treatments-xml/data/03/E1/87/03E187FBFFE7FF92FF34FF149E59FAE7.xml
2024-06-21 12:22:17 +02:00

413 lines
46 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="A0255410682DDD2665566BF693C2FC5F" ID-DOI="10.11646/zootaxa.3709.1.2" ID-GBIF-Dataset="64d5b399-d948-4189-8af8-75f5ffc6b62d" ID-ISSN="1175-5326" ID-Zenodo-Dep="222914" ID-ZooBank="A610056E-AE10-483A-B3E6-7D0C2AA168A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460673260853" checkinUser="plazi" docAuthor="Guéorguiev, Borislav" docDate="2013" docId="03E187FBFFE7FF92FF34FF149E59FAE7" docLanguage="en" docName="zt03709p070.pdf" docOrigin="Zootaxa 3709 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Oreophilus) podgoricensis Guéorguiev, 2013, sp. n." docType="treatment" docUuid="DB40A55B-7D70-4158-A1A8-BA038D02B1C1" docUuidSource="ZooBank" docVersion="10" masterDocId="FFD8FF83FFE0FF9BFFA3FF839C02FFD7" masterDocTitle="Taxonomic, nomenclatural, and faunistic records for species in tribes Melaenini, Moriomorphini, Pterostichini, Licinini, and Sphodrini (Coleoptera: Carabidae)" updateTime="1698348865835" updateUser="plazi">
<mods:mods id="8331C22049ECD402FB9A0669DBFD90E0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="590888C3C837B02B1B643AD7CECCD5A8">
<mods:title id="836353420A54B0B804BCF0AA8D0A8917">Taxonomic, nomenclatural, and faunistic records for species in tribes Melaenini, Moriomorphini, Pterostichini, Licinini, and Sphodrini (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="B2156614DE6D8059E14AC18136578BCA" type="personal">
<mods:role id="285AC2954988F473C2F9AAFE5A1EEDF6">
<mods:roleTerm id="CAA0DD6E5ACDCA044F741D69C8500285">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F397E0B04761E981D487A6BD992038A2">Guéorguiev, Borislav</mods:namePart>
</mods:name>
<mods:typeOfResource id="3BE2C9FA7A5203F031C305200D6A19AE">text</mods:typeOfResource>
<mods:relatedItem id="D3B1B42372568CE77BF31EDABBADCEE1" type="host">
<mods:titleInfo id="CA1C4346FD602E4C7F2D394D57B0FF07">
<mods:title id="A2167920A051D3E04C97CB7C453AAA1A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E13A2F28BF435603D1337425107EE5AD">
<mods:date id="D8D712D0692C9C3A0B43EE88D91F026A">2013</mods:date>
<mods:detail id="C5FF47CD7FE4C95DD4115C4CB6E3B015" type="volume">
<mods:number id="8972C36153CCA4805C38DE27218A2ACF">3709</mods:number>
</mods:detail>
<mods:detail id="A4B20B00B093FAF568B540396F5B59A0" type="issue">
<mods:number id="DD9E3FC9C6E428FD175B723A904D0B66">1</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="E64E47A854FB500671779EFB45F82CC6">journal volume</mods:classification>
<mods:identifier id="AB237432EE5D028F5BB6F9C13A9BE6BA" type="DOI">10.11646/zootaxa.3709.1.2</mods:identifier>
<mods:identifier id="753A3E6F49415709829DC38783EE12CD" type="GBIF-Dataset">64d5b399-d948-4189-8af8-75f5ffc6b62d</mods:identifier>
<mods:identifier id="E3B28E2D065E6452BB349D3A509B572E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DA818AA97E3523A6EF98F766B9798A1F" type="Zenodo-Dep">222914</mods:identifier>
<mods:identifier id="E9BB2186BCFA951A17AA52F3F1F0B175" type="ZooBank">A610056E-AE10-483A-B3E6-7D0C2AA168A1</mods:identifier>
</mods:mods>
<treatment id="03E187FBFFE7FF92FF34FF149E59FAE7" ID-DOI="http://doi.org/10.5281/zenodo.6150464" ID-GBIF-Taxon="119577705" ID-Zenodo-Dep="6150464" LSID="urn:lsid:zoobank.org:act:DB40A55B-7D70-4158-A1A8-BA038D02B1C1" httpUri="http://treatment.plazi.org/id/03E187FBFFE7FF92FF34FF149E59FAE7" lastPageId="9" pageId="7">
<subSubSection id="C3526566FFE7FF9CFF34FF149E87FF04" pageId="7" type="nomenclature">
<paragraph id="8BF736EDFFE7FF9CFF34FF149EC4FF65" blockId="7.[151,710,151,211]" box="[151,710,151,178]" pageId="7">
<heading id="D0BF8181FFE7FF9CFF34FF149EC4FF65" bold="true" box="[151,710,151,178]" fontSize="11" level="1" pageId="7" reason="1">
<emphasis id="B93CEAFFFFE7FF9CFF34FF149EC4FF65" bold="true" box="[151,710,151,178]" pageId="7">
<taxonomicName id="4C484D6EFFE7FF9CFF34FF149E7EFF66" ID-CoL="8MGX9" box="[151,636,151,178]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="podgoricensis" status="sp. nov." subGenus="Oreophilus">
<emphasis id="B93CEAFFFFE7FF9CFF34FF149D2FFF66" bold="true" box="[151,301,151,177]" italics="true" pageId="7">Pterostichus</emphasis>
(
<emphasis id="B93CEAFFFFE7FF9CFE9EFF149DC6FF66" bold="true" box="[317,452,151,177]" italics="true" pageId="7">Oreophilus</emphasis>
)
<emphasis id="B93CEAFFFFE7FF9CFE75FF149E7EFF66" bold="true" box="[470,636,151,177]" italics="true" pageId="7">podgoricensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5784FFE7FF9CFD20FF1B9EC4FF65" box="[643,710,152,178]" pageId="7" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EDFFE7FF9CFF34FF399E87FF04" blockId="7.[151,710,151,211]" box="[151,645,186,211]" pageId="7">
Habitus:
<figureCitation id="13732A68FFE7FF9CFF5EFF399D42FF04" box="[253,320,186,211]" captionStart="FIGURE 9" captionStartId="7.[151,250,1691,1713]" captionText="FIGURE 9. Holotype of Pterostichus (Oreophilus) podgoricensis sp. n. Photo by Assen Ihnatov." pageId="7">Fig. 9</figureCitation>
; male genitalia:
