treatments-xml/data/03/D6/13/03D61304343DFFC2FEC6FA09FD0BFBC1.xml
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<document id="13E64E15C6F90DD1B5FD606EDEE10A3D" ID-DOI="10.11646/zootaxa.1214.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5064559" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625282476021" checkinUser="felipe" docAuthor="Komai, Tomoyuki &amp; Osawa, Masayuki" docDate="2006" docId="03D61304343DFFC2FEC6FA09FD0BFBC1" docLanguage="en" docName="zt01214p107.pdf" docOrigin="Zootaxa 1214 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1214.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Pagurixus brachydactylus Komai &amp; Osawa 2006, n. sp." docType="treatment" docVersion="4" lastPageNumber="71" masterDocId="FFEF6B7C3402FF84FFCEFFC5FFFCFFC7" masterDocTitle="A review of the Pagurixus boninensis species group, with descriptions of six new species (Crustacea: Decapoda: Anomura: Paguridae)" masterLastPageNumber="107" masterPageNumber="1" pageNumber="64" updateTime="1698986104405" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3A471F7E5876812BB52B9993404BE798">A review of the Pagurixus boninensis species group, with descriptions of six new species (Crustacea: Decapoda: Anomura: Paguridae)</mods:title>
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<mods:namePart id="A9595D14AA4EF24AA5BDF371D6520592">Komai, Tomoyuki</mods:namePart>
<mods:affiliation id="2CADADA17464482155A92AE4362502D2">Natural History Museum and Institute, Chiba, 955 - 2 Aoba-cho, Chuo-ku, Chiba 260 - 8682, Japan.</mods:affiliation>
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<mods:namePart id="54B56E2341565D00D9A704764365DE21">Osawa, Masayuki</mods:namePart>
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<subSubSection id="C365F199343DFFBBFEC6FA09FD71FA21" box="[264,653,1484,1510]" pageId="63" pageNumber="64" type="nomenclature">
<paragraph id="8BC0A212343DFFBBFEC6FA09FD71FA21" blockId="63.[264,653,1484,1545]" box="[264,653,1484,1510]" pageId="63" pageNumber="64">
<heading id="D088157E343DFFBBFEC6FA09FD71FA21" bold="true" box="[264,653,1484,1510]" fontSize="11" level="1" pageId="63" pageNumber="64" reason="1">
<emphasis id="B90B7E00343DFFBBFEC6FA09FD71FA21" bold="true" box="[264,653,1484,1510]" pageId="63" pageNumber="64">
<taxonomicName id="4C7FD991343DFFBBFEC6FA09FDBEFA21" ID-CoL="c5261e57-7aa6-4025-96d3-8fe2e18f3269" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" box="[264,578,1484,1510]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brachydactylus" status="sp. nov.">
<emphasis id="B90B7E00343DFFBBFEC6FA09FDBEFA21" bold="true" box="[264,578,1484,1510]" italics="true" pageId="63" pageNumber="64">Pagurixus brachydactylus</emphasis>
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<taxonomicNameLabel id="A238C37B343DFFBBFD85FA09FD71FA21" box="[587,653,1484,1510]" pageId="63" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C365F199343DFFBBFEC6FA2AFDC8F9CE" box="[264,564,1519,1545]" pageId="63" pageNumber="64" type="description">
<paragraph id="8BC0A212343DFFBBFEC6FA2AFDC8F9CE" blockId="63.[264,653,1484,1545]" box="[264,564,1519,1545]" pageId="63" pageNumber="64">
(
<figureCitation id="1344BE97343DFFBBFEDFFA2AFE62F9CE" box="[273,414,1519,1545]" captionStart-0="FIGURE 26" captionStart-1="FIGURE 27" captionStart-2="FIGURE 28" captionStartId-0="66.[264,368,1637,1661]" captionStartId-1="67.[264,367,1690,1714]" captionStartId-2="68.[264,368,1437,1461]" captionTargetBox-0="[296,1304,475,1605]" captionTargetBox-1="[310,1250,321,1679]" captionTargetBox-2="[299,1292,316,1423]" captionTargetId-0="figure-190@66.[264,1323,451,1613]" captionTargetId-1="figure-81@67.[286,1311,290,1680]" captionTargetId-2="figure-156@68.[266,1325,300,1438]" captionTargetPageId-0="66" captionTargetPageId-1="67" captionTargetPageId-2="68" captionText-0="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." captionText-1="FIGURE 27. Pagurixus brachydactylus n. sp. Holotype male (SL 2.4 mm) from Chichi­jima Island, Ogasawara Islands (KMNH­Iv 000,021). A, right cheliped, chela and carpus, dorsal view; B, entire right cheliped, mesial view; C, same, lateral view; D, left cheliped, chela, dorsal view; E, entire left cheliped, mesial view; F, same, lateral view; G, left cheliped, carpus, dorsal view. Scale bar: 1 mm." captionText-2="FIGURE 28. Pagurixus brachydactylus n. sp. Holotype male (SL 2.4 mm) from Chichi­jima Island, Ogasawara Islands (KMNH­Iv 000,021). A, right second pereopod, lateral view; B, left third pereopod, lateral view; C, right second pereopod, dactylus and distal part of propodus, mesial view; D, left third pereopod, dactylus and distal part of propodus, mesial view. Scale bar: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5064638" figureDoi-1="http://doi.org/10.5281/zenodo.5064642" figureDoi-2="http://doi.org/10.5281/zenodo.5064644" httpUri-0="https://zenodo.org/record/5064638/files/figure.png" httpUri-1="https://zenodo.org/record/5064642/files/figure.png" httpUri-2="https://zenodo.org/record/5064644/files/figure.png" pageId="63" pageNumber="64">Figs. 2628</figureCitation>
,
<figureCitation id="1344BE97343DFFBBFE64FA2AFDFEF9CE" box="[426,514,1519,1545]" captionStart="FIGURE 45" captionStartId="100.[264,367,1516,1540]" captionTargetBox="[285,1310,300,1456]" captionTargetId="figure-133@100.[285,1310,300,1456]" captionTargetPageId="100" captionText="FIGURE 45. A, Pagurixus boninensis (Melin, 1939), ovigerous female (SL 2.7 mm) from Hachijo Island, Izu Islands, Japan (CMNH­ZC 1688), photo taken by J. Okuno; B, Pagurixus carinimanus n. sp., paratype female (SL 1.6 mm) from Okinawa Island, Ryukyu Islands (CBM­ZC 8481), photo taken by Y. Fujita; C, D, Pagurixus brachydactylus n. sp., male (SL 2.6 mm) from Chichi­jima Island, Ogasawara Islands, photo taken by M. Osawa." figureDoi="http://doi.org/10.5281/zenodo.5064702" httpUri="https://zenodo.org/record/5064702/files/figure.png" pageId="63" pageNumber="64">45C, D</figureCitation>
,
<figureCitation id="1344BE97343DFFBBFDC0FA2AFDD0F9CE" box="[526,556,1519,1545]" captionStart="FIGURE 48" captionStartId="102.[264,367,1798,1822]" captionTargetBox="[269,1278,1196,1754]" captionTargetId="figure-121@102.[264,1289,1158,1780]" captionTargetPageId="102" captionText="FIGURE 48. Geographic distribution of Pagurixus boninensis (), P. festinus (), P. brachydactylus n. sp. (), P. carinimanus n. sp. (), P. paulayi n. sp. (Π), P. pseliophorus n. sp ()., and P. ruber n. sp. ()." figureDoi="http://doi.org/10.5281/zenodo.5064708" httpUri="https://zenodo.org/record/5064708/files/figure.png" pageId="63" pageNumber="64">48</figureCitation>
)
</paragraph>
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<taxonomicName id="4C7FD991343DFFBBFEC6F9FAFE12F991" ID-CoL="4e0dd3de-c691-4112-a001-45ed9e02e133" baseAuthorityName="Melin" baseAuthorityYear="1939" box="[264,494,1599,1622]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="boninensis">
<emphasis id="B90B7E00343DFFBBFEC6F9FAFE12F991" box="[264,494,1599,1622]" italics="true" pageId="63" pageNumber="64">Pagurixus boninensis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0ADE8403343DFFBBFDC1F9FBFD39F992" author="Ooishi, S." box="[527,709,1598,1621]" page="90" pageId="63" pageNumber="64" year="1970">
<bibRefCitation id="EFEEDFE3343DFFBBFDC1F9FBFD39F992" author="Ooishi, S." box="[527,709,1598,1621]" pageId="63" pageNumber="64" pagination="75 - 104" refId="ref42933" refString="Ooishi, S. (1970) Marine invertebrates fauna of the Ogasawara and Volcano Islands collected by S. Ooishi, Y. Tomida, K. Izawa and S. Manabe. In: Report on the Marine Biological Expedition to the Ogasawara (Bonin) Islands, 1968, pp. 75 - 104." type="journal article" year="1970">Ooishi, 1970: 90</bibRefCitation>
</treatmentCitation>
, pl. 12: fig. 13.
