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<document id="C291E8D23D9A9B557D352A03D44F158A" ID-DOI="10.5281/zenodo.5360449" ID-ISSN="0024-4082" ID-Zenodo-Dep="5360449" ID-ZooBank="591555C6-EF37-495B-A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630232463663" checkinUser="marcus" docAuthor="Rajabizadeh, Mahdi, Nagy, Zoltán T., Adriaens, Dominique, Avci, Aziz, Masroor, Rafaqat, Schmidtler, Josef, Nazarov, Roman, Esmaeili, Hamid Reza &amp; Christiaens, Joachim" docDate="2016" docId="03D39A7AFFC19F37FF5DA46812D06BC7" docLanguage="en" docName="zoj12342.pdf" docOrigin="Zoological Journal of the Linnean Society 176" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Contia persica Boulenger 1893" docType="treatment" docVersion="3" lastPageNumber="900" masterDocId="FFEAE202FFD79F21FFA5A128115C6F53" masterDocTitle="Alpine-Himalayan orogeny drove correlated morphological, molecular, and ecological diversification in the Persian dwarf snake (Squamata: Serpentes: Eirenis persicus)" masterLastPageNumber="913" masterPageNumber="878" pageNumber="900" updateTime="1699193229270" updateUser="plazi">
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<mods:title id="5C3B930151A166745A9720108BA05974">Alpine-Himalayan orogeny drove correlated morphological, molecular, and ecological diversification in the Persian dwarf snake (Squamata: Serpentes: Eirenis persicus)</mods:title>
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<mods:namePart id="BC555DA02A51EB04D73B1D433F651EBA">Rajabizadeh, Mahdi</mods:namePart>
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<mods:namePart id="33114EF82916C05E05715197B5D7B313">Nagy, Zoltán T.</mods:namePart>
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<mods:namePart id="6ED903CF82AD6C6F44856C45E9EE5F5A">Adriaens, Dominique</mods:namePart>
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<mods:namePart id="99E1EBF5A380F038106FCEF9D72F9BC0">Avci, Aziz</mods:namePart>
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<mods:namePart id="E6661AF9F77054CED7AB77A4B41C2AEC">Masroor, Rafaqat</mods:namePart>
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<mods:namePart id="BFE159D65EA2E15CC61A3B50A99C6E1E">Schmidtler, Josef</mods:namePart>
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<mods:namePart id="2144F9ABCF427A2604D3D1ED87637251">Nazarov, Roman</mods:namePart>
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<mods:title id="C945A89F73CEF42B1D14D6C2EE43E673">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="800B3E7B1AEA899E003B64BCB8AC27E8">2016</mods:date>
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<treatment id="03D39A7AFFC19F37FF5DA46812D06BC7" LSID="urn:lsid:plazi:treatment:03D39A7AFFC19F37FF5DA46812D06BC7" httpUri="http://treatment.plazi.org/id/03D39A7AFFC19F37FF5DA46812D06BC7" lastPageNumber="900" pageId="22" pageNumber="900">
<subSubSection id="C36078E7FFC19F37FF5DA468136E6A26" pageId="22" pageNumber="900" type="nomenclature">
<paragraph id="8BC52B6CFFC19F37FF5DA468136E6A26" blockId="22.[248,699,1344,1397]" pageId="22" pageNumber="900">
<heading id="D08D9C00FFC19F37FF5DA468136E6A26" centered="true" fontSize="8" level="2" pageId="22" pageNumber="900" reason="8">
<emphasis id="B90EF77EFFC19F37FF5DA468134A6A07" box="[248,534,1344,1365]" italics="true" pageId="22" pageNumber="900">
<taxonomicName id="4C7A50EFFFC19F37FF5DA46810ED6A07" authorityName="Boulenger" authorityYear="1893" box="[248,433,1344,1365]" class="Reptilia" family="Dipsadidae" genus="Contia" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="882" phylum="Chordata" rank="species" species="persica">CONTIA PERSICA</taxonomicName>
PERSICA
</emphasis>