<figureCitation id="13732A68FFE7FF9CFE5AFF399E83FF04" box="[505,641,186,211]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1721,1743]" captionTargetBox="[161,1417,957,1660]" captionTargetId="figure@7.[151,1436,950,1666]" captionTargetPageId="7" captionText="FIGURES 10 15. Male genitalia of Pterostichus (Oreophilus) podgoricensis sp. n., holotype: Fig. 10, median lobe of aedeagus, left lateral aspect; Fig. 11, median lobe of aedeagus, right lateral aspect; Fig. 12, median lobe of aedeagus, ventral aspect; Fig. 13, right paramere, internal face; Fig. 14, right paramere, external face; Fig. 15, left paramere, internal face. Scale bar: 1 mm (Figs. 10 12); 0.5 mm (Figs. 13 15)." httpUri="https://zenodo.org/record/222919/files/figure.png" pageId="7">Figs. 1015</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3526566FFE7FF9CFF34FE819E08FD70" pageId="7" type="materials_examined">
<paragraph id="8BF736EDFFE7FF9CFF34FE819FB5FE24" blockId="7.[151,1436,258,895]" pageId="7">
<emphasis id="B93CEAFFFFE7FF9CFF34FE819D46FECC" bold="true" box="[151,324,258,283]" pageId="7">
<typeStatus id="54F3884FFFE7FF9CFF34FE819CD0FECC" box="[151,210,258,283]" pageId="7">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54F3884FFFE7FF9CFEFBFE819DC2FECC" box="[344,448,258,283]" pageId="7" type="holotype">Holotype</typeStatus>
♂, well preserved (no part to missing), mounted on card, genitalia dissected and deposited in euparal on separate card pinned below the first card, “NHRS-JLKB 000020046” [black print on white label] / “Titograd
<collectingCountry id="F35F767DFFE7FF9CFEFCFEC99DDCFEB4" box="[351,478,330,355]" name="Serbia" pageId="7">Yugoslavia</collectingCountry>
” [handwritten on white label] / “coll. J. Ferrer” [handwritten on white label] / “H O L O T Y P E
<taxonomicName id="4C484D6EFFE7FF9CFEB1FEED9E43FE51" box="[274,577,366,390]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="podgoricensis" status="sp. nov.">
<emphasis id="B93CEAFFFFE7FF9CFEB1FEED9E43FE51" box="[274,577,366,390]" italics="true" pageId="7">Pterostichus podgoricensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5784FFE7FF9CFDE4FEEC9E80FE51" box="[583,642,367,390]" pageId="7" rank="species">sp. n.</taxonomicNameLabel>
Guéorguiev des. 2012” [black print on red label] (
<collectionCode id="ED59AE28FFE7FF9CFB17FEEC9907FE50" box="[1204,1285,367,391]" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="7">NHRS</collectionCode>
).
<typeStatus id="54F3884FFFE7FF9CFAB4FEEC997BFE50" box="[1303,1401,367,391]" pageId="7" type="paratype">Paratype</typeStatus>
♀, slightly damaged (segments 911 of right antennae and fore legs missing), pinned, genitalia not dissected, wrongly labelled “Teneriffa Coll. O. Thieme” [black print on white label] “P A R A T Y P E
<taxonomicName id="4C484D6EFFE7FF9CFBE5FE359977FE19" box="[1094,1397,438,462]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="podgoricensis" status="sp. nov.">
<emphasis id="B93CEAFFFFE7FF9CFBE5FE359977FE19" box="[1094,1397,438,462]" italics="true" pageId="7">Pterostichus podgoricensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5784FFE7FF9CFADFFE349CAEFE25" pageId="7" rank="species">sp. n.</taxonomicNameLabel>
Guéorguiev des. 2012” [black print on red label] [
<figureCitation id="13732A68FFE7FF9CFD42FE599F24FE24" box="[737,806,474,499]" captionStart="FIGURE 9" captionStartId="7.[151,250,1691,1713]" captionText="FIGURE 9. Holotype of Pterostichus (Oreophilus) podgoricensis sp. n. Photo by Assen Ihnatov." pageId="7">Fig. 9</figureCitation>
] (
<collectionCode id="ED59AE28FFE7FF9CFC9FFE589FA9FE24" box="[828,939,475,499]" pageId="7">MNHUB</collectionCode>
).
</paragraph>
<paragraph id="8BF736EDFFE7FF9CFF64FE7D9E08FD70" blockId="7.[151,1436,258,895]" pageId="7">
<emphasis id="B93CEAFFFFE7FF9CFF64FE7D9DF9FDC0" bold="true" box="[199,507,510,535]" pageId="7">Other material examined</emphasis>
. Specimens of different European taxa of
<taxonomicName id="4C484D6EFFE7FF9CFC49FE7D98D6FDC0" authority="Bonelli" authorityName="Bonelli" box="[1002,1236,510,535]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFC49FE7D9877FDC1" box="[1002,1141,510,534]" italics="true" pageId="7">Pterostichus</emphasis>
Bonelli
</taxonomicName>
, namely those of
<taxonomicName id="4C484D6EFFE7FF9CFF34FDA09D7FFDEC" authority="Reitter" authorityName="Reitter" box="[151,381,546,571]" class="Insecta" family="Carabidae" genus="Adelopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFF34FDA09D1CFDEC" box="[151,286,547,571]" italics="true" pageId="7">Adelopterus</emphasis>
Reitter
</taxonomicName>
,
<taxonomicName id="4C484D6EFFE7FF9CFE32FDA09E51FDEC" authority="Heyden" authorityName="Heyden" box="[401,595,546,571]" class="Insecta" family="Carabidae" genus="Aphaon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFE32FDA09DEAFDEC" box="[401,488,547,571]" italics="true" pageId="7">Aphaon</emphasis>
Heyden
</taxonomicName>
,
<taxonomicName id="4C484D6EFFE7FF9CFDCBFDA09F5DFDEC" authority="Chaudoir" authorityName="Chaudoir" box="[616,863,546,571]" class="Insecta" family="Carabidae" genus="Calopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFDCBFDA09EE6FDEC" box="[616,740,547,571]" italics="true" pageId="7">Calopterus</emphasis>
