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<document id="4258D196F4FC9A9D5CF23A0C005075EE" ID-CLB-Dataset="54097" ID-DOI="10.11646/megataxa.4.1.1" ID-GBIF-Dataset="7c9bc39d-9910-46f1-9558-c0647b9cbbdb" ID-ISSN="2703-3090" ID-Zenodo-Dep="4591138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614033145335" checkinUser="plazi" docAuthor="Kim, Il-Hoi &amp; Boxshall, Geoff A." docDate="2020" docId="03C487CBEE933BF0FCEFFAFFFD82F9FB" docLanguage="en" docName="megataxa.4.1.1.pdf.imf" docOrigin="Megataxa 4 (1)" docSource="http://zoobank.org/3fdd970e-62f1-4f67-8cce-10870bdb3c01" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Lonchidiopsis replicata Kim &amp; Boxshall 2020, sp. nov." docType="treatment" docVersion="10" lastPageNumber="454" masterDocId="FFFDFFB3EF513A35FFC5FFCEFFC5FFC1" masterDocTitle="Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)" masterLastPageNumber="6" masterPageNumber="1" pageNumber="451" updateTime="1698930853666" updateUser="ExternalLinkService">
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<mods:titleInfo id="CBA0BEAA13DE85A838C585419FAA89A6">
<mods:title id="8A62C8EB19845FE59BE73CF931DD574C">Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)</mods:title>
</mods:titleInfo>
<mods:name id="11163EC2CAA0BF22536458264323D037" type="personal">
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<mods:namePart id="CEA8CCF848B8A3A9D86C7CF495C8A3C4">Kim, Il-Hoi</mods:namePart>
<mods:nameIdentifier id="F2FB88736D139CCDADB46B3242262A87" type="ORCID">0000-0002-7332-0043</mods:nameIdentifier>
<mods:affiliation id="14C6AE3EDB2601D7F09D0CA2405C684D">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
<mods:nameIdentifier id="7E9A783B845605D6E21B4ADB0592FFBD" type="email">ihkim@gwnu.ac.kr</mods:nameIdentifier>
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<mods:namePart id="1F89B8C3D073A25F849053396DC6A57E">Boxshall, Geoff A.</mods:namePart>
<mods:nameIdentifier id="43637275E7888B98823D737E6A4E61C8" type="ORCID">0000-0002-7332-0043</mods:nameIdentifier>
<mods:affiliation id="A2E41E120EB4339109E6E190B105CC79">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
<mods:nameIdentifier id="1FC946202BF7BB9010BF01D3DB7B3C17" type="email">ihkim@gwnu.ac.kr</mods:nameIdentifier>
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<mods:title id="799A72159DDFE7D415E64EE09633B85E">Megataxa</mods:title>
</mods:titleInfo>
<mods:part id="B6C0B3BEF57111097A4D7AEA271B7E13">
<mods:date id="A9CE5BC20C8688CA025E5EB8C9A6A124">2020</mods:date>
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<mods:number id="6337C19292D9046863405F15695A7E42">2020-12-24</mods:number>
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<mods:number id="C579808987F6A5FDBB27901A45946D96">4</mods:number>
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<mods:number id="70E1383E2A1B7E7B35F0C5C371DF94A6">1</mods:number>
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<mods:classification id="0E944FA4F63B962CA6B1A86751FBFBA4">journal article</mods:classification>
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<treatment id="03C487CBEE933BF0FCEFFAFFFD82F9FB" ID-DOI="http://doi.org/10.5281/zenodo.5699887" ID-GBIF-Taxon="194475077" ID-Zenodo-Dep="5699887" LSID="urn:lsid:plazi:treatment:03C487CBEE933BF0FCEFFAFFFD82F9FB" httpUri="http://treatment.plazi.org/id/03C487CBEE933BF0FCEFFAFFFD82F9FB" lastPageId="453" lastPageNumber="454" pageId="450" pageNumber="451">
<subSubSection id="C3776556EE933BF7FCEFFAFFFB6CFA8A" box="[810,1193,1329,1356]" pageId="450" pageNumber="451" type="nomenclature">
<paragraph id="8BD236DDEE933BF7FCEFFAFFFB6CFA8A" blockId="450.[810,1193,1329,1389]" box="[810,1193,1329,1356]" pageId="450" pageNumber="451">
<heading id="D09A81B1EE933BF7FCEFFAFFFB6CFA8A" box="[810,1193,1329,1356]" fontSize="11" level="2" pageId="450" pageNumber="451" reason="2">
<emphasis id="B919EACFEE933BF7FCEFFAFFFB6CFA8A" bold="true" box="[810,1193,1329,1356]" pageId="450" pageNumber="451">
<taxonomicName id="4C6D4D5EEE933BF7FCEFFAFFFB80FA8D" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[810,1093,1329,1356]" class="Maxillopoda" family="Notodelphyidae" genus="Lonchidiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="450" pageNumber="451" phylum="Arthropoda" rank="species" species="replicata" status="sp. nov.">
