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<document id="CEF6BA886724BEA85B13AAA8BEE26A4A" ID-CLB-Dataset="54097" ID-DOI="10.11646/megataxa.4.1.1" ID-GBIF-Dataset="7c9bc39d-9910-46f1-9558-c0647b9cbbdb" ID-ISSN="2703-3090" ID-Zenodo-Dep="4591138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614033145335" checkinUser="plazi" docAuthor="Kim, Il-Hoi &amp; Boxshall, Geoff A." docDate="2020" docId="03C487CBED7E3807FF4DF8C3FAE2F8C7" docLanguage="en" docName="megataxa.4.1.1.pdf.imf" docOrigin="Megataxa 4 (1)" docSource="http://zoobank.org/3fdd970e-62f1-4f67-8cce-10870bdb3c01" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Scoliosoma haplomerosum Kim &amp; Boxshall 2020, gen. et sp. nov." docType="treatment" docVersion="10" lastPageNumber="563" masterDocId="FFFDFFB3EF513A35FFC5FFCEFFC5FFC1" masterDocTitle="Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)" masterLastPageNumber="6" masterPageNumber="1" pageNumber="560" updateTime="1698930853666" updateUser="ExternalLinkService">
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<mods:title id="04902BCBD76F2F878B1BC64FC18A89FA">Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)</mods:title>
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<mods:namePart id="EE09D10F903C434CFC00AA6D6116EE52">Kim, Il-Hoi</mods:namePart>
<mods:nameIdentifier id="E8048B3364D35C45AB50D2988CCDF9D8" type="ORCID">0000-0002-7332-0043</mods:nameIdentifier>
<mods:affiliation id="FBE413FF46070B40D823A1C2F3BB4B39">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
<mods:nameIdentifier id="3191413EF7782496DE907B19418851FD" type="email">ihkim@gwnu.ac.kr</mods:nameIdentifier>
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<mods:namePart id="4365FD885AF30E2374E58FF98ECF673D">Boxshall, Geoff A.</mods:namePart>
<mods:nameIdentifier id="CB2C427723FF4D32AF7EEA75D0CCECAB" type="ORCID">0000-0002-7332-0043</mods:nameIdentifier>
<mods:affiliation id="015E5721672185E6075771990010F206">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
<mods:nameIdentifier id="9E31CA8F894E78644FA683709B90FDA3" type="email">ihkim@gwnu.ac.kr</mods:nameIdentifier>
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<mods:title id="1B438A640815D33BD88F10F87440D8C8">Megataxa</mods:title>
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<mods:date id="E3DEC4A4482FCB9F27008735F3C35DCA">2020</mods:date>
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<mods:number id="19AF516ADB23BF524D8110D954B19F33">2020-12-24</mods:number>
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<treatment id="03C487CBED7E3807FF4DF8C3FAE2F8C7" ID-DOI="http://doi.org/10.5281/zenodo.5699945" ID-GBIF-Taxon="194474947" ID-Zenodo-Dep="5699945" LSID="urn:lsid:plazi:treatment:03C487CBED7E3807FF4DF8C3FAE2F8C7" httpUri="http://treatment.plazi.org/id/03C487CBED7E3807FF4DF8C3FAE2F8C7" lastPageId="562" lastPageNumber="563" pageId="559" pageNumber="560">
<subSubSection id="C3776556ED7E381AFF4DF8C3FD48F8E9" box="[136,653,1805,1832]" pageId="559" pageNumber="560" type="nomenclature">
<paragraph id="8BD236DDED7E381AFF4DF8C3FD48F8E9" blockId="559.[136,653,1805,1866]" box="[136,653,1805,1832]" pageId="559" pageNumber="560">
<heading id="D09A81B1ED7E381AFF4DF8C3FD48F8E9" box="[136,653,1805,1832]" fontSize="11" level="2" pageId="559" pageNumber="560" reason="2">