Chaudoir
</taxonomicName>
,
<taxonomicName id="4C484D6EFFE7FF9CFCD7FDA09851FDEC" authority="Latreille" authorityName="Latreille" box="[884,1107,546,571]" class="Insecta" family="Carabidae" genus="Cheporus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFCD7FDA09FE3FDEC" box="[884,993,547,571]" italics="true" pageId="7">Cheporus</emphasis>
Latreille
</taxonomicName>
,
<taxonomicName id="4C484D6EFFE7FF9CFBCBFDA09908FDEC" authority="Gozis" authorityName="Gozis" box="[1128,1290,546,571]" class="Insecta" family="Carabidae" genus="Lianoe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFBCBFDA098B4FDED" box="[1128,1206,547,570]" italics="true" pageId="7">Lianoe</emphasis>
Gozis
</taxonomicName>
,
<taxonomicName id="4C484D6EFFE7FF9CFABCFDA09D03FD88" authority="Chaudoir" authorityName="Chaudoir" class="Insecta" family="Carabidae" genus="Oreophilus" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFABCFDA0999EFDEC" box="[1311,1436,547,571]" italics="true" pageId="7">Oreophilus</emphasis>
Chaudoir
</taxonomicName>
,
<taxonomicName id="4C484D6EFFE7FF9CFEAEFDC59E62FD88" authority="Desbrochers" authorityName="Desbrochers" box="[269,608,582,607]" class="Insecta" family="Carabidae" genus="Parapterostichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFEAEFDC59DCFFD89" box="[269,461,582,606]" italics="true" pageId="7">Parapterostichus</emphasis>
Desbrochers
</taxonomicName>
des Loges,
<taxonomicName id="4C484D6EFFE7FF9CFD4AFDC59FD4FD88" authority="Reitter" authorityName="Reitter" box="[745,982,582,607]" class="Insecta" family="Carabidae" genus="Platypterinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFD4AFDC59F82FD89" box="[745,896,582,606]" italics="true" pageId="7">Platypterinus</emphasis>
Reitter
</taxonomicName>
,
<taxonomicName id="4C484D6EFFE7FF9CFC41FDC598D7FD88" authority="Chaudoir" authorityName="Chaudoir" box="[994,1237,582,607]" class="Insecta" family="Carabidae" genus="Platypterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFC41FDC59861FD89" box="[994,1123,582,606]" italics="true" pageId="7">Platypterus</emphasis>
Chaudoir
</taxonomicName>
, and
<emphasis id="B93CEAFFFFE7FF9CFAB2FDC59CA0FD55" italics="true" pageId="7">
<taxonomicName id="4C484D6EFFE7FF9CFAB2FDC5999EFD89" box="[1297,1436,582,606]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus" sensu="stricto" status="s">Pterostichus</taxonomicName>
<taxonomicNameLabel id="A20F5784FFE7FF9CFF34FDE89CA0FD55" box="[151,162,619,642]" pageId="7" sensu="stricto">s</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B93CEAFFFFE7FF9CFF12FDE89CCFFD55" box="[177,205,619,642]" italics="true" pageId="7">str</emphasis>
. (in the sense of
<bibRefCitation id="EFD94B1CFFE7FF9CFE37FDE89E40FD54" author="Bousquet" box="[404,578,619,643]" pageId="7" refString="Bousquet, Y. (2003) Tribe Pterostichini Bonelli, 1810. In: Lobl I, Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga, Apollo Books, Stenstrup, pp. 469 - 521." type="book chapter" year="2003">Bousquet 2003</bibRefCitation>
). All the above mentioned subgenera include taxa having sternum VII with sexual modification in the males.
</paragraph>
</subSubSection>
<subSubSection id="C3526566FFE7FF9CFF64FD319E2CFCA9" pageId="7" type="diagnosis">
<paragraph id="8BF736EDFFE7FF9CFF64FD319E2CFCA9" blockId="7.[151,1436,258,895]" pageId="7">
<emphasis id="B93CEAFFFFE7FF9CFF64FD319D38FD1C" bold="true" box="[199,314,690,715]" pageId="7">Diagnosis</emphasis>
. Large, wingless, distinctly two-colored beetle, characterized by presence of sub-quadrate pronotum with dentiform basal angles, interval 3 with three foveolate setiferous punctures, apical end of stria 7 with four-five small setiferous punctures, sternum VII in male strongly depressed at sides and apical part, with stout medial, longitudinal projection and fine transverse subapical sulcus broken at carina (sternum VII in female with slight medial projection and fine transverse subapical sulcus broken at middle), median lobe of aedeagus stout, with apex clearly bent to right at dorsal aspect.
</paragraph>
</subSubSection>
<caption id="DF376665FFE7FF9CFF34F918986CF966" box="[151,1134,1691,1713]" pageId="7">
<paragraph id="8BF736EDFFE7FF9CFF34F918986CF966" blockId="7.[151,1436,1691,1835]" box="[151,1134,1691,1713]" pageId="7">
<emphasis id="B93CEAFFFFE7FF9CFF34F9189D0EF966" bold="true" box="[151,268,1691,1713]" pageId="7">FIGURE 9</emphasis>
. Holotype of
<taxonomicName id="4C484D6EFFE7FF9CFE3AF9189F31F966" box="[409,819,1691,1713]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="podgoricensis" status="sp. nov." subGenus="Oreophilus">
<emphasis id="B93CEAFFFFE7FF9CFE3AF9189E14F966" box="[409,534,1691,1713]" italics="true" pageId="7">Pterostichus</emphasis>
(
<emphasis id="B93CEAFFFFE7FF9CFD86F9189E94F966" box="[549,662,1691,1713]" italics="true" pageId="7">Oreophilus</emphasis>
)
<emphasis id="B93CEAFFFFE7FF9CFD06F9189F31F966" box="[677,819,1691,1713]" italics="true" pageId="7">podgoricensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5784FFE7FF9CFC9AF91F9F72F966" box="[825,880,1692,1713]" pageId="7" rank="species">sp. n.</taxonomicNameLabel>
Photo by Assen Ihnatov.