<emphasis id="B919EACFEE933BF7FCEFFAFFFB80FA8D" bold="true" box="[810,1093,1329,1356]" italics="true" pageId="450" pageNumber="451">Lonchidiopsis replicata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EE933BF7FB89FAFFFB6CFA8A" box="[1100,1193,1329,1355]" pageId="450" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3776556EE933BF7FCEFFA9DFC18FAAC" box="[810,989,1363,1389]" pageId="450" pageNumber="451" type="description">
<paragraph id="8BD236DDEE933BF7FCEFFA9DFC18FAAC" blockId="450.[810,1193,1329,1389]" box="[810,989,1363,1389]" pageId="450" pageNumber="451">
(
<figureCitation id="13562A58EE933BF7FCF7FA9DFC5BFAAC" box="[818,926,1363,1389]" captionStart="FIGURE 299" captionStartId="451.[136,245,1907,1933]" captionTargetBox="[231,1322,200,1837]" captionTargetId="figure-18@451.[202,1385,185,1870]" captionTargetPageId="451" captionText="FIGURE 299. Lochidiopsis replicata sp. nov., female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="450" pageNumber="451">Figs. 299</figureCitation>
,
<figureCitation id="13562A58EE933BF7FC6CFA9DFC10FAAC" box="[937,981,1363,1389]" captionStart="FIGURE 300" captionStartId="452.[136,245,1431,1457]" captionTargetBox="[173,1425,246,1341]" captionTargetId="figure-20@452.[136,1451,219,1358]" captionTargetPageId="452" captionText="FIGURE 300. Lochidiopsis replicata sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 5. Scale bars: A–D, 0.02 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591895" httpUri="https://zenodo.org/record/4591895/files/figure.png" pageId="450" pageNumber="451">300</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3776556EE933BF7FCEFFA56FCACF9BF" pageId="450" pageNumber="451" type="materials_examined">
<paragraph id="8BD236DDEE933BF7FCEFFA56FCACF9BF" blockId="450.[810,1452,1431,2070]" pageId="450" pageNumber="451">
<emphasis id="B919EACFEE933BF7FCEFFA56FC16FA70" bold="true" box="[810,979,1431,1458]" pageId="450" pageNumber="451">Type material</emphasis>
.
<materialsCitation id="3B053C80EE933BF7FC20FA59FCACF9BF" ID-GBIF-Occurrence="3748446509" collectedFrom="from Plurella kottae Monniot F. &amp; Monniot C., 1996" collectingDate="1994-02-07" country="Philippines" elevation="27" latitude="10.2865" location="Buyong" longLatPrecision="13" longitude="124.0025" municipality="Buyong" pageId="450" pageNumber="451" specimenCode="MNHN-IU-2014-21358" specimenCount="1" specimenCount-female="1" stateProvince="Mactan Island" typeStatus="holotype">
<typeStatus id="54D6887FEE933BF7FC20FA59FB88FA70" box="[997,1101,1431,1457]" pageId="450" pageNumber="451" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6BFD54EE933BF7FB92FA56FBA9FA73" box="[1111,1132,1432,1458]" pageId="450" pageNumber="451" type="female">♀</specimenCount>
(dissected and mounted on a slide,
<specimenCode id="DBCB9EA6EE933BF7FC54FA77FB65FA15" box="[913,1184,1465,1492]" pageId="450" pageNumber="451">MNHN-IU-2014-21358</specimenCode>
)
<collectedFrom id="38717520EE933BF7FB72FA77FB79FA34" pageId="450" pageNumber="451">
from
<taxonomicName id="4C6D4D5EEE933BF7FB38FA77FB79FA34" authority="Monniot F. &amp; Monniot C., 1996" authorityName="Monniot F. &amp; Monniot C." authorityYear="1996" class="Ascidiacea" family="Plurellidae" genus="Plurella" kingdom="Animalia" order="Enterogona" pageId="450" pageNumber="451" phylum="Chordata" rank="species" species="kottae">
<emphasis id="B919EACFEE933BF7FB38FA77FA6EFA12" box="[1277,1451,1465,1491]" italics="true" pageId="450" pageNumber="451">Plurella kottae</emphasis>
Monniot F. &amp; Monniot C., 1996
</taxonomicName>
</collectedFrom>
(Holotype MNHN-IT-2008-6248 = MNHN P7/PLU.A/1), CRRFOCDN 2088-F,
<collectingMunicipality id="6BB6ACA7EE933BF7FC49F9EEFC22F9FB" box="[908,999,1568,1594]" pageId="450" pageNumber="451">Buyong</collectingMunicipality>
,
<collectingRegion id="49A9F83FEE933BF7FC3DF9EEFB5AF9F8" box="[1016,1183,1567,1594]" pageId="450" pageNumber="451">Mactan Island</collectingRegion>
,
<collectingRegion id="49A9F83FEE933BF7FB75F9D1FB2BF9F8" box="[1200,1262,1567,1593]" country="Philippines" name="Cebu" pageId="450" pageNumber="451">Cebu</collectingRegion>
, the
<collectingCountry id="F37A764DEE933BF7FAE8F9D1FA6EF9F8" box="[1325,1451,1567,1593]" name="Philippines" pageId="450" pageNumber="451">Philippines</collectingCountry>