<emphasis id="B919EACFED7E381AFF4DF8C3FD48F8E9" bold="true" box="[136,653,1805,1832]" pageId="559" pageNumber="560">
<taxonomicName id="4C6D4D5EED7E381AFF4DF8C3FE0AF8E9" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[136,463,1805,1832]" class="Maxillopoda" family="Notodelphyidae" genus="Scoliosoma" kingdom="Animalia" order="Cyclopoida" pageId="559" pageNumber="560" phylum="Arthropoda" rank="species" species="haplomerosum" status="gen. et sp. nov.">
<emphasis id="B919EACFED7E381AFF4DF8C3FE0AF8E9" bold="true" box="[136,463,1805,1832]" italics="true" pageId="559" pageNumber="560">Scoliosoma haplomerosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4ED7E381AFE13F8C0FD48F8E9" box="[470,653,1806,1832]" pageId="559" pageNumber="560" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3776556ED7E381AFF4DF8FEFEFEF88B" box="[136,315,1839,1866]" pageId="559" pageNumber="560" type="description">
<paragraph id="8BD236DDED7E381AFF4DF8FEFEFEF88B" blockId="559.[136,653,1805,1866]" box="[136,315,1839,1866]" pageId="559" pageNumber="560">
(
<figureCitation id="13562A58ED7E381AFF55F8FEFF38F88B" box="[144,253,1839,1866]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="559" pageNumber="560">Figs. 375</figureCitation>
,
<figureCitation id="13562A58ED7E381AFECDF8E1FEF1F88B" box="[264,308,1839,1866]" captionStart="FIGURE 376" captionStartId="561.[136,245,1907,1933]" captionTargetBox="[228,1334,212,1845]" captionTargetId="figure-17@561.[195,1393,185,1870]" captionTargetPageId="561" captionText="FIGURE 376. Scoliosoma haplomerosum gen. et sp. nov., female: A, leg 3; B, leg 4. Copepodid V female: C, habitus, dorsal; D, habitus, left; E, urosome, dorsal; F, antennule; G, leg 1; H, leg 2; I, leg 3; J, leg 4. Scale bars: A, B, F–J, 0.02 mm; C, D, 0.1 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592066" httpUri="https://zenodo.org/record/4592066/files/figure.png" pageId="559" pageNumber="560">376</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3776556ED7E381AFF4DF8BAFAFEFAE8" pageId="559" pageNumber="560" type="materials_examined">
<paragraph id="8BD236DDED7E381AFF4DF8BAFAFEFAE8" blockId="559.[136,778,1907,2071]" lastBlockId="559.[810,1452,1159,2070]" pageId="559" pageNumber="560">
<emphasis id="B919EACFED7E381AFF4DF8BAFEEBF84C" bold="true" box="[136,302,1907,1934]" pageId="559" pageNumber="560">Type material</emphasis>
.
<materialsCitation id="3B053C80ED7E381AFEFEF8BAFE67F871" ID-GBIF-Occurrence="3748446476" pageId="559" pageNumber="560" specimenCode="MNHN-IU-2014-21413" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54D6887FED7E381AFEFEF8BAFE66F84F" box="[315,419,1908,1934]" pageId="559" pageNumber="560" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D6BFD54ED7E381AFE71F8BAFE02F84F" box="[436,455,1908,1934]" pageId="559" pageNumber="560" type="female">♀</specimenCount>
, urosome partially damaged,
<specimenCode id="DBCB9EA6ED7E381AFF4DF858FE5FF871" box="[136,410,1942,1968]" pageId="559" pageNumber="560">MNHN-IU-2014-21413</specimenCode>
)
</materialsCitation>
,
<typeStatus id="54D6887FED7E381AFE75F858FDDEF871" box="[432,539,1942,1968]" pageId="559" pageNumber="560" type="paratype">paratypes</typeStatus>
(1 damaged
