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<document id="056DBF99DF7E729725CE80FC9E8EC988" ID-CLB-Dataset="54097" ID-DOI="10.11646/megataxa.4.1.1" ID-GBIF-Dataset="7c9bc39d-9910-46f1-9558-c0647b9cbbdb" ID-ISSN="2703-3090" ID-Zenodo-Dep="4591138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614033145335" checkinUser="plazi" docAuthor="Kim, Il-Hoi &amp; Boxshall, Geoff A." docDate="2020" docId="03C487CBEECE3BAAFF4DFC57FBCEFCB1" docLanguage="en" docName="megataxa.4.1.1.pdf.imf" docOrigin="Megataxa 4 (1)" docSource="http://zoobank.org/3fdd970e-62f1-4f67-8cce-10870bdb3c01" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Vaoda Kim &amp; Boxshall 2020, gen. nov." docType="treatment" docVersion="8" lastPageNumber="416" masterDocId="FFFDFFB3EF513A35FFC5FFCEFFC5FFC1" masterDocTitle="Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)" masterLastPageNumber="6" masterPageNumber="1" pageNumber="416" updateTime="1698930853666" updateUser="ExternalLinkService">
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<mods:title id="19B466E1E0971490FC3415A06AC975FE">Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)</mods:title>
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<mods:namePart id="2609DA0EA57D7AE11C361A573EB144CB">Kim, Il-Hoi</mods:namePart>
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<mods:affiliation id="B80F86120F3EFD92EEFC34BFCAB96FB6">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<mods:namePart id="87DE08F3B3CAC56F217594DEA6782F27">Boxshall, Geoff A.</mods:namePart>
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<mods:affiliation id="94D81FB5E055F6DFFCB044E95DC547F6">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<treatment id="03C487CBEECE3BAAFF4DFC57FBCEFCB1" ID-DOI="http://doi.org/10.5281/zenodo.5699871" ID-GBIF-Taxon="194475153" ID-Zenodo-Dep="5699871" LSID="urn:lsid:plazi:treatment:03C487CBEECE3BAAFF4DFC57FBCEFCB1" httpUri="http://treatment.plazi.org/id/03C487CBEECE3BAAFF4DFC57FBCEFCB1" lastPageNumber="416" pageId="415" pageNumber="416">
<subSubSection id="C3776556EECE3BAAFF4DFC57FE8DFC72" box="[136,328,921,948]" pageId="415" pageNumber="416" type="nomenclature">
<paragraph id="8BD236DDEECE3BAAFF4DFC57FE8DFC72" blockId="415.[136,328,921,948]" box="[136,328,921,948]" pageId="415" pageNumber="416">
<heading id="D09A81B1EECE3BAAFF4DFC57FE8DFC72" bold="true" box="[136,328,921,948]" fontSize="11" level="1" pageId="415" pageNumber="416" reason="1">
<emphasis id="B919EACFEECE3BAAFF4DFC57FE8DFC72" bold="true" box="[136,328,921,948]" pageId="415" pageNumber="416">
<taxonomicName id="4C6D4D5EEECE3BAAFF4DFC57FF16FC75" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[136,211,921,948]" class="Maxillopoda" family="Notodelphyidae" genus="Vaoda" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEECE3BAAFF4DFC57FF16FC75" bold="true" box="[136,211,921,948]" italics="true" pageId="415" pageNumber="416">Vaoda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EECE3BAAFF1FFC57FE8DFC72" box="[218,328,921,947]" pageId="415" pageNumber="416" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C3776556EECE3BAAFF4DFC13FF03F813" pageId="415" pageNumber="416" type="diagnosis">
<paragraph id="8BD236DDEECE3BAAFF4DFC13FF03F813" blockId="415.[136,778,989,2070]" pageId="415" pageNumber="416">
<emphasis id="B919EACFEECE3BAAFF4DFC13FF3EFC36" bold="true" box="[136,251,989,1015]" pageId="415" pageNumber="416">Diagnosis</emphasis>
. Body dorsoventrally depressed in female, narrower in male. Prosome consisting of cephalosome and 4 pedigerous somites: dorsal cephalic shield and second and third pedigerous somites of female each with welldeveloped epimera. Fourth pedigerous somite forming brood pouch, much longer than other pedigerous somites. Free urosome 5-segmented in female and 6-segmented in male. Rostrum well-developed. Antennule 9-segmented. Antenna 4-segmented; endopod 2-segmented with small terminal claw, exopod represented by 2 vestigial setae. Mandible with 5 teeth on coxal gnathobase; basis with 1 seta; exopod of female with 4 