<collectingRegion id="49BB6CF0343DFFBBFCA3F9FBFC69F992" box="[877,917,1598,1621]" country="Maldives" name="Uthuru" pageId="63" pageNumber="64">Not</collectingRegion>
<taxonomicName id="4C7FD991343DFFBBFC52F9FAFADCF992" ID-CoL="4e0dd3de-c691-4112-a001-45ed9e02e133" authority="(Melin, 1939)" baseAuthorityName="Melin" baseAuthorityYear="1939" box="[924,1312,1598,1622]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="boninensis">
<emphasis id="B90B7E00343DFFBBFC52F9FAFB7EF991" box="[924,1154,1599,1622]" italics="true" pageId="63" pageNumber="64">Pagurixus boninensis</emphasis>
(
<bibRefCitation id="EFEEDFE3343DFFBBFB5EF9FBFAE4F992" author="Melin, G." box="[1168,1304,1598,1621]" pageId="63" pageNumber="64" pagination="1 - 119" refId="ref42341" refString="Melin, G. (1939) Paguriden und Galatheiden von Prof. Dr. Sixten Bocks Expedition nach den Bonin-Inseln 1914. Kongliga Svenska Vetenskapsakademiens Handlinger, Series 3, 18 (2), 1 - 119." type="journal article" year="1939">Melin, 1939</bibRefCitation>
)
</taxonomicName>
.
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<emphasis id="B90B7E00343DFFBBFEC6F942FE55F967" box="[264,425,1671,1696]" italics="true" pageId="63" pageNumber="64">Type material</emphasis>
</paragraph>
<paragraph id="8BC0A212343DFFBBFEF6F96AFAD8F936" blockId="63.[264,1323,1671,1897]" pageId="63" pageNumber="64">
<materialsCitation id="3B17A84F343DFFBBFEF6F96AFAD8F936" collectingDate="1968-08-07" collectorName="S. Ooishi." county="Ogasawara Islands" elevation="3" location="Minami-saki" municipality="Chichi-jima Island" pageId="63" pageNumber="64" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C41CB0343DFFBBFEF6F96AFE2BF90E" box="[312,471,1711,1737]" pageId="63" pageNumber="64" type="holotype">HOLOTYPE</typeStatus>
: KMNH­Iv 000,021, male (SL
<quantity id="4C870FF7343DFFBBFCA1F96AFC2AF90E" box="[879,982,1711,1737]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="63" pageNumber="64" unit="mm" value="2.4">2.4 mm</quantity>
),
<location id="8EA0F4C9343DFFBBFC3CF96AFB77F90E" LSID="urn:lsid:plazi:treatment:03D61304343DFFC2FEC6FA09FD0BFBC1:8EA0F4C9343DFFBBFC3CF96AFB77F90E" box="[1010,1163,1711,1737]" county="Ogasawara Islands" municipality="Chichi-jima Island" name="Minami-saki" pageId="63" pageNumber="64">Minami­saki</location>
,
<collectingMunicipality id="6BA43868343DFFBBFB51F96AFEAEF936" pageId="63" pageNumber="64">Chichi­jima Island</collectingMunicipality>
,
<collectingCounty id="62A1DA9E343DFFBBFE90F912FDC0F936" box="[350,572,1751,1777]" pageId="63" pageNumber="64">Ogasawara Islands</collectingCounty>
,
<quantity id="4C870FF7343DFFBBFD86F912FD8AF936" box="[584,630,1751,1777]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="64" unit="m" value="3.0">
<elevation id="00524521343DFFBBFD86F912FD8AF936" box="[584,630,1751,1777]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="64" unit="m" value="3.0">3 m</elevation>
</quantity>
,
<collectedFrom id="3863E1EF343DFFBBFD4CF912FC51F937" box="[642,941,1751,1777]" pageId="63" pageNumber="64">
on
<taxonomicName id="4C7FD991343DFFBBFD68F912FC51F937" baseAuthorityName="Ellis &amp; Solander" baseAuthorityYear="1786" box="[678,941,1751,1776]" class="Anthozoa" family="Pocilloporidae" genus="Pocillopora" kingdom="Animalia" order="Scleractinia" pageId="63" pageNumber="64" phylum="Cnidaria" rank="species" species="verrucosa">
<emphasis id="B90B7E00343DFFBBFD68F912FC51F937" box="[678,941,1751,1776]" italics="true" pageId="63" pageNumber="64">Pocillopora verrucosa</emphasis>
</taxonomicName>
</collectedFrom>
,
<date id="FFC184D2343DFFBBFC72F912FB96F936" box="[956,1130,1751,1777]" pageId="63" pageNumber="64" value="1968-08-07">
<collectingDate id="EF857D3A343DFFBBFC72F912FB96F936" box="[956,1130,1751,1777]" pageId="63" pageNumber="64" value="1968-08-07">7 August 1968</collectingDate>
</date>
, coll.
<collectorName id="268AC7C4343DFFBBFB7EF912FAD8F936" box="[1200,1316,1751,1777]" pageId="63" pageNumber="64">S. Ooishi.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BC0A212343DFFBBFEC6F93AFB3FF8AE" blockId="63.[264,1323,1671,1897]" pageId="63" pageNumber="64">
<materialsCitation id="3B17A84F343DFFBBFEC6F93AFB58F8DE" box="[264,1188,1791,1817]" collectingDate="1968-08-07" collectorName="S. Ooishi." county="Ogasawara Islands" elevation="3" location="Minami-saki" municipality="Chichi-jima Island" pageId="63" pageNumber="64" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54C41CB0343DFFBBFEC6F93AFE52F8DE" box="[264,430,1791,1817]" pageId="63" pageNumber="64" type="paratype">PARATYPES</typeStatus>
: KMNH­Iv 000,022,
<specimenCount id="9D79699B343DFFBBFD66F93AFD0AF8DE" box="[680,758,1791,1817]" pageId="63" pageNumber="64" type="male">1 male</specimenCount>
(SL
<quantity id="4C870FF7343DFFBBFCFEF93AFC71F8DE" box="[816,909,1791,1817]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="63" pageNumber="64" unit="mm" value="2.2">2.2 mm</quantity>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B17A84F343DFFBBFB61F93AFC5EF886" collectingDate="1968-08-07" collectorName="S. Ooishi." county="Ogasawara Islands" elevation="3" location="Minami-saki" municipality="Chichi-jima Island" pageId="63" pageNumber="64" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
KMNH­Iv 000,023,
<specimenCount id="9D79699B343DFFBBFEB6F8E2FE1CF886" box="[376,480,1831,1857]" pageId="63" pageNumber="64" type="female">1 female</specimenCount>
(SL
<quantity id="4C870FF7343DFFBBFDD1F8E2FD83F886" box="[543,639,1831,1857]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="63" pageNumber="64" unit="mm" value="2.3">2.3 mm</quantity>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B17A84F343DFFBBFC7EF8E2FB42F8AE" collectingDate="1998-10-19" collectorName="M. Osawa" location="Haha-jima Island" pageId="63" pageNumber="64" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
CBM­ZC 8478,
<specimenCount id="9D79699B343DFFBBFBB8F8E2FB3BF886" box="[1142,1223,1831,1857]" pageId="63" pageNumber="64" type="male">1 male</specimenCount>
(SL
<quantity id="4C870FF7343DFFBBFAC8F8E2FEC6F8AE" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="63" pageNumber="64" unit="mm" value="2.4">2.4 mm</quantity>
), Higashi Port,
<location id="8EA0F4C9343DFFBBFE3EF88AFD41F8AE" LSID="urn:lsid:plazi:treatment:03D61304343DFFC2FEC6FA09FD0BFBC1:8EA0F4C9343DFFBBFE3EF88AFD41F8AE" box="[496,701,1871,1897]" name="Haha-jima Island" pageId="63" pageNumber="64">Haha­jima Island</location>
, subtidal, coll.
