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<document id="C450E6F631A7CB829C847B79230AF6CE" ID-DOI="10.11646/zootaxa.4999.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5119209" ID-ZooBank="08A81188-B063-4391-B91F-000CE4D03313" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626872050065" checkinUser="plazi" docAuthor="Strümpher, Werner P. &amp; Daniel, Gimo M." docDate="2021" docId="03D664610A2EFFBFF6F170CEFBBF58C2" docLanguage="en" docName="zootaxa.4999.4.3.pdf" docOrigin="Zootaxa 4999 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stiptopodius brancoi Strumpher &amp; Daniel 2021, new species" docType="treatment" docUuid="28B9704C-5D11-4BFA-AC09-5558AC5242EE" docUuidSource="ZooBank" docVersion="3" lastPageNumber="340" masterDocId="FFEF1C190A2FFFBAF6667105FFCA5817" masterDocTitle="A new dung beetle species of the genus Stiptopodius Harold, 1871 (Coleoptera Scarabaeidae: Scarabaeinae) recorded from xeric savanna in Namibia" masterLastPageNumber="342" masterPageNumber="335" pageNumber="336" updateTime="1699011889599" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="1A37354B2190427C3E28D4AA9EC3EBBB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BC982829D523C637D31177B616B11BEC">
<mods:title id="38B9109371345FE642BA92D49048C44F">A new dung beetle species of the genus Stiptopodius Harold, 1871 (Coleoptera Scarabaeidae: Scarabaeinae) recorded from xeric savanna in Namibia</mods:title>
</mods:titleInfo>
<mods:name id="E2EE2EA7A4D00EBF7FF73734C27D811F" type="personal">
<mods:role id="DA385103579A21BD0459AC88B6C426CB">
<mods:roleTerm id="9ECEB8FDC69358B1F0623280CE2A376B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="801F6E7B8FEBEA2593A40520DABB1969">Strümpher, Werner P.</mods:namePart>
<mods:affiliation id="6E0E046FE7BBEBE79B4473080AAD5856">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
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<mods:role id="5F68640DBD3C3AB8DBF4511A0D02FEFF">
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</mods:role>
<mods:namePart id="7B34FBE53F31B2EDCA3AF44C6597E42A">Daniel, Gimo M.</mods:namePart>
<mods:nameIdentifier id="12F9B2FE4E86DA98BA4C7A37BF3AB986" type="ORCID">0000-0003-3602-5034</mods:nameIdentifier>
<mods:affiliation id="0F57F15A1CC3028474D5F743C148665E">Department of Terrestrial Invertebrates, The National Museum, 36 Aliwal Street, Bloemfontein, South Africa. gimo. daniel @ nasmus. co. za; https: // orcid. org / 0000 - 0003 - 3602 - 5034 &amp; Department of Biological &amp; Environmental Sciences, Walter Sisulu University, Mthatha, South Africa</mods:affiliation>
<mods:nameIdentifier id="D812E34B4C2FF41EE239FB2C65760966" type="email">gimo.daniel@nasmus.co.za</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1362A1A1255749340370CFE180F006FA">text</mods:typeOfResource>
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<mods:titleInfo id="19EDAF62A418EFAD850EDCD7948AA55B">
<mods:title id="CBA1AC9252E5B942002DF316F13FBA83">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="841EF87BEACE1DA86FEC3996A6E800FD">
<mods:date id="CE3F52BD665C39E3FE2E8543FC93F71C">2021</mods:date>
<mods:detail id="EC68C4E7891107B417CFB9ACE7054BA0" type="pubDate">
<mods:number id="631C332CC9A9D2751E335EBA61D0D266">2021-07-13</mods:number>
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<mods:detail id="FD7FBFCEECCA8DCA9FD0AC82FBDCD477" type="volume">
<mods:number id="FAB484917F66782D6D5E429A5E077484">4999</mods:number>
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<mods:number id="7E7FE00CBFAF37267CB356E383E02BE6">4</mods:number>
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<mods:classification id="132FE843AAF54727DA20F982D66182A5">journal article</mods:classification>
<mods:identifier id="06987064B195D58594E06039BC7FBB4B" type="DOI">10.11646/zootaxa.4999.4.3</mods:identifier>
<mods:identifier id="296FAF9B2D3639EC5E1E0C7BE68397AC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9EB8F2B80B57D5265DC7DF6CF885259B" type="Zenodo-Dep">5119209</mods:identifier>
<mods:identifier id="81C450037C1A1A791B685AC7C201AE05" type="ZooBank">08A81188-B063-4391-B91F-000CE4D03313</mods:identifier>
</mods:mods>
<treatment id="03D664610A2EFFBFF6F170CEFBBF58C2" LSID="urn:lsid:zoobank.org:act:28B9704C-5D11-4BFA-AC09-5558AC5242EE" httpUri="http://treatment.plazi.org/id/03D664610A2EFFBFF6F170CEFBBF58C2" lastPageId="5" lastPageNumber="340" pageId="1" pageNumber="336">
<subSubSection id="C36586FC0A2EFFBBF6F170CEFCF059F2" box="[151,826,459,486]" pageId="1" pageNumber="336" type="nomenclature">
<paragraph id="8BC0D5770A2EFFBBF6F170CEFCF059F2" blockId="1.[151,969,459,558]" box="[151,826,459,486]" pageId="1" pageNumber="336">
<heading id="D088621B0A2EFFBBF6F170CEFCF059F2" bold="true" box="[151,826,459,486]" fontSize="11" level="1" pageId="1" pageNumber="336" reason="1">
<emphasis id="B90B09650A2EFFBBF6F170CEFCF059F2" bold="true" box="[151,826,459,486]" pageId="1" pageNumber="336">
<taxonomicName id="4C7FAEF40A2EFFBBF6F170CEFD5759F2" ID-CoL="8LZ32" authority="Strumpher &amp; Daniel" authorityName="Strumpher &amp; Daniel" authorityYear="2021" box="[151,669,459,486]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="brancoi" status="sp. nov.">
<emphasis id="B90B09650A2EFFBBF6F170CEFE5B59F1" bold="true" box="[151,401,459,486]" italics="true" pageId="1" pageNumber="336">Stiptopodius brancoi</emphasis>
Strümpher &amp; Daniel
</taxonomicName>
,
