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<document id="C00C495AE1DB650E9E431B1773884835" ID-CLB-Dataset="38932" ID-DOI="10.11646/zootaxa.4120.1.1" ID-GBIF-Dataset="606b81b7-45af-4821-a885-ebf237fd4cc4" ID-ISSN="1175-5326" ID-Zenodo-Dep="266831" ID-ZooBank="EC979B80-9468-4E56-B5C0-90A7F1C78CB3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464936090671" checkinUser="plazi" docAuthor="Fu, Zhao, Toda, Masanori J., Li, Nan-Nan, Zhang, Ya-Ping &amp; Gao, Jian-Jun" docDate="2016" docId="03DD4E62FFEB7235149FFA55E5C5FD7C" docLanguage="en" docName="zootaxa.4120.pdf.imf" docOrigin="Zootaxa 4120 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Impatiophila quadrangulata Fu &amp; Gao, sp. nov." docType="treatment" docVersion="11" lastPageNumber="60" masterDocId="FFE4361AFFD2720E1408FF84E61DFFB0" masterDocTitle="A new genus of anthophilous drosophilids, Impatiophila (Diptera, Drosophilidae): morphology, DNA barcoding and molecular phylogeny, with descriptions of thirty-nine new species" masterLastPageNumber="100" masterPageNumber="1" pageNumber="58" updateTime="1698662698461" updateUser="ExternalLinkService">
<mods:mods id="72C99F800DDA308792552B40B9F865B7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2FA20507BA6085BD817DC9DB10C56E97">A new genus of anthophilous drosophilids, Impatiophila (Diptera, Drosophilidae): morphology, DNA barcoding and molecular phylogeny, with descriptions of thirty-nine new species</mods:title>
</mods:titleInfo>
<mods:name id="BB6307751D6758815801607862B53996" type="personal">
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<mods:namePart id="E6525DFE7E291287B24177C94D55CE86">Fu, Zhao</mods:namePart>
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<mods:namePart id="42B44CAEBDF17EA8113BC9B8FAE02598">Toda, Masanori J.</mods:namePart>
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<mods:namePart id="1EFAE1366101DA45E5674D3870B7AA5C">Li, Nan-Nan</mods:namePart>
</mods:name>
<mods:name id="CB1AFE528CA44B3EC96298C87E249E3A" type="personal">
<mods:role id="9A18A6D6FD16C9E601603E57F3EC6DE4">
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<mods:namePart id="8CC11EA5AF3BBB6A7D1CFF2B0861CA73">Zhang, Ya-Ping</mods:namePart>
</mods:name>
<mods:name id="20F5324D215BA5DAC05DEA2EF0AB82B8" type="personal">
<mods:role id="DC0A260BE9C22749D5EE3743C3B18600">
<mods:roleTerm id="A5667502653B39262B5AF02B74F23F12">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FA3A89DB3352ACBC3BD1A6C1A97C925B">Gao, Jian-Jun</mods:namePart>
</mods:name>
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<mods:title id="3455D16E4202DD2B54EED97D4A8E6648">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="09FC6B7B055C433D9D78EFFECFBC563A">
<mods:date id="B745BA5457DF149A1DDE21DAD03E4E1D">2016</mods:date>
<mods:detail id="5133D6F4D809A91EE02AA1E6939F4E1B" type="volume">
<mods:number id="C5E90E88A65190A1B3922837E6E78BF2">4120</mods:number>
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<mods:classification id="F8A1D82E8F9E6B3F366F5DE531E279E3">journal article</mods:classification>
<mods:identifier id="617D20AD5673C28E32A18AB4957CC4C5" type="CLB-Dataset">38932</mods:identifier>
<mods:identifier id="965C973AF789E4752E3472B30CA43CE2" type="DOI">10.11646/zootaxa.4120.1.1</mods:identifier>
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<treatment id="03DD4E62FFEB7235149FFA55E5C5FD7C" ID-DOI="http://doi.org/10.5281/zenodo.6060679" ID-GBIF-Taxon="120812876" ID-Zenodo-Dep="6060679" LSID="urn:lsid:plazi:treatment:03DD4E62FFEB7235149FFA55E5C5FD7C" httpUri="http://treatment.plazi.org/id/03DD4E62FFEB7235149FFA55E5C5FD7C" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58">