<specimenCount id="9D6BFD54ED7E381AFD7DF858FD08F871" box="[696,717,1942,1968]" pageId="559" pageNumber="560">♀</specimenCount>
with maxillae dissected and 5 intact copepodids, MNHN-IU-2014-21414), and dissected
<typeStatus id="54D6887FED7E381AFE03F814FDF4F835" box="[454,561,2010,2036]" pageId="559" pageNumber="560" type="paratype">paratypes</typeStatus>
(1 damaged
<specimenCount id="9D6BFD54ED7E381AFD00F814FD1FF835" box="[709,730,2010,2036]" pageId="559" pageNumber="560">♀</specimenCount>
and 1 copepodid, figured) from galls on the surface of brachial wall of
<taxonomicName id="4C6D4D5EED7E381AFCBAFB49FADEFB63" authority="Sluiter, 1885" authorityName="Sluiter" authorityYear="1885" box="[895,1307,1159,1186]" class="Ascidiacea" family="Styelidae" genus="Polycarpa" kingdom="Animalia" order="Pleurogona" pageId="559" pageNumber="560" phylum="Chordata" rank="species" species="cryptocarpa">
<emphasis id="B919EACFED7E381AFCBAFB49FB41FB60" box="[895,1156,1159,1185]" italics="true" pageId="559" pageNumber="560">Polycarpa cryptocarpa</emphasis>
Sluiter, 1885
</taxonomicName>
(MNHN-IT-2008-6441 =
<collectionCode id="ED7CAE18ED7E381AFC04FB64FBD2FB05" box="[961,1047,1194,1220]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="559" pageNumber="560">MNHN</collectionCode>
S1 POL.B 454); 6 copepodids from fibrillar networks on outer surface of one of galls,
<collectionCode id="ED7CAE18ED7E381AFAA1FB05FCBCFAC6" pageId="559" pageNumber="560">CRRF OCDN</collectionCode>
7521-K, near Port Vila,
<collectingCountry id="F37A764DED7E381AFB77FB20FACBFAC9" box="[1202,1294,1262,1288]" name="Vanuatu" pageId="559" pageNumber="560">Vanuatu</collectingCountry>
(
<geoCoordinate id="EE59501AED7E381AFAE1FB23FA62FAC9" box="[1316,1447,1261,1288]" degrees="17" direction="south" minutes="45.41" orientation="latitude" pageId="559" pageNumber="560" precision="9" value="-17.756834">17°45.41 Ś</geoCoordinate>
,
<geoCoordinate id="EE59501AED7E381AFCEFFAC1FC79FAEB" box="[810,956,1295,1322]" degrees="168" direction="east" minutes="16.90" orientation="longitude" pageId="559" pageNumber="560" precision="9" value="168.28166">168°16.90 É</geoCoordinate>
), depth
<quantity id="4C959B38ED7E381AFBD0FAC1FB95FAEB" box="[1045,1104,1295,1322]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="559" pageNumber="560" unit="m" value="12.0">12 m</quantity>
,
<date id="FFD3101DED7E381AFB99FAC1FAFDFAE8" box="[1116,1336,1295,1321]" pageId="559" pageNumber="560" value="2000-11-16">16 November 2000</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3776556ED7E381AFC9AFAFFFC46FA70" pageId="559" pageNumber="560" type="etymology">
<paragraph id="8BD236DDED7E381AFC9AFAFFFC46FA70" blockId="559.[810,1452,1159,2070]" pageId="559" pageNumber="560">
<emphasis id="B919EACFED7E381AFC9AFAFFFC27FA8A" bold="true" box="[863,994,1329,1355]" pageId="559" pageNumber="560">Etymology</emphasis>
. The specific name is derived from the Greek
<emphasis id="B919EACFED7E381AFC62FA9DFC2DFAAC" box="[935,1000,1363,1389]" italics="true" pageId="559" pageNumber="560">haplo</emphasis>
(=single) and
<emphasis id="B919EACFED7E381AFB5DFA9AFB01FAAC" box="[1176,1220,1364,1389]" italics="true" pageId="559" pageNumber="560">mer</emphasis>
(=apart), referring to the unsegmented endopod of the maxilla in the new species.