</paragraph>
</caption>
<caption id="DF376665FFE7FF9CFF34F93A9E7CF8FC" httpUri="https://zenodo.org/record/222919/files/figure.png" pageId="7" targetBox="[161,1417,957,1660]" targetPageId="7">
<paragraph id="8BF736EDFFE7FF9CFF34F93A9E7CF8FC" blockId="7.[151,1436,1691,1835]" pageId="7">
<emphasis id="B93CEAFFFFE7FF9CFF34F93A9D51F918" bold="true" box="[151,339,1721,1743]" pageId="7">FIGURES 1015</emphasis>
. Male genitalia of
<taxonomicName id="4C484D6EFFE7FF9CFD88F93A9FD3F918" box="[555,977,1721,1743]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="podgoricensis" status="sp. nov." subGenus="Oreophilus">
<emphasis id="B93CEAFFFFE7FF9CFD88F93A9EABF918" box="[555,681,1721,1743]" italics="true" pageId="7">Pterostichus</emphasis>
(
<emphasis id="B93CEAFFFFE7FF9CFD1EF93A9F2CF918" box="[701,814,1721,1743]" italics="true" pageId="7">Oreophilus</emphasis>
)
<emphasis id="B93CEAFFFFE7FF9CFCE0F93A9FD3F918" box="[835,977,1721,1743]" italics="true" pageId="7">podgoricensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5784FFE7FF9CFC7EF9399819F918" box="[989,1051,1722,1743]" pageId="7" rank="species">sp. n.</taxonomicNameLabel>
, holotype: Fig. 10, median lobe of aedeagus, left lateral aspect; Fig. 11, median lobe of aedeagus, right lateral aspect; Fig. 12, median lobe of aedeagus, ventral aspect; Fig. 13, right paramere, internal face; Fig. 14, right paramere, external face; Fig. 15, left paramere, internal face. Scale bar: 1 mm (Figs. 10
<emphasis id="B93CEAFFFFE7FF9CFEFCF8969D6EF8FC" bold="true" box="[351,364,1813,1835]" pageId="7"></emphasis>
12); 0.5 mm (Figs. 13
<emphasis id="B93CEAFFFFE7FF9CFDEAF8969E57F8FC" bold="true" box="[585,597,1813,1835]" pageId="7"></emphasis>
15).
</paragraph>
</caption>
<subSubSection id="C3526566FFE7FF92FF64F8D6999FFCC2" lastPageId="9" pageId="7" type="description">
<paragraph id="8BF736EDFFE7FF93FF64F8D69D6DF89A" blockId="7.[151,1437,1877,2010]" lastBlockId="8.[151,1436,151,2013]" lastPageId="8" pageId="7">
<emphasis id="B93CEAFFFFE7FF9CFF64F8D69D53F8B9" bold="true" box="[199,337,1877,1902]" pageId="7">Description</emphasis>
.
<emphasis id="B93CEAFFFFE7FF9CFEC0F8D69DBEF8BA" box="[355,444,1877,1901]" italics="true" pageId="7">Habitus</emphasis>
. Large-sized, bicoloured species of
<taxonomicName id="4C484D6EFFE7FF9CFCC3F8D69FE9F8BA" box="[864,1003,1877,1901]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE7FF9CFCC3F8D69FE9F8BA" box="[864,1003,1877,1901]" italics="true" pageId="7">Pterostichus</emphasis>
</taxonomicName>
, with sub-oval, slightly convex body and sides of pronotum distinctly oblique and sinuate in front of dentiform basal angles.
<emphasis id="B93CEAFFFFE7FF9CFBDBF8F99918F846" box="[1144,1306,1914,1937]" italics="true" pageId="7">Measurements</emphasis>
. BL:
<quantity id="4CB09B08FFE7FF9CFAFEF8F99CFCF862" metricMagnitude="-2" metricUnit="m" metricValue="1.46" metricValueMax="1.51" metricValueMin="1.41" pageId="7" unit="mm" value="14.6" valueMax="15.1" valueMin="14.1">14.1 15.1 mm</quantity>
(
<quantity id="4CB09B08FFE7FF9CFEACF81D9D7AF862" box="[271,376,1950,1974]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="7" unit="mm" value="14.1">14.1 mm</quantity>
in
<typeStatus id="54F3884FFFE7FF9CFE02F81E9E0AF861" box="[417,520,1949,1974]" pageId="7" type="holotype">holotype</typeStatus>
); BW:
<quantity id="4CB09B08FFE7FF9CFDFFF81E9EE7F862" box="[604,741,1949,1974]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.2" metricValueMin="5.5" pageId="7" unit="mm" value="5.85" valueMax="6.2" valueMin="5.5">5.56.2 mm</quantity>
(
<quantity id="4CB09B08FFE7FF9CFD54F81E9F53F862" box="[759,849,1949,1974]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="7" unit="mm" value="5.5">5.5 mm</quantity>
in
<typeStatus id="54F3884FFFE7FF9CFCD9F81E9FE3F861" box="[890,993,1949,1974]" pageId="7" type="holotype">holotype</typeStatus>
).
<emphasis id="B93CEAFFFFE7FF9CFC55F81D983FF862" box="[1014,1085,1950,1973]" italics="true" pageId="7">Ratios</emphasis>
. PW/HW: 1.481.55; PW/PL: 1.321.37; PbW/PaW: 1.061.1; EW/PW: 1.271.31; EL/EW: 1.381.39.
<emphasis id="B93CEAFFFFE7FF9CFC4AF8429835F80E" box="[1001,1079,1985,2009]" italics="true" pageId="7">Colour</emphasis>
. Piceous on dorsal and ventral surface as well as on appendages, femora red-yellow, excluding piceous extremities (proximal tips and knees) in
<typeStatus id="54F3884FFFE8FF93FF34FF3F9CFFFF02" box="[151,253,188,213]" pageId="8" type="holotype">holotype</typeStatus>
; same in
<typeStatus id="54F3884FFFE8FF93FECCFF3E9DD0FF02" box="[367,466,189,213]" pageId="8" type="paratype">paratype</typeStatus>
, but palpomeres and segments 511 of antennae testaceous, coxae, trochanteri, and proximal tips of femora red-yellow.
<emphasis id="B93CEAFFFFE8FF93FD9EFF639EEAFF2F" box="[573,744,224,248]" italics="true" pageId="8">Microsculpture</emphasis>
. Entire body and appendages with very fine, hardly visible isodiametric microreticulation, somewhere partially reduced.
<emphasis id="B93CEAFFFFE8FF93FCC2FE869FA5FECB" box="[865,935,261,284]" italics="true" pageId="8">Lustre</emphasis>
. Strong dorsally and ventrally, glossy, not iridescent.