<bibRefCitation id="EFEB569DFFC19F37FDBAA468136E6A26" author="Terent ev PV &amp; Chernov SA" pageId="22" pageNumber="900" refId="ref17793" refString="Terent ev PV, Chernov SA. 1965. Key to amphibian and reptiles. Jerusalem: Israel Program for Scientific translation [English translation of Terent ev &amp; Chernov, 1949]." type="book" year="1965">TERENT ÉV &amp; CHERNOV, 1965</bibRefCitation>
</heading>
</paragraph>
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<subSubSection id="C36078E7FFC19F37FF03A4AE101A6AAB" pageId="22" pageNumber="900" type="materials_examined">
<paragraph id="8BC52B6CFFC19F37FF03A4AE10556AC8" blockId="22.[166,780,1414,1528]" box="[166,265,1414,1435]" pageId="22" pageNumber="900">
<emphasis id="B90EF77EFFC19F37FF03A4AE10556AC8" box="[166,265,1414,1435]" italics="true" pageId="22" pageNumber="900">
<typeStatus id="54C195CEFFC19F37FF03A4AE10556AC8" box="[166,265,1414,1435]" pageId="22" pageNumber="900" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC52B6CFFC19F37FF03A48D101A6AAB" blockId="22.[166,780,1414,1528]" pageId="22" pageNumber="900">
<collectionCode id="ED6BB3A9FFC19F37FF03A48D11936AE9" box="[166,207,1445,1466]" pageId="22" pageNumber="900">ZSI</collectionCode>
4828 (collection of the Zoological Survey of
<collectingCountry id="F36D6BFCFFC19F37FD6CA48D12546AE8" box="[713,776,1445,1467]" name="India" pageId="22" pageNumber="900">India</collectingCountry>
, Calcutta,
<collectingCountry id="F36D6BFCFFC19F37FEB8A4EB10076A8A" box="[285,347,1475,1497]" name="India" pageId="22" pageNumber="900">India</collectingCountry>
) (
<bibRefCitation id="EFEB569DFFC19F37FED2A4EB13496A8A" author="Sclater WL" box="[375,533,1475,1497]" pageId="22" pageNumber="900" refId="ref17495" refString="Sclater WL. 1891. List of snakes in the Indian Museum. Calcutta: Baptist Mission Press. X + 79 pp." type="book" year="1891">Sclater, 1891</bibRefCitation>
;
<bibRefCitation id="EFEB569DFFC19F37FD83A4EC10666AAB" author="Das I &amp; Dattagupta B &amp; Gayen NC" pageId="22" pageNumber="900" pagination="121 - 172" refId="ref15527" refString="Das I, Dattagupta B, Gayen NC. 1998. History and catalogue of reptile types in the collection of the Zoological Survey of India. Journal of South Asian Natural History 3: 121 - 172." type="journal article" year="1998">Das, Dattagupta &amp; Gayen, 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36078E7FFC19F37FF03A70B12D06BC7" pageId="22" pageNumber="900" type="description">
<paragraph id="8BC52B6CFFC19F37FF03A70B1073696B" blockId="22.[166,781,1571,1684]" box="[166,303,1571,1592]" pageId="22" pageNumber="900">
<taxonomicName id="4C7A50EFFFC19F37FF03A70B1073696B" box="[166,303,1571,1592]" class="Insecta" family="Lycaenidae" genus="Terra" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="900" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="B90EF77EFFC19F37FF03A70B1073696B" box="[166,303,1571,1592]" italics="true" pageId="22" pageNumber="900">Terra typica</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BC52B6CFFC19F37FF03A769135E69C7" blockId="22.[166,781,1571,1684]" pageId="22" pageNumber="900">
<materialsCitation id="3B122131FFC19F37FF0FA76910A369C7" country="Iran" latitude="28.920277" location="Bushehr" longLatPrecision="20" longitude="50.83583" pageId="22" pageNumber="900" specimenCount="1" stateProvince="Bushehr">
Bushire, Persia [
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city (
<geoCoordinate id="EE4E4DABFFC19F37FE5AA76913276904" box="[511,635,1601,1623]" degrees="28" direction="north" minutes="55" orientation="latitude" pageId="22" pageNumber="900" precision="15" seconds="13" value="28.920277">28°5513″N</geoCoordinate>
,
<geoCoordinate id="EE4E4DABFFC19F37FD22A769125C6904" box="[647,768,1601,1623]" degrees="50" direction="east" minutes="50" orientation="longitude" pageId="22" pageNumber="900" precision="15" seconds="09" value="50.83583">50°5009″E</geoCoordinate>
),