<subSubSection id="C36EACFFFFEB7237149FFA55E4ECFA5A" box="[151,753,1489,1514]" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="8BCBFF74FFEB7237149FFA55E4ECFA5A" blockId="57.[151,753,1489,1548]" box="[151,753,1489,1514]" pageId="57" pageNumber="58">
<heading id="D0834818FFEB7237149FFA55E4ECFA5A" bold="true" box="[151,753,1489,1514]" fontSize="10" level="2" pageId="57" pageNumber="58" reason="2">
<emphasis id="B9002366FFEB7237149FFA55E4ECFA5A" bold="true" box="[151,753,1489,1514]" pageId="57" pageNumber="58">
23)
<taxonomicName id="4C7484F7FFEB723714CBFA55E497FA5A" ID-CoL="8CFBF" authority="Fu &amp; Gao" authorityName="Fu &amp; Gao" box="[195,650,1489,1514]" class="Insecta" family="Drosophilidae" genus="Impatiophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="quadrangulata" status="sp. nov.">
<emphasis id="B9002366FFEB723714CBFA55E417FA5A" bold="true" box="[195,522,1489,1514]" italics="true" pageId="57" pageNumber="58">Impatiophila quadrangulata</emphasis>
Fu &amp; Gao
</taxonomicName>
,
<taxonomicNameLabel id="A2339E1DFFEB7237169FFA55E4ECFA5A" box="[663,753,1489,1514]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36EACFFFFEB7237149FFA71E79AF9BC" box="[151,391,1525,1548]" pageId="57" pageNumber="58" type="description">
<paragraph id="8BCBFF74FFEB7237149FFA71E79AF9BC" blockId="57.[151,753,1489,1548]" box="[151,391,1525,1548]" pageId="57" pageNumber="58">
(
<figureCitation id="134FE3F1FFEB72371496FA71E6F7F9BB" box="[158,234,1525,1547]" captionStart="FIGURE 33" captionStartId="58.[151,244,1637,1658]" captionTargetBox="[302,1284,300,1615]" captionTargetId="figure@58.[302,1284,299,1615]" captionTargetPageId="58" captionText="FIGURE 33. Impatiophila quadrangulata Fu &amp; Gao, sp. nov. Adult male (holotype, # 02574) and female (paratype, # 02575): A, periphallic organs (caudolateral view); B, caudoventral part of epandrium; C, surstylus (caudal view); D, tenth sternite; E, phallic organs (dorsal view); F, phallic organs (lateral view); G, oviscapt (lateral view); H, oviscapt (ventral view)." httpUri="https://zenodo.org/record/266865/files/figure.png" pageId="57" pageNumber="58">Fig. 33</figureCitation>
; Pl. 3F; Pl. 7A)
</paragraph>
</subSubSection>
<subSubSection id="C36EACFFFFEB7237149FF9BDE5B6F953" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="8BCBFF74FFEB7237149FF9BDE5B6F953" blockId="57.[151,1436,1592,2018]" pageId="57" pageNumber="58">
<emphasis id="B9002366FFEB7237149FF9BDE70DF9E0" box="[151,272,1593,1616]" italics="true" pageId="57" pageNumber="58">Diagnosis.</emphasis>
Abdominal tergites III and IV each with broad, blackish brown, caudal band medially somewhat protruded; V with caudal band medially neither interrupted/constricted nor protruded (Pl. 3E). Setae of middle row on 2nd costal section all heavy, peg-like setae, present beyond tip of R2+3. Paramere somewhat quadrangular in lateral view, with apically diverging process ventrosubbasally (at continuum between paramere and hypandrium) and 7 sensilla arranged in chevroned row on lateral surface (
<figureCitation id="134FE3F1FFEB72371732F94FE590F953" box="[826,909,1739,1764]" captionStart="FIGURE 33" captionStartId="58.[151,244,1637,1658]" captionTargetBox="[302,1284,300,1615]" captionTargetId="figure@58.[302,1284,299,1615]" captionTargetPageId="58" captionText="FIGURE 33. Impatiophila quadrangulata Fu &amp; Gao, sp. nov. Adult male (holotype, # 02574) and female (paratype, # 02575): A, periphallic organs (caudolateral view); B, caudoventral part of epandrium; C, surstylus (caudal view); D, tenth sternite; E, phallic organs (dorsal view); F, phallic organs (lateral view); G, oviscapt (lateral view); H, oviscapt (ventral view)." httpUri="https://zenodo.org/record/266865/files/figure.png" pageId="57" pageNumber="58">Fig. 33</figureCitation>
F).