<emphasis id="B93CEAFFFFE8FF93FEB7FEAB9D52FE97" box="[276,336,296,320]" italics="true" pageId="8">Head</emphasis>
. Moderately large in relation to prothorax, impunctate; eyes moderately large, convex, laterally rather protruded in
<typeStatus id="54F3884FFFE8FF93FED2FECF9DD1FEB2" box="[369,467,332,357]" pageId="8" type="holotype">holotype</typeStatus>
(smaller, slightly convex, laterally slightly protruded in
<typeStatus id="54F3884FFFE8FF93FBEEFECF98B0FEB3" box="[1101,1202,332,356]" pageId="8" type="paratype">paratype</typeStatus>
), temporae short and steep in
<typeStatus id="54F3884FFFE8FF93FF50FEEC9D57FE5F" box="[243,341,367,392]" pageId="8" type="holotype">holotype</typeStatus>
(longer and slanting in
<typeStatus id="54F3884FFFE8FF93FDFDFEF39EC6FE5F" box="[606,708,368,392]" pageId="8" type="paratype">paratype</typeStatus>
); paraorbital sulci deep, surpassing posterior margin of eye; frons with indistinct, sub-linear frontal grooves, divergent posteriorly, terminate before level of posterior supraorbital pore; apex of clypeus concave, clypeal suture clear, but degrade; labrum transverse, slightly concave in middle; mandibles moderately elongate, apically suddenly curved; antennae moderately long, distal margin of penultimate segment exceeding posterior border of pronotum; labial pits distinct, mentum tooth long and bifid; both medial and lateral setae of submentum present.
<emphasis id="B93CEAFFFFE8FF93FD90FDA69EA0FDEB" box="[563,674,549,572]" italics="true" pageId="8">Pronotum</emphasis>
. Sub-quadrate, wider than long; widest diameter before middle, in second fifth; sides evenly curved and convergent, more posteriorly, less anteriorly, almost straight along second third, subsequently convex and sinuate in front of dentiform basal angles; sides bordered, with thick bead, bordering continue on anterior margin (disappearing before midline), absent along basal margin; apex only slightly shorter than base, concave, apical angles projecting, rounded off; base almost straight, slightly concave in middle, basal angles dentiform, somewhat projecting outside; anterior transverse sulcus vanished, posterior transverse sulcus deep, coinciding with anterior edge of basal foveae; midline well-impressed, long, not reaching anterior margin, reaching posterior one; disk slightly to moderately convex; basal part with two sub-linear impressions, which form together deep basal fovea from each side, external impressions short, convex outside, internal impressions long and straight, each fovea more or less coarsely punctate inside, central area of basis wrinkled close to margin; anterior marginal pore in second fifth of pronotum, at point of maximum width of pronotum, posterior marginal pore at basal angles.
<emphasis id="B93CEAFFFFE8FF93FE5BFC339E3CFC1F" box="[504,574,944,968]" italics="true" pageId="8">Elytra</emphasis>
. Oval, relatively short and wide, dorsally slightly convex, widest diameter behind middle; basal margin distinct, slightly sinuate, reaching parascutellar stria; humeri obtusely angled; lateral margin almost parallel in
<typeStatus id="54F3884FFFE8FF93FE15FC749E19FBC7" box="[438,539,1015,1040]" pageId="8" type="holotype">holotype</typeStatus>
, more round in
<typeStatus id="54F3884FFFE8FF93FD6FFC7B9F2DFBC7" box="[716,815,1016,1040]" pageId="8" type="paratype">paratype</typeStatus>
, bead moderately wide, subapical sinuation very slight, apex slightly round (in
<typeStatus id="54F3884FFFE8FF93FE04FB9F9E09FBE2" box="[423,523,1052,1077]" pageId="8" type="holotype">holotype</typeStatus>
) to almost truncate (in
<typeStatus id="54F3884FFFE8FF93FCBFFB9F9F80FBE3" box="[796,898,1052,1076]" pageId="8" type="paratype">paratype</typeStatus>
); elytral internal plica indistinct; scutellar pore present; all striae well-impressed, impunctate, continuous from basal margin to apex, parascutellar stria distinct; intervals plane in
<typeStatus id="54F3884FFFE8FF93FECEFBE79DD0FBAA" box="[365,466,1124,1149]" pageId="8" type="holotype">holotype</typeStatus>
, moderately convex in
<typeStatus id="54F3884FFFE8FF93FD48FBE69F4DFBAA" box="[747,847,1125,1149]" pageId="8" type="paratype">paratype</typeStatus>
; three foveolate setiferous punctures in middle of interval 3 or so; four-five small setiferous punctures in apical end of stria 7 close to subapical sinuation; umbilicate setiferous series in stria 8 with a row of 25 pores on left elytron and 29 pores on right elytron in
<typeStatus id="54F3884FFFE8FF93FA99FB2F999EFB12" box="[1338,1436,1196,1221]" pageId="8" type="holotype">holotype</typeStatus>
(respectively, 32 and 30 pores in
<typeStatus id="54F3884FFFE8FF93FDB2FB539E74FB3F" box="[529,630,1232,1256]" pageId="8" type="paratype">paratype</typeStatus>
), two rows uninterrupted; hind wings vestigial.
<emphasis id="B93CEAFFFFE8FF93FB36FB539940FB3F" box="[1173,1346,1232,1256]" italics="true" pageId="8">Ventral surface</emphasis>
(
<emphasis id="B93CEAFFFFE8FF93FAF0FB539D36FADB" italics="true" pageId="8">thorax and abdomen</emphasis>
). Prosternum, mesosternum, metasternum, and first two abdominal sternites punctate laterally, impunctate in middle; morphological sternites VVII (visible ones IVVI) of abdomen impunctate, only laterally superficially wrinkled; prosternal process unbordered; metepisternum subquadrate, slightly wider than long; morphological sternum VII in male bisetose, wrinkled distally, strongly depressed at sides and at apical part, with stout projection forming a longitudinal, sub-pyramidal carina at middle and with an incomplete, fine transverse subapical sulcus broken at carina (same in female quadrisetose, less wrinkled distally, slightly depressed at sides, with very slight, but evident central projection and incomplete, fine transverse subapical sulcus broken at projection).
<emphasis id="B93CEAFFFFE8FF93FEBCFA729D56F9DF" box="[287,340,1521,1544]" italics="true" pageId="8">Legs</emphasis>
. Basal tarsomeres 13 of male protarsi expanded and biseriately squamose underneath; front tibia with three clip setae; mesotibial ctenidium well-differentiated; seta on middle trochanteri present; middle femur with two posterior setae on ventral side; metatarsal carina present on basal tarsomeres 13; seta of hind trochanteri absent; hind coxae without medial seta; onychium setose or not ventrally (if setose then with single short and thick seta unilaterally or a pair of such setae bilaterally).
