF4F0B69FECCFE7F" box="[189,280,382,408]" pageId="9" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
Unfortunately, the sample from Kveda Tlughi is considered lost in the
<collectionCode id="ED59AE2FFFC0696CFB930B69FB78FE7F" box="[1121,1196,382,408]" pageId="9" pageNumber="48">IZISU</collectionCode>
collection. From the available pictures of the gonopods alone (
<figureCitation id="13732A6FFFC0696CFD2A0BB6FC83FE5C" box="[728,855,417,443]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,829,1844]" captionTargetId="figure-162@8.[189,1398,829,1844]" captionTargetPageId="8" captionText="Fig. 4. AB. Acanthophorella aurita Antić sp. nov., holotype, ♂ (NHMW MY10363). CD. Acanthophorella cf. A. aurita from Kveda Tlughi. A, C. Anterior gonopods, anterior and posterior views, respectively. B, D. Posterior gonopods, anterior views. Scale bars = 0.2 mm; CD not to scale." figureDoi="http://doi.org/10.5281/zenodo.10149308" httpUri="https://zenodo.org/record/10149308/files/figure.png" pageId="9" pageNumber="48">Fig. 4CD</figureCitation>
) it is evident that both males are very similar, although the shape of the angiocoxites of the posterior gonopods is slightly different. Additional males from both localities, as well as in between, could clarify whether it is only a variation in the shape of the angiocoxites of the posterior gonopods, which seems most likely. The locality Kveda Tlughi is marked with a question mark on the map for
<taxonomicName id="4C484D69FFC0696CFD9C0839FD03FDA0" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[622,727,558,584]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="aurita">
<emphasis id="B93CEAF8FFC0696CFD9C0839FD03FDA0" box="[622,727,558,584]" italics="true" pageId="9" pageNumber="48">A. aurita</emphasis>
</taxonomicName>
for the time being.
</paragraph>
<caption id="DF376662FFC0696CFF4F0D09FEBAF80E" ID-DOI="http://doi.org/10.5281/zenodo.10149310" ID-Zenodo-Dep="10149310" httpUri="https://zenodo.org/record/10149310/files/figure.png" pageId="9" pageNumber="48" startId="9.[189,232,1822,1848]" targetBox="[199,1388,793,1738]" targetPageId="9">
<paragraph id="8BF736EAFFC0696CFF4F0D09FEBAF80E" blockId="9.[187,1399,1820,2025]" pageId="9" pageNumber="48">
<emphasis id="B93CEAF8FFC0696CFF4F0D09FEDFF8DF" bold="true" box="[189,267,1822,1848]" pageId="9" pageNumber="48">Fig. 5.</emphasis>
<taxonomicName id="4C484D69FFC0696CFEE10D09FDB8F8DF" authority="Antic" authorityName="Antic" authorityYear="2023" box="[275,620,1822,1848]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="aurita" status="sp. nov.">
<emphasis id="B93CEAF8FFC0696CFEE10D09FDF2F8DF" box="[275,550,1822,1848]" italics="true" pageId="9" pageNumber="48">Acanthophorella aurita</emphasis>
Antić
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC0696CFD860D08FD05F8DE" box="[628,721,1823,1849]" pageId="9" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMW MY10363), gonopods.
<emphasis id="B93CEAF8FFC0696CFB030D09FAD2F8DF" bold="true" box="[1265,1286,1822,1848]" pageId="9" pageNumber="48">A</emphasis>
. Anterior gonopods, anterior view.
<emphasis id="B93CEAF8FFC0696CFE010D55FDD3F8BB" bold="true" box="[499,519,1858,1884]" pageId="9" pageNumber="48">B</emphasis>
. Anterior gonopods, posterior view.
<emphasis id="B93CEAF8FFC0696CFC3B0D56FC0AF8BC" bold="true" box="[969,990,1857,1883]" pageId="9" pageNumber="48">C</emphasis>
. Anterior gonopods, lateral view.
<emphasis id="B93CEAF8FFC0696CFF4F0D72FF06F898" bold="true" box="[189,210,1893,1919]" pageId="9" pageNumber="48">D</emphasis>
. Posterior gonopods, anterior view.
<emphasis id="B93CEAF8FFC0696CFD600D72FD72F898" bold="true" box="[658,678,1893,1919]" pageId="9" pageNumber="48">E</emphasis>
. Right posterior gonopod, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; cv = coxal vesicle; ll = lateral lamella; mp = medial part; s = sternum; sl = sternal lamella; t = telopodital pigment remnants; tf = angiocoxal tuft. Drawings by DA. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BF736EAFFC3696FFF4F0B1EFAD7FE8E" blockId="10.[189,1398,265,362]" pageId="10" pageNumber="49">
Near the
<typeStatus id="54F38848FFC3696FFEC10B1EFEB2FEC4" box="[307,358,265,291]" pageId="10" pageNumber="49">type</typeStatus>
locality, in Urta (= Jegeta 1) Cave, we recorded a larger number of juveniles (five collected,
<collectionCode id="ED59AE2FFFC3696FFEC50B3BFE50FEA1" box="[311,388,300,326]" pageId="10" pageNumber="49">IZISU</collectionCode>
) during the 2022 expedition, but in the absence of an adult male we shall not assign these specimens to
<taxonomicName id="4C484D69FFC3696FFE520B47FDDDFE8E" authority="Antić &amp; Šević &amp; Barjadze &amp; Makarov, 2023" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[416,521,336,362]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="aurita" status="sp. nov.">
<emphasis id="B93CEAF8FFC3696FFE520B47FDDDFE8E" box="[416,521,336,362]" italics="true" pageId="10" pageNumber="49">A. aurita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC3696FFDE20B47FDB9FE8D" box="[528,621,336,362]" pageId="10" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
, although it is very likely that they are the same species.