(
<geoCoordinate id="EE59501AEE933BF7FCF7F98CFC7CF99D" box="[818,953,1602,1628]" degrees="10" direction="north" minutes="17.19" orientation="latitude" pageId="450" pageNumber="451" precision="9" value="10.2865">10°17.19’N</geoCoordinate>
,
<geoCoordinate id="EE59501AEE933BF7FC09F98FFB9AF99D" box="[972,1119,1601,1628]" degrees="124" direction="east" minutes="00.15" orientation="longitude" pageId="450" pageNumber="451" precision="9" value="124.0025">124°00.15’E</geoCoordinate>
), depth
<quantity id="4C959B38EE933BF7FB03F98CFACCF99D" box="[1222,1289,1602,1628]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="450" pageNumber="451" unit="m" value="27.0">
<elevation id="0040D1EEEE933BF7FB03F98CFACCF99D" box="[1222,1289,1602,1628]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="450" pageNumber="451" unit="m" value="27.0">27 m</elevation>
</quantity>
,
<date id="FFD3101DEE933BF7FAD9F98CFCA0F9BF" pageId="450" pageNumber="451" value="1994-02-07">
<collectingDate id="EF97E9F5EE933BF7FAD9F98CFCA0F9BF" pageId="450" pageNumber="451" value="1994-02-07">07 February 1994</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3776556EE933BF7FC9AF948FC7BF922" pageId="450" pageNumber="451" type="etymology">
<paragraph id="8BD236DDEE933BF7FC9AF948FC7BF922" blockId="450.[810,1452,1431,2070]" pageId="450" pageNumber="451">
<emphasis id="B919EACFEE933BF7FC9AF948FC25F961" bold="true" box="[863,992,1670,1696]" pageId="450" pageNumber="451">Etymology</emphasis>
. The specific name is derived from the Latin
<emphasis id="B919EACFEE933BF7FCB5F969FC75F900" box="[880,944,1703,1729]" italics="true" pageId="450" pageNumber="451">replic</emphasis>
(=fold back), referring to the shape of the brood pouch.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EE933BF0FC9AF925FEBBFB02" lastPageId="453" lastPageNumber="454" pageId="450" pageNumber="451" type="description">
<paragraph id="8BD236DDEE933BF1FC9AF925FD0AF925" blockId="450.[810,1452,1431,2070]" lastBlockId="452.[136,778,1567,2070]" lastPageId="452" lastPageNumber="453" pageId="450" pageNumber="451">
<emphasis id="B919EACFEE933BF7FC9AF925FBAEF8C4" bold="true" box="[863,1131,1771,1797]" pageId="450" pageNumber="451">Descriptionoffemale</emphasis>
. Body (
<figureCitation id="13562A58EE933BF7FB16F925FA8BF8C7" box="[1235,1358,1771,1798]" captionStart="FIGURE 299" captionStartId="451.[136,245,1907,1933]" captionTargetBox="[231,1322,200,1837]" captionTargetId="figure-18@451.[202,1385,185,1870]" captionTargetPageId="451" captionText="FIGURE 299. Lochidiopsis replicata sp. nov., female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="450" pageNumber="451">Fig. 299A</figureCitation>
) highly transformed, laterally compressed, consisting of cephalothorax, short neck, greatly expanded brood pouch, and 4-segmented freeurosome. Bodylength 1.70 mm. Cephalic shield expanded ventrolaterally, concealing part of surface extending from mouthparts to third pedigerous somite in lateral view. Short neck region without appendages. Fourth pedigerous somite forming brood pouch, extremely compressed but expanded in vertical planeto form paireddiscoid lateralfolds separatedby deep dorsal furrow. Freeurosome (
<figureCitation id="13562A58EE953BF1FE2DF98FFDA5F99D" box="[488,608,1601,1628]" captionStart="FIGURE 299" captionStartId="451.[136,245,1907,1933]" captionTargetBox="[231,1322,200,1837]" captionTargetId="figure-18@451.[202,1385,185,1870]" captionTargetPageId="451" captionText="FIGURE 299. Lochidiopsis replicata sp. nov., female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="452" pageNumber="453">Fig. 299B</figureCitation>
) 4-segmented, consisting of genital double-somite and 3-segmented abdomen, Caudal rami fused to anal somite without any trace of articulation, slightly longer than wide; armed with 6 minute setae on convex distal margin (
<figureCitation id="13562A58EE953BF1FD88F907FD06F925" box="[589,707,1737,1764]" captionStart="FIGURE 299" captionStartId="451.[136,245,1907,1933]" captionTargetBox="[231,1322,200,1837]" captionTargetId="figure-18@451.[202,1385,185,1870]" captionTargetPageId="451" captionText="FIGURE 299. Lochidiopsis replicata sp. nov., female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="452" pageNumber="453">Fig. 299C</figureCitation>
).