<taxonomicNameLabel id="A238B41E0A2EFFBBF4CC70CEFCF059F2" box="[682,826,459,485]" pageId="1" pageNumber="336" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36586FC0A2EFFBBF6F170EAFC035A39" pageId="1" pageNumber="336" type="description">
<paragraph id="8BC0D5770A2EFFBBF6F170EAFEE45A1E" blockId="1.[151,969,459,558]" box="[151,302,495,522]" pageId="1" pageNumber="336">
(
<figureCitation id="1344C9F20A2EFFBBF6F970EAFEC15A1D" box="[159,267,495,522]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Figs. 1–5</figureCitation>
,
<figureCitation id="1344C9F20A2EFFBBF77170EAFEEC5A1E" box="[279,294,495,521]" captionStart="FIGURE 9" captionStartId="4.[151,250,1999,2023]" captionTargetBox="[221,1362,748,1964]" captionTargetId="figure-293@4.[213,1375,741,1975]" captionTargetPageId="4" captionText="FIGURE 9. Type locality of Stiptopodius brancoi Strümpher &amp; Daniel, new species in Namibia." figureDoi="http://doi.org/10.5281/zenodo.5119215" httpUri="https://zenodo.org/record/5119215/files/figure.png" pageId="1" pageNumber="336">9</figureCitation>
)
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6F17316FC035A39" blockId="1.[151,969,459,558]" box="[151,969,531,558]" pageId="1" pageNumber="336">
<uri id="FFEED9750A2EFFBBF6F17316FC035A39" box="[151,969,531,558]" pageId="1" pageNumber="336">
urn:lsid:zoobank.org:act:
<uuid id="FFD9EFA20A2EFFBBF7C07316FC035A39" box="[422,969,531,558]" pageId="1" pageNumber="336">28B9704C-5D11-4BFA-AC09-5558AC5242EE</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C36586FC0A2EFFBBF6F1735EFA4D5B12" pageId="1" pageNumber="336" type="materials_examined">
<paragraph id="8BC0D5770A2EFFBBF6F1735EFCCC5A62" blockId="1.[151,1437,603,2034]" box="[151,774,603,629]" pageId="1" pageNumber="336">
<materialsCitation id="3B17DF2A0A2EFFBBF6F1735EFCCB5A62" box="[151,769,603,629]" country="Namibia" location="Gocheganas" pageId="1" pageNumber="336" specimenCount="1" stateProvince="Khomas" typeStatus="holotype">
<emphasis id="B90B09650A2EFFBBF6F1735EFEF35A62" bold="true" box="[151,313,603,629]" pageId="1" pageNumber="336">
<typeStatus id="54C46BD50A2EFFBBF6F1735EFF185A62" box="[151,210,603,629]" pageId="1" pageNumber="336">Type</typeStatus>
locality:
</emphasis>
<location id="8EA083AC0A2EFFBBF726735EFE065A62" LSID="urn:lsid:plazi:treatment:03D664610A2EFFBFF6F170CEFBBF58C2:8EA083AC0A2EFFBBF726735EFE065A62" box="[320,460,603,629]" country="Namibia" name="Gocheganas" pageId="1" pageNumber="336" stateProvince="Khomas">Gocheganas</location>
,
<collectingRegion id="49BB1B950A2EFFBBF7B1735EFD585A62" box="[471,658,603,629]" country="Namibia" name="Khomas" pageId="1" pageNumber="336">Khomas District</collectingRegion>
,
<collectingCountry id="F36895E70A2EFFBBF4FB735EFCCB5A62" box="[669,769,603,629]" name="Namibia" pageId="1" pageNumber="336">Namibia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A1737AFA4D5B12" blockId="1.[151,1437,603,2034]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A1737AFEBF5A8E" bold="true" box="[199,373,639,665]" pageId="1" pageNumber="336">Type material.</emphasis>
<materialsCitation id="3B17DF2A0A2EFFBBF71A737AFA4D5B12" accessionNumber="H56379" collectingDate="1982-02-22" collectingDateMax="1982-03-20" collectingDateMin="1982-02-22" collectionCode="TMSA" collectorName="M. - L. Perinth" country="Namibia" latitude="-22.833334" location="Rectangle" longLatPrecision="1258" longitude="17.183332" pageId="1" pageNumber="336" specimenCount="1" specimenCount-male="1" stateProvince="Khomas" typeStatus="holotype">
<typeStatus id="54C46BD50A2EFFBBF71A737AFE2D5A8E" box="[380,487,639,665]" pageId="1" pageNumber="336" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D791EFE0A2EFFBBF7947385FDCD5A8D" box="[498,519,640,666]" pageId="1" pageNumber="336" type="male">♂</specimenCount>
(
<collectionCode id="ED6E4DB20A2EFFBBF470737AFDA25A8E" box="[534,616,639,665]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="1" pageNumber="336">TMSA</collectionCode>
, aedeagus extracted) (
<figureCitation id="1344C9F20A2EFFBBF506737AFC065A8D" box="[864,972,639,666]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Figs. 1–5</figureCitation>
):
<location id="8EA083AC0A2EFFBBF584737AFB9F5A8E" LSID="urn:lsid:plazi:treatment:03D664610A2EFFBFF6F170CEFBBF58C2:8EA083AC0A2EFFBBF584737AFB9F5A8E" box="[994,1109,639,665]" country="Namibia" latitude="-22.833334" longLatPrecision="1258" longitude="17.183332" name="Rectangle" pageId="1" pageNumber="336" stateProvince="Khomas">Rectangle</location>
, faded white cardstock: “Go- cheganas [Nature Reserve] 26 |
<geoCoordinate id="EE4BB3B00A2EFFBBF79E73A0FD9D5AD7" box="[504,599,677,704]" degrees="22" direction="south" minutes="50" orientation="latitude" pageId="1" pageNumber="336" precision="925" value="-22.833334">22°50′S</geoCoordinate>
.,
<geoCoordinate id="EE4BB3B00A2EFFBBF40373A0FD085AD7" box="[613,706,677,704]" degrees="17" direction="east" minutes="11" orientation="longitude" pageId="1" pageNumber="336" precision="925" value="17.183332">17°11′E</geoCoordinate>
. | WINDHOEK DISTRICT [
<collectingRegion id="49BB1B950A2EFFBBF26473A6FB0A5AAA" box="[1026,1216,675,701]" country="Namibia" name="Khomas" pageId="1" pageNumber="336">Khomas District</collectingRegion>
,
<collectingCountry id="F36895E70A2EFFBBF2AC73A7FAE45AAA" box="[1226,1326,674,701]" name="Namibia" pageId="1" pageNumber="336">Namibia</collectingCountry>
] |
<date id="FFC1F3B70A2EFFBBF32073A6FEF75AF6" pageId="1" pageNumber="336" value="1982-02-22" valueMax="1982-03-20" valueMin="1982-02-22">
<collectingDate id="EF850A5F0A2EFFBBF32073A6FEF75AF6" pageId="1" pageNumber="336" value="1982-02-22" valueMax="1982-03-20" valueMin="1982-02-22">22Feb.- 20March 1982</collectingDate>
