154 lines
19 KiB
XML
154 lines
19 KiB
XML
<document id="F3BC156593B6B68C7C4E93DE6DC83793" ID-DOI="10.5281/zenodo.186646" ID-GBIF-Dataset="e80d7f6c-b367-43d7-80a8-1403082fb549" ID-ISSN="1175-5326" ID-Zenodo-Dep="186646" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461054379412" checkinUser="plazi" docAuthor="Wu, Min & Zheng, Wei" docDate="2009" docId="03E18793FF9AFF96FF16FDF170F272D5" docLanguage="en" docName="zt02053p031.pdf" docOrigin="Zootaxa 2053" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pupinidius obrutschewi subsp. contractus Moellendorff 1901" docType="treatment" docVersion="7" lastPageNumber="17" masterDocId="FFD8FFEBFF95FF86FF81FF937359701A" masterDocTitle="A review of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), with the description of a new species" masterLastPageNumber="31" masterPageNumber="1" pageNumber="16" updateTime="1698580698258" updateUser="plazi">
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<mods:titleInfo id="D248DDDEE1948CF81E709B3DCCC74E25">
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<mods:title id="A483272AE0E061DFEF16FC8E5197A0DD">A review of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), with the description of a new species</mods:title>
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<mods:name id="48734840119AE1F88609FD520E1DB702" type="personal">
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<mods:namePart id="1DB2A3D15372066421C9ADA8A43E7355">Wu, Min</mods:namePart>
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<mods:roleTerm id="282F765063BE5B4431A52743F4EFB07A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BA4FCE4695AED6548E6F6E6AE134562D">Zheng, Wei</mods:namePart>
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<mods:typeOfResource id="39926F34C067D15230A723BCA8617020">text</mods:typeOfResource>
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<mods:title id="E71F7906A3EC92FCEAAD51C19EFB10C9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="07948E12DE671713B10E1CA3F8D65349">
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<mods:date id="DA0C7E1C3ACF9B17FFB9E12CF1D34F3D">2009</mods:date>
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<mods:detail id="AE4235A98A0A78289AEE6875F62819C4" type="volume">
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<mods:number id="359007539FF17D1D15B0A1EE5192C8D9">2053</mods:number>
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<mods:start id="E6B02B6884BC1C576F6B149935327026">1</mods:start>
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<mods:end id="3CF9F4B3F10C9B1BBBD768E80BA73B73">31</mods:end>
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<mods:classification id="0C646191EAAE651AE97A790305D3A3A9">journal article</mods:classification>
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<mods:identifier id="176869EDAEC9FD24B44FEF8FD61438CE" type="DOI">10.5281/zenodo.186646</mods:identifier>
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<mods:identifier id="38472A845E8E1F4B8375430E565D029F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F558C2AFA2F2BC2D7926C26AD1CB7872" type="Zenodo-Dep">186646</mods:identifier>
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<treatment id="03E18793FF9AFF96FF16FDF170F272D5" ID-DOI="http://doi.org/10.5281/zenodo.6212429" ID-GBIF-Taxon="119612768" ID-Zenodo-Dep="6212429" LSID="urn:lsid:plazi:treatment:03E18793FF9AFF96FF16FDF170F272D5" httpUri="http://treatment.plazi.org/id/03E18793FF9AFF96FF16FDF170F272D5" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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<subSubSection id="C352650EFF9AFF89FF16FDF170367266" box="[151,879,610,636]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="8BF73685FF9AFF89FF16FDF170367266" blockId="15.[151,879,610,671]" box="[151,879,610,636]" pageId="15" pageNumber="16">
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<heading id="D0BF81E9FF9AFF89FF16FDF170367266" bold="true" box="[151,879,610,636]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<emphasis id="B93CEA97FF9AFF89FF16FDF170367266" bold="true" box="[151,879,610,636]" pageId="15" pageNumber="16">
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10.