</paragraph>
<caption id="DF126655EE923BF6FF4DF8BDFA53F813" ID-DOI="http://doi.org/10.5281/zenodo.4591893" ID-Zenodo-Dep="4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="451" pageNumber="452" startId="451.[136,245,1907,1933]" targetBox="[231,1322,200,1837]" targetPageId="451">
<paragraph id="8BD236DDEE923BF6FF4DF8BDFA53F813" blockId="451.[136,1451,1907,2002]" pageId="451" pageNumber="452">
<emphasis id="B919EACFEE923BF6FF4DF8BDFEE9F84F" bold="true" box="[136,300,1907,1934]" pageId="451" pageNumber="452">FIGURE 299.</emphasis>
<taxonomicName id="4C6D4D5EEE923BF6FEF4F8BAFDEFF84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[305,554,1908,1934]" class="Maxillopoda" family="Notodelphyidae" genus="Lochidiopsis" kingdom="Animalia" order="Cyclopoida" pageId="451" pageNumber="452" phylum="Arthropoda" rank="species" species="replicata" status="sp. nov.">
<emphasis id="B919EACFEE923BF6FEF4F8BAFDEFF84F" box="[305,554,1908,1934]" italics="true" pageId="451" pageNumber="452">Lochidiopsis replicata</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE923BF6FDF5F8BAFD4DF84F" bold="true" box="[560,648,1908,1934]" pageId="451" pageNumber="452">
<taxonomicNameLabel id="A22A57B4EE923BF6FDF5F8BAFD4DF84F" box="[560,648,1908,1934]" pageId="451" pageNumber="452" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm.
</paragraph>
</caption>
<caption id="DF126655EE953BF1FF4DFA59FDC0FA15" ID-DOI="http://doi.org/10.5281/zenodo.4591895" ID-Zenodo-Dep="4591895" httpUri="https://zenodo.org/record/4591895/files/figure.png" pageId="452" pageNumber="453" startId="452.[136,245,1431,1457]" targetBox="[173,1425,246,1341]" targetPageId="452">
<paragraph id="8BD236DDEE953BF1FF4DFA59FDC0FA15" blockId="452.[136,1452,1431,1492]" pageId="452" pageNumber="453">
<emphasis id="B919EACFEE953BF1FF4DFA59FEEBFA70" bold="true" box="[136,302,1431,1457]" pageId="452" pageNumber="453">FIGURE 300.</emphasis>
<taxonomicName id="4C6D4D5EEE953BF1FEF2FA59FDF7FA70" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[311,562,1431,1457]" class="Maxillopoda" family="Notodelphyidae" genus="Lochidiopsis" kingdom="Animalia" order="Cyclopoida" pageId="452" pageNumber="453" phylum="Arthropoda" rank="species" species="replicata" status="sp. nov.">
<emphasis id="B919EACFEE953BF1FEF2FA59FDF7FA70" box="[311,562,1431,1457]" italics="true" pageId="452" pageNumber="453">Lochidiopsis replicata</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE953BF1FDFFFA56FD50FA73" bold="true" box="[570,661,1432,1458]" pageId="452" pageNumber="453">
<taxonomicNameLabel id="A22A57B4EE953BF1FDFFFA56FD50FA73" box="[570,661,1432,1458]" pageId="452" pageNumber="453" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 5. Scale bars: A–D, 0.02 mm; E, 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BD236DDEE953BF1FF78F922FB7DF9BC" blockId="452.[136,778,1567,2070]" lastBlockId="452.[810,1452,1567,2070]" pageId="452" pageNumber="453">
Rostrum (
<figureCitation id="13562A58EE953BF1FEFCF925FE7DF8C7" box="[313,440,1771,1798]" captionStart="FIGURE 299" captionStartId="451.[136,245,1907,1933]" captionTargetBox="[231,1322,200,1837]" captionTargetId="figure-18@451.[202,1385,185,1870]" captionTargetPageId="451" captionText="FIGURE 299. Lochidiopsis replicata sp. nov., female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="452" pageNumber="453">Fig. 299D</figureCitation>
) small, 20×22 μm, highly sclerotized with rounded apex. Antennule (
<figureCitation id="13562A58EE953BF1FD49F8C3FCC7F8E9" box="[652,770,1805,1832]" captionStart="FIGURE 299" captionStartId="451.[136,245,1907,1933]" captionTargetBox="[231,1322,200,1837]" captionTargetId="figure-18@451.[202,1385,185,1870]" captionTargetPageId="451" captionText="FIGURE 299. Lochidiopsis replicata sp. nov., female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="452" pageNumber="453">Fig. 299F</figureCitation>
) about120 μmlong, 6-segmented; firstandsecondsegments extremely expanded; third segment inserting into middle of secondsegment; armatureformula 2, 14, 3, 2, 2, and 13. Antenna (
<figureCitation id="13562A58EE953BF1FED9F858FE55F871" box="[284,400,1942,1968]" captionStart="FIGURE 299" captionStartId="451.[136,245,1907,1933]" captionTargetBox="[231,1322,200,1837]" captionTargetId="figure-18@451.[202,1385,185,1870]" captionTargetPageId="451" captionText="FIGURE 299. Lochidiopsis replicata sp. nov., female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="452" pageNumber="453">Fig. 299G</figureCitation>
) 3-segmented, consistingof coxa, allobasis, and 1-segmented endopod; allobasis 57×23 μm, unarmed, with unsclerotized area near middle as trace of articulation; freeendopodal segment 32×15 μm, armed with 2 setae in middle of inner margin, and ornamented with 4 or 5 spinules on distal outer margin; terminal claw about 19 μm long, attenuated at tip.