<emphasis id="B93CEAFFFFE8FF93FD73F9039F76F94F" box="[720,884,1664,1688]" italics="true" pageId="8">Male genitalia</emphasis>
. Median lobe of aedeagus stout and curved down at medial third in lateral aspect (
<figureCitation id="13732A68FFE8FF93FDA7F9279E8EF96B" box="[516,652,1700,1725]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1721,1743]" captionTargetBox="[161,1417,957,1660]" captionTargetId="figure@7.[151,1436,950,1666]" captionTargetPageId="7" captionText="FIGURES 10 15. Male genitalia of Pterostichus (Oreophilus) podgoricensis sp. n., holotype: Fig. 10, median lobe of aedeagus, left lateral aspect; Fig. 11, median lobe of aedeagus, right lateral aspect; Fig. 12, median lobe of aedeagus, ventral aspect; Fig. 13, right paramere, internal face; Fig. 14, right paramere, external face; Fig. 15, left paramere, internal face. Scale bar: 1 mm (Figs. 10 12); 0.5 mm (Figs. 13 15)." httpUri="https://zenodo.org/record/222919/files/figure.png" pageId="8">Figs. 1011</figureCitation>
), bent to right ventrally (
<figureCitation id="13732A68FFE8FF93FC04F9279FF9F96B" box="[935,1019,1700,1725]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1721,1743]" captionTargetBox="[161,1417,957,1660]" captionTargetId="figure@7.[151,1436,950,1666]" captionTargetPageId="7" captionText="FIGURES 10 15. Male genitalia of Pterostichus (Oreophilus) podgoricensis sp. n., holotype: Fig. 10, median lobe of aedeagus, left lateral aspect; Fig. 11, median lobe of aedeagus, right lateral aspect; Fig. 12, median lobe of aedeagus, ventral aspect; Fig. 13, right paramere, internal face; Fig. 14, right paramere, external face; Fig. 15, left paramere, internal face. Scale bar: 1 mm (Figs. 10 12); 0.5 mm (Figs. 13 15)." httpUri="https://zenodo.org/record/222919/files/figure.png" pageId="8">Fig. 12</figureCitation>
); basal part with developed bulb and concave basal orifice; apex straight, wide, laterally compressed, with strongly sclerotized tip; ostium fully deflected to left; internal sac with numerous chitinized prickles and a moderately sclerotized plate, visible from below. Right paramere elongate, slightly curve down in medial part (
<figureCitation id="13732A68FFE8FF93FC71F88C9863F8FF" box="[978,1121,1807,1832]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1721,1743]" captionTargetBox="[161,1417,957,1660]" captionTargetId="figure@7.[151,1436,950,1666]" captionTargetPageId="7" captionText="FIGURES 10 15. Male genitalia of Pterostichus (Oreophilus) podgoricensis sp. n., holotype: Fig. 10, median lobe of aedeagus, left lateral aspect; Fig. 11, median lobe of aedeagus, right lateral aspect; Fig. 12, median lobe of aedeagus, ventral aspect; Fig. 13, right paramere, internal face; Fig. 14, right paramere, external face; Fig. 15, left paramere, internal face. Scale bar: 1 mm (Figs. 10 12); 0.5 mm (Figs. 13 15)." httpUri="https://zenodo.org/record/222919/files/figure.png" pageId="8">Figs. 1314</figureCitation>
); left paramere wide, subquadratic (
<figureCitation id="13732A68FFE8FF93FEADF8B79D60F89A" box="[270,354,1844,1869]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1721,1743]" captionTargetBox="[161,1417,957,1660]" captionTargetId="figure@7.[151,1436,950,1666]" captionTargetPageId="7" captionText="FIGURES 10 15. Male genitalia of Pterostichus (Oreophilus) podgoricensis sp. n., holotype: Fig. 10, median lobe of aedeagus, left lateral aspect; Fig. 11, median lobe of aedeagus, right lateral aspect; Fig. 12, median lobe of aedeagus, ventral aspect; Fig. 13, right paramere, internal face; Fig. 14, right paramere, external face; Fig. 15, left paramere, internal face. Scale bar: 1 mm (Figs. 10 12); 0.5 mm (Figs. 13 15)." httpUri="https://zenodo.org/record/222919/files/figure.png" pageId="8">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BF736EDFFE8FF92FF64F8D49886FE5F" blockId="8.[151,1436,151,2013]" lastBlockId="9.[151,1437,151,1328]" lastPageId="9" pageId="8">
<emphasis id="B93CEAFFFFE8FF93FF64F8D49D37F8A7" bold="true" box="[199,309,1879,1904]" pageId="8">Affinities</emphasis>
. The new species is best placed within subgenus
<taxonomicName id="4C484D6EFFE8FF93FCD7F8DB9FF3F8A7" box="[884,1009,1880,1904]" class="Insecta" family="Carabidae" genus="Oreophilus" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE8FF93FCD7F8DB9FF3F8A7" box="[884,1009,1880,1904]" italics="true" pageId="8">Oreophilus</emphasis>
</taxonomicName>
due to the presence of the following combination of characters: 1/ pronotum with sides straight to convex towards the dentiform basal angles and with relatively deep basal foveae; 2/ elytral interval 3 with foveolate setiferous punctures in middle; 3/ end of elytral stria 7 with four-five small setiferous punctures; 4/ elytral apices of elytra somewhat truncate; 5/ onychium without or with single (or with a pair of) short and thick setae ventrally; 6/ sternum VII with elevated longitudinal carina in the males; 7/ apical lamella of the median lobe of aedeagus bent to left at dorsal view; 8/ right paramere elongate, nearly straight. As a rule, the species of
<taxonomicName id="4C484D6EFFE9FF92FDF0FF639ED2FF2F" box="[595,720,224,248]" class="Insecta" family="Carabidae" genus="Oreophilus" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="B93CEAFFFFE9FF92FDF0FF639ED2FF2F" box="[595,720,224,248]" italics="true" pageId="9">Oreophilus</emphasis>
</taxonomicName>
possess a glabrous onychium, viewed from below, in contrast to the taxa of
<emphasis id="B93CEAFFFFE9FF92FE86FE879DCAFECB" box="[293,456,260,284]" italics="true" pageId="9">
<taxonomicName id="4C484D6EFFE9FF92FE86FE879DB2FECB" box="[293,432,260,284]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus" sensu="stricto" status="s">Pterostichus</taxonomicName>
<taxonomicNameLabel id="A20F5784FFE9FF92FE1EFE869DCAFECB" box="[445,456,261,284]" pageId="9" sensu="stricto">s</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B93CEAFFFFE9FF92FE78FE869DF5FECB" box="[475,503,261,284]" italics="true" pageId="9">str</emphasis>
., with an onychium bearing two or more pairs of long setae. However, some individuals of
<taxonomicName id="4C484D6EFFE9FF92FE9AFEAA9F28FE97" authority="Fairmaire &amp; Laboulbene" authorityName="Fairmaire &amp; Laboulbene" box="[313,810,295,320]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="impressus" subGenus="Oreophilus">
<emphasis id="B93CEAFFFFE9FF92FE9AFEAA9D45FE97" box="[313,327,297,320]" italics="true" pageId="9">P</emphasis>
. (
<emphasis id="B93CEAFFFFE9FF92FEFDFEAB9D70FEE8" box="[350,370,296,319]" italics="true" pageId="9">O</emphasis>
.)