<collectingRegion id="49BEE58EFFC19F37FF03A748102E6925" box="[166,370,1632,1654]" country="Iran" name="Bushehr" pageId="22" pageNumber="900">Bushehr province</collectingRegion>
, south-western
<collectingCountry id="F36D6BFCFFC19F37FD83A74813066926" box="[550,602,1632,1653]" name="Iran" pageId="22" pageNumber="900">Iran</collectingCountry>
, the locality indicated by circle
<quantity id="4C828689FFC19F37FEC8A75610F569C7" box="[365,425,1662,1684]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" pageId="22" pageNumber="900" unit="in" value="29.0">29 in</quantity>
<figureCitation id="134137E9FFC19F37FE16A75610AB69C7" box="[435,503,1662,1684]" captionStart="Figure 2" captionStartId="4.[46,125,1401,1422]" captionTargetBox="[104,1695,472,1368]" captionTargetId="figure-0@4.[97,1697,465,1373]" captionTargetPageId="4" captionText="Figure 2. All available distribution records of the Eirenis persicus species group. Circles indicate the E. persicus specimens, with the persicus morph bearing unicoloured dorsal scales; squires indicate specimens from eastern Iran, southern Turkmenistan, and southern and western Pakistan, having both persicus morph that base of their anterior dorsal scales are darker, and walteri morph; stars indicate specimens referred to Eirenis mcmahoni (Wall, 1911); plus symbols indicate specimens in north-eastern Pakistan, having both persicus morphs that base of their anterior dorsal scales are darker, and with walteri morph; asterisks indicate specimens with the novum pattern; triangles indicate specimens of the nigrofasciatus morph. Circle 29 indicates the type locality of Cyclophis persicus Anderson, 1872; squire 2 indicates the type locality of Pseudocyclophis walteri Boettger, 1888; squire 7 indicates the type locality of Contia zebrina Wall, 1923; triangle 6 indicates the type locality of Contia persica var. nigrofasciata Nikolsky, 1907; star 3 indicates the type locality of Contia angusticeps Boulenger, 1894; plus symbols 25 indicate the type series localities of Contia mcmahoni Wall, 1911. For more details, see Appendix 1." figureDoi="http://doi.org/10.5281/zenodo.5360455" httpUri="https://zenodo.org/record/5360455/files/figure.png" pageId="22" pageNumber="900">Fig. 2</figureCitation>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC52B6CFFC19F37FF03A797107B6987" blockId="22.[166,781,1727,1902]" box="[166,295,1727,1748]" pageId="22" pageNumber="900">
<emphasis id="B90EF77EFFC19F37FF03A797107B6987" box="[166,295,1727,1748]" italics="true" pageId="22" pageNumber="900">Description</emphasis>
</paragraph>
<paragraph id="8BC52B6CFFC19F37FF03A7F614F86EC1" blockId="22.[166,781,1727,1902]" lastBlockId="22.[828,1444,197,892]" pageId="22" pageNumber="900">
<emphasis id="B90EF77EFFC19F37FF03A7F611E669A0" box="[166,186,1758,1779]" italics="true" pageId="22" pageNumber="900">N</emphasis>
= 9;
<specimenCount id="9D7CE0E5FFC19F37FF54A7F5104B69A1" box="[241,279,1757,1779]" pageId="22" pageNumber="900" type="male">5♂</specimenCount>
,
<specimenCount id="9D7CE0E5FFC19F37FE83A7F6101B69A1" box="[294,327,1758,1779]" pageId="22" pageNumber="900" type="female">4♀</specimenCount>
. 190203 (
<emphasis id="B90EF77EFFC19F37FE6FA7F5108A69A1" box="[458,470,1757,1778]" italics="true" pageId="22" pageNumber="900">x</emphasis>
= 195.6) ventrals in
<specimenCount id="9D7CE0E5FFC19F37FD67A7C8138769A1" box="[706,731,1760,1778]" pageId="22" pageNumber="900" type="male"></specimenCount>
and 198214 (
<emphasis id="B90EF77EFFC19F37FEBCA7D410796842" box="[281,293,1788,1809]" italics="true" pageId="22" pageNumber="900">x</emphasis>
= 205.0) in
<specimenCount id="9D7CE0E5FFC19F37FE08A7D7109D6842" box="[429,449,1791,1809]" pageId="22" pageNumber="900" type="female"></specimenCount>
; 6483 subcaudals (
<emphasis id="B90EF77EFFC19F37FD09A7D413E46842" box="[684,696,1788,1809]" italics="true" pageId="22" pageNumber="900">x</emphasis>
= 75.0) in