</date>
| Preserv. traps |
<collectorName id="268AB0A10A2EFFBBF46073C2FD515AF6" box="[518,667,711,737]" pageId="1" pageNumber="336">M.-L.Perinth</collectorName>
” || Rectangle, white cardstock: “Line 1 | Banana bait” || Rectangle, white: “H56379” || Rectangle, red cardstock: “
<emphasis id="B90B09650A2EFFBBF4C673EEFCF55B12" bold="true" box="[672,831,747,773]" pageId="1" pageNumber="336">
<typeStatus id="54C46BD50A2EFFBBF4C673EEFCF55B12" box="[672,831,747,773]" pageId="1" pageNumber="336" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
|
<taxonomicName id="4C7FAEF40A2EFFBBF53473EEFBF65B12" authorityName="Strumpher &amp; Daniel" authorityYear="2021" box="[850,1084,747,773]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="brancoi">
<emphasis id="B90B09650A2EFFBBF53473EEFBF65B12" box="[850,1084,747,773]" italics="true" pageId="1" pageNumber="336">Stiptopodius brancoi</emphasis>
</taxonomicName>
| Strümpher &amp; Daniel 2021”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36586FC0A2EFFB9F6A1720AFE6A5B02" lastPageId="3" lastPageNumber="338" pageId="1" pageNumber="336" type="description">
<paragraph id="8BC0D5770A2EFFBBF6A1720AFC665B3D" blockId="1.[151,1437,603,2034]" box="[199,940,783,810]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A1720AFE935B3E" bold="true" box="[199,345,783,809]" pageId="1" pageNumber="336">Description.</emphasis>
<emphasis id="B90B09650A2EFFBBF706720AFE475B3E" box="[352,397,783,809]" italics="true" pageId="1" pageNumber="336">Size</emphasis>
: Length:
<quantity id="4C8778920A2EFFBBF79C720AFD985B3D" box="[506,594,783,810]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="336" unit="mm" value="4.4">4.4 mm</quantity>
, width across humeri:
<quantity id="4C8778920A2EFFBBF529720AFC625B3D" box="[847,936,783,810]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="336" unit="mm" value="2.3">2.3 mm</quantity>
.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A17236FA8D5B66" blockId="1.[151,1437,603,2034]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A17236FEDC5B5A" box="[199,278,819,845]" italics="true" pageId="1" pageNumber="336">Colour</emphasis>
(
<figureCitation id="1344C9F20A2EFFBBF7417236FE5F5B5A" box="[295,405,819,845]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Figs. 1–3</figureCitation>
): Head and pronotum metallic green, elytra brown with brassy reflection, ventral side dark brown with a green metallic reflection. Legs dark brown. Body setae yellow unless stated otherwise below.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A1727EFC025C7A" blockId="1.[151,1437,603,2034]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A1727EFECE5B82" box="[199,260,891,917]" italics="true" pageId="1" pageNumber="336">Head</emphasis>
(
<figureCitation id="1344C9F20A2EFFBBF770727EFE4C5B82" box="[278,390,891,917]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Figs. 1–3</figureCitation>
): Surface sculpted with discrete punctures, punctation varying from fine to large, surfaces between punctures smooth with shiny reflection, sparsely setose. Clypeus broadly rounded, anteriorly sinuated, strongly bidentate, teeth slightly recurved, apically rounded, notch between medial clypeal teeth deeply concave, clypeal edge laterad of teeth recurved and somewhat sinuate; frontoclypeal carina well-defined (
<figureCitation id="1344C9F20A2EFFBBF2A872E2FAD85C16" box="[1230,1298,999,1025]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Fig. 3</figureCitation>
, red arrow); clypeogenal suture visible. Genal protuberance roughly triangular, angles acute. Frontal carina distinct, entire, not interrupted medially (
<figureCitation id="1344C9F20A2EFFBBF7EE752AFE015C5E" box="[392,459,1071,1097]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Fig. 3</figureCitation>
, yellow arrow). Antennae with eight antennomeres; labium setose. Setae around mouthparts dense, yellow to rust brown. Eyes in dorsal view small, slit-shaped.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A1757DFE2B5D36" blockId="1.[151,1437,603,2034]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A1757DFEFC5C86" box="[199,310,1144,1169]" italics="true" pageId="1" pageNumber="336">Pronotum</emphasis>
(
<figureCitation id="1344C9F20A2EFFBBF72E7572FE7D5C86" box="[328,439,1143,1169]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Figs. 1–2</figureCitation>
): Convex, slightly wider than long; surface densely punctate, punctation varying from fine (dorsally) to large (laterally), surfaces between punctures smooth with shiny reflection. Pronotum marginate all around, anterolateral sides setose, with central disc mostly glabrous; anterior margin as wide as posterior; lateral edges strongly sinuate before posterolateral angles (in lateral view), lateral sides of pronotum strongly concave from posterolateral angles onward.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A1742EFAA85DEE" blockId="1.[151,1437,603,2034]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A1742EFEC75D52" box="[199,269,1323,1349]" italics="true" pageId="1" pageNumber="336">Elytra</emphasis>
(
<figureCitation id="1344C9F20A2EFFBBF779742EFEAE5D52" box="[287,356,1323,1349]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Fig. 1</figureCitation>