</paragraph>
<paragraph id="8BD236DDEE953BF0FC9AF94BFEA9FD43" blockId="452.[810,1452,1567,2070]" lastBlockId="453.[136,778,173,880]" lastPageId="453" lastPageNumber="454" pageId="452" pageNumber="453">
Labrum (
<figureCitation id="13562A58EE953BF1FC18F94BFBA0F961" box="[989,1125,1669,1696]" captionStart="FIGURE 299" captionStartId="451.[136,245,1907,1933]" captionTargetBox="[231,1322,200,1837]" captionTargetId="figure-18@451.[202,1385,185,1870]" captionTargetPageId="451" captionText="FIGURE 299. Lochidiopsis replicata sp. nov., female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="452" pageNumber="453">Fig. 299E</figureCitation>
) with large, linguiform posteromedian lobe and setulose posterior margin. Mandible (
<figureCitation id="13562A58EE953BF1FC6AF907FBEAF925" box="[943,1071,1737,1764]" captionStart="FIGURE 299" captionStartId="451.[136,245,1907,1933]" captionTargetBox="[231,1322,200,1837]" captionTargetId="figure-18@451.[202,1385,185,1870]" captionTargetPageId="451" captionText="FIGURE 299. Lochidiopsis replicata sp. nov., female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="452" pageNumber="453">Fig. 299H</figureCitation>
) with broad coxalgnathobase bearing 5 teeth on medial margin and 1 setaon proximal margin: basis with seta at mediodistal corner: exopod with 5 large setae, medial seta pinnate proximally but with spinulesalongdistalpartof medial margin; firstendopodal segment almost fused to basis, with trace of articulation on medial side, armed with 4 setae, distalmost seta broad and pinnate, with long setules; second endopodal segment armedwith 9 setae, 5 medial setae spinulose along distal half of medial margin. Maxillule (
<figureCitation id="13562A58EE953BF1FB66F832FACEF7D7" box="[1187,1291,2044,2070]" captionStart="FIGURE 299" captionStartId="451.[136,245,1907,1933]" captionTargetBox="[231,1322,200,1837]" captionTargetId="figure-18@451.[202,1385,185,1870]" captionTargetPageId="451" captionText="FIGURE 299. Lochidiopsis replicata sp. nov., female.A, habitus, left; B, urosome, right; C, posterior part of urosome, dorsal view showing rami fused to anal somite, with inset showing detail of caudal setae; D, rostrum; E, labrum; F, antennule; G, antenna; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, C, 0.1 mm; D, 0.01 mm; E–I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591893" httpUri="https://zenodo.org/record/4591893/files/figure.png" pageId="452" pageNumber="453">Fig. 299I</figureCitation>
) with 10 setae on arthrite; coxa with 1 setaon endite and 2 very unequal setae on epipodite; basis with 4 setae (second and fourth setae shorter than first and third); exopod with 4 setae on distal margin; endopod with 2 medial and 3 distal setae (second medial seta broad, curved outwards, with thick setules along convex margin; outermost distal seta longest). Maxilla (
<figureCitation id="13562A58EE943BF0FE93FEB7FE0FFE52" box="[342,458,377,403]" captionStart="FIGURE 300" captionStartId="452.[136,245,1431,1457]" captionTargetBox="[173,1425,246,1341]" captionTargetId="figure-20@452.[136,1451,219,1358]" captionTargetPageId="452" captionText="FIGURE 300. Lochidiopsis replicata sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 5. Scale bars: A–D, 0.02 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591895" httpUri="https://zenodo.org/record/4591895/files/figure.png" pageId="453" pageNumber="454">Fig. 300A</figureCitation>
) 4-segmented; syncoxawith 4, 1, 2, and 3 setae on first to fourth endites, respectively; basis with large claw bearing serrations along distal half of convex margin plus 2 unequal setae; endopod atrophied, 2-segmented, first segment unarmed, second with 5 small setae. Maxilliped (
<figureCitation id="13562A58EE943BF0FE90FDEDFE03FDFC" box="[341,454,547,573]" captionStart="FIGURE 300" captionStartId="452.[136,245,1431,1457]" captionTargetBox="[173,1425,246,1341]" captionTargetId="figure-20@452.[136,1451,219,1358]" captionTargetPageId="452" captionText="FIGURE 300. Lochidiopsis replicata sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 5. Scale bars: A–D, 0.02 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591895" httpUri="https://zenodo.org/record/4591895/files/figure.png" pageId="453" pageNumber="454">Fig. 300B</figureCitation>
) distinctly 2-segmented; first segment with 6 setaeon medial margin; secondsegment narrow with 3 setae.