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<taxonomicName id="4C484D06FF9AFF89FF42FDF170367266" ID-CoL="7V29Y" authority="Moellendorff, 1901" authorityName="Moellendorff" authorityYear="1901" box="[195,879,610,636]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="subSpecies" species="obrutschewi" subSpecies="contractus">
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<emphasis id="B93CEA97FF9AFF89FF42FDF1713F7266" bold="true" box="[195,614,610,636]" italics="true" pageId="15" pageNumber="16">Pupinidius obrutschewi contractus</emphasis>
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(
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<bibRefCitation id="EFD94B74FF9AFF89FDF4FDF1703E7266" author="Moellendorff" box="[629,871,610,636]" pageId="15" pageNumber="31" refString="Moellendorff, O. F. (1901) Binnen-Mollusken aus Westchina und Centralasien. II. Annuaire du Musee Zoologique de l'Academie Imperiale des St. - Petersburg, 6, 299 - 404, Taf. XII - XVII." type="journal article" year="1901">Moellendorff, 1901</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C352650EFF9AFF89FF16FD1672D27285" box="[151,395,645,671]" pageId="15" pageNumber="16" type="description">
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<paragraph id="8BF73685FF9AFF89FF16FD1672D27285" blockId="15.[151,879,610,671]" box="[151,395,645,671]" pageId="15" pageNumber="16">
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<figureCitation id="13732A00FF9AFF89FF16FD1673B97285" box="[151,224,645,671]" captionStart="FIGURE 1" captionStartId="3.[151,255,1609,1633]" captionTargetBox="[161,1429,545,1583]" captionTargetId="figure@3.[151,1436,527,1584]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of Pupinidius spp. Red dot, Pupinidius anocamptus, P. nanpingensis nanpingensis and P. nanpingensis ambigua; red circle, P. porrectus; black dot, P. pupinella altispirus; black circle, P. chrysalis; blue dot, P. obrutschewi contractus and P. pupinella pupinella; blue circle: P. latilabrum; green dot, P. melinostoma, P. nanpingensis nanpingensis and P. wenxian sp. nov.; yellow dot, P. obrutschewi obrutschewi and P. streptaxis; yellow circle, P. porrecta pygmaea. P. pupinidius and P. gregorii have not been mapped here due to the unrecognizable locality names. Although the original description of the type locality of P. chrysalis is not accurate and geographically wrong, mapping of the species is possible here based only on the recognized information therein." httpUri="https://zenodo.org/record/186647/files/figure.png" pageId="15" pageNumber="16">Figs 1</figureCitation>
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,
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<figureCitation id="13732A00FF9AFF89FF6FFD1673A57285" box="[238,252,645,671]" captionStart="FIGURE 5. A – B" captionStartId="14.[151,260,1827,1851]" captionTargetBox="[212,1373,178,1801]" captionTargetId="figure@14.[212,1374,174,1803]" captionTargetPageId="14" captionText="FIGURE 5. A – B, Pupinidius obrutschewi obrutschewi (Moellendorff, 1901), 2 paratypes of SMF- 41983. C – D, Pupinidius obrutschewi contractus (Moellendorff, 1901). C, HBUMM- 04282 - specimen 11; D, Lectotype, SMF- 41981. E – F, Pupinidius porrectus (Moellendorff, 1901). E, Lectotype, SMF- 41998; F, NHM- 1902.5.13.20 Reg." httpUri="https://zenodo.org/record/186651/files/figure.png" pageId="15" pageNumber="16">5</figureCitation>
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C–D, 8F, 14
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C352650EFF9AFF89FF16FD4170647313" pageId="15" pageNumber="16" type="reference_group">
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<paragraph id="8BF73685FF9AFF89FF16FD4170647313" blockId="15.[151,1378,722,777]" pageId="15" pageNumber="16">
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<treatmentCitationGroup id="AB5811ABFF9AFF89FF16FD4170647313" pageId="15" pageNumber="16">
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<taxonomicName id="4C484D06FF9AFF89FF16FD4170C772F3" ID-CoL="7TR76" authority="Moellendorff, 1901: 343" authorityName="Moellendorff" authorityPageNumber="343" authorityYear="1901" box="[151,926,722,745]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="subSpecies" species="obrutschewi" subGenus="Pupinidius" subSpecies="contractus">
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<emphasis id="B93CEA97FF9AFF89FF16FD41725A72F3" box="[151,259,722,745]" italics="true" pageId="15" pageNumber="16">Buliminus</emphasis>
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(
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<emphasis id="B93CEA97FF9AFF89FE93FD4172DC72F3" box="[274,389,722,745]" italics="true" pageId="15" pageNumber="16">Pupinidius</emphasis>
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)
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<emphasis id="B93CEA97FF9AFF89FE14FD4171D672F3" box="[405,655,722,745]" italics="true" pageId="15" pageNumber="16">obrutschewi contractus</emphasis>
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<treatmentCitation id="0AE91094FF9AFF89FD14FD4170C772F3" author="Moellendorff" box="[661,926,722,745]" page="343" pageId="15" pageNumber="16" year="1901">