<collectorName id="268AC7C4343DFFBBFCA1F88AFC17F8AE" box="[879,1003,1871,1897]" pageId="63" pageNumber="64">M. Osawa</collectorName>
,
<date id="FFC184D2343DFFBBFC38F88AFB42F8AE" box="[1014,1214,1871,1897]" pageId="63" pageNumber="64" value="1998-10-19">
<collectingDate id="EF857D3A343DFFBBFC38F88AFB42F8AE" box="[1014,1214,1871,1897]" pageId="63" pageNumber="64" value="1998-10-19">19 October 1998</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0A2123442FFC4FEC6FED9FE6EFEF2" blockId="64.[264,1325,284,1892]" box="[264,402,284,309]" pageId="64" pageNumber="65">
<emphasis id="B90B7E003442FFC4FEC6FED9FE6EFEF2" box="[264,402,284,309]" italics="true" pageId="64" pageNumber="65">Description</emphasis>
</paragraph>
<paragraph id="8BC0A2123442FFC4FEF6FE81FE00FE39" blockId="64.[264,1325,284,1892]" pageId="64" pageNumber="65">
Shield (
<figureCitation id="1344BE973442FFC4FE5CFE81FE00FE99" box="[402,508,324,350]" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="64" pageNumber="65">Fig. 26A</figureCitation>
) 1.01.1 times as long as broad; anterior margin between rostrum and lateral projections slightly concave; anterolateral margins sloping; dorsal surface with few tufts of short setae laterally. Rostrum triangular, reaching midlength of ocular acicles, moderately slender, terminating acutely. Lateral projections only slightly produced, with submarginal spinule.
</paragraph>
<paragraph id="8BC0A2123442FFC4FEF6FDC9FC0AFD59" blockId="64.[264,1325,284,1892]" pageId="64" pageNumber="65">
Ocular peduncle (
<figureCitation id="1344BE973442FFC4FDC5FDC9FD8BFDE1" box="[523,631,524,550]" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="64" pageNumber="65">Fig. 26A</figureCitation>
) moderately long and stout, 0.60.7 length of shield, each with row of tufts of short setae on dorsal surface mesially; corneas not dilated, corneal width 0.30.4 of peduncular length; basal part slightly broader than corneal width. Ocular acicles subovate or subtriangular, with small submarginal spine.
</paragraph>
<paragraph id="8BC0A2123442FFC4FEF6FD69FE04FC49" blockId="64.[264,1325,284,1892]" pageId="64" pageNumber="65">
Antennular peduncle (
<figureCitation id="1344BE973442FFC4FD8DFD69FD29FD01" box="[579,725,684,710]" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="64" pageNumber="65">Fig. 26A, B</figureCitation>
) overreaching distal margin of cornea by 0.10.2 length of ultimate segment. Ultimate segment with tufts of long setae at dorsolateral angle; 2 longitudinal rows of setae on ventral surface, each consisting of mixture of dense short setae and tufts of longer setae directed ventrolaterally (
<figureCitation id="1344BE973442FFC4FC7CFCE1FBDFFCF9" box="[946,1059,804,830]" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="64" pageNumber="65">Fig. 26C</figureCitation>
). Basal segment with small lateral spine on statocyst lobe. Ventral flagellum with numerous long setae on lateral and ventral margins.
</paragraph>
<paragraph id="8BC0A2123442FFC4FEF6FC59FE3AFB61" blockId="64.[264,1325,284,1892]" pageId="64" pageNumber="65">
Antennal peduncle (
<figureCitation id="1344BE973442FFC4FDE6FC59FD69FC71" box="[552,661,924,950]" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="64" pageNumber="65">Fig. 26A</figureCitation>
) just reaching or slightly overreaching distal margin of cornea. Second segment with small spine at dorsomesial distal angle; laterodistal projection short, not reaching midlength of fourth segment. First segment with small laterodistal spine; ventromesial distal margin produced, unarmed or with spinule just lateral to antennal gland opening. Antennal acicle short, arcuate, reaching base of cornea; mesial margin with row of short stiff setae. Flagellum relatively short, exceeding 2.0 length of shield.
</paragraph>
<paragraph id="8BC0A2123442FFC4FEF6FB71FC37F8A3" blockId="64.[264,1325,284,1892]" pageId="64" pageNumber="65">
Right cheliped of males (
<figureCitation id="1344BE973442FFC4FDACFB71FD13FB09" box="[610,751,1204,1230]" captionStart="FIGURE 27" captionStartId="67.[264,367,1690,1714]" captionTargetBox="[310,1250,321,1679]" captionTargetId="figure-81@67.[286,1311,290,1680]" captionTargetPageId="67" captionText="FIGURE 27. Pagurixus brachydactylus n. sp. Holotype male (SL 2.4 mm) from Chichi­jima Island, Ogasawara Islands (KMNH­Iv 000,021). A, right cheliped, chela and carpus, dorsal view; B, entire right cheliped, mesial view; C, same, lateral view; D, left cheliped, chela, dorsal view; E, entire left cheliped, mesial view; F, same, lateral view; G, left cheliped, carpus, dorsal view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5064642" httpUri="https://zenodo.org/record/5064642/files/figure.png" pageId="64" pageNumber="65">Fig. 27AC</figureCitation>
) not particularly elongate, stout. Chela subovate in dorsal view, 1.51.7 times longer than broad. Dactylus 0.60.7 length of palm; dorsomesial margin not delimited; surfaces covered with coarse granules; cutting edge with 4 or 5 low calcareous teeth, terminating in small calcareous claw. Palm slightly longer than carpus; convex dorsal surface covered with coarse granules, dorsolateral and dorsomesial margins not delimited; lateral, mesial and ventral surfaces granular, with few tufts of short setae on ventral surface. Cutting edge of fixed finger with 1 or 2 low calcareous teeth, terminating in calcareous claw. Carpus slightly longer than merus; dorsolateral and dorsomesial margins not delimited; all surfaces coarsely granular (granules occasionally arranged in short to long transverse rows), lateral surface lacking median ridge. Meral­carpal articulation lacking any pronounced clockwise rotation; dorsal surface of merus with few low transverse ridges distally, otherwise almost smooth, and with few tufts of setae, dorsodistal margin with row of short bristles, but without spines; lateral face with low, rounded protuberances in ventral half, otherwise nearly smooth, ventrolateral margin with row of moderately small spines in distal half; mesial face nearly smooth, with scattered long stiff setae, ventromesial margin with row of rounded tubercles; ventral faces with scattered small tubercles and long setae. Ischium with row of small denticles on ventromesial margin, otherwise unarmed.