</paragraph>
</subSubSection>
<subSubSection id="C3776556ED7E3807FC9AFA77FA47FEB0" lastPageId="562" lastPageNumber="563" pageId="559" pageNumber="560" type="description">
<paragraph id="8BD236DDED7E381AFC9AFA77FB59F870" blockId="559.[810,1452,1159,2070]" pageId="559" pageNumber="560">
<emphasis id="B919EACFED7E381AFC9AFA77FBA0FA12" bold="true" box="[863,1125,1465,1491]" pageId="559" pageNumber="560">Descriptionoffemale</emphasis>
. Body (
<figureCitation id="13562A58ED7E381AFB03FA77FAF8FA15" box="[1222,1341,1465,1492]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="559" pageNumber="560">Fig. 375A</figureCitation>
) strongly curved ventrally, with weak exoskeleton; body length 2.12 mm. Prosome tapering anteriorly, divisible by 3 contrictionsinto cephalosome, firstand secondpedigerous somites; lateral margins of dorsal cephalic shield extended ventrally and concealing entire mouthparts in lateral view. Brood pouch formed by fused third and fourth pedigerous somites. Urosome slightly shorter than prosome, 6-segmented but first urosomite not articulated from prosome. Genitalsomite (second urosomite) short. Posterior part of anal somite ornamented with minute spinules. Caudal rami (
<figureCitation id="13562A58ED7E381AFB82F8FEFB07F88B" box="[1095,1218,1839,1866]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="559" pageNumber="560">Fig. 375B</figureCitation>
) small, 1.27 times longerthanwide (42×33 μm), rounded distally, covered with minute spinules; armed with 3 smallsetae (outer lateral, medioventral, and distal).
</paragraph>
<paragraph id="8BD236DDED7E3807FC9AF876FE1FFE16" blockId="559.[810,1452,1159,2070]" lastBlockId="562.[136,778,173,2070]" lastPageId="562" lastPageNumber="563" pageId="559" pageNumber="560">
Rostrum (
<figureCitation id="13562A58ED7E381AFC0AF876FB86F813" box="[975,1091,1975,2002]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="559" pageNumber="560">Fig. 375C</figureCitation>
) semicircular, rounded apically, wider than long, and covered with minute spinules. Antennule (
<figureCitation id="13562A58ED7E381AFC78F832FB85F7D7" box="[957,1088,2043,2070]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="559" pageNumber="560">Fig. 375D</figureCitation>
) shortand broad, leaf-like, 122×102 μm, unsegmented but with trace of articulation on posterior side; armed with several small setae plus minute surface spinules distally. Antenna (
<figureCitation id="13562A58ED633807FD4CFF3FFCC7FECD" box="[649,770,241,268]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="562" pageNumber="563">Fig. 375E</figureCitation>
) stout, 4-segmented; coxa and basis unarmed, each much wider than long; first endopodal segment also unarmed, as long as wide, its outer margin much shorter than inner margin; second endopodal segment about twice as long as wide; armed with 4 small setae plus small terminal claw, about half as long as segment.
</paragraph>
<caption id="DF126655ED613805FF4DF8BDFD4BF813" ID-DOI="http://doi.org/10.5281/zenodo.4592064" ID-Zenodo-Dep="4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="560" pageNumber="561" startId="560.[136,245,1907,1933]" targetBox="[233,1333,210,1848]" targetPageId="560">
<paragraph id="8BD236DDED613805FF4DF8BDFD4BF813" blockId="560.[136,1452,1907,2002]" pageId="560" pageNumber="561">
<emphasis id="B919EACFED613805FF4DF8BDFEEEF84F" bold="true" box="[136,299,1907,1934]" pageId="560" pageNumber="561">FIGURE 375.</emphasis>
<taxonomicName id="4C6D4D5EED613805FEF4F8BAFD99F84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[305,604,1908,1934]" class="Maxillopoda" family="Notodelphyidae" genus="Scoliosoma" kingdom="Animalia" order="Cyclopoida" pageId="560" pageNumber="561" phylum="Arthropoda" rank="species" species="haplomerosum" status="gen. et sp. nov.">
<emphasis id="B919EACFED613805FEF4F8BAFD99F84F" box="[305,604,1908,1934]" italics="true" pageId="560" pageNumber="561">Scoliosoma haplomerosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4ED613805FDA7F8BAFCCEF84F" box="[610,779,1908,1934]" pageId="560" pageNumber="561" rank="species">
<emphasis id="B919EACFED613805FDA7F8BAFCC3F84F" bold="true" box="[610,774,1908,1934]" pageId="560" pageNumber="561">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm.