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<bibRefCitation id="EFD94B74FF9AFF89FD14FD41703072F3" author="Moellendorff" box="[661,873,722,745]" pageId="15" pageNumber="31" refString="Moellendorff, O. F. (1901) Binnen-Mollusken aus Westchina und Centralasien. II. Annuaire du Musee Zoologique de l'Academie Imperiale des St. - Petersburg, 6, 299 - 404, Taf. XII - XVII." type="journal article" year="1901">Moellendorff, 1901</bibRefCitation>
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: 343
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</treatmentCitation>
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</taxonomicName>
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, pl. 13, figs 25, 26, 27;
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<treatmentCitation id="0AE91094FF9AFF89FB1BFD41760772F3" author="Kobelt" box="[1178,1374,722,745]" page="847" pageId="15" pageNumber="16" year="1902">
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<bibRefCitation id="EFD94B74FF9AFF89FB1BFD41767072F3" author="Kobelt" box="[1178,1321,722,745]" pageId="15" pageNumber="31" refString="Kobelt, W. (1902) Die Familie Buliminidae. Systematisches Conchylien-Cabinet, 397 - 1051, 133 pls." type="book chapter" year="1902">Kobelt, 1902</bibRefCitation>
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: 847
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</treatmentCitation>
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.
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<taxonomicName id="4C484D06FF9AFF89FF16FD6171F17313" ID-CoL="7V29Y" authority="Yen, 1939: 83" authorityName="Yen" authorityPageNumber="83" authorityYear="1939" box="[151,680,754,777]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="subSpecies" species="obrutschewi" subSpecies="contractus">
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<emphasis id="B93CEA97FF9AFF89FF16FD6171527313" box="[151,523,754,777]" italics="true" pageId="15" pageNumber="16">Pupinidius obrutschewi contractus</emphasis>
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<treatmentCitation id="0AE91094FF9AFF89FD90FD6171F17313" author="Yen" box="[529,680,754,777]" page="83" pageId="15" pageNumber="16" year="1939">
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<bibRefCitation id="EFD94B74FF9AFF89FD90FD6171D87313" author="Yen" box="[529,641,754,777]" pageId="15" pageNumber="31" refString="Yen, T. C. (1939) Die chinesischen Land-und Sußwasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 444, 1 - 234, pls. 1 - 16." type="journal article" year="1939">Yen, 1939</bibRefCitation>
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: 83
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</treatmentCitation>
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</taxonomicName>
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, pl. 7, fig. 47.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C352650EFF9AFF89FF16FCAB72067478" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="8BF73685FF9AFF89FF16FCAB72067478" blockId="15.[151,1438,824,2011]" pageId="15" pageNumber="16">
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<emphasis id="B93CEA97FF9AFF89FF16FCAB72CD7348" bold="true" box="[151,404,824,850]" pageId="15" pageNumber="16">Material examined.</emphasis>
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HBUMM-04282 (19 fmss. All measured; specimen-2 for dissection):
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<quantity id="4CB09B60FF9AFF89FB77FCAA760B7349" box="[1270,1362,825,851]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="16" unit="m" value="1250.0">1250 m</quantity>
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a.s.l., Nanyu, Zhouqu County (33°48΄ N, 104°18΄ E), Gansu,
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<date id="FFF61045FF9AFF89FCB9FCF3709E7360" box="[824,967,864,890]" pageId="15" pageNumber="16" value="2004-04-30">2004-IV-30</date>
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, coll. M.Wu. HBUMM-00333 (8 fmss without striations):
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<quantity id="4CB09B60FF9AFF89FE05FC1572B973BA" box="[388,480,902,928]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="16" unit="m" value="1250.0">1250 m</quantity>
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a.s.l., Nanyu, Zhouqu County (33°48΄ N, 104°18΄ E), Gansu,
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<date id="FFF61045FF9AFF89FB4CFC15760273BA" box="[1229,1371,902,928]" pageId="15" pageNumber="16" value="2004-04-30">2004-IV-30</date>
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, coll. M.Wu. SMF-41981:
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<typeStatus id="54F38827FF9AFF89FE12FC3E715773DD" box="[403,526,941,967]" pageId="15" pageNumber="16" type="lectotype">Lectotype</typeStatus>
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. Zw. Li-dshia-pu u. Hsi-ku-tsheng (today’s Zhouqu County), Kansu,
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<collectingCountry id="F35F7615FF9AFF89FACFFC3E76C073DD" box="[1358,1433,941,967]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
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. Slg.