<specimenCount id="9D7CE0E5FFC19F37FF63A6351183687C" box="[198,223,1821,1839]" pageId="22" pageNumber="900" type="male"></specimenCount>
, 5477 (
<emphasis id="B90EF77EFFC19F37FEEEA633100B6863" box="[331,343,1819,1840]" italics="true" pageId="22" pageNumber="900">x</emphasis>
= 66.25) in
<specimenCount id="9D7CE0E5FFC19F37FE47A63510AB687C" box="[482,503,1821,1839]" pageId="22" pageNumber="900" type="female"></specimenCount>
; ratio of the tail length over total length 21.9329.09 (
<emphasis id="B90EF77EFFC19F37FDA2A611134F681D" box="[519,531,1849,1870]" italics="true" pageId="22" pageNumber="900">x</emphasis>
= 23.80) in
<specimenCount id="9D7CE0E5FFC19F37FD3EA61413EF681D" box="[667,691,1852,1870]" pageId="22" pageNumber="900" type="male"></specimenCount>
, 18.92 22.60 (
<emphasis id="B90EF77EFFC19F37FF5EA670105B683E" box="[251,263,1880,1901]" italics="true" pageId="22" pageNumber="900">x</emphasis>
= 20.42) in
<specimenCount id="9D7CE0E5FFC19F37FE37A67310FB683E" box="[402,423,1883,1901]" pageId="22" pageNumber="900" type="female"></specimenCount>
. 15/15/15 dorsal scale rows on the anterior, mid, and posterior portion of the body; usually seven (88.9%), rarely eight, supralabials and usually eight (88.9%), rarely seven, infralabials on both sides of the head; posterior chin shields attached or separated by a single scale; no loreal scale; usually ten (66.7%), rarely eight or nine, scales between the last infralabials and nine to 12 scales behind the parietals.
</paragraph>
<paragraph id="8BC52B6CFFC19F37FCF0A0B315CE6C2F" blockId="22.[828,1444,197,892]" pageId="22" pageNumber="900">
Dorsal background colour yellowish, light brown or pale olive, without dorsal pattern; usually three dark transverse markings on the head and the nape, one on the posterior snout, another one on the third/ fourth anterior parietals, and the last one on the nape, separated from each other by a narrow light band; alternatively the dorsal head (except at the tip of the snout) and the nape are completely blackish; lateral and ventral head may be uniform cream or with dark markings on snout, upper temporal, below or around eye, and adjacent to the supralabials and anterior to the infralabials; lower side of body unicolour cream; the napes band is five to eight scales wide dorsally, shrinks over lateral neck, interrupted over the ventral or preventral scales or rarely (only in
<specimenCount id="9D7CE0E5FFC19F37FB5AA26014C06C0E" box="[1279,1436,839,861]" pageId="22" pageNumber="900" type="generic">one specimen</specimenCount>
) connected ventrally (
<figureCitation id="134137E9FFC19F37FB95A24E15D96C2F" box="[1072,1157,870,892]" captionStart="Figure 1" captionStartId="2.[166,245,1333,1352]" captionTargetBox="[325,1285,195,1303]" captionTargetId="figure-170@2.[325,1285,195,1303]" captionTargetPageId="2" captionText="Figure 1. Different morphs of the Eirenis persicus species group: A, persicus morph from Dasht-e Arjan, Fars province, south-western Iran (photo by F. Hidary); B, walteri morph from Dehbakri, Kerman province, south-eastern Iran (photo by R. Nazarov); C, new morph from Sisakht, Yasuj province, central Zagros mountains, Iran (photo by H. Esmaeili); D, persicus morph from Kafir Kot, Khyber Pakhtunkhwa Province, Pakistan (photo by R. Masroor); E, nigrofasciatus morph from Dezful, Khuzestan province, south-western Iran (photo by F. Hidary)." figureDoi="http://doi.org/10.5281/zenodo.5360453" httpUri="https://zenodo.org/record/5360453/files/figure.png" pageId="22" pageNumber="900">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="8BC52B6CFFC19F37FC99A28F129B6CEF" blockId="22.[828,1443,935,1172]" box="[828,967,935,956]" pageId="22" pageNumber="900">
<emphasis id="B90EF77EFFC19F37FC99A28F129B6CEF" box="[828,967,935,956]" italics="true" pageId="22" pageNumber="900">Distribution</emphasis>
</paragraph>