</paragraph>
<paragraph id="8BC0A2123443FFC5FEF6FED9FD38FEF1" blockId="65.[264,1325,284,1867]" box="[312,708,284,310]" pageId="65" pageNumber="66">Right cheliped of female missing.</paragraph>
<paragraph id="8BC0A2123443FFC5FEF6FE86FC78FBAF" blockId="65.[264,1325,284,1867]" pageId="65" pageNumber="66">
Left cheliped (
<figureCitation id="1344BE973443FFC5FE25FE86FD82FE9A" box="[491,638,323,349]" captionStart="FIGURE 27" captionStartId="67.[264,367,1690,1714]" captionTargetBox="[310,1250,321,1679]" captionTargetId="figure-81@67.[286,1311,290,1680]" captionTargetPageId="67" captionText="FIGURE 27. Pagurixus brachydactylus n. sp. Holotype male (SL 2.4 mm) from Chichi­jima Island, Ogasawara Islands (KMNH­Iv 000,021). A, right cheliped, chela and carpus, dorsal view; B, entire right cheliped, mesial view; C, same, lateral view; D, left cheliped, chela, dorsal view; E, entire left cheliped, mesial view; F, same, lateral view; G, left cheliped, carpus, dorsal view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5064642" httpUri="https://zenodo.org/record/5064642/files/figure.png" pageId="65" pageNumber="66">Fig. 27DG</figureCitation>
) moderately slender, similar between males and female. Chela 2.22.5 times longer than broad. Dactylus slightly longer than palm, with sparse tufts of setae on surfaces (setae on ventral surface longest); dorsal surface unarmed; cutting edge with row of small corneous teeth, terminating in small corneous claw. Palm about half length of carpus; dorsal surface weakly elevated in mid­line and bearing scattered very small tubercles, dorsolateral margin faintly delimited by row of small tubercles, dorsomesial margins not delimited; lateral, mesial and ventral surfaces with small tubercles or low protuberances; scattered long setae present on mesial and ventral surfaces. Cutting edge of fixed finger with row of small corneous teeth. Carpus moderately stout, somewhat compressed laterally, 0.80.9 length of chela and slightly shorter than merus; length 2.12.3 of distal width and 1.82.0 of greatest height; dorsolateral and dorsomesial margins each with row of low, small protuberances and short bristles, but no conspicuous spines present; surfaces with few very small tubercles or granules; sparse long setae on mesial and ventral surfaces; lateral surface lacking median longitudinal ridge, thus nearly perpendicular. Merus with 1 subdistal transverse ridge extending onto lateral and mesial surfaces, otherwise nearly smooth, dorsodistal margin with row of short stiff setae; lateral surface with scattered, very small low protuberances, ventrolateral margin with row of moderately small spines; mesial face nearly smooth, ventromesial margin faintly granular and with row of short to long bristles or stiff setae; ventral surface weakly convex, with few small low tubercles and long stiff setae. Ischium with row of tiny denticles on ventromesial margin, otherwise unarmed.
</paragraph>
<paragraph id="8BC0A2123443FFC5FEF6FBB1FD8DF977" blockId="65.[264,1325,284,1867]" pageId="65" pageNumber="66">
Ambulatory legs (
<figureCitation id="1344BE973443FFC5FDC5FBB1FD6AFB49" box="[523,662,1140,1166]" captionStart="FIGURE 28" captionStartId="68.[264,368,1437,1461]" captionTargetBox="[299,1292,316,1423]" captionTargetId="figure-156@68.[266,1325,300,1438]" captionTargetPageId="68" captionText="FIGURE 28. Pagurixus brachydactylus n. sp. Holotype male (SL 2.4 mm) from Chichi­jima Island, Ogasawara Islands (KMNH­Iv 000,021). A, right second pereopod, lateral view; B, left third pereopod, lateral view; C, right second pereopod, dactylus and distal part of propodus, mesial view; D, left third pereopod, dactylus and distal part of propodus, mesial view. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064644" httpUri="https://zenodo.org/record/5064644/files/figure.png" pageId="65" pageNumber="66">Fig. 28A, B</figureCitation>
) relatively short, stout, similar from right to left. Dactyli (
<figureCitation id="1344BE973443FFC5FEDFFB5EFE58FB72" box="[273,420,1179,1205]" captionStart="FIGURE 28" captionStartId="68.[264,368,1437,1461]" captionTargetBox="[299,1292,316,1423]" captionTargetId="figure-156@68.[266,1325,300,1438]" captionTargetPageId="68" captionText="FIGURE 28. Pagurixus brachydactylus n. sp. Holotype male (SL 2.4 mm) from Chichi­jima Island, Ogasawara Islands (KMNH­Iv 000,021). A, right second pereopod, lateral view; B, left third pereopod, lateral view; C, right second pereopod, dactylus and distal part of propodus, mesial view; D, left third pereopod, dactylus and distal part of propodus, mesial view. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5064644" httpUri="https://zenodo.org/record/5064644/files/figure.png" pageId="65" pageNumber="66">Fig. 28C, D</figureCitation>
) 0.50.8 length of propodi, 3.03.7 times longer than high, terminating in large corneous claws; dorsal surfaces each with row of sparse short setae; lateral and mesial faces faintly sulcate medially, each with few tufts of short setae, mesial faces unarmed; ventral margins each with 36 moderately long corneous spines notably increasing in size distally. Propodi not tapering distally, 3.24.0 times longer than high; dorsal surfaces each with row of low protuberances and short stiff setae or bristles; lateral faces smooth or with scattered very low protuberances; ventral margins each with 13 corneous spinules, ventrodistal margins each with paired corneous spines. Carpi each with small dorsodistal spine, dorsal surfaces nearly smooth, but with row of short bristles; lateral faces with sparse granules and few short bristles. Meri each with moderately long setae and short bristles on nearly smooth dorsal surface; lateral surfaces nearly smooth; ventrolateral distal margins each with small subdistal spine (second) or unarmed (third), ventral surfaces with row of low protuberances (second) or smooth (third), all with sparse short to moderately long setae.
</paragraph>
<paragraph id="8BC0A2123443FFC6FEF6F979FBCAFE6C" blockId="65.[264,1325,284,1867]" lastBlockId="66.[264,1323,284,427]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
Fourth pereopods generally similar from right to left in males (
<figureCitation id="1344BE973443FFC5FBD8F979FB5BF911" box="[1046,1191,1724,1750]" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="65" pageNumber="66">Fig. 26DG</figureCitation>
), markedly unequal with left larger in female (
<figureCitation id="1344BE973443FFC5FD7BF926FCB6F93A" box="[693,842,1763,1789]" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="65" pageNumber="66">Fig. 26HK</figureCitation>
). Dactyli terminating in small corneous claws, with long setae on dorsal margins (setae longer and more numerous in female than in male). Propodi of males each with tuft of several setae on nearly flat mesial face at base of fixed finger. Left propodus of female with prominent tuft of long setae on weakly convex mesial face at base of ventrodistal extremity, right propodus with tuft of some shorter setae on flat mesial face at base of ventrodistal extremity. Left carpus of female with prominent tuft of long setae at ventrodistal angle on mesial face.
</paragraph>
<caption id="DF00F29A3440FFC6FEC6F9A0FCB0F8A8" ID-DOI="http://doi.org/10.5281/zenodo.5064638" ID-Zenodo-Dep="5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="66" pageNumber="67" startId="66.[264,368,1637,1661]" targetBox="[296,1304,475,1605]" targetPageId="66">
<paragraph id="8BC0A2123440FFC6FEC6F9A0FCB0F8A8" blockId="66.[264,1323,1637,1903]" pageId="66" pageNumber="67">
<emphasis id="B90B7E003440FFC6FEC6F9A0FE6BF9BA" bold="true" box="[264,407,1637,1661]" pageId="66" pageNumber="67">FIGURE 26.</emphasis>
<taxonomicName id="4C7FD9913440FFC6FE53F9A3FD48F9BA" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" box="[413,692,1638,1661]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="brachydactylus" status="sp. nov.">
<emphasis id="B90B7E003440FFC6FE53F9A3FD48F9BA" box="[413,692,1638,1661]" italics="true" pageId="66" pageNumber="67">Pagurixus brachydactylus</emphasis>
</taxonomicName>
<emphasis id="B90B7E003440FFC6FD74F9A0FD09F9BA" bold="true" box="[698,757,1637,1661]" pageId="66" pageNumber="67">
<taxonomicNameLabel id="A238C37B3440FFC6FD74F9A0FD09F9BA" box="[698,757,1637,1661]" pageId="66" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN.