): With distinct narrow and shallow striae, striae weakly punctate, punctures regularly spaced. Interstriae very slightly convex, interstria I–VI with two rows of punctures aligned on either side leaving middle of interstria impunctate, surface smooth, punctation on interstria VII and VIII irregularly and not aligned in two rows; all interstriae sparsely setose with only some puncture bearing a short seta; all interstriae starting from the basis of elytra except interstria VII which starts from the middle of the humeral umbone. Humeral umbone weakly defined. Lateral margins straight, marginate, bearing row of setae; elytral margin concave proximally, somewhat convex distally.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A17706FC565E55" blockId="1.[151,1437,603,2034]" pageId="1" pageNumber="336">
<taxonomicName id="4C7FAEF40A2EFFBBF6A17706FE7D5E0A" authority=": Marginate" authorityName="Marginate" box="[199,439,1539,1566]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="336" phylum="Chordata" rank="genus">
<emphasis id="B90B09650A2EFFBBF6A17706FEF85E0A" box="[199,306,1539,1565]" italics="true" pageId="1" pageNumber="336">Pygidium</emphasis>
: Marginate
</taxonomicName>
, moderately convex, surface with discrete, round punctures, punctation ocellate, surfaces between punctures smooth, shiny, each puncture bearing a short seta.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A1774EFDE35E72" blockId="1.[151,1437,603,2034]" box="[199,553,1611,1638]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A1774EFE6C5E72" box="[199,422,1611,1637]" italics="true" pageId="1" pageNumber="336">Metathoracic wings</emphasis>
: Complete.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A17775FB5E5E9E" blockId="1.[151,1437,603,2034]" box="[199,1172,1647,1674]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A17775FE805E9E" box="[199,330,1648,1673]" italics="true" pageId="1" pageNumber="336">Prosternum</emphasis>
: Surface setose, anterior angles of prosternum slanted forward and inward.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A17796FF125EC6" blockId="1.[151,1437,603,2034]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A17796FE8C5EBA" box="[199,326,1683,1709]" italics="true" pageId="1" pageNumber="336">Propleuron</emphasis>
(
<figureCitation id="1344C9F20A2EFFBBF73F7796FE6B5EBA" box="[345,417,1683,1709]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Fig. 2</figureCitation>
): Dull, finely punctate, punctures with long seta, surface between punctures faintly textured.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A177D9FB455EE1" blockId="1.[151,1437,603,2034]" box="[199,1167,1755,1782]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A177D9FE935EE2" box="[199,345,1756,1781]" italics="true" pageId="1" pageNumber="336">Mesoventrite</emphasis>
(
<figureCitation id="1344C9F20A2EFFBBF70F77DEFE645EE2" box="[361,430,1755,1781]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Fig. 2</figureCitation>
): Dull, finely setose, with closely set large but shallow punctures.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A17605FEBB5F91" blockId="1.[151,1437,603,2034]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A17605FE9C5F0E" box="[199,342,1792,1817]" italics="true" pageId="1" pageNumber="336">Metaventrite</emphasis>
(
<figureCitation id="1344C9F20A2EFFBBF70077FAFE615F0E" box="[358,427,1791,1817]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Fig. 2</figureCitation>
): Shiny, with metallic green reflection, gibbose anteriorly then sloping forwards; disc glabrous, slightly convex; fairly densely punctate, punctures irregular in size, punctures small or fine medially and on anterior slope, laterally punctures larger on sides and along metaventral margins, larger punctures setiferous, each bearing a long seta.
</paragraph>
<paragraph id="8BC0D5770A2EFFBBF6A1768AFDE55FE5" blockId="1.[151,1437,603,2034]" pageId="1" pageNumber="336">
<emphasis id="B90B09650A2EFFBBF6A1768AFE615FBE" box="[199,427,1935,1961]" italics="true" pageId="1" pageNumber="336">Abdominal ventrites</emphasis>
(
<figureCitation id="1344C9F20A2EFFBBF7DA768AFDC95FBE" box="[444,515,1935,1961]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="1" pageNumber="336">Fig. 2</figureCitation>
): Shiny, with faint metallic green reflection, ventrites with ocellate punctures arranged in rows laterally, punctures setiferous, each with a long seta, glabrous medially; last abdominal ventrite narrowed medially (male character).
</paragraph>
<caption id="DF0085FF0A2DFFB8F6F17692FD6E5FEF" ID-DOI="http://doi.org/10.5281/zenodo.5119211" ID-Zenodo-Dep="5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="2" pageNumber="337" startId="2.[151,264,1943,1967]" targetBox="[205,1360,208,1920]" targetPageId="2">
<paragraph id="8BC0D5770A2DFFB8F6F17692FD6E5FEF" blockId="2.[151,1437,1943,2040]" pageId="2" pageNumber="337">
<emphasis id="B90B09650A2DFFB8F6F17692FEF65FB8" bold="true" box="[151,316,1943,1967]" pageId="2" pageNumber="337">FIGURES 1–5.</emphasis>
<taxonomicName id="4C7FAEF40A2DFFB8F723769DFD325FA7" authority="Strumpher &amp; Daniel" authorityName="Strumpher &amp; Daniel" authorityYear="2021" box="[325,760,1944,1968]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="brancoi" status="sp. nov.">
<emphasis id="B90B09650A2DFFB8F723769DFDD15FA7" box="[325,539,1944,1968]" italics="true" pageId="2" pageNumber="337">Stiptopodius brancoi</emphasis>
Strümpher &amp; Daniel
</taxonomicName>
,
<taxonomicNameLabel id="A238B41E0A2DFFB8F562769DFC495FA7" box="[772,899,1944,1968]" pageId="2" pageNumber="337" rank="species">new species</taxonomicNameLabel>
, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels.