<emphasis id="B93CEAFFFFE9FF92FE2AFEAB9DFAFEE8" box="[393,504,296,319]" italics="true" pageId="9">impressus</emphasis>
(Fairmaire &amp; Laboulbéne)
</taxonomicName>
,
<emphasis id="B93CEAFFFFE9FF92FC96FEAA9F41FE97" box="[821,835,297,320]" italics="true" pageId="9">P</emphasis>
. (O.)
<taxonomicName id="4C484D6EFFE9FF92FC25FEAB989BFE97" authority="Dejean" authorityName="Dejean" box="[902,1177,295,320]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="flavofemoratus">
<emphasis id="B93CEAFFFFE9FF92FC25FEAB982CFE97" box="[902,1070,296,320]" italics="true" pageId="9">flavofemoratus</emphasis>
(Dejean)
</taxonomicName>
,
<taxonomicName id="4C484D6EFFE9FF92FB07FEAA9CFEFEB2" authority="Dejean" authorityName="Dejean" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="multipunctatus" subGenus="Oreophilus">
<emphasis id="B93CEAFFFFE9FF92FB07FEAA98B0FE97" box="[1188,1202,297,320]" italics="true" pageId="9">P</emphasis>
. (
<emphasis id="B93CEAFFFFE9FF92FB69FEAB98DCFEE8" box="[1226,1246,296,319]" italics="true" pageId="9">O</emphasis>
.)
<emphasis id="B93CEAFFFFE9FF92FB56FEAB9999FE97" box="[1269,1435,296,320]" italics="true" pageId="9">multipunctatus</emphasis>
(Dejean)
</taxonomicName>
, as well as
<taxonomicName id="4C484D6EFFE9FF92FE26FECE9E7AFEB3" box="[389,632,332,356]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="podgoricensis" status="sp. nov." subGenus="Oreophilus">
<emphasis id="B93CEAFFFFE9FF92FE26FECE9D91FEB3" box="[389,403,333,356]" italics="true" pageId="9">P</emphasis>
. (
<emphasis id="B93CEAFFFFE9FF92FE0EFECF9DC3FEB4" box="[429,449,332,355]" italics="true" pageId="9">O</emphasis>
.)
<emphasis id="B93CEAFFFFE9FF92FE78FECF9E7AFEB3" box="[475,632,332,356]" italics="true" pageId="9">podgoricensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5784FFE9FF92FD21FECE9EC2FEB3" box="[642,704,333,356]" pageId="9" rank="species">sp. n.</taxonomicNameLabel>
possess an onychium ventrally with single short and thick seta unilaterally, or with a pair of such setae bilaterally especially on the middle and hind legs.
</paragraph>
<paragraph id="8BF736EDFFE9FF92FF64FE16999FFCC2" blockId="9.[151,1437,151,1328]" pageId="9">
<taxonomicName id="4C484D6EFFE9FF92FF64FE169D86FE7B" box="[199,388,404,428]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="podgoricensis" status="sp. nov.">
<emphasis id="B93CEAFFFFE9FF92FF64FE169CD7FE7B" box="[199,213,405,428]" italics="true" pageId="9">P</emphasis>
.
<emphasis id="B93CEAFFFFE9FF92FF44FE179D86FE7B" box="[231,388,404,428]" italics="true" pageId="9">podgoricensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5784FFE9FF92FE33FE169DD3FE7B" box="[400,465,405,428]" pageId="9" rank="species">sp. n.</taxonomicNameLabel>
somewhat resembles
<taxonomicName id="4C484D6EFFE9FF92FD76FE169809FE7A" authority="Dejean" authorityName="Dejean" box="[725,1035,404,429]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="flavofemoratus">
<emphasis id="B93CEAFFFFE9FF92FD76FE169EE1FE7B" box="[725,739,405,428]" italics="true" pageId="9">P</emphasis>
.
<emphasis id="B93CEAFFFFE9FF92FD55FE179F9CFE7B" box="[758,926,404,428]" italics="true" pageId="9">flavofemoratus</emphasis>
(Dejean)
</taxonomicName>
from the Western Alps in: 1/ the overall structure of the pronotum (i.e., basal impressions united in the same manner, deep basal foveae and dentiform basal angles); 2/ yellowish femora; 3/ elytra with somewhat truncate apices; 4/ elytral interval 7 without foveolate setiferous punctures; 5/ apical end of stria 7 with three-five small setiferous punctures. However, the habitus of the new species is more oval and robust (more elongate in
<taxonomicName id="4C484D6EFFE9FF92FC01FDA69864FDEB" box="[930,1126,548,572]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="flavofemoratus">
<emphasis id="B93CEAFFFFE9FF92FC01FDA69FB2FDEB" box="[930,944,549,572]" italics="true" pageId="9">P</emphasis>
.
<emphasis id="B93CEAFFFFE9FF92FC1DFDA79864FDEB" box="[958,1126,548,572]" italics="true" pageId="9">flavofemoratus</emphasis>
</taxonomicName>
), the sides of the pronotum are strongly convex towards the basal angles (vs. sides of pronotum almost straight towards angles), and the apices of the elytra are less truncate (vs. apices of elytra reversely truncate towards suture). The new species is similar also to
<taxonomicName id="4C484D6EFFE9FF92FF1BFD129DA7FD7F" authority="Dejean" authorityName="Dejean" box="[184,421,655,680]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="spinolae">
<emphasis id="B93CEAFFFFE9FF92FF1BFD129CC4FD7F" box="[184,198,657,680]" italics="true" pageId="9">P</emphasis>
.