</paragraph>
</caption>
<caption id="DF00F29A3441FFC7FEC6F95FFE81F8FB" ID-DOI="http://doi.org/10.5281/zenodo.5064642" ID-Zenodo-Dep="5064642" httpUri="https://zenodo.org/record/5064642/files/figure.png" pageId="67" pageNumber="68" startId="67.[264,367,1690,1714]" targetBox="[310,1250,321,1679]" targetPageId="67">
<paragraph id="8BC0A2123441FFC7FEC6F95FFE81F8FB" blockId="67.[264,1325,1690,1852]" pageId="67" pageNumber="68">
<emphasis id="B90B7E003441FFC7FEC6F95FFE61F975" bold="true" box="[264,413,1690,1714]" pageId="67" pageNumber="68">FIGURE 27.</emphasis>
<taxonomicName id="4C7FD9913441FFC7FE67F95EFD39F975" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" box="[425,709,1691,1714]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="brachydactylus" status="sp. nov.">
<emphasis id="B90B7E003441FFC7FE67F95EFD39F975" box="[425,709,1691,1714]" italics="true" pageId="67" pageNumber="68">Pagurixus brachydactylus</emphasis>
</taxonomicName>
<emphasis id="B90B7E003441FFC7FD1FF95FFCEDF975" bold="true" box="[721,785,1690,1714]" pageId="67" pageNumber="68">
<taxonomicNameLabel id="A238C37B3441FFC7FD1FF95FFCEDF975" box="[721,785,1690,1714]" pageId="67" pageNumber="68" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype male (SL 2.4 mm) from Chichi­jima Island, Ogasawara Islands (KMNH­Iv 000,021). A, right cheliped, chela and carpus, dorsal view; B, entire right cheliped, mesial view; C, same, lateral view; D, left cheliped, chela, dorsal view; E, entire left cheliped, mesial view; F, same, lateral view; G, left cheliped, carpus, dorsal view. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF00F29A3446FFC0FEC6FA58FAE2F9DB" ID-DOI="http://doi.org/10.5281/zenodo.5064644" ID-Zenodo-Dep="5064644" httpUri="https://zenodo.org/record/5064644/files/figure.png" pageId="68" pageNumber="69" startId="68.[264,368,1437,1461]" targetBox="[299,1292,316,1423]" targetPageId="68">
<paragraph id="8BC0A2123446FFC0FEC6FA58FAE2F9DB" blockId="68.[264,1323,1437,1564]" pageId="68" pageNumber="69">
<emphasis id="B90B7E003446FFC0FEC6FA58FE61FA72" bold="true" box="[264,413,1437,1461]" pageId="68" pageNumber="69">FIGURE 28.</emphasis>
<taxonomicName id="4C7FD9913446FFC0FE67FA5BFD39FA72" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" box="[425,709,1438,1461]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="brachydactylus" status="sp. nov.">
<emphasis id="B90B7E003446FFC0FE67FA5BFD39FA72" box="[425,709,1438,1461]" italics="true" pageId="68" pageNumber="69">Pagurixus brachydactylus</emphasis>
</taxonomicName>
<emphasis id="B90B7E003446FFC0FD1FFA58FCEDFA72" bold="true" box="[721,785,1437,1461]" pageId="68" pageNumber="69">
<taxonomicNameLabel id="A238C37B3446FFC0FD1FFA58FCEDFA72" box="[721,785,1437,1461]" pageId="68" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype male (SL 2.4 mm) from Chichi­jima Island, Ogasawara Islands (KMNH­Iv 000,021). A, right second pereopod, lateral view; B, left third pereopod, lateral view; C, right second pereopod, dactylus and distal part of propodus, mesial view; D, left third pereopod, dactylus and distal part of propodus, mesial view. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BC0A2123446FFC0FEF6F98EFEBCF91A" blockId="68.[264,1322,1611,1877]" pageId="68" pageNumber="69">
Anterior lobe of sixth thoracic sternite (
<figureCitation id="1344BE973446FFC0FCC1F98EFC8BF9A2" box="[783,887,1611,1637]" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="68" pageNumber="69">Fig. 26L</figureCitation>
) subrectangular, anterolateral angles rounded, anterior margin with row of moderately short setae. Eighth thoracic sternite (
<figureCitation id="1344BE973446FFC0FB34F9B6FEBDF972" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="68" pageNumber="69">Fig. 26M</figureCitation>
) composed of two unequal, narrowly separated, rounded lobes (right lobe larger than left).
</paragraph>
<paragraph id="8BC0A2123446FFC1FEF6F92EFC73FE99" blockId="68.[264,1322,1611,1877]" lastBlockId="69.[264,1325,284,990]" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
Males with slightly unequal coxae of fifth pereopods (
<figureCitation id="1344BE973446FFC0FC78F92EFBD6F8C2" box="[950,1066,1771,1797]" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="68" pageNumber="69">Fig. 26M</figureCitation>
). Right coxa with tuft of numerous long stiff setae directed toward left and passing across left coxa; ventromesial protrusion absent, but papilla­like very short sexual tube or protrusion of vas deferens occasionally present. Left coxa also with gonopore partially masked by tuft of setae directed ventrally. Female with unpaired left gonopore.
</paragraph>
<paragraph id="8BC0A2123447FFC1FEF6FEA9FE0EFE69" blockId="69.[264,1325,284,990]" pageId="69" pageNumber="70">
Telson (
<figureCitation id="1344BE973447FFC1FE59FEA9FDFFFE41" box="[407,515,364,390]" captionStart="FIGURE 26" captionStartId="66.[264,368,1637,1661]" captionTargetBox="[296,1304,475,1605]" captionTargetId="figure-190@66.[264,1323,451,1613]" captionTargetPageId="66" captionText="FIGURE 26. Pagurixus brachydactylus n. sp. AG, LN, holotype male (SL 2.4 mm) from Chichijima Island, Ogasawara Islands (KMNH­Iv 000,021); HK, paratype female (SL 2.3 mm) from same locality (KMNH­Iv 000,023). A, shield and cephalic appendages, dorsal view; B, left antennule, lateral view; C, ultimate segment and ventral flagellum of antennule, ventral view; D, H, left fourth pereopod, dactylus and propodus, lateral view; E, I, same, mesial view; F, J, right fourth pereopod, dactylus and propodus, lateral view; G, K, same, mesial view; L, sixth thoracic sternite, ventral view; M, coxae of fifth pereopods and eighth thoracic sternite, ventral view; N, telson, dorsal view. Scale bars: 1 mm for A; 0.5 mm for BN." figureDoi="http://doi.org/10.5281/zenodo.5064638" httpUri="https://zenodo.org/record/5064638/files/figure.png" pageId="69" pageNumber="70">Fig. 26N</figureCitation>
) with terminal margins oblique, minutely denticulate or with 4 or 5 very small spinules.