</paragraph>
</caption>
<paragraph id="8BC0D5770A2CFFB9F6A17192FDB85ABE" blockId="3.[151,1436,151,789]" pageId="3" pageNumber="338">
<emphasis id="B90B09650A2CFFB9F6A17192FF3658A7" box="[199,252,151,176]" italics="true" pageId="3" pageNumber="338">Legs</emphasis>
(
<figureCitation id="1344C9F20A2CFFB9F7687192FEB658A6" box="[270,380,151,177]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="3" pageNumber="338">Figs. 1–2</figureCitation>
): Profemora dorsally and ventrally sparsely punctate, punctures bearing long setae, surfaces smooth; upper fore edge crenulated by closely set punctures bearing very long stiff setae; lower fore edge strongly marginate; hind edge weakly marginate. Meso- and metafemora dorsally and ventrally sparsely punctate, punctures fine, surfaces shiny; a row of widely spaced punctures present between the upper and lower hind edges, punctures bearing long setae; fore and lower hind edges not marginate, upper hind edge marginate. Protibiae with four prominent teeth on outer margin, margin distinctly crenulate basally and between prominent teeth, first two apical teeth very strongly developed, the third tooth intermediate between the second and forth, the forth tooth only slightly larger than basal crenulations; a single row of aligned setae running parallel to outer margin visible in dorsal view; ventrally with two carinae, a median one smooth, running nearly parallel to the outer margin, the other close to the inner margin, crenulated by setiferous closely set punctures. Protibial spur long and inwardly curved. Meso- and metatibiae triangular and strongly widened apically, with strong spur internally projected. Protarsi with five tarsomeres and clearly visible pair of claws; protarsomere V at least 3x longer than others (each individually). Meso- and metatarsomeres I–IV triangular, longer than wide, with internal line of strong setae on tarsomere I, strongly concave medially, bilobate distally, external lobe strong, internal lobe weak; tarsomere V cylindrical and longer than others, followed by pair of very tiny apical claws.
</paragraph>
<paragraph id="8BC0D5770A2CFFB9F6A173B6FA935AE6" blockId="3.[151,1436,151,789]" pageId="3" pageNumber="338">
<emphasis id="B90B09650A2CFFB9F6A173B6FEA45ADA" box="[199,366,691,717]" italics="true" pageId="3" pageNumber="338">Male genitalia</emphasis>
(
<figureCitation id="1344C9F20A2CFFB9F7E673B6FE0D5ADA" box="[384,455,691,717]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="3" pageNumber="338">Fig. 4</figureCitation>
): Dissection of the aedeagus by previous workers left the phallobase distorted, fortunately parameres were left intact. Parameres symmetrical, somewhat extended, distal portion with acute tips.
</paragraph>
<paragraph id="8BC0D5770A2CFFB9F6A173FEFE6A5B02" blockId="3.[151,1436,151,789]" box="[199,416,763,789]" pageId="3" pageNumber="338">
<emphasis id="B90B09650A2CFFB9F6A173FEFED15B02" box="[199,283,763,789]" italics="true" pageId="3" pageNumber="338">Female</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<caption id="DF0085FF0A2CFFB9F6F176B4FC485FF9" ID-DOI="http://doi.org/10.5281/zenodo.5119213" ID-Zenodo-Dep="5119213" httpUri="https://zenodo.org/record/5119213/files/figure.png" pageId="3" pageNumber="338" startId="3.[151,264,1969,1993]" targetBox="[157,1436,853,1943]" targetPageId="3">
<paragraph id="8BC0D5770A2CFFB9F6F176B4FC485FF9" blockId="3.[151,1437,1969,2030]" pageId="3" pageNumber="338">
<emphasis id="B90B09650A2CFFB9F6F176B4FEF45FDD" bold="true" box="[151,318,1969,1994]" pageId="3" pageNumber="338">FIGURES 6–8.</emphasis>
<taxonomicName id="4C7FAEF40A2CFFB9F72E76B7FD335FDD" authority="(Peringuey, 1901)" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[328,761,1970,1994]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B90B09650A2CFFB9F72E76B7FDFC5FDD" box="[328,566,1970,1994]" italics="true" pageId="3" pageNumber="338">Stiptopodius singularis</emphasis>
(
<bibRefCitation id="EFEEA8860A2CFFB9F42176B7FD385FDD" author="Peringuey, L." box="[583,754,1970,1994]" pageId="3" pageNumber="338" pagination="1 - 920" refId="ref5645" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12, 1 - 920. https: // doi. org / 10.1080 / 21560382.1901.9525977" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
)
</taxonomicName>
(SANC: South African National Collection of Insects, Pretoria, South Africa). 1, habitus, dorsal view; 2, aedeagus, lateral view; 3, labels.
</paragraph>
</caption>
<subSubSection id="C36586FC0A2BFFBEF6A17193FEBB5ABE" pageId="4" pageNumber="339" type="diagnosis">
<paragraph id="8BC0D5770A2BFFBEF6A17193FA4459C6" blockId="4.[151,1437,150,717]" pageId="4" pageNumber="339">
<emphasis id="B90B09650A2BFFBEF6A17193FE8858A7" bold="true" box="[199,322,150,176]" pageId="4" pageNumber="339">Diagnosis.</emphasis>
<taxonomicName id="4C7FAEF40A2BFFBEF72E7192FDF858A6" authority="Strümpher &amp; Daniel, 2021" authorityName="Strümpher &amp; Daniel" authorityYear="2021" box="[328,562,151,177]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="brancoi" status="sp. nov.">
<emphasis id="B90B09650A2BFFBEF72E7192FDF858A6" box="[328,562,151,177]" italics="true" pageId="4" pageNumber="339">Stiptopodius brancoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238B41E0A2BFFBEF45F7192FD0A58A6" box="[569,704,151,177]" pageId="4" pageNumber="339" rank="species">new species</taxonomicNameLabel>
belongs to the
<taxonomicName id="4C7FAEF40A2BFFBEF50D7192FC1158A6" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[875,987,151,177]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B90B09650A2BFFBEF50D7192FC1158A6" box="[875,987,151,177]" italics="true" pageId="4" pageNumber="339">singularis</emphasis>
</taxonomicName>
species group based on the shape of the lateral edge of the pronotum, which is strongly sinuate before posterolateral angle (in lateral view; for details see