</paragraph>
</caption>
<caption id="DF126655ED603804FF4DF8BDFE31F813" ID-DOI="http://doi.org/10.5281/zenodo.4592066" ID-Zenodo-Dep="4592066" httpUri="https://zenodo.org/record/4592066/files/figure.png" pageId="561" pageNumber="562" startId="561.[136,245,1907,1933]" targetBox="[228,1334,212,1845]" targetPageId="561">
<paragraph id="8BD236DDED603804FF4DF8BDFE31F813" blockId="561.[136,1451,1907,2002]" pageId="561" pageNumber="562">
<emphasis id="B919EACFED603804FF4DF8BDFEE9F84F" bold="true" box="[136,300,1907,1934]" pageId="561" pageNumber="562">FIGURE 376.</emphasis>
<taxonomicName id="4C6D4D5EED603804FEF7F8BAFD9BF84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[306,606,1908,1934]" class="Maxillopoda" family="Notodelphyidae" genus="Scoliosoma" kingdom="Animalia" order="Cyclopoida" pageId="561" pageNumber="562" phylum="Arthropoda" rank="species" species="haplomerosum" status="gen. et sp. nov.">
<emphasis id="B919EACFED603804FEF7F8BAFD9BF84F" box="[306,606,1908,1934]" italics="true" pageId="561" pageNumber="562">Scoliosoma haplomerosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4ED603804FDA0F8BAFCD4F84F" box="[613,785,1908,1934]" pageId="561" pageNumber="562" rank="species">
<emphasis id="B919EACFED603804FDA0F8BAFCCEF84F" bold="true" box="[613,779,1908,1934]" pageId="561" pageNumber="562">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, female: A, leg 3; B, leg 4. Copepodid V female: C, habitus, dorsal; D, habitus, left; E, urosome, dorsal; F, antennule; G, leg 1; H, leg 2; I, leg 3; J, leg 4. Scale bars: A, B, F–J, 0.02 mm; C, D, 0.1 mm; E, 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BD236DDED633807FC4FFE55FABFFDA1" pageId="562" pageNumber="563">
<table id="F96DC47DED63C5CAFCEFFE55FABFFDA1" box="[810,1402,411,608]" gridcols="5" gridrows="6" inLine="true" pageId="562" pageNumber="563">
<tr id="355D349FED63C5CAFCEFFE55FABFFE74" box="[810,1402,411,437]" gridrow="0" pageId="562" pageNumber="563" rowspan-0="1">
<th id="768C5DE3ED63C5CAFC4FFE55FC00FE74" box="[906,965,411,437]" gridcol="1" gridrow="0" pageId="562" pageNumber="563">Coxa</th>
<th id="768C5DE3ED63C5CAFC2FFE55FBE2FE74" box="[1002,1063,411,437]" gridcol="2" gridrow="0" pageId="562" pageNumber="563">Basis</th>
<th id="768C5DE3ED63C5CAFB8FFE55FB01FE74" box="[1098,1220,411,437]" gridcol="3" gridrow="0" pageId="562" pageNumber="563">Exopod</th>
<th id="768C5DE3ED63C5CAFACFFE55FABFFE74" box="[1290,1402,411,437]" gridcol="4" gridrow="0" pageId="562" pageNumber="563">Endopod</th>
</tr>
<tr id="355D349FED63C5CAFCEFFE73FABFFE19" box="[810,1402,445,472]" gridrow="1" pageId="562" pageNumber="563">
<th id="768C5DE3ED63C5CAFCEFFE73FCAFFE19" box="[810,874,445,472]" gridcol="0" gridrow="1" pageId="562" pageNumber="563">Leg 1</th>
<td id="768C5DE3ED63C5CAFC4FFE73FC00FE19" box="[906,965,445,472]" gridcol="1" gridrow="1" pageId="562" pageNumber="563">0-0</td>