</paragraph>
<paragraph id="8BC0A2123447FFC1FEF6FE79FD6FFD29" blockId="69.[264,1325,284,990]" pageId="69" pageNumber="70">
Colour in life. (
<figureCitation id="1344BE973447FFC1FE38FE79FD6EFE11" box="[502,658,444,470]" captionStart="FIGURE 45" captionStartId="100.[264,367,1516,1540]" captionTargetBox="[285,1310,300,1456]" captionTargetId="figure-133@100.[285,1310,300,1456]" captionTargetPageId="100" captionText="FIGURE 45. A, Pagurixus boninensis (Melin, 1939), ovigerous female (SL 2.7 mm) from Hachijo Island, Izu Islands, Japan (CMNH­ZC 1688), photo taken by J. Okuno; B, Pagurixus carinimanus n. sp., paratype female (SL 1.6 mm) from Okinawa Island, Ryukyu Islands (CBM­ZC 8481), photo taken by Y. Fujita; C, D, Pagurixus brachydactylus n. sp., male (SL 2.6 mm) from Chichi­jima Island, Ogasawara Islands, photo taken by M. Osawa." figureDoi="http://doi.org/10.5281/zenodo.5064702" httpUri="https://zenodo.org/record/5064702/files/figure.png" pageId="69" pageNumber="70">Fig. 45 C, D</figureCitation>
) Shield generally white. Ocular peduncles pale brown, becoming darker distally. Right cheliped generally pale brownish purple; carpus with brownish purple blotches. Left cheliped white in ground color; chela, carpi and meri with brownish purple blotches. Ambulatory legs generally white, with brownish purple markings; dactyli each with blotch on proximal part, extending along midline toward base of claw on lateral face and along dorsal face; propodi each with broad band in proximal half, spotted by white; carpi each with irregular blotch on dorsolateral surface; meri each with 2 blotches on dorsal surface.
</paragraph>
<paragraph id="8BC0A2123447FFC1FEF6FD39FD78FC19" blockId="69.[264,1325,284,990]" pageId="69" pageNumber="70">Colour in preservative. Color mostly faded away, but orange markings on ambulatory legs still evident. Dactyli each with blotch on proximal part, extending along midline toward base of claw on lateral face; propodi white in distal half and orange in proximal half, latter occasionally interrupted by white blotch on dorsal surface; carpi with 2 orange blotches dorsally, connected by median stripe on lateral surface; meri each with orange blotch on dorsal surface distally.</paragraph>
<paragraph id="8BC0A2123447FFC2FEC6FBCAFE92FEF2" blockId="69.[264,1186,1039,1105]" lastBlockId="70.[264,1323,284,870]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
<emphasis id="B90B7E003447FFC1FEC6FBCAFE6BFBEF" box="[264,407,1039,1064]" italics="true" pageId="69" pageNumber="70">Distribution</emphasis>
So far known only from the Ogasawara Islands (
<figureCitation id="1344BE973447FFC1FCBFFBF2FC35FB96" box="[881,969,1079,1105]" captionStart="FIGURE 48" captionStartId="102.[264,367,1798,1822]" captionTargetBox="[269,1278,1196,1754]" captionTargetId="figure-121@102.[264,1289,1158,1780]" captionTargetPageId="102" captionText="FIGURE 48. Geographic distribution of Pagurixus boninensis (), P. festinus (), P. brachydactylus n. sp. (), P. carinimanus n. sp. (), P. paulayi n. sp. (Π), P. pseliophorus n. sp ()., and P. ruber n. sp. ()." figureDoi="http://doi.org/10.5281/zenodo.5064708" httpUri="https://zenodo.org/record/5064708/files/figure.png" pageId="69" pageNumber="70">Fig. 48</figureCitation>
); intertidal to
<specimenCount id="9D79699B3447FFC1FBA0FBF2FB5EFB96" box="[1134,1186,1079,1105]" pageId="69" pageNumber="70" type="generic">
<quantity id="4C870FF73447FFC1FBA0FBF2FB62FB96" box="[1134,1182,1079,1105]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="69" pageNumber="70" unit="m" value="3.0">3 m</quantity>
.
</specimenCount>
<emphasis id="B90B7E003444FFC2FEC6FED9FE92FEF2" box="[264,366,284,309]" italics="true" pageId="70" pageNumber="71">Remarks</emphasis>
</paragraph>
<caption id="DF00F29A3447FFC1FEC6FBBEFB22FB54" box="[264,1246,1147,1171]" pageId="69" pageNumber="70" startId="69.[264,352,1147,1171]">
<paragraph id="8BC0A2123447FFC1FEC6FBBEFB22FB54" blockId="69.[264,1246,1147,1171]" box="[264,1246,1147,1171]" pageId="69" pageNumber="70">
<emphasis id="B90B7E003447FFC1FEC6FBBEFE87FB54" bold="true" box="[264,379,1147,1171]" pageId="69" pageNumber="70">TABLE 2.</emphasis>
Comparison between
<taxonomicName id="4C7FD9913447FFC1FDA4FBB9FC7DFB54" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" box="[618,897,1148,1171]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="brachydactylus" status="sp. nov.">
<emphasis id="B90B7E003447FFC1FDA4FBB9FC7DFB54" box="[618,897,1148,1171]" italics="true" pageId="69" pageNumber="70">Pagurixus brachydactylus</emphasis>
</taxonomicName>
<emphasis id="B90B7E003447FFC1FC46FBBEFC38FB54" bold="true" box="[904,964,1147,1171]" pageId="69" pageNumber="70">
<taxonomicNameLabel id="A238C37B3447FFC1FC46FBBEFC38FB54" box="[904,964,1147,1171]" pageId="69" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C7FD9913447FFC1FC36FBB9FB60FB54" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" box="[1016,1180,1148,1171]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pseliophorus" status="sp. nov.">
<emphasis id="B90B7E003447FFC1FC36FBB9FB60FB54" box="[1016,1180,1148,1171]" italics="true" pageId="69" pageNumber="70">P. pseliophorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238C37B3447FFC1FB6CFBBEFB22FB54" box="[1186,1246,1147,1171]" pageId="69" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BC0A2123447FFC1FDB8FB71FBCAF8A7" pageId="69" pageNumber="70">
<table id="F97F50B23447007BFEDBFB76FAE1F8A7" box="[277,1309,1203,1888]" gridcols="7" gridrows="11" pageId="69" pageNumber="70">
<tr id="354FA0503447007BFEDBFB76FAE1FB0C" box="[277,1309,1203,1227]" gridrow="0" pageId="69" pageNumber="70" rowspan-0="1" rowspan-3="1" rowspan-6="1">
<th id="769EC92C3447007BFDB8FB76FC8AFB0C" box="[630,886,1203,1227]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="69" pageNumber="70">
<taxonomicName id="4C7FD9913447FFC1FDB8FB71FCC8FB0C" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" box="[630,820,1204,1227]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="brachydactylus" status="sp. nov.">
<emphasis id="B90B7E003447FFC1FDB8FB71FCC8FB0C" box="[630,820,1204,1227]" italics="true" pageId="69" pageNumber="70">P. brachydactylus</emphasis>
</taxonomicName>
<emphasis id="B90B7E003447FFC1FCF4FB76FC8AFB0C" bold="true" box="[826,886,1203,1227]" pageId="69" pageNumber="70">
<taxonomicNameLabel id="A238C37B3447FFC1FCF4FB76FC8AFB0C" box="[826,886,1203,1227]" pageId="69" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
<th id="769EC92C3447007BFC19FB76FB19FB0C" box="[983,1253,1203,1227]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="69" pageNumber="70">
<taxonomicName id="4C7FD9913447FFC1FC19FB71FB87FB0C" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" box="[983,1147,1204,1227]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pseliophorus" status="sp. nov.">