<bibRefCitation id="EFEEA8860A2BFFBEF6F171DAFEED58EE" author="Branco, T." box="[151,295,223,249]" pageId="4" pageNumber="339" pagination="265 - 285" refId="ref5160" refString="Branco, T. (1991) Revision des genres du &quot; groupe &quot; Stiptopodius: Les genres Stiptopodius Harold et Stiptotarsus Branco (Coleoptera: Scarabaeidae). Annales de la Societe Entomologique de France, Nouvelle Serie, 27, 265 - 285." type="journal article" year="1991">Branco 1991</bibRefCitation>
and
<bibRefCitation id="EFEEA8860A2BFFBEF73A71DAFDE358EE" author="Daniel, G. M. &amp; Deschodt, C. M. &amp; Davis, A. L. V. &amp; Sole, C. L." box="[348,553,223,249]" pageId="4" pageNumber="339" pagination="394 - 404" refId="ref5355" refString="Daniel, G. M., Deschodt, C. M., Davis, A. L. V. &amp; Sole, C. L. (2020) The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera: Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa. Zootaxa, 4763 (3), 394 - 404. https: // doi. org / 10.11646 / zootaxa. 4763.3.4" type="journal article" year="2020">
Daniel
<emphasis id="B90B09650A2BFFBEF7CB71DAFE1458EE" box="[429,478,223,249]" italics="true" pageId="4" pageNumber="339">et al</emphasis>
. 2020
</bibRefCitation>
). In the
<taxonomicName id="4C7FAEF40A2BFFBEF4E771DAFD3B58EE" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[641,753,223,249]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B90B09650A2BFFBEF4E771DAFD3B58EE" box="[641,753,223,249]" italics="true" pageId="4" pageNumber="339">singularis</emphasis>
</taxonomicName>
species group the new species is morphologically most similar to
<taxonomicName id="4C7FAEF40A2BFFBEF6BA7006FD79590A" authority="(Peringuey, 1901)" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[220,691,259,285]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B90B09650A2BFFBEF6BA7006FE2A590A" box="[220,480,259,285]" italics="true" pageId="4" pageNumber="339">Stiptopodius singularis</emphasis>
(
<bibRefCitation id="EFEEA8860A2BFFBEF7967006FD60590A" author="Peringuey, L." box="[496,682,259,285]" pageId="4" pageNumber="339" pagination="1 - 920" refId="ref5645" refString="Peringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 12, 1 - 920. https: // doi. org / 10.1080 / 21560382.1901.9525977" type="journal article" year="1901">Péringuey, 1901</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="1344C9F20A2BFFBEF4A27006FCFA5909" box="[708,816,259,286]" captionStart="FIGURES 6–8" captionStartId="3.[151,264,1969,1993]" captionTargetBox="[157,1436,853,1943]" captionTargetId="figure-299@3.[151,1436,813,1946]" captionTargetPageId="3" captionText="FIGURES 6–8. Stiptopodius singularis (Péringuey, 1901) (SANC: South African National Collection of Insects, Pretoria, South Africa). 1, habitus, dorsal view; 2, aedeagus, lateral view; 3, labels." figureDoi="http://doi.org/10.5281/zenodo.5119213" httpUri="https://zenodo.org/record/5119213/files/figure.png" pageId="4" pageNumber="339">Figs. 6–8</figureCitation>
) however, the new species can easily be distinguished from the latter by the distinctive metallic green sheen on the pronotum (
<figureCitation id="1344C9F20A2BFFBEF5A77022FBC95956" box="[961,1027,295,321]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="4" pageNumber="339">Fig. 1</figureCitation>
), whereas
<taxonomicName id="4C7FAEF40A2BFFBEF2197022FA485956" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[1151,1410,295,321]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B90B09650A2BFFBEF2197022FA485956" box="[1151,1410,295,321]" italics="true" pageId="4" pageNumber="339">Stiptopodius singularis</emphasis>
</taxonomicName>
is predominantly black/brown with metallic sheen (
<figureCitation id="1344C9F20A2BFFBEF4D8704EFCC85971" box="[702,770,331,358]" captionStart="FIGURES 6–8" captionStartId="3.[151,264,1969,1993]" captionTargetBox="[157,1436,853,1943]" captionTargetId="figure-299@3.[151,1436,813,1946]" captionTargetPageId="3" captionText="FIGURES 6–8. Stiptopodius singularis (Péringuey, 1901) (SANC: South African National Collection of Insects, Pretoria, South Africa). 1, habitus, dorsal view; 2, aedeagus, lateral view; 3, labels." figureDoi="http://doi.org/10.5281/zenodo.5119213" httpUri="https://zenodo.org/record/5119213/files/figure.png" pageId="4" pageNumber="339">Fig. 6</figureCitation>
). Furthermore,
<taxonomicName id="4C7FAEF40A2BFFBEF5D4704EFB565972" authorityName="Strümpher &amp; Daniel" authorityYear="2021" box="[946,1180,331,357]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="brancoi">
<emphasis id="B90B09650A2BFFBEF5D4704EFB565972" box="[946,1180,331,357]" italics="true" pageId="4" pageNumber="339">Stiptopodius brancoi</emphasis>
</taxonomicName>
has well-defined frontal carina (
<figureCitation id="1344C9F20A2BFFBEF76B706AFE9A599E" box="[269,336,367,393]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="4" pageNumber="339">Fig. 4</figureCitation>
), whereas the frontal carina is weak to obsolete in
<taxonomicName id="4C7FAEF40A2BFFBEF5E4706AFB4E599E" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[898,1156,367,393]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B90B09650A2BFFBEF5E4706AFB4E599E" box="[898,1156,367,393]" italics="true" pageId="4" pageNumber="339">Stiptopodius singularis</emphasis>
</taxonomicName>
(
<figureCitation id="1344C9F20A2BFFBEF2F5706AFB1D599D" box="[1171,1239,367,394]" captionStart="FIGURES 6–8" captionStartId="3.[151,264,1969,1993]" captionTargetBox="[157,1436,853,1943]" captionTargetId="figure-299@3.[151,1436,813,1946]" captionTargetPageId="3" captionText="FIGURES 6–8. Stiptopodius singularis (Péringuey, 1901) (SANC: South African National Collection of Insects, Pretoria, South Africa). 1, habitus, dorsal view; 2, aedeagus, lateral view; 3, labels." figureDoi="http://doi.org/10.5281/zenodo.5119213" httpUri="https://zenodo.org/record/5119213/files/figure.png" pageId="4" pageNumber="339">Fig. 6</figureCitation>
). Lastly, the male genitalia of the two species are distinctive.