</paragraph>
<paragraph id="8BD236DDEE943BF0FF78FD44FD70FCAE" blockId="453.[136,778,173,880]" pageId="453" pageNumber="454">
Legs 1–3 biramous with 2-segmented rami (
<figureCitation id="13562A58EE943BF0FD19FD47FF2AFD07" captionStart="FIGURE 300" captionStartId="452.[136,245,1431,1457]" captionTargetBox="[173,1425,246,1341]" captionTargetId="figure-20@452.[136,1451,219,1358]" captionTargetPageId="452" captionText="FIGURE 300. Lochidiopsis replicata sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 5. Scale bars: A–D, 0.02 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591895" httpUri="https://zenodo.org/record/4591895/files/figure.png" pageId="453" pageNumber="454">Fig. 300C, D</figureCitation>
); innercoxalsetaabsent; outersetaonbasis large inleg 1, butsmall in legs 2 and 3. Inner margin of basis and outer margin of endopod setulose in leg 1, but smooth in legs 2 and 3. Compound second endopodal segment of leg 1 ovate with spinulose margins. Leg 4 absent. Armature formula for legs 1–3 as follows:
</paragraph>
<paragraph id="8BD236DDEE943BF0FC4FFD47FA41FCEA" pageId="453" pageNumber="454">
<table id="F96DC47DEE94C5CAFCEFFD47FA58FCED" box="[810,1437,649,812]" gridcols="5" gridrows="5" inLine="true" pageId="453" pageNumber="454">
<tr id="355D349FEE94C5CAFCEFFD47FA58FD62" box="[810,1437,649,675]" gridrow="0" pageId="453" pageNumber="454" rowspan-0="1">
<th id="768C5DE3EE94C5CAFC4FFD47FC00FD62" box="[906,965,649,675]" gridcol="1" gridrow="0" pageId="453" pageNumber="454">Coxa</th>
<th id="768C5DE3EE94C5CAFC2FFD47FBE2FD62" box="[1002,1063,649,675]" gridcol="2" gridrow="0" pageId="453" pageNumber="454">Basis</th>
<th id="768C5DE3EE94C5CAFB8FFD47FB38FD62" box="[1098,1277,649,675]" gridcol="3" gridrow="0" pageId="453" pageNumber="454">Exopod</th>
<th id="768C5DE3EE94C5CAFACFFD47FA58FD62" box="[1290,1437,649,675]" gridcol="4" gridrow="0" pageId="453" pageNumber="454">Endopod</th>
</tr>
<tr id="355D349FEE94C5CAFCEFFD65FA58FD07" box="[810,1437,683,710]" gridrow="1" pageId="453" pageNumber="454">
<th id="768C5DE3EE94C5CAFCEFFD65FCAFFD07" box="[810,874,683,710]" gridcol="0" gridrow="1" pageId="453" pageNumber="454">Leg 1</th>
<td id="768C5DE3EE94C5CAFC4FFD65FC00FD07" box="[906,965,683,710]" gridcol="1" gridrow="1" pageId="453" pageNumber="454">0-0</td>
<td id="768C5DE3EE94C5CAFC2FFD65FBE2FD07" box="[1002,1063,683,710]" gridcol="2" gridrow="1" pageId="453" pageNumber="454">1-1</td>
<td id="768C5DE3EE94C5CAFB8FFD65FB38FD07" box="[1098,1277,683,710]" gridcol="3" gridrow="1" pageId="453" pageNumber="454">I-1; I-0; II, I, I</td>
<td id="768C5DE3EE94C5CAFACFFD65FA58FD07" box="[1290,1437,683,710]" gridcol="4" gridrow="1" pageId="453" pageNumber="454">0-1; 1, 2, 2</td>
</tr>
<tr id="355D349FEE94C5CAFCEFFD03FA58FD29" box="[810,1437,717,744]" gridrow="2" pageId="453" pageNumber="454">
<th id="768C5DE3EE94C5CAFCEFFD03FCAFFD29" box="[810,874,717,744]" gridcol="0" gridrow="2" pageId="453" pageNumber="454">Leg 2</th>
<td id="768C5DE3EE94C5CAFC4FFD03FC00FD29" box="[906,965,717,744]" gridcol="1" gridrow="2" pageId="453" pageNumber="454">0-0</td>
<td id="768C5DE3EE94C5CAFC2FFD03FBE2FD29" box="[1002,1063,717,744]" gridcol="2" gridrow="2" pageId="453" pageNumber="454">1-0</td>
<td id="768C5DE3EE94C5CAFB8FFD03FB38FD29" box="[1098,1277,717,744]" gridcol="3" gridrow="2" pageId="453" pageNumber="454">
1-1; 1-0; 2, II,
<td id="768C5DE3EE94C5CAFACFFD03FA58FD29" box="[1290,1437,717,744]" gridcol="4" gridrow="2" pageId="453" pageNumber="454">10-1; II, I, I+1</td>
</td>
</tr>
<tr id="355D349FEE94C5CAFCEFFD21FA58FCCB" box="[810,1437,751,778]" gridrow="3" pageId="453" pageNumber="454">
<th id="768C5DE3EE94C5CAFCEFFD21FCAFFCCB" box="[810,874,751,778]" gridcol="0" gridrow="3" pageId="453" pageNumber="454">Leg 3</th>
<td id="768C5DE3EE94C5CAFC4FFD21FC00FCCB" box="[906,965,751,778]" gridcol="1" gridrow="3" pageId="453" pageNumber="454">0-0</td>
<td id="768C5DE3EE94C5CAFC2FFD21FBE2FCCB" box="[1002,1063,751,778]" gridcol="2" gridrow="3" pageId="453" pageNumber="454">1-0</td>
<td id="768C5DE3EE94C5CAFB8FFD21FB38FCCB" box="[1098,1277,751,778]" gridcol="3" gridrow="3" pageId="453" pageNumber="454">1-0; 0-0; 1, 1, 2</td>
<td id="768C5DE3EE94C5CAFACFFD21FA58FCCB" box="[1290,1437,751,778]" gridcol="4" gridrow="3" pageId="453" pageNumber="454">0-1; 1, 2, 1</td>