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<collectingCountry id="F35F7615FF9AFF89FF4FFC4773BA73F4" box="[206,227,980,1006]" name="Iceland" pageId="15" pageNumber="16">O</collectingCountry>
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.v.Moellendorff. SMF-41985:
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<typeStatus id="54F38827FF9AFF89FDCFFC4771E373F4" box="[590,698,980,1006]" pageId="15" pageNumber="16" type="paratype">Paratype</typeStatus>
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. 1 fms. Zw. Li-dshia-pu u. Hsi-ku-tsheng, Kansu,
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<collectingCountry id="F35F7615FF9AFF89FA97FC47763873F4" box="[1302,1377,980,1006]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
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. Slg.
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<collectingCountry id="F35F7615FF9AFF89FF16FC6973F2740E" box="[151,171,1018,1044]" name="Iceland" pageId="15" pageNumber="16">O</collectingCountry>
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.v.Moellendorff (measurement of
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<typeStatus id="54F38827FF9AFF89FDB7FC697121740E" box="[566,632,1018,1044]" pageId="15" pageNumber="16">types</typeStatus>
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: ewh 1.500–1.563–1.625, wh 6.375–6.500–6.625, h 18.1–18.7–19.3, diam. maj. 11.3–11.5–11.6, aph 9.3–9.7–10.1, apw 7.2–7.3–7.4, h/diam. maj. 1.56–1.63–1.70, h/aph 1.91–1.93–1.95).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C352650EFF9AFF89FF44FBFD71357492" box="[197,620,1134,1160]" pageId="15" pageNumber="16" type="distribution">
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||
<paragraph id="8BF73685FF9AFF89FF44FBFD71357492" blockId="15.[151,1438,824,2011]" box="[197,620,1134,1160]" pageId="15" pageNumber="16">
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<emphasis id="B93CEA97FF9AFF89FF44FBFD723C7492" bold="true" box="[197,357,1134,1160]" pageId="15" pageNumber="16">Distribution.</emphasis>
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Gansu (
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<typeStatus id="54F38827FF9AFF89FE46FBFD72A27492" box="[455,507,1134,1160]" pageId="15" pageNumber="16">type</typeStatus>
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locality).
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</paragraph>
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</subSubSection>
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<subSubSection id="C352650EFF9AFF89FF44FB06703F74B5" box="[197,870,1173,1199]" pageId="15" pageNumber="16" type="materials_examined">
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||
<paragraph id="8BF73685FF9AFF89FF44FB06703F74B5" blockId="15.[151,1438,824,2011]" box="[197,870,1173,1199]" pageId="15" pageNumber="16">
|
||
<emphasis id="B93CEA97FF9AFF89FF44FB06723374B5" bold="true" box="[197,362,1173,1199]" pageId="15" pageNumber="16">
|
||
<typeStatus id="54F38827FF9AFF89FF44FB06725A74B5" box="[197,259,1173,1199]" pageId="15" pageNumber="16">Type</typeStatus>
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locality.
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</emphasis>
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Zw. Li-dshia-pu u. Hsi-ku-tsheng (Gansu).
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</paragraph>
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</subSubSection>
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<subSubSection id="C352650EFF9AFF96FF44FB2870F272D5" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="diagnosis">
|
||
<paragraph id="8BF73685FF9AFF89FF44FB28725D756A" blockId="15.[151,1438,824,2011]" pageId="15" pageNumber="16">
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||
<emphasis id="B93CEA97FF9AFF89FF44FB28721D74CF" bold="true" box="[197,324,1211,1237]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
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Shell bandless; striate. Aperture faintly with or without angular tubercle. Most swollen part at body whorl. With 6.000–6.875 whorls; height 17.1–20.6, diam. maj. 11.1–12.9. Penis without penial caecum; internally without fine pilaster. Papilla developed by fused proximal pilasters forming V-shaped structure absent. Epiphallus straight. Epiphallic caecum present. Free oviduct much shorter than vagina, almost invisible.