<taxonomicName id="4C7FAEF40A2BFFBEF41C7096FCAE59BA" authorityName="Strümpher &amp; Daniel" authorityYear="2021" box="[634,868,403,429]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="brancoi">
<emphasis id="B90B09650A2BFFBEF41C7096FCAE59BA" box="[634,868,403,429]" italics="true" pageId="4" pageNumber="339">Stiptopodius brancoi</emphasis>
</taxonomicName>
have the tips of the distal portion of the parameres more acute, while in
<taxonomicName id="4C7FAEF40A2BFFBEF7E570B2FD4C59C6" baseAuthorityName="Peringuey" baseAuthorityYear="1901" box="[387,646,439,465]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B90B09650A2BFFBEF7E570B2FD4C59C6" box="[387,646,439,465]" italics="true" pageId="4" pageNumber="339">Stiptopodius singularis</emphasis>
</taxonomicName>
the tips are more rounded or blunt (as shown in
<figureCitation id="1344C9F20A2BFFBEF2C370B2FB2D59C6" box="[1189,1255,439,465]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1360,208,1920]" captionTargetId="figure-18@2.[185,1401,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–5. Stiptopodius brancoi Strümpher &amp; Daniel, new species, holotype male (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, head and clypeus, dorsal view; red arrow indicate frontoclypeal carina, yellow arrow indicate frontal carina; 4, aedeagus, lateral view; 5, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5119211" httpUri="https://zenodo.org/record/5119211/files/figure.png" pageId="4" pageNumber="339">Fig. 4</figureCitation>
versus
<figureCitation id="1344C9F20A2BFFBEF35A70B2FA4A59C6" box="[1340,1408,439,465]" captionStart="FIGURES 6–8" captionStartId="3.[151,264,1969,1993]" captionTargetBox="[157,1436,853,1943]" captionTargetId="figure-299@3.[151,1436,813,1946]" captionTargetPageId="3" captionText="FIGURES 6–8. Stiptopodius singularis (Péringuey, 1901) (SANC: South African National Collection of Insects, Pretoria, South Africa). 1, habitus, dorsal view; 2, aedeagus, lateral view; 3, labels." figureDoi="http://doi.org/10.5281/zenodo.5119213" httpUri="https://zenodo.org/record/5119213/files/figure.png" pageId="4" pageNumber="339">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BC0D5770A2BFFBEF6A170DEFEBB5ABE" blockId="4.[151,1437,150,717]" pageId="4" pageNumber="339">
<taxonomicName id="4C7FAEF40A2BFFBEF6A170DEFE7E59E2" authority="Strümpher &amp; Daniel, 2021" authorityName="Strümpher &amp; Daniel" authorityYear="2021" box="[199,436,475,501]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="brancoi" status="sp. nov.">
<emphasis id="B90B09650A2BFFBEF6A170DEFE7E59E2" box="[199,436,475,501]" italics="true" pageId="4" pageNumber="339">Stiptopodius brancoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238B41E0A2BFFBEF7D970DEFD8259E2" box="[447,584,475,501]" pageId="4" pageNumber="339" rank="species">new species</taxonomicNameLabel>
also bear superficial resemblance to
<taxonomicName id="4C7FAEF40A2BFFBEF59C70DEFEE35A0E" authority="Daniel &amp; Deschodt, 2020" authorityName="Daniel &amp; Deschodt" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="mullerae">
<emphasis id="B90B09650A2BFFBEF59C70DEFB3E59E2" box="[1018,1268,475,501]" italics="true" pageId="4" pageNumber="339">Stiptopodius mullerae</emphasis>
<bibRefCitation id="EFEEA8860A2BFFBEF29870DEFEE35A0E" author="Daniel, G. M. &amp; Deschodt, C. M. &amp; Davis, A. L. V. &amp; Sole, C. L." pageId="4" pageNumber="339" pagination="394 - 404" refId="ref5355" refString="Daniel, G. M., Deschodt, C. M., Davis, A. L. V. &amp; Sole, C. L. (2020) The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera: Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa. Zootaxa, 4763 (3), 394 - 404. https: // doi. org / 10.11646 / zootaxa. 4763.3.4" type="journal article" year="2020">Daniel &amp; Deschodt, 2020</bibRefCitation>
</taxonomicName>
. These are the only two species in the group with green metallic pronota. However, the two species can easily be separated as follows:
<taxonomicName id="4C7FAEF40A2BFFBEF7977326FD2D5A2A" authorityName="Daniel &amp; Deschodt" authorityYear="2020" box="[497,743,547,573]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="mullerae">
<emphasis id="B90B09650A2BFFBEF7977326FD2D5A2A" box="[497,743,547,573]" italics="true" pageId="4" pageNumber="339">Stiptopodius mullerae</emphasis>
</taxonomicName>
is much larger (&gt;
<quantity id="4C8778920A2BFFBEF5D47326FC3E5A2A" box="[946,1012,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="339" unit="mm" value="7.0">7 mm</quantity>
) and is the single species of the genus which have the head, pronotum and elytra fully/densely setose, whereas
<emphasis id="B90B09650A2BFFBEF5A57342FB555A76" box="[963,1183,583,609]" italics="true" pageId="4" pageNumber="339">
Stitopodius
<taxonomicName id="4C7FAEF40A2BFFBEF22E7342FB555A76" authorityName="Strümpher &amp; Daniel" authorityYear="2021" box="[1096,1183,583,609]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="brancoi">brancoi</taxonomicName>
</emphasis>
is small (&lt;
<quantity id="4C8778920A2BFFBEF3457343FAAF5A76" box="[1315,1381,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="339" unit="mm" value="5.0">5 mm</quantity>
) and mostly glabrous in comparison. Male genitalia of both species are distinctive (for a more detailed treatment see