</paragraph>
</subSubSection>
<subSubSection id="C36EACFFFFEB723514CFF974E268FDA8" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58" type="description">
<paragraph id="8BCBFF74FFEB723714CFF974E2B5F8B8" blockId="57.[151,1436,1592,2018]" box="[199,1192,1775,1800]" pageId="57" pageNumber="58">
<emphasis id="B9002366FFEB723714CFF974E751F8B7" box="[199,332,1776,1799]" italics="true" pageId="57" pageNumber="58">Description</emphasis>
(♂, ♀). Head: Cibarium with 34 medial and 67 posterior sensilla per side.
</paragraph>
<paragraph id="8BCBFF74FFEB723714CFF897E321F8E0" blockId="57.[151,1436,1592,2018]" pageId="57" pageNumber="58">Thorax (Pl. 3F): Postpronotum yellow; scutum blackish brown, with 3 pale, longitudinal stripes and pale posterior margin; scutellum blackish brown, pale anteriorly and laterally; pleura brown to blackish brown.</paragraph>
<paragraph id="8BCBFF74FFEB723714CFF8DFE6F4F82A" blockId="57.[151,1436,1592,2018]" pageId="57" pageNumber="58">Wing (Pl. 3F) hyaline, yellowish; veins pale brown. C1 setae 2, ventral one shorter and thinner. Haltere whitish yellow.</paragraph>
<paragraph id="8BCBFF74FFEB723714CFF823E716F852" blockId="57.[151,1436,1592,2018]" pageId="57" pageNumber="58">Legs (Pl. 3F) yellowish brown. Hindleg tibia without black, apically blunt, stout spine at outer apex on underside.</paragraph>
<paragraph id="8BCBFF74FFE8723414CFFF13E745FF48" blockId="58.[151,1437,151,248]" pageId="58" pageNumber="59">Abdomen (Pl. 3F): Tergites yellowish brown; II VI each with broad, blackish brown, caudal band, notched medially on II. Sternites grayish yellow; male VI somewhat quadrate, much wider than long, nearly straight on posterior margin.</paragraph>
<caption id="DF0BAFFCFFE87234149FF9E1E285F905" httpUri="https://zenodo.org/record/266865/files/figure.png" pageId="58" pageNumber="59" targetBox="[302,1284,300,1615]" targetPageId="58">
<paragraph id="8BCBFF74FFE87234149FF9E1E285F905" blockId="58.[151,1436,1636,1717]" pageId="58" pageNumber="59">
<emphasis id="B9002366FFE87234149FF9E1E516F9CA" bold="true" box="[151,779,1636,1658]" pageId="58" pageNumber="59">
FIGURE 33.
<taxonomicName id="4C7484F7FFE87234152AF9E0E4B2F9CA" authority="Fu &amp; Gao" authorityName="Fu &amp; Gao" box="[290,687,1636,1658]" class="Insecta" family="Drosophilidae" genus="Impatiophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="quadrangulata" status="sp. nov.">
<emphasis id="B9002366FFE87234152AF9E0E421F9CA" bold="true" box="[290,572,1636,1658]" italics="true" pageId="58" pageNumber="59">Impatiophila quadrangulata</emphasis>
Fu &amp; Gao
</taxonomicName>
,
<taxonomicNameLabel id="A2339E1DFFE8723416B4F9E1E516F9CA" box="[700,779,1637,1658]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Adult male (holotype, #02574) and female (paratype, #02575): A, periphallic organs (caudolateral view); B, caudoventral part of epandrium; C, surstylus (caudal view); D, tenth sternite; E, phallic organs (dorsal view); F, phallic organs (lateral view); G, oviscapt (lateral view); H, oviscapt (ventral view).