|
||
</paragraph>
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<paragraph id="8BF73685FF9AFF89FF44FAEE77C6777D" blockId="15.[151,1438,824,2011]" pageId="15" pageNumber="16">
|
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Re-description of shell. Pointed-ovate; apex protruded. Most swollen part at body whorl. Shell. Shell dextral; thin; solid; almost opaque; glossy; speckled or not; not spirally grooved; with 6.000–6.329–6.875 whorls. Whorls not shouldered; weakly convex. Growth lines usually not very clear. Embryonic shell smooth; polished; with 1.375–1.559–1.750 whorls. Postnuclear whorls smooth. Suture normal, without narrow defined zone on beneath whorl. Last whorl gradually ascending towards aperture; rounded at periphery; without smoothed spiral peripheral depression nor rugate region. Shell 17.1–18.5–20.6 high; 11.1–11.9–
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||
<quantity id="4CB09B60FF9AFF89FA80F9AD760B7642" box="[1281,1362,1598,1624]" metricMagnitude="-1" metricUnit="m" metricValue="3.2765999999999997" pageId="15" pageNumber="16" unit="in" value="12.9">12.9 in</quantity>
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diam. maj. Height/width ratio 1.44–1.55–1.71. Aperture in a plane; completely attached to body whorl; truncateovate; not adnate; oblique; not armed; weakly with angular tubercle, or without angular tubercle; 9.0–9.7–10.8 high; 6.8–7.3–7.8 wide. Ratio of shell height to aperture height 1.78–1.90–2.03. Peristome reflexed, without distinct cuff; expanded; sharp. Parietal callus indistinct. Columellar margin slightly reflexed. Columella vertical. Outer edge of columellar lip vertical. Umbilicus very narrowly open. Apical whorls normally coloured. Shell multicoloured; light-chestnut with more or less evenly distributed whitish radial striation except apex. Aperture white; not brightly banded; spirally without colour band.
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</paragraph>
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<paragraph id="8BF73685FF9AFF96FF44F8E7711F7298" blockId="15.[151,1438,824,2011]" lastBlockId="16.[151,1437,152,719]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">Genitalia. Vas deferens short; swollen proximally; entering epiphallus apically with distinct demarcation. Epiphallus short; cylindrical; of uniform thickness; normally straight; externally smooth. Epiphallic caecum located near vas deferens entrance; blunt apically. Flagellum short; subglobular; proximally normal; with tip blunt. Penis with terminal entrance of epiphallus; clavate; without penial caecum; distally enlarged; thinwalled; internally without fine longitudinal pilaster, with 1 V-shaped pilaster. V-shaped structure distally without a papilla. Other penial inner structure absent. Penis without process. Penial verge absent. Penial appendix short; branched off from proximal portion of penis; divided into sections including A-1+A-2, A-3 and A-4+A-5. A-1 and A-2 fused. A-2 and A-3 not fused. Boundary between A-4 and A-5 indistinct. A-5 very short; straight. Penial retractor biramous; attaching to penis in middle part and to A-1+ A-2; with penial retractor arms arising from diaphragm separately. Additional retractor rather than penial or appendicular absent. Muscular band connecting vagina and epiphallus absent. Atrium short; without atrial retractor. Free oviduct much shorter than vagina, almost invisible. Vagina moderately long; not swollen; straight; not lined with loose, spongy tissue; unpigmented. Bursa copulatrix duct short; proximally straight. Bursa copulatrix with stalk; without apical ligament; small; with long neck; poorly defined. Diverticle normally present; shorter than reservoir; expanded. Bursa copulatrix and diverticle distinguishable; forked immediately near their base. Spermatophore unknown.</paragraph>
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<paragraph id="8BF73685FF85FF96FF44FD1D70F272D5" blockId="16.[151,1437,152,719]" pageId="16" pageNumber="17">Measurement of genitalia. P: 5.2; Ep: 3.3; Fl: 0.6; VD: 5.3; Va: 4.7; FO: 0; BCD: 3.8; BC: 1.2; D: 9.1; A- 1+A-2: 3.4; A3: 1; A-4+A-5: 6.8 (in HBUMM-04282-specimen 2).</paragraph>
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</subSubSection>
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</treatment>
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</document> |