<emphasis id="B90B7E003447FFC1FC19FB71FB87FB0C" box="[983,1147,1204,1227]" italics="true" pageId="69" pageNumber="70">P. pseliophorus</emphasis>
</taxonomicName>
<emphasis id="B90B7E003447FFC1FB4CFB76FB42FB0C" bold="true" box="[1154,1214,1203,1227]" pageId="69" pageNumber="70">
<taxonomicNameLabel id="A238C37B3447FFC1FB4CFB76FB42FB0C" box="[1154,1214,1203,1227]" pageId="69" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="354FA0503447007BFEDBFB1BFAE1FB32" box="[277,1309,1246,1269]" gridrow="1" pageId="69" pageNumber="70" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="769EC92C3447007BFEDBFB1BFDA0FB32" box="[277,604,1246,1269]" gridcol="0" gridrow="1" pageId="69" pageNumber="70">Shield length/width</th>
<td id="769EC92C3447007BFDB8FB1BFCF8FB32" box="[630,772,1246,1269]" gridcol="1" gridrow="1" pageId="69" pageNumber="70">1.01.1</td>
<td id="769EC92C3447007BFC19FB1BFB5AFB32" box="[983,1190,1246,1269]" gridcol="4" gridrow="1" pageId="69" pageNumber="70">1.11.3</td>
</tr>
<tr id="354FA0503447007BFEDBFACCFAE1FAE7" box="[277,1309,1289,1312]" gridrow="2" pageId="69" pageNumber="70">
<th id="769EC92C3447007BFEDBFACCFDA0FAE7" box="[277,604,1289,1312]" gridcol="0" gridrow="2" pageId="69" pageNumber="70">Setal rows on ventral surface</th>
<td id="769EC92C3447007BFDB8FACCFC8AFAE7" box="[630,886,1289,1312]" colspan="2" colspanRight="1" gridcol="1" gridrow="2" pageId="69" pageNumber="70">mixture of numerous</td>
<td id="769EC92C3447007BFC43FACCFC41FAE7" box="[909,957,1289,1312]" gridcol="3" gridrow="2" pageId="69" pageNumber="70">very</td>
<td id="769EC92C3447007BFC19FACCFB5AFAE7" box="[983,1190,1289,1312]" gridcol="4" gridrow="2" pageId="69" pageNumber="70">irregular rows of</td>
<td id="769EC92C3447007BFB78FACCFAE1FAE7" box="[1206,1309,1289,1312]" colspan="2" colspanRight="1" gridcol="5" gridrow="2" pageId="69" pageNumber="70">relatively</td>
</tr>
<tr id="354FA0503447007BFEDBFAE9FAE1FAA2" box="[277,1309,1324,1381]" gridrow="3" pageId="69" pageNumber="70" rowspan-5="1" rowspan-6="1">
<th id="769EC92C3447007BFEDBFAE9FDA0FAA2" box="[277,604,1324,1381]" gridcol="0" gridrow="3" pageId="69" pageNumber="70">of ultimate segment of antennular peduncle</th>
<td id="769EC92C3447007BFDB8FAE9FC41FAA2" box="[630,957,1324,1381]" colspan="3" colspanRight="2" gridcol="1" gridrow="3" pageId="69" pageNumber="70">short setae and tufts of longer setae</td>
<td id="769EC92C3447007BFC19FAE9FB5AFAA2" box="[983,1190,1324,1381]" gridcol="4" gridrow="3" pageId="69" pageNumber="70">long setae</td>
</tr>
<tr id="354FA0503447007BFEDBFABCFAE1FA74" box="[277,1309,1401,1459]" gridrow="4" pageId="69" pageNumber="70" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="769EC92C3447007BFEDBFABCFDA0FA74" box="[277,604,1401,1459]" gridcol="0" gridrow="4" pageId="69" pageNumber="70">Dorsolateral margin of palm of male right cheliped</th>
<td id="769EC92C3447007BFDB8FABCFCF8FA74" box="[630,772,1401,1459]" gridcol="1" gridrow="4" pageId="69" pageNumber="70">not delimited</td>
<td id="769EC92C3447007BFC19FABCFB5AFA74" box="[983,1190,1401,1459]" gridcol="4" gridrow="4" pageId="69" pageNumber="70">faintly delimited</td>
</tr>
<tr id="354FA0503447007BFEDBFA03FAE1F9C7" box="[277,1309,1478,1536]" gridrow="5" pageId="69" pageNumber="70" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="769EC92C3447007BFEDBFA03FDA0F9C7" box="[277,604,1478,1536]" gridcol="0" gridrow="5" pageId="69" pageNumber="70">Left cheliped, carpus length/ width</th>
<td id="769EC92C3447007BFDB8FA03FCF8F9C7" box="[630,772,1478,1536]" gridcol="1" gridrow="5" pageId="69" pageNumber="70">2.22.4</td>
<td id="769EC92C3447007BFC19FA03FB5AF9C7" box="[983,1190,1478,1536]" gridcol="4" gridrow="5" pageId="69" pageNumber="70">2.52.8</td>
</tr>
<tr id="354FA0503447007BFEDBF9D1FAE1F9B7" box="[277,1309,1556,1648]" gridrow="6" pageId="69" pageNumber="70">
<th id="769EC92C3447007BFEDBF9D1FDA0F9B7" box="[277,604,1556,1648]" gridcol="0" gridrow="6" pageId="69" pageNumber="70">Dorsolateral and dorsomesial margins of carpus of left cheliped</th>
<td id="769EC92C3447007BFDB8F9D1FC41F9B7" box="[630,957,1556,1648]" colspan="3" colspanRight="2" gridcol="1" gridrow="6" pageId="69" pageNumber="70">with row of small spines only on dorsomesial margin, dorsolateral margin unarmed</td>
<td id="769EC92C3447007BFC19F9D1FB5AF9B7" box="[983,1190,1556,1648]" gridcol="4" gridrow="6" pageId="69" pageNumber="70">each with row of protuberances</td>
<td id="769EC92C3447007BFB78F9D1FB19F9B7" box="[1206,1253,1556,1648]" gridcol="5" gridrow="6" pageId="69" pageNumber="70">very</td>
<td id="769EC92C3447007BFB3AF9D1FAE1F9B7" box="[1268,1309,1556,1648]" gridcol="6" gridrow="6" pageId="69" pageNumber="70">low</td>
</tr>
<tr id="354FA0503447007BFEDBF941FAE1F97A" box="[277,1309,1668,1725]" gridrow="7" pageId="69" pageNumber="70" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="769EC92C3447007BFEDBF941FDA0F97A" box="[277,604,1668,1725]" gridcol="0" gridrow="7" pageId="69" pageNumber="70">Ambulatory legs, dactyli length/height</th>
<td id="769EC92C3447007BFDB8F941FCF8F97A" box="[630,772,1668,1725]" gridcol="1" gridrow="7" pageId="69" pageNumber="70">3.03.7</td>
<td id="769EC92C3447007BFC19F941FB5AF97A" box="[983,1190,1668,1725]" gridcol="4" gridrow="7" pageId="69" pageNumber="70">3.84.1</td>
</tr>
<tr id="354FA0503447007BFEDBF914FAE1F92F" box="[277,1309,1745,1768]" gridrow="8" pageId="69" pageNumber="70" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="769EC92C3447007BFEDBF914FDA0F92F" box="[277,604,1745,1768]" gridcol="0" gridrow="8" pageId="69" pageNumber="70">Eighth thoracic sternite</th>
<td id="769EC92C3447007BFDB8F914FC8AF92F" box="[630,886,1745,1768]" colspan="2" colspanRight="1" gridcol="1" gridrow="8" pageId="69" pageNumber="70">2 unequal lobes</td>
<td id="769EC92C3447007BFC19F914FB5AF92F" box="[983,1190,1745,1768]" gridcol="4" gridrow="8" pageId="69" pageNumber="70">2 subequal lobes</td>
</tr>
<tr id="354FA0503447007BFEDBF939FAE1F8F2" box="[277,1309,1788,1845]" gridrow="9" pageId="69" pageNumber="70">
<th id="769EC92C3447007BFEDBF939FDA0F8F2" box="[277,604,1788,1845]" gridcol="0" gridrow="9" pageId="69" pageNumber="70">Right coxae of fifth pereopod in male</th>
<td id="769EC92C3447007BFDB8F939FCF8F8F2" box="[630,772,1788,1845]" gridcol="1" gridrow="9" pageId="69" pageNumber="70">without protrusion</td>
<td id="769EC92C3447007BFCFDF939FC41F8F2" box="[819,957,1788,1845]" colspan="2" colspanRight="1" gridcol="2" gridrow="9" pageId="69" pageNumber="70">ventromesial</td>
<td id="769EC92C3447007BFC19F939FAE1F8F2" box="[983,1309,1788,1845]" colspan="3" colspanRight="2" gridcol="4" gridrow="9" pageId="69" pageNumber="70">with prominent ventromesial protrusion</td>
</tr>