</paragraph>
</caption>
<paragraph id="8BCBFF74FFE8723414CFF965E20EF845" blockId="58.[151,1437,1761,2038]" pageId="58" pageNumber="59">
Male terminalia (
<figureCitation id="134FE3F1FFE87234158FF965E7C5F94A" box="[391,472,1761,1786]" captionStart="FIGURE 33" captionStartId="58.[151,244,1637,1658]" captionTargetBox="[302,1284,300,1615]" captionTargetId="figure@58.[302,1284,299,1615]" captionTargetPageId="58" captionText="FIGURE 33. Impatiophila quadrangulata Fu &amp; Gao, sp. nov. Adult male (holotype, # 02574) and female (paratype, # 02575): A, periphallic organs (caudolateral view); B, caudoventral part of epandrium; C, surstylus (caudal view); D, tenth sternite; E, phallic organs (dorsal view); F, phallic organs (lateral view); G, oviscapt (lateral view); H, oviscapt (ventral view)." httpUri="https://zenodo.org/record/266865/files/figure.png" pageId="58" pageNumber="59">Fig. 33</figureCitation>
A F): Epandrium pubescent in small patches on posteromedial portion only, obliquely truncate on ventral margin, ventrally with ca. 12 long setae; caudoventral process somewhat triangular, with ca. 40 short spine-like setulae. Surstylus moderately concave in ventral 2/3 of distal margin, with 1 long, thick seta in addition to 2 stout, upturned prensiseta around caudodorsal corner, 5 apically converging prensisetae along distal margin and 1 short seta near ventral apex. Tenth sternite somewhat triangularly expanded on dorsolateral margins, convex on dorsal margin. Cercus pubescent on anteromedial portion, with ca. 28 long setae. Hypandrium slightly narrowing anteriorly, narrower than 1/2 length. Gonopod apically right-angled in lateral view. Paramere only slightly longer than wide. Aedeagal basal process about 3/4 length of paramere.
</paragraph>
<paragraph id="8BCBFF74FFE9723514CFFF13E23CFEAD" blockId="59.[151,1437,151,717]" pageId="59" pageNumber="60">
Female terminalia (
<figureCitation id="134FE3F1FFE9723515ACFF13E7E8FF00" box="[420,501,151,176]" captionStart="FIGURE 33" captionStartId="58.[151,244,1637,1658]" captionTargetBox="[302,1284,300,1615]" captionTargetId="figure@58.[302,1284,299,1615]" captionTargetPageId="58" captionText="FIGURE 33. Impatiophila quadrangulata Fu &amp; Gao, sp. nov. Adult male (holotype, # 02574) and female (paratype, # 02575): A, periphallic organs (caudolateral view); B, caudoventral part of epandrium; C, surstylus (caudal view); D, tenth sternite; E, phallic organs (dorsal view); F, phallic organs (lateral view); G, oviscapt (lateral view); H, oviscapt (ventral view)." httpUri="https://zenodo.org/record/266865/files/figure.png" pageId="59" pageNumber="60">Fig. 33</figureCitation>
G, H; Pl. 7A): Oviscapt valve brown, gently curved on dorsosubapical margin and less expanded dorsomedially in lateral view, with 3 dorsal ovisensilla, 1921 lateral ones on nearly straight (in ventral view) margin of apicolateral flap and 10 distally increasing in size on slightly convex, ventral margin. Spermathecal capsule brown; introvert depth of duct about 1/4 of capsule height.
</paragraph>
<paragraph id="8BCBFF74FFE9723514CFFEACE2FFFED4" blockId="59.[151,1437,151,717]" pageId="59" pageNumber="60">
Measurements: BL = 2.23 (
<specimenCount id="9D7234FDFFE972351609FEACE438FEF0" box="[513,549,296,320]" pageId="59" pageNumber="60" type="generic">2♂</specimenCount>
<typeStatus id="54CF41D6FFE972351625FEACE480FEF0" box="[557,669,296,320]" pageId="59" pageNumber="60" type="paratype">paratypes</typeStatus>
: 2.202.47, 2♀
<typeStatus id="54CF41D6FFE97235175DFEACE5D8FEF0" box="[853,965,296,320]" pageId="59" pageNumber="60" type="paratype">paratypes</typeStatus>
: 2.632.70) mm, ThL = 1.13 (1.101.17, 1.131.16) mm, WL = 2.48 (2.602.60, 2.632.83) mm, WW = 1.14 (1.171.20, 1.141.23) mm.