setae, distalmost seta enlarged; exopod of male with 5 subequal setae; endopod with 3 and 7 setae on first and second segments, respectively. Maxillulewith well-defined arthrite, coxal endite and epipodite; basis fused with exopod and endopod forming complex armed with reduced number of setae. Maxilla 3-segmented with unsegmentedendopod bearing 3 setae. Maxilliped unsegmented, armedwith 7 setae medially and 2 setae apically in female, but only 5 setae medially and 2 setae apically in male. Leg 1 with 3-segmented rami; inner coxal seta large; basis with inner distal spine; first endopodal segment with rudimentary inner seta. Legs 24 with 3-segmented exopods and 2- segmented endopods; coxalacking inner seta. Leg 4 without inner seta on first segment of both rami. Leg 5 small, protopod slender with outer naked seta, exopodal segment with 2 unequal apical setae. Leg 6 absent in female; represented by 3 setae on genital operculum in male.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EECE3BAAFF78F814FE16F7D7" pageId="415" pageNumber="416" type="type_taxon">
<paragraph id="8BD236DDEECE3BAAFF78F814FE16F7D7" blockId="415.[136,778,989,2070]" pageId="415" pageNumber="416">
<emphasis id="B919EACFEECE3BAAFF78F814FE00F832" bold="true" box="[189,453,2009,2036]" pageId="415" pageNumber="416">
<typeStatus id="54D6887FEECE3BAAFF78F814FF3DF835" box="[189,248,2010,2036]" pageId="415" pageNumber="416">Type</typeStatus>
and only species
</emphasis>
.
<taxonomicName id="4C6D4D5EEECE3BAAFE10F814FD42F835" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[469,647,2010,2036]" class="Maxillopoda" family="Notodelphyidae" genus="Vaoda" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="species" species="depressa" status="gen. et sp. nov.">
<emphasis id="B919EACFEECE3BAAFE10F814FD42F835" box="[469,647,2010,2036]" italics="true" pageId="415" pageNumber="416">Vaoda depressa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EECE3BAAFD55F814FF7FF7D7" pageId="415" pageNumber="416" rank="species">
<emphasis id="B919EACFEECE3BAAFD55F814FF71F7D7" bold="true" pageId="415" pageNumber="416">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EECE3BAAFC9AFF63FC50FECA" pageId="415" pageNumber="416" type="etymology">
<paragraph id="8BD236DDEECE3BAAFC9AFF63FC50FECA" blockId="415.[810,1452,173,880]" pageId="415" pageNumber="416">
<emphasis id="B919EACFEECE3BAAFC9AFF63FC25FF06" bold="true" box="[863,992,173,199]" pageId="415" pageNumber="416">Etymology</emphasis>
. The new generic name is an anagram of
<collectingRegion id="49A9F83FEECE3BAAFC89FF01FC5DFF28" box="[844,920,207,233]" country="Philippines" name="Davao" pageId="415" pageNumber="416">Davao</collectingRegion>
, the
<typeStatus id="54D6887FEECE3BAAFC16FF01FBC6FF28" box="[979,1027,207,233]" pageId="415" pageNumber="416">type</typeStatus>
locality of the
<typeStatus id="54D6887FEECE3BAAFB79FF01FB29FF28" box="[1212,1260,207,233]" pageId="415" pageNumber="416">type</typeStatus>
species. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EECE3BAAFC9AFEDDFBCEFCB1" pageId="415" pageNumber="416" type="discussion">
<paragraph id="8BD236DDEECE3BAAFC9AFEDDFBCEFCB1" blockId="415.[810,1452,173,880]" pageId="415" pageNumber="416">
<emphasis id="B919EACFEECE3BAAFC9AFEDDFC09FEEC" bold="true" box="[863,972,275,301]" pageId="415" pageNumber="416">Remarks</emphasis>
. The new genus shares the fusion of the basis, exopod andendopod of themaxillulewith
<taxonomicName id="4C6D4D5EEECE3BAAFAFEFEFBFA6EFE8E" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1339,1451,309,335]" class="Maxillopoda" family="Notodelphyidae" genus="Sympygus" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEECE3BAAFAFEFEFBFA6EFE8E" box="[1339,1451,309,335]" italics="true" pageId="415" pageNumber="416">Sympygus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EECE3BAAFCEFFE99FC5CFEB0" box="[810,921,343,369]" pageId="415" pageNumber="416" rank="genus">
<emphasis id="B919EACFEECE3BAAFCEFFE99FC56FEB0" bold="true" box="[810,915,343,369]" pageId="415" pageNumber="416">gen. nov</emphasis>
.