<paragraph id="8BC52B6CFFC19F37FC99A2EE12D06BC7" blockId="22.[828,1443,935,1172]" pageId="22" pageNumber="900">
<materialsCitation id="3B122131FFC19F37FC99A2EE12D46BC7" country="Iran" location="western Zagros" municipality="southern Zagros" pageId="22" pageNumber="900" specimenCount="1" stateProvince="Fars">
Occurs in the
<collectingMunicipality id="6BA1B116FFC19F37FC47A2EE15C26C88" box="[994,1182,966,988]" pageId="22" pageNumber="900">southern Zagros</collectingMunicipality>
and in adjacent areas to the
<location id="8EA57DB7FFC19F37FC2EA2CD15606CA9" LSID="urn:lsid:plazi:treatment:03D39A7AFFC19F37FF5DA46812D06BC7:8EA57DB7FFC19F37FC2EA2CD15606CA9" box="[907,1084,997,1018]" country="Iran" municipality="southern Zagros" name="western Zagros" pageId="22" pageNumber="900" stateProvince="Fars">western Zagros</location>
foothills and the central plain of
<collectingCountry id="F36D6BFCFFC19F37FCFEA52C12D36B4A" box="[859,911,1028,1049]" name="Iran" pageId="22" pageNumber="900">Iran</collectingCountry>
, from the south the species range extends to the Persian Gulf coastline. The species has also been reported from
<collectingRegion id="49BEE58EFFC19F37FC45A569154A6B05" box="[992,1046,1089,1110]" country="Iran" name="Fars" pageId="22" pageNumber="900">Fars</collectingRegion>
,
<collectingRegion id="49BEE58EFFC19F37FB86A56815206B05" box="[1059,1148,1088,1110]" country="Iran" name="Esfahan" pageId="22" pageNumber="900">Isfahan</collectingRegion>
,
<collectingRegion id="49BEE58EFFC19F37FB22A568145F6B05" box="[1159,1283,1088,1110]" country="Iran" name="Khuzestan" pageId="22" pageNumber="900">Khuzestan</collectingRegion>
, and
<collectingRegion id="49BEE58EFFC19F37FAE7A56814FF6B05" box="[1346,1443,1088,1110]" country="Iran" name="Bushehr" pageId="22" pageNumber="900">Bushehr</collectingRegion>
provinces (localities indicated by circles
<quantity id="4C828689FFC19F37FAB8A57714FF6B26" box="[1309,1443,1119,1141]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" metricValueMax="7.874" metricValueMin="5.334" pageId="22" pageNumber="900" unit="in" value="26.0" valueMax="31.0" valueMin="21.0">21 to 31 in</quantity>
<figureCitation id="134137E9FFC19F37FC99A55612236BC7" box="[828,895,1150,1172]" captionStart="Figure 2" captionStartId="4.[46,125,1401,1422]" captionTargetBox="[104,1695,472,1368]" captionTargetId="figure-0@4.[97,1697,465,1373]" captionTargetPageId="4" captionText="Figure 2. All available distribution records of the Eirenis persicus species group. Circles indicate the E. persicus specimens, with the persicus morph bearing unicoloured dorsal scales; squires indicate specimens from eastern Iran, southern Turkmenistan, and southern and western Pakistan, having both persicus morph that base of their anterior dorsal scales are darker, and walteri morph; stars indicate specimens referred to Eirenis mcmahoni (Wall, 1911); plus symbols indicate specimens in north-eastern Pakistan, having both persicus morphs that base of their anterior dorsal scales are darker, and with walteri morph; asterisks indicate specimens with the novum pattern; triangles indicate specimens of the nigrofasciatus morph. Circle 29 indicates the type locality of Cyclophis persicus Anderson, 1872; squire 2 indicates the type locality of Pseudocyclophis walteri Boettger, 1888; squire 7 indicates the type locality of Contia zebrina Wall, 1923; triangle 6 indicates the type locality of Contia persica var. nigrofasciata Nikolsky, 1907; star 3 indicates the type locality of Contia angusticeps Boulenger, 1894; plus symbols 25 indicate the type series localities of Contia mcmahoni Wall, 1911. For more details, see Appendix 1." figureDoi="http://doi.org/10.5281/zenodo.5360455" httpUri="https://zenodo.org/record/5360455/files/figure.png" pageId="22" pageNumber="900">Fig. 2</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>