</paragraph>
<paragraph id="8BCBFF74FFE9723514CFFEEBE268FDA8" blockId="59.[151,1437,151,717]" pageId="59" pageNumber="60">
Indices: arb = 3/1 (
<specimenCount id="9D7234FDFFE9723515AAFEF4E7DBFE38" box="[418,454,368,392]" pageId="59" pageNumber="60" type="generic">2♂</specimenCount>
, 2♀
<typeStatus id="54CF41D6FFE97235160AFEF4E46FFE38" box="[514,626,368,392]" pageId="59" pageNumber="60" type="paratype">paratypes</typeStatus>
: 23/1), FW/HW = 0.44 (0.410.44), ch/o = 0.11 (0.080.10), prorb = 0.91 (0.911.07), rcorb = 0.48 (0.650.78), orbito = 0.95 (1.071.35), vb = 0.35 (
<specimenCount id="9D7234FDFFE972351048FE10E27EFE1C" box="[1088,1123,404,428]" pageId="59" pageNumber="60" type="generic">1♂</specimenCount>
, 2♀
<typeStatus id="54CF41D6FFE972351096FE11E313FE1D" box="[1182,1294,405,429]" pageId="59" pageNumber="60" type="paratype">paratypes</typeStatus>
: 0.390.47), dcl = 0.61 (0.550.60), dcp = 0.40 (0.320.37), sterno = 0.71 (0.690.80), m-sterno = 0.75 (0.750.83), sctl = 1.06 (0.991.05), sctlp = 1.13 (1.001.28), C = 2.19 (2.372.50), 4c = 0.98 (0.870.94),
<date id="FFCAD9B4FFE972351054FE58E265FE44" box="[1116,1144,476,500]" pageId="59" pageNumber="60">4v</date>
= 1.85 (1.621.68),
<date id="FFCAD9B4FFE97235116FFE58E39EFE45" box="[1383,1411,476,501]" pageId="59" pageNumber="60">5x</date>
= 1.81 (1.501.73), ac = 3.16 (2.873.12), M = 0.55 (0.440.50), C3F = 0.06 (0.060.09).
</paragraph>
</subSubSection>
<subSubSection id="C36EACFFFFE9723514CFFDA0E5B3FD35" pageId="59" pageNumber="60" type="materials_examined">
<paragraph id="8BCBFF74FFE9723514CFFDA0E380FD8C" blockId="59.[151,1437,151,717]" box="[199,1437,548,573]" pageId="59" pageNumber="60">
<typeStatus id="54CF41D6FFE9723514CFFDA0E733FD8D" box="[199,302,548,573]" pageId="59" pageNumber="60" type="holotype">
<emphasis id="B9002366FFE9723514CFFDA0E733FD8D" box="[199,302,548,573]" italics="true" pageId="59" pageNumber="60">Holotype</emphasis>
</typeStatus>
. ♂ (#02574), Mt. Emei, Sichuan,
<collectingCountry id="F363BFE4FFE9723516BCFDA0E4E7FD8D" box="[692,762,548,573]" name="China" pageId="59" pageNumber="60">China</collectingCountry>
, ca.
<quantity id="4C8C5291FFE972351723FDA0E59CFD8C" box="[811,897,548,573]" metricMagnitude="3" metricUnit="m" metricValue="2.54" pageId="59" pageNumber="60" unit="m" value="2540.0">2540 m</quantity>
,
<date id="FFCAD9B4FFE972351785FDA0E212FD8D" box="[909,1039,548,573]" pageId="59" pageNumber="60" value="1992-07-17">17.vii.1992</date>
, by net sweeping, M.J. Toda (
<collectionCode id="ED6567B1FFE972351154FDA0E38DFD8C" box="[1372,1424,548,572]" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="59" pageNumber="60">KIZ</collectionCode>
).