</tr>
<tr id="355D349FEE94C5CAFCEFFCDFFA58FCED" box="[810,1437,785,812]" gridrow="4" pageId="453" pageNumber="454">
<th id="768C5DE3EE94C5CAFCEFFCDFFCAFFCED" box="[810,874,785,812]" gridcol="0" gridrow="4" pageId="453" pageNumber="454">Leg 4</th>
<td id="768C5DE3EE94C5CAFC4FFCDFFC00FCED" box="[906,965,785,812]" gridcol="1" gridrow="4" pageId="453" pageNumber="454">0-0</td>
<td id="768C5DE3EE94C5CAFC2FFCDFFBE2FCED" box="[1002,1063,785,812]" gridcol="2" gridrow="4" pageId="453" pageNumber="454">1-0</td>
<td id="768C5DE3EE94C5CAFB8FFCDFFB38FCED" box="[1098,1277,785,812]" gridcol="3" gridrow="4" pageId="453" pageNumber="454">1-0; 0-0; 0, 3, 0</td>
<td id="768C5DE3EE94C5CAFACFFCDFFA58FCED" box="[1290,1437,785,812]" gridcol="4" gridrow="4" pageId="453" pageNumber="454">0-0; 0, 2, 0</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD236DDEE943BF0FEE0FC57FD26FC39" pageId="453" pageNumber="454">
<table id="F96DC47DEE94C5CAFF4DFC57FD3DFC39" box="[136,760,921,1016]" gridcols="5" gridrows="3" inLine="true" pageId="453" pageNumber="454">
<tr id="355D349FEE94C5CAFF4DFC57FD3DFC72" box="[136,760,921,947]" gridrow="0" pageId="453" pageNumber="454" rowspan-0="1">
<th id="768C5DE3EE94C5CAFEE1FC57FE9AFC72" box="[292,351,921,947]" gridcol="1" gridrow="0" pageId="453" pageNumber="454">Coxa</th>
<th id="768C5DE3EE94C5CAFEBBFC57FE7EFC72" box="[382,443,921,947]" gridcol="2" gridrow="0" pageId="453" pageNumber="454">Basis</th>
<th id="768C5DE3EE94C5CAFE18FC57FD8AFC72" box="[477,591,921,947]" gridcol="3" gridrow="0" pageId="453" pageNumber="454">Exopod</th>
<th id="768C5DE3EE94C5CAFDADFC57FD3DFC72" box="[616,760,921,947]" gridcol="4" gridrow="0" pageId="453" pageNumber="454">Endopod</th>
</tr>
<tr id="355D349FEE94C5CAFF4DFC75FD3DFC17" box="[136,760,955,982]" gridrow="1" pageId="453" pageNumber="454">
<th id="768C5DE3EE94C5CAFF4DFC75FEC6FC17" box="[136,259,955,982]" gridcol="0" gridrow="1" pageId="453" pageNumber="454">Leg 1</th>
<td id="768C5DE3EE94C5CAFEE1FC75FE9AFC17" box="[292,351,955,982]" gridcol="1" gridrow="1" pageId="453" pageNumber="454">0-0</td>
<td id="768C5DE3EE94C5CAFEBBFC75FE7EFC17" box="[382,443,955,982]" gridcol="2" gridrow="1" pageId="453" pageNumber="454">1-I</td>
<td id="768C5DE3EE94C5CAFE18FC75FD8AFC17" box="[477,591,955,982]" gridcol="3" gridrow="1" pageId="453" pageNumber="454">I-1; II, I, I</td>
<td id="768C5DE3EE94C5CAFDADFC75FD3DFC17" box="[616,760,955,982]" gridcol="4" gridrow="1" pageId="453" pageNumber="454">0-1; 1, 2, I+1</td>
</tr>
<tr id="355D349FEE94C5CAFF4DFC13FD3DFC39" box="[136,760,989,1016]" gridrow="2" pageId="453" pageNumber="454">
<th id="768C5DE3EE94C5CAFF4DFC13FEC6FC39" box="[136,259,989,1016]" gridcol="0" gridrow="2" pageId="453" pageNumber="454">Legs 2 &amp; 3</th>
<td id="768C5DE3EE94C5CAFEE1FC13FE9AFC39" box="[292,351,989,1016]" gridcol="1" gridrow="2" pageId="453" pageNumber="454">0-0</td>
<td id="768C5DE3EE94C5CAFEBBFC13FE7EFC39" box="[382,443,989,1016]" gridcol="2" gridrow="2" pageId="453" pageNumber="454">1-0</td>
<td id="768C5DE3EE94C5CAFE18FC13FD8AFC39" box="[477,591,989,1016]" gridcol="3" gridrow="2" pageId="453" pageNumber="454">I-1; I, I, I</td>
<td id="768C5DE3EE94C5CAFDADFC13FD3DFC39" box="[616,760,989,1016]" gridcol="4" gridrow="2" pageId="453" pageNumber="454">0-0; I, II, II</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD236DDEE943BF0FF78FBECFDD5FB60" blockId="453.[136,778,1057,1594]" pageId="453" pageNumber="454">
Leg 5 (
<figureCitation id="13562A58EE943BF0FEC9FBEFFEBEFBFA" box="[268,379,1057,1083]" captionStart="FIGURE 300" captionStartId="452.[136,245,1431,1457]" captionTargetBox="[173,1425,246,1341]" captionTargetId="figure-20@452.[136,1451,219,1358]" captionTargetPageId="452" captionText="FIGURE 300. Lochidiopsis replicata sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 5. Scale bars: A–D, 0.02 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591895" httpUri="https://zenodo.org/record/4591895/files/figure.png" pageId="453" pageNumber="454">Fig. 300E</figureCitation>
) represented by small free exopodal segment about 3.6 times longer than wide (101×28 μm), with narrow proximal third and broader distal two-thirds; armed with 2 unequal setae distally.