<tr id="354FA0503447007BFEDBF88CFAE1F8A7" box="[277,1309,1865,1888]" gridrow="10" pageId="69" pageNumber="70" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="769EC92C3447007BFEDBF88CFDA0F8A7" box="[277,604,1865,1888]" gridcol="0" gridrow="10" pageId="69" pageNumber="70">Coloration in life</th>
<td id="769EC92C3447007BFDB8F88CFCF8F8A7" box="[630,772,1865,1888]" gridcol="1" gridrow="10" pageId="69" pageNumber="70">Fig. 45C, D</td>
<td id="769EC92C3447007BFC19F88CFB5AF8A7" box="[983,1190,1865,1888]" gridcol="4" gridrow="10" pageId="69" pageNumber="70">Fig. 46A</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC0A2123444FFC2FEF6FE81FE5AFD29" blockId="70.[264,1323,284,870]" pageId="70" pageNumber="71">
Among the seven species of the subgroup B of the
<taxonomicName id="4C7FD9913444FFC2FC30FE81FB58FE9A" baseAuthorityName="Melin" baseAuthorityYear="1939" box="[1022,1188,324,349]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="boninensis">
<emphasis id="B90B7E003444FFC2FC30FE81FB58FE9A" box="[1022,1188,324,349]" italics="true" pageId="70" pageNumber="71">P. boninensis</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C7FD9913444FFC2FADBFE81FE47FE42" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="brachydactylus" status="sp. nov.">
<emphasis id="B90B7E003444FFC2FADBFE81FE47FE42" italics="true" pageId="70" pageNumber="71">P. brachydactylus</emphasis>
</taxonomicName>
<emphasis id="B90B7E003444FFC2FE08FEAEFDF1FE42" bold="true" box="[454,525,363,389]" pageId="70" pageNumber="71">
<taxonomicNameLabel id="A238C37B3444FFC2FE08FEAEFDF1FE42" box="[454,525,363,389]" pageId="70" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
most closely resembles
<taxonomicName id="4C7FD9913444FFC2FCF4FEA9FC0DFE42" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" box="[826,1009,364,389]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="pseliophorus" status="sp. nov.">
<emphasis id="B90B7E003444FFC2FCF4FEA9FC0DFE42" box="[826,1009,364,389]" italics="true" pageId="70" pageNumber="71">P. pseliophorus</emphasis>
</taxonomicName>
<emphasis id="B90B7E003444FFC2FC33FEAEFBBFFE42" bold="true" box="[1021,1091,363,389]" pageId="70" pageNumber="71">
<taxonomicNameLabel id="A238C37B3444FFC2FC33FEAEFBBFFE42" box="[1021,1091,363,389]" pageId="70" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
These two species share short and stout dactyli of the ambulatory legs and the absence of conspicuous spines on the dorsal surface of the carpus of the male right cheliped. Morphological differences between the two species are summarized in
<tableCitation id="C6FD97A93444FFC2FCDBFE21FC8DFE39" box="[789,881,484,510]" captionStart="TABLE 2" captionStartId="69.[264,352,1147,1171]" captionText="TABLE 2. Comparison between Pagurixus brachydactylus n. sp. and P. pseliophorus n. sp." pageId="70" pageNumber="71">Table 2</tableCitation>
. The shape of the shield, structure of the setal series on the ultimate segment of the antennular peduncle, armament of the carpus of the left cheliped, and structure of the coxa of right fifth pereopod in males are useful for species discrimination. The coloration in life is also different between the two species. For example, the propodi of the ambulatory legs are white in the distal half and purplish brown in the proximal half in
<taxonomicName id="4C7FD9913444FFC2FDD5FD69FD10FD02" authorityName="Komai &amp; Osawa" authorityYear="2006" box="[539,748,684,709]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="brachydactylus">
<emphasis id="B90B7E003444FFC2FDD5FD69FD10FD02" box="[539,748,684,709]" italics="true" pageId="70" pageNumber="71">P. brachydactylus</emphasis>
</taxonomicName>
, while they have middle dark brown bands in
<taxonomicName id="4C7FD9913444FFC2FADBFD69FE63FD2A" authorityName="Komai &amp; Osawa" authorityYear="2006" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="pseliophorus">
<emphasis id="B90B7E003444FFC2FADBFD69FE63FD2A" italics="true" pageId="70" pageNumber="71">P. pseliophorus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC0A2123444FFC2FEF6FD39FDF9FCA1" blockId="70.[264,1323,284,870]" pageId="70" pageNumber="71">
As mentioned before, reexamination of the specimens from the Ogasawara Islands, reported by
<bibRefCitation id="EFEEDFE33444FFC2FE59FCE1FDC3FCF9" author="Ooishi, S." box="[407,575,804,830]" pageId="70" pageNumber="71" pagination="75 - 104" refId="ref42933" refString="Ooishi, S. (1970) Marine invertebrates fauna of the Ogasawara and Volcano Islands collected by S. Ooishi, Y. Tomida, K. Izawa and S. Manabe. In: Report on the Marine Biological Expedition to the Ogasawara (Bonin) Islands, 1968, pp. 75 - 104." type="journal article" year="1970">Ooishi (1970)</bibRefCitation>
, has shown that they do not represent
<taxonomicName id="4C7FD9913444FFC2FBC8FCE1FB5DFCFA" baseAuthorityName="Melin" baseAuthorityYear="1939" box="[1030,1185,804,829]" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" sensu="stricto" species="boninensis">
<emphasis id="B90B7E003444FFC2FBC8FCE1FB5DFCFA" box="[1030,1185,804,829]" italics="true" pageId="70" pageNumber="71">P. boninensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238C37B3444FFC2FB64FCE1FB24FCF9" box="[1194,1240,804,830]" pageId="70" pageNumber="71" sensu="stricto">s. s.</taxonomicNameLabel>
, but
<taxonomicName id="4C7FD9913444FFC2FADBFCE1FE47FCA2" authority="Komai &amp; Osawa, 2006" authorityName="Komai &amp; Osawa" authorityYear="2006" class="Malacostraca" family="Paguridae" genus="Pagurixus" kingdom="Animalia" order="Decapoda" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="brachydactylus" status="sp. nov.">
<emphasis id="B90B7E003444FFC2FADBFCE1FE47FCA2" italics="true" pageId="70" pageNumber="71">P. brachydactylus</emphasis>
</taxonomicName>
<emphasis id="B90B7E003444FFC2FE0DFC89FDF9FCA1" bold="true" box="[451,517,844,870]" pageId="70" pageNumber="71">
<taxonomicNameLabel id="A238C37B3444FFC2FE0DFC89FDF9FCA1" box="[451,517,844,870]" pageId="70" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BC0A2123444FFC2FEC6FC59FE79FC72" blockId="70.[264,1322,924,1030]" box="[264,389,924,949]" pageId="70" pageNumber="71">
<emphasis id="B90B7E003444FFC2FEC6FC59FE79FC72" box="[264,389,924,949]" italics="true" pageId="70" pageNumber="71">Etymology</emphasis>
</paragraph>
<paragraph id="8BC0A2123444FFC2FEF6FC01FD0BFBC1" blockId="70.[264,1322,924,1030]" pageId="70" pageNumber="71">
From the Greek
<emphasis id="B90B7E003444FFC2FE34FC01FDAAFC1A" box="[506,598,964,989]" italics="true" pageId="70" pageNumber="71">brachys</emphasis>
, meaning short, and
<emphasis id="B90B7E003444FFC2FCAFFC01FC3EFC1A" box="[865,962,964,989]" italics="true" pageId="70" pageNumber="71">dactylos</emphasis>
, meaning “claw”, in reference to the short dactyli of the ambulatory legs.
</paragraph>
</subSubSection>
</treatment>
</document>