</taxonomicNameLabel>
and with
<taxonomicName id="4C6D4D5EEECE3BAAFBD2FE99FB8AFEB0" authorityName="Stock" authorityYear="1967" box="[1047,1103,343,369]" class="Maxillopoda" family="Notodelphyidae" genus="Sesir" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEECE3BAAFBD2FE99FB8AFEB0" box="[1047,1103,343,369]" italics="true" pageId="415" pageNumber="416">Sesir</emphasis>
</taxonomicName>
, and all three genera share a swimming leg setation pattern which is not significantly reduced.
<taxonomicName id="4C6D4D5EEECE3BAAFC56FE55FC1CFE74" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[915,985,411,437]" class="Maxillopoda" family="Notodelphyidae" genus="Vaoda" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEECE3BAAFC56FE55FC1CFE74" box="[915,985,411,437]" italics="true" pageId="415" pageNumber="416">Vaoda</emphasis>
</taxonomicName>
<emphasis id="B919EACFEECE3BAAFC27FE55FB88FE74" bold="true" box="[994,1101,411,437]" pageId="415" pageNumber="416">
<taxonomicNameLabel id="A22A57B4EECE3BAAFC27FE55FB88FE74" box="[994,1101,411,437]" pageId="415" pageNumber="416" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from the other two genera by a number of features: (1) the body of the female is dorsoventrally depressed, whereas in
<taxonomicName id="4C6D4D5EEECE3BAAFCEFFDCFFC5FFDDA" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[810,922,513,539]" class="Maxillopoda" family="Notodelphyidae" genus="Sympygus" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEECE3BAAFCEFFDCFFC5FFDDA" box="[810,922,513,539]" italics="true" pageId="415" pageNumber="416">Sympygus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EECE3BAAFC61FDCFFBD6FDDD" box="[932,1043,513,540]" pageId="415" pageNumber="416" rank="genus">
<emphasis id="B919EACFEECE3BAAFC61FDCFFBC9FDDA" bold="true" box="[932,1036,513,539]" pageId="415" pageNumber="416">gen. nov</emphasis>
.
</taxonomicNameLabel>
the body is bilaterally compressed and in
<taxonomicName id="4C6D4D5EEECE3BAAFCBAFDEDFC72FDFC" authorityName="Stock" authorityYear="1967" box="[895,951,547,573]" class="Maxillopoda" family="Notodelphyidae" genus="Sesir" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEECE3BAAFCBAFDEDFC72FDFC" box="[895,951,547,573]" italics="true" pageId="415" pageNumber="416">Sesir</emphasis>
</taxonomicName>
it is cylindrical and there is no expressed segmentation in the urosome; (2) the mandibular exopod is armed with 4 setae in
<taxonomicName id="4C6D4D5EEECE3BAAFB83FDA9FB49FD40" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1094,1164,615,641]" class="Maxillopoda" family="Notodelphyidae" genus="Vaoda" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEECE3BAAFB83FDA9FB49FD40" box="[1094,1164,615,641]" italics="true" pageId="415" pageNumber="416">Vaoda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EECE3BAAFB50FDA9FAC4FD43" box="[1173,1281,615,642]" pageId="415" pageNumber="416" rank="genus">
<emphasis id="B919EACFEECE3BAAFB50FDA9FB3FFD40" bold="true" box="[1173,1274,615,641]" pageId="415" pageNumber="416">gen. nov</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="4C6D4D5EEECE3BAAFAFEFDA9FA6EFD40" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1339,1451,615,641]" class="Maxillopoda" family="Notodelphyidae" genus="Sympygus" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEECE3BAAFAFEFDA9FA6EFD40" box="[1339,1451,615,641]" italics="true" pageId="415" pageNumber="416">Sympygus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EECE3BAAFCEFFD47FC50FD65" box="[810,917,649,676]" pageId="415" pageNumber="416" rank="genus">
<emphasis id="B919EACFEECE3BAAFCEFFD47FC4AFD62" bold="true" box="[810,911,649,675]" pageId="415" pageNumber="416">gen. nov</emphasis>
.