<bibRefCitation id="EFEEA8860A2BFFBEF6F1738AFEAF5ABE" author="Daniel, G. M. &amp; Deschodt, C. M. &amp; Davis, A. L. V. &amp; Sole, C. L." box="[151,357,655,681]" pageId="4" pageNumber="339" pagination="394 - 404" refId="ref5355" refString="Daniel, G. M., Deschodt, C. M., Davis, A. L. V. &amp; Sole, C. L. (2020) The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera: Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa. Zootaxa, 4763 (3), 394 - 404. https: // doi. org / 10.11646 / zootaxa. 4763.3.4" type="journal article" year="2020">
Daniel
<emphasis id="B90B09650A2BFFBEF68F738AFED05ABE" box="[233,282,655,681]" italics="true" pageId="4" pageNumber="339">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36586FC0A2BFFBEF6A173B7FAF05ADA" box="[199,1338,690,717]" pageId="4" pageNumber="339" type="distribution">
<paragraph id="8BC0D5770A2BFFBEF6A173B7FAF05ADA" blockId="4.[151,1437,150,717]" box="[199,1338,690,717]" pageId="4" pageNumber="339">
<emphasis id="B90B09650A2BFFBEF6A173B7FEAB5ADB" bold="true" box="[199,353,690,716]" pageId="4" pageNumber="339">Distribution.</emphasis>
<taxonomicName id="4C7FAEF40A2BFFBEF70173B6FD9B5ADA" authority="Strümpher &amp; Daniel, 2021" authorityName="Strümpher &amp; Daniel" authorityYear="2021" box="[359,593,691,717]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="brancoi" status="sp. nov.">
<emphasis id="B90B09650A2BFFBEF70173B6FD9B5ADA" box="[359,593,691,717]" italics="true" pageId="4" pageNumber="339">Stiptopodius brancoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238B41E0A2BFFBEF43F73B6FD155ADA" box="[601,735,691,717]" pageId="4" pageNumber="339" rank="species">new species</taxonomicNameLabel>
is only known from
<typeStatus id="54C46BD50A2BFFBEF5AF73B6FC335ADA" box="[969,1017,691,717]" pageId="4" pageNumber="339">type</typeStatus>
locality in
<collectingCountry id="F36895E70A2BFFBEF21E73B6FB125ADA" box="[1144,1240,691,717]" name="Namibia" pageId="4" pageNumber="339">Namibia</collectingCountry>
(
<figureCitation id="1344C9F20A2BFFBEF28E73B6FAE65ADA" box="[1256,1324,691,717]" captionStart="FIGURE 9" captionStartId="4.[151,250,1999,2023]" captionTargetBox="[221,1362,748,1964]" captionTargetId="figure-293@4.[213,1375,741,1975]" captionTargetPageId="4" captionText="FIGURE 9. Type locality of Stiptopodius brancoi Strümpher &amp; Daniel, new species in Namibia." figureDoi="http://doi.org/10.5281/zenodo.5119215" httpUri="https://zenodo.org/record/5119215/files/figure.png" pageId="4" pageNumber="339">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0085FF0A2BFFBEF6F176CAFBA45FF0" ID-DOI="http://doi.org/10.5281/zenodo.5119215" ID-Zenodo-Dep="5119215" box="[151,1134,1999,2024]" httpUri="https://zenodo.org/record/5119215/files/figure.png" pageId="4" pageNumber="339" startId="4.[151,250,1999,2023]" targetBox="[221,1362,748,1964]" targetPageId="4">
<paragraph id="8BC0D5770A2BFFBEF6F176CAFBA45FF0" blockId="4.[151,1134,1999,2024]" box="[151,1134,1999,2024]" pageId="4" pageNumber="339">
<emphasis id="B90B09650A2BFFBEF6F176CAFED95FF0" bold="true" box="[151,275,1999,2023]" pageId="4" pageNumber="339">FIGURE 9.</emphasis>
Type locality of
<taxonomicName id="4C7FAEF40A2BFFBEF7D976CAFCA15FF0" authority="Strumpher &amp; Daniel" authorityName="Strumpher &amp; Daniel" authorityYear="2021" box="[447,875,1999,2023]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="brancoi" status="sp. nov.">
<emphasis id="B90B09650A2BFFBEF7D976CAFD595FF0" box="[447,659,1999,2023]" italics="true" pageId="4" pageNumber="339">Stiptopodius brancoi</emphasis>
Strümpher &amp; Daniel
</taxonomicName>
,
<taxonomicNameLabel id="A238B41E0A2BFFBEF51376D5FC255FF0" box="[885,1007,1999,2023]" pageId="4" pageNumber="339" rank="species">new species</taxonomicNameLabel>
in Namibia.
</paragraph>
</caption>
<subSubSection id="C36586FC0A2AFFBFF6A17193FBBF58C2" pageId="5" pageNumber="340" type="etymology">
<paragraph id="8BC0D5770A2AFFBFF6A17193FA5658A7" blockId="5.[151,1436,150,213]" box="[199,1436,150,177]" pageId="5" pageNumber="340">
<emphasis id="B90B09650A2AFFBFF6A17193FE8458A7" bold="true" box="[199,334,150,176]" pageId="5" pageNumber="340">Etymology.</emphasis>
The new species is dedicated to
<collectingCountry id="F36895E70A2AFFBFF4AD7193FCD358A7" box="[715,793,150,176]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="5" pageNumber="340">Tristão</collectingCountry>
Branco (
<collectingRegion id="49BB1B950A2AFFBFF5E27193FC0F58A7" box="[900,965,150,176]" country="Portugal" name="Porto" pageId="5" pageNumber="340">Porto</collectingRegion>
,
<collectingCountry id="F36895E70A2AFFBFF5B47193FBFF58A6" box="[978,1077,150,177]" name="Portugal" pageId="5" pageNumber="340">Portugal</collectingCountry>
), a specialist in Palaearctic and
</paragraph>
<paragraph id="8BC0D5770A2AFFBFF6F171BFFBBF58C2" blockId="5.[151,1436,150,213]" box="[151,1141,186,213]" pageId="5" pageNumber="340">
Afrotropical Scarabaeoidea who worked extensively on
<taxonomicName id="4C7FAEF40A2AFFBFF56B71BFFC5358C3" authorityName="Harold" authorityYear="1871" box="[781,921,186,212]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B90B09650A2AFFBFF56B71BFFC5358C3" box="[781,921,186,212]" italics="true" pageId="5" pageNumber="340">Stiptopodius</emphasis>
</taxonomicName>
and related genera.
</paragraph>
</subSubSection>
</treatment>
</document>
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