</paragraph>
<paragraph id="8BD236DDEE943BF0FF78FB67FEBBFB02" blockId="453.[136,778,1057,1594]" box="[189,382,1193,1220]" pageId="453" pageNumber="454">
<emphasis id="B919EACFEE943BF0FF78FB67FF3FFB02" bold="true" box="[189,250,1193,1219]" pageId="453" pageNumber="454">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EE943BF0FF78FB05FD82F9FB" pageId="453" pageNumber="454" type="discussion">
<paragraph id="8BD236DDEE943BF0FF78FB05FD82F9FB" blockId="453.[136,778,1057,1594]" pageId="453" pageNumber="454">
<emphasis id="B919EACFEE943BF0FF78FB05FEEFFB24" bold="true" box="[189,298,1227,1253]" pageId="453" pageNumber="454">Remarks</emphasis>
. Despite the highly transformed body form which makes
<taxonomicName id="4C6D4D5EEE943BF0FEE7FB20FE61FAC6" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[290,420,1261,1288]" class="Maxillopoda" family="Notodelphyidae" genus="Lochidiopsis" kingdom="Animalia" order="Cyclopoida" pageId="453" pageNumber="454" phylum="Arthropoda" rank="species" species="replicata" status="sp. nov.">
<emphasis id="B919EACFEE943BF0FEE7FB20FEF7FAC6" box="[290,306,1262,1287]" italics="true" pageId="453" pageNumber="454">L</emphasis>
.
<emphasis id="B919EACFEE943BF0FE85FB23FE61FAC6" box="[320,420,1261,1287]" italics="true" pageId="453" pageNumber="454">replicata</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE943BF0FE6EFB23FDC0FAC6" bold="true" box="[427,517,1261,1287]" pageId="453" pageNumber="454">
<taxonomicNameLabel id="A22A57B4EE943BF0FE6EFB23FDC0FAC6" box="[427,517,1261,1287]" pageId="453" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
instantly recognizable, its cephalic appendages and legs share various similarities with those of its three congeners. The unique features of the new species within the genus include; the caudal rami are completely fused to the anal somite (vs. the caudal rami are free in all congeneric species), the distal endopodal segment of leg 1 is armed with 1 spine and 4 setae (vs. 1 spineand 5 setaein congeners), and leg 4 is absent (vs. this leg is present in congeners, although it is rudimentary in
<taxonomicName id="4C6D4D5EEE943BF0FEF0F9EEFE66F9F8" authorityName="Jones &amp; Montez Moreno" authorityYear="1981" box="[309,419,1568,1594]" class="Maxillopoda" family="Notodelphyidae" genus="Lonchidiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="453" pageNumber="454" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="B919EACFEE943BF0FEF0F9EEFE80F9F8" box="[309,325,1568,1593]" italics="true" pageId="453" pageNumber="454">L</emphasis>
.
<emphasis id="B919EACFEE943BF0FE97F9EEFE66F9F8" box="[338,419,1568,1593]" italics="true" pageId="453" pageNumber="454">setosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6D4D5EEE943BF0FE1FF9EEFDF2F9F8" authorityName="Stock" authorityYear="1967" box="[474,567,1568,1594]" class="Maxillopoda" family="Notodelphyidae" genus="Lonchidiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="453" pageNumber="454" phylum="Arthropoda" rank="species" species="tripes">
<emphasis id="B919EACFEE943BF0FE1FF9EEFE2FF9F8" box="[474,490,1568,1593]" italics="true" pageId="453" pageNumber="454">L</emphasis>
.
<emphasis id="B919EACFEE943BF0FE32F9EEFDF2F9F8" box="[503,567,1568,1593]" italics="true" pageId="453" pageNumber="454">tripes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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