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFC3696FFF4F0B87FC24FCB0" pageId="10" pageNumber="49" type="biology_ecology">
<paragraph id="8BF736EAFFC3696FFF4F0B87FE05FE4C" blockId="10.[189,465,400,427]" box="[189,465,400,427]" pageId="10" pageNumber="49">
<heading id="D0BF8186FFC3696FFF4F0B87FE05FE4C" bold="true" box="[189,465,400,427]" fontSize="11" level="3" pageId="10" pageNumber="49" reason="3">
<emphasis id="B93CEAF8FFC3696FFF4F0B87FE05FE4C" bold="true" box="[189,465,400,427]" pageId="10" pageNumber="49">Localities and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFC3696FFF4F0BA8FC5DFD14" blockId="10.[188,1399,447,755]" pageId="10" pageNumber="49">
The
<typeStatus id="54F38848FFC3696FFF000BA8FE8DFE3E" box="[242,345,447,473]" pageId="10" pageNumber="49" type="holotype">holotype</typeStatus>
male was collected in a small, slightly descending cavity with a small entrance, and with a hall about
<quantity id="4CB09B0FFFC3696FFEA80BF5FE01FE1A" box="[346,469,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="10" pageNumber="49" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">33.5 mm</quantity>
long and
<quantity id="4CB09B0FFFC3696FFDA70BF5FD6DFE1A" box="[597,697,482,509]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="10" pageNumber="49" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">1.52 m</quantity>
wide. This cavity is characterised by an additional small opening on the ceiling, causing the floor to be covered with moist, black, humic soil with leaf litter and a thin layer of dead leaves. The specimen was found in this substrate (Jozef Grego pers. com.). Another millipede found in the cavity was
<taxonomicName id="4C484D69FFC3696FFDBD085BFBFBFD81" authority="Golovatch, 1977" authorityName="Golovatch" authorityYear="1977" box="[591,1071,588,614]" class="Diplopoda" family="Julidae" genus="Pachyiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="krivolutskyi">
<emphasis id="B93CEAF8FFC3696FFDBD085BFCB6FD81" box="[591,866,588,614]" italics="true" pageId="10" pageNumber="49">Pachyiulus krivolutskyi</emphasis>
Golovatch, 1977
</taxonomicName>
, a common epigean species in the western Caucasus. The cavity is tentatively named Jegeta 2 Cave or Cave 2 near the ruins of Jegeta church. The sample from Kveda Tlughi, which is most likely
<taxonomicName id="4C484D69FFC3696FFC590884FBC3FD4B" authority="Antić &amp; Šević &amp; Barjadze &amp; Makarov, 2023" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[939,1047,659,685]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="aurita" status="sp. nov.">
<emphasis id="B93CEAF8FFC3696FFC590884FBC3FD4B" box="[939,1047,659,685]" italics="true" pageId="10" pageNumber="49">A. aurita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC3696FFBD20884FB54FD4A" box="[1056,1152,659,685]" pageId="10" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
, as well, came from sieving leaf litter. In the Urta (= Jegeta 1) cave, juveniles of possibly this species were observed only in and on dead wood in the cave, but very close to the entrance.
</paragraph>
<paragraph id="8BF736EAFFC3696FFF4F090DFC24FCB0" blockId="10.[188,1399,794,856]" pageId="10" pageNumber="49">
This species is an epigean member of the genus
<taxonomicName id="4C484D69FFC3696FFD1C090DFC66FCD3" authorityName="Antic &amp; Makarov" authorityYear="2016" box="[750,946,794,820]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAF8FFC3696FFD1C090DFC66FCD3" box="[750,946,794,820]" italics="true" pageId="10" pageNumber="49">Acanthophorella</emphasis>
</taxonomicName>
and as juveniles possibly belonging to this species were found in the cave, it may be considered troglophilic.
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFC3696FFF4F0968FCBDFC0C" pageId="10" pageNumber="49" type="distribution">
<paragraph id="8BF736EAFFC3696FFF4F0968FE82FC7E" blockId="10.[189,342,895,921]" box="[189,342,895,921]" pageId="10" pageNumber="49">
<heading id="D0BF8186FFC3696FFF4F0968FE82FC7E" bold="true" box="[189,342,895,921]" fontSize="11" level="3" pageId="10" pageNumber="49" reason="3">
<emphasis id="B93CEAF8FFC3696FFF4F0968FE82FC7E" bold="true" box="[189,342,895,921]" pageId="10" pageNumber="49">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFC3696FFF4F09B9FCBDFC0C" blockId="10.[189,1399,941,1003]" pageId="10" pageNumber="49">
A Georgian endemic known from the Urta Mountains (Khobi Municipality) and possibly from Kveda Tlughi (Ambrolauri Municipality) (
<figureCitation id="13732A6FFFC3696FFDAD09C6FD62FC0C" box="[607,694,977,1003]" captionStart="Fig" captionStartId="35.[189,232,1998,2024]" captionTargetBox="[189,1398,1153,1959]" captionTargetId="figure-355@35.[189,1398,1150,1959]" captionTargetPageId="35" captionText="Fig. 22. Distribution of the genus Acanthophorella Antić &amp; Makarov, 2016." figureDoi="http://doi.org/10.5281/zenodo.10149347" httpUri="https://zenodo.org/record/10149347/files/figure.png" pageId="10" pageNumber="49">Fig. 22</figureCitation>
, green circles).
</paragraph>
</subSubSection>
</treatment>
</document>