</taxonomicNameLabel>
comparedto 5 setaein
<taxonomicName id="4C6D4D5EEECE3BAAFB63FD47FB1BFD62" authorityName="Stock" authorityYear="1967" box="[1190,1246,649,675]" class="Maxillopoda" family="Notodelphyidae" genus="Sesir" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEECE3BAAFB63FD47FB1BFD62" box="[1190,1246,649,675]" italics="true" pageId="415" pageNumber="416">Sesir</emphasis>
</taxonomicName>
; (3) themaxillule possessesa coxalendite which islacking inboth
<taxonomicName id="4C6D4D5EEECE3BAAFAFEFD65FA6EFD04" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1339,1451,683,709]" class="Maxillopoda" family="Notodelphyidae" genus="Sympygus" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEECE3BAAFAFEFD65FA6EFD04" box="[1339,1451,683,709]" italics="true" pageId="415" pageNumber="416">Sympygus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EECE3BAAFCEFFD03FC5CFD29" box="[810,921,717,744]" pageId="415" pageNumber="416" rank="genus">
<emphasis id="B919EACFEECE3BAAFCEFFD03FC57FD26" bold="true" box="[810,914,717,743]" pageId="415" pageNumber="416">gen. nov</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="4C6D4D5EEECE3BAAFC1DFD03FBD5FD26" authorityName="Stock" authorityYear="1967" box="[984,1040,717,743]" class="Maxillopoda" family="Notodelphyidae" genus="Sesir" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEECE3BAAFC1DFD03FBD5FD26" box="[984,1040,717,743]" italics="true" pageId="415" pageNumber="416">Sesir</emphasis>
</taxonomicName>
; (4) the endopod of the maxilla is unsegmented, comparedto 3-segmented in
<taxonomicName id="4C6D4D5EEECE3BAAFAFEFD21FA6EFCC8" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1339,1451,751,777]" class="Maxillopoda" family="Notodelphyidae" genus="Sympygus" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEECE3BAAFAFEFD21FA6EFCC8" box="[1339,1451,751,777]" italics="true" pageId="415" pageNumber="416">Sympygus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EECE3BAAFCEFFCDFFC5DFCED" box="[810,920,785,812]" pageId="415" pageNumber="416" rank="genus">
<emphasis id="B919EACFEECE3BAAFCEFFCDFFC54FCEA" bold="true" box="[810,913,785,811]" pageId="415" pageNumber="416">gen. nov</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="4C6D4D5EEECE3BAAFC10FCDFFBC8FCEA" authorityName="Stock" authorityYear="1967" box="[981,1037,785,811]" class="Maxillopoda" family="Notodelphyidae" genus="Sesir" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEECE3BAAFC10FCDFFBC8FCEA" box="[981,1037,785,811]" italics="true" pageId="415" pageNumber="416">Sesir</emphasis>
</taxonomicName>
; and (5) the setation of legs 14 is better developed than in
<taxonomicName id="4C6D4D5EEECE3BAAFB85FCFDFBBDFC8C" authorityName="Stock" authorityYear="1967" box="[1088,1144,819,845]" class="Maxillopoda" family="Notodelphyidae" genus="Sesir" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEECE3BAAFB85FCFDFBBDFC8C" box="[1088,1144,819,845]" italics="true" pageId="415" pageNumber="416">Sesir</emphasis>
</taxonomicName>
, but less developed than in
<taxonomicName id="4C6D4D5EEECE3BAAFCEFFC98FC5FFCB1" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[810,922,854,880]" class="Maxillopoda" family="Notodelphyidae" genus="Sympygus" kingdom="Animalia" order="Cyclopoida" pageId="415" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEECE3BAAFCEFFC98FC5FFCB1" box="[810,922,854,880]" italics="true" pageId="415" pageNumber="416">Sympygus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EECE3BAAFC64FC9BFBCEFCB1" box="[929,1035,853,880]" pageId="415" pageNumber="416" rank="genus">
<emphasis id="B919EACFEECE3BAAFC64FC9BFBC0FCAE" bold="true" box="[929,1029,853,879]" pageId="415" pageNumber="416">gen. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>