<emphasis id="B93CEAFFFFE9FF92FF7AFD139D3AFD7F" box="[217,312,656,680]" italics="true" pageId="9">spinolae</emphasis>
(Dejean)
</taxonomicName>
from the Western Alps. Both species possess red-yellow femora, an elytral interval 7 without large foveolate setiferous punctures and stria 7 with more than two small setiferous punctures. But, the former species is distinguished from the latter in the more robust habitus and pronotum having sides somewhat convex towards basal angles, outer and inner basal impressions united in a basal fovea, and basal angles dentiform.
</paragraph>
</subSubSection>
<subSubSection id="C3526566FFE9FF92FF64FC9C994AFB12" pageId="9" type="distribution">
<paragraph id="8BF736EDFFE9FF92FF64FC9C9FBDFBC7" blockId="9.[151,1437,151,1328]" pageId="9">
<emphasis id="B93CEAFFFFE9FF92FF64FC9C9D5BFCEF" bold="true" box="[199,345,799,824]" pageId="9">Distribution</emphasis>
.
<collectingCountry id="F35F767DFFE9FF92FECAFCA39DF7FCEF" box="[361,501,800,824]" name="Serbia" pageId="9">Montenegro</collectingCountry>
, the vicinity of Podgorica City. According to the data, the precise
<typeStatus id="54F3884FFFE9FF92FB50FCA39921FCEF" box="[1267,1315,800,824]" pageId="9">type</typeStatus>
locality is unknown. The
<typeStatus id="54F3884FFFE9FF92FEECFCC79DB3FC8A" box="[335,433,836,861]" pageId="9" type="holotype">holotype</typeStatus>
was collected in the neighborhood of Podgorica (former Titograd), the capital of
<collectingCountry id="F35F767DFFE9FF92FF34FCEB9D26FC57" box="[151,292,872,896]" name="Serbia" pageId="9">Montenegro</collectingCountry>
; see “Collecting circumstances”. Undoubtedly the only
<typeStatus id="54F3884FFFE9FF92FC6EFCEB982CFC57" box="[973,1070,872,896]" pageId="9" type="paratype">paratype</typeStatus>
is wrongly labeled being not collected in the Tenerife Island. The following fact proves that. Except for the
<typeStatus id="54F3884FFFE9FF92FBFEFC0F98C0FC72" box="[1117,1218,908,933]" pageId="9" type="holotype">holotype</typeStatus>
, a male specimen
<taxonomicName id="4C484D6EFFE9FF92FF34FC339DD8FC1F" authority="Schaum" authorityName="Schaum" box="[151,474,943,968]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="brucki">
<emphasis id="B93CEAFFFFE9FF92FF34FC339D71FC1F" box="[151,371,944,968]" italics="true" pageId="9">Pterostichus brucki</emphasis>
Schaum
</taxonomicName>
, “NHRS-JLKB 000020045” [print] / “Titograd
<collectingCountry id="F35F767DFFE9FF92FBA1FC2C987CFC1F" box="[1026,1150,943,968]" name="Serbia" pageId="9">Yugoslavia</collectingCountry>
<date id="FFF6102DFFE9FF92FB2BFC2C98DDFC1F" box="[1160,1247,943,968]" pageId="9" value="80-06-26">26.6.80</date>
” [handwritten] / “
<date id="FFF6102DFFE9FF92FF01FC579D33FC3B" box="[162,305,980,1005]" pageId="9" value="1980-06-26">26. VI. 1980</date>
Jōrgen Nielsen” [handwritten] / “coll. J. Ferrer” [handwritten], is present in NHRS. The labels in the two samples have serial numbers consecutive and identical handwritten.
</paragraph>
<paragraph id="8BF736EDFFE9FF92FF64FB9F994AFB12" blockId="9.[151,1437,151,1328]" pageId="9">
<emphasis id="B93CEAFFFFE9FF92FF64FB9F9DF3FBE2" bold="true" box="[199,497,1052,1077]" pageId="9">Collecting circumstances</emphasis>
. Dr. Julio Ferrer (NHRS) kindly informed me that the
<typeStatus id="54F3884FFFE9FF92FBC3FB9F98C0FBE2" box="[1120,1218,1052,1077]" pageId="9" type="holotype">holotype</typeStatus>
of the new species and the male of
<taxonomicName id="4C484D6EFFE9FF92FEEEFBC29DB3FB8F" box="[333,433,1088,1112]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="brucki">
<emphasis id="B93CEAFFFFE9FF92FEEEFBC29D59FB8F" box="[333,347,1089,1112]" italics="true" pageId="9">P</emphasis>
.
<emphasis id="B93CEAFFFFE9FF92FEC9FBC39DB3FB8F" box="[362,433,1088,1112]" italics="true" pageId="9">brucki</emphasis>
</taxonomicName>
have been donated to the entomological collection of NHRS by Prof. Dr. Jørgen Nielsen (Zoological Museum, the Natural History Museum of
<collectingCountry id="F35F767DFFE9FF92FD54FBE79F60FBAA" box="[759,866,1124,1149]" name="Denmark" pageId="9">Denmark</collectingCountry>
, Copenhagen,
<collectingCountry id="F35F767DFFE9FF92FBA4FBE79877FBAA" box="[1031,1141,1124,1149]" name="Denmark" pageId="9">Denmark</collectingCountry>
). The two specimens were collected by the latter during “a short tourist visit arranged by a charter from
<collectingCountry id="F35F767DFFE9FF92FBA9FB049861FB77" box="[1034,1123,1159,1184]" name="Sweden" pageId="9">Sweden</collectingCountry>
and as far I know, the visit was strictly confined to the city of Titograd and local suburbs.” (Dr. Julio Ferrer, personal communication).
</paragraph>
</subSubSection>
<subSubSection id="C3526566FFE9FF92FF64FB4C9E59FAE7" pageId="9" type="etymology">
<paragraph id="8BF736EDFFE9FF92FF64FB4C9E59FAE7" blockId="9.[151,1437,151,1328]" pageId="9">
<emphasis id="B93CEAFFFFE9FF92FF64FB4C9D44FB3F" bold="true" box="[199,326,1231,1256]" pageId="9">Etymology</emphasis>
. The specific epithet is a noun in nominative singular standing in preposition. It honors Podgorica (a South Slavic word meaning “under the small hill” in English), the capital of
<collectingCountry id="F35F767DFFE9FF92FB8CFB7698BAFADA" box="[1071,1208,1269,1293]" name="Serbia" pageId="9">Montenegro</collectingCountry>
and place in which vicinities the new species was collected.
</paragraph>
</subSubSection>
</treatment>
</document>