</paragraph>
<paragraph id="8BCBFF74FFE9723514CFFDCDE5B3FD35" blockId="59.[151,1437,151,717]" pageId="59" pageNumber="60">
<typeStatus id="54CF41D6FFE9723514CFFDCDE724FDD0" box="[199,313,585,608]" pageId="59" pageNumber="60" type="paratype">
<emphasis id="B9002366FFE9723514CFFDCDE724FDD0" box="[199,313,585,608]" italics="true" pageId="59" pageNumber="60">Paratypes</emphasis>
</typeStatus>
:
<collectingCountry id="F363BFE4FFE972351541FDCCE7BCFDD0" box="[329,417,584,608]" name="China" pageId="59" pageNumber="60">CHINA</collectingCountry>
:
<specimenCount id="9D7234FDFFE9723515BAFDCCE7C8FDD0" box="[434,469,584,608]" pageId="59" pageNumber="60" type="generic">1♂</specimenCount>
, 2♀ (
<collectionCode id="ED6567B1FFE972351611FDCCE450FDD0" box="[537,589,584,608]" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="59" pageNumber="60">KIZ</collectionCode>
: #02575, #02883, #02884),
<specimenCount id="9D7234FDFFE972351785FDCCE5ADFDD0" box="[909,944,584,608]" pageId="59" pageNumber="60" type="generic">1♂</specimenCount>
(
<collectionCode id="ED6567B1FFE9723517C9FDCCE212FDD0" box="[961,1039,584,608]" pageId="59" pageNumber="60">SEHU</collectionCode>
), same data as
<typeStatus id="54CF41D6FFE9723510B6FDC3E33EFDD0" box="[1214,1315,583,608]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D7234FDFFE972351138FDCCE34EFDD0" box="[1328,1363,584,608]" pageId="59" pageNumber="60" type="generic">1♂</specimenCount>
(
<collectionCode id="ED6567B1FFE97235116CFDCCE385FDD0" box="[1380,1432,584,608]" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="59" pageNumber="60">KIZ</collectionCode>
: #02885) Mt. Emei, Sichuan,
<date id="FFCAD9B4FFE9723515EDFDE8E47AFD35" box="[485,615,620,645]" pageId="59" pageNumber="60" value="1992-07-19">19.vii.1992</date>
, by net sweeping, M.J. Toda.
</paragraph>
</subSubSection>
<subSubSection id="C36EACFFFFE9723514CFFD0BE401FD18" box="[199,540,655,680]" pageId="59" pageNumber="60" type="distribution">
<paragraph id="8BCBFF74FFE9723514CFFD0BE401FD18" blockId="59.[151,1437,151,717]" box="[199,540,655,680]" pageId="59" pageNumber="60">
<emphasis id="B9002366FFE9723514CFFD0BE74DFD18" box="[199,336,655,680]" italics="true" pageId="59" pageNumber="60">Distribution</emphasis>
.
<collectingCountry id="F363BFE4FFE972351556FD0BE7BCFD18" box="[350,417,655,680]" name="China" pageId="59" pageNumber="60">China</collectingCountry>
(Sichuan).
</paragraph>
</subSubSection>
<subSubSection id="C36EACFFFFE9723514CFFD30E5C5FD7C" box="[199,984,692,717]" pageId="59" pageNumber="60" type="etymology">
<paragraph id="8BCBFF74FFE9723514CFFD30E5C5FD7C" blockId="59.[151,1437,151,717]" box="[199,984,692,717]" pageId="59" pageNumber="60">
<emphasis id="B9002366FFE9723514CFFD30E720FD7D" box="[199,317,692,717]" italics="true" pageId="59" pageNumber="60">Etymology</emphasis>
. Referring to the quadrangular (
<emphasis id="B9002366FFE9723516AFFD30E545FD7D" box="[679,856,692,717]" italics="true" pageId="59" pageNumber="60">quadrangulatus</emphasis>
) paramere.
</paragraph>
</subSubSection>
</treatment>
</document>