<td id="768C5DE3ED63C5CAFC2FFE73FBE2FE19" box="[1002,1063,445,472]" gridcol="2" gridrow="1" pageId="562" pageNumber="563">1-0</td>
<td id="768C5DE3ED63C5CAFB8FFE73FB01FE19" box="[1098,1220,445,472]" gridcol="3" gridrow="1" pageId="562" pageNumber="563">1-0; 3, 1, 5</td>
<td id="768C5DE3ED63C5CAFACFFE73FABFFE19" box="[1290,1402,445,472]" gridcol="4" gridrow="1" pageId="562" pageNumber="563">1, 2, 4</td>
</tr>
<tr id="355D349FED63C5CAFCEFFE11FABFFE3B" box="[810,1402,479,506]" gridrow="2" pageId="562" pageNumber="563">
<th id="768C5DE3ED63C5CAFCEFFE11FCAFFE3B" box="[810,874,479,506]" gridcol="0" gridrow="2" pageId="562" pageNumber="563">Leg 2</th>
<td id="768C5DE3ED63C5CAFC4FFE11FC00FE3B" box="[906,965,479,506]" gridcol="1" gridrow="2" pageId="562" pageNumber="563">0-0</td>
<td id="768C5DE3ED63C5CAFC2FFE11FBE2FE3B" box="[1002,1063,479,506]" gridcol="2" gridrow="2" pageId="562" pageNumber="563">1-0</td>
<td id="768C5DE3ED63C5CAFB8FFE11FB01FE3B" box="[1098,1220,479,506]" gridcol="3" gridrow="2" pageId="562" pageNumber="563">1-0; 4, 1, 5</td>
<td id="768C5DE3ED63C5CAFACFFE11FABFFE3B" box="[1290,1402,479,506]" gridcol="4" gridrow="2" pageId="562" pageNumber="563">1, 2, 4</td>
</tr>
<tr id="355D349FED63C5CAFCEFFDCFFABFFDDD" box="[810,1402,513,540]" gridrow="3" pageId="562" pageNumber="563">
<th id="768C5DE3ED63C5CAFCEFFDCFFCAFFDDD" box="[810,874,513,540]" gridcol="0" gridrow="3" pageId="562" pageNumber="563">Leg 3</th>
<td id="768C5DE3ED63C5CAFC4FFDCFFC00FDDD" box="[906,965,513,540]" gridcol="1" gridrow="3" pageId="562" pageNumber="563">0-0</td>
<td id="768C5DE3ED63C5CAFC2FFDCFFBE2FDDD" box="[1002,1063,513,540]" gridcol="2" gridrow="3" pageId="562" pageNumber="563">1-0</td>
<td id="768C5DE3ED63C5CAFB8FFDCFFB01FDDD" box="[1098,1220,513,540]" gridcol="3" gridrow="3" pageId="562" pageNumber="563">1-0; 4, 1, 5</td>
<td id="768C5DE3ED63C5CAFACFFDCFFABFFDDD" box="[1290,1402,513,540]" gridcol="4" gridrow="3" pageId="562" pageNumber="563">1, 2, 3</td>
</tr>
<tr id="355D349FED63C5CAFCEFFDEDFABFFDFF" box="[810,1402,547,574]" gridrow="4" pageId="562" pageNumber="563">
<th id="768C5DE3ED63C5CAFCEFFDEDFCAFFDFF" box="[810,874,547,574]" gridcol="0" gridrow="4" pageId="562" pageNumber="563">Leg 4</th>
<td id="768C5DE3ED63C5CAFC4FFDEDFC00FDFF" box="[906,965,547,574]" gridcol="1" gridrow="4" pageId="562" pageNumber="563">0-0</td>
<td id="768C5DE3ED63C5CAFC2FFDEDFBE2FDFF" box="[1002,1063,547,574]" gridcol="2" gridrow="4" pageId="562" pageNumber="563">1-0</td>
<td id="768C5DE3ED63C5CAFB8FFDEDFB01FDFF" box="[1098,1220,547,574]" gridcol="3" gridrow="4" pageId="562" pageNumber="563">1-0; 4, 1, 5</td>
<td id="768C5DE3ED63C5CAFACFFDEDFABFFDFF" box="[1290,1402,547,574]" gridcol="4" gridrow="4" pageId="562" pageNumber="563">0, 2, 3</td>
</tr>
<tr id="355D349FED63C5CAFCEFFD8BFABFFDA1" box="[810,1402,581,608]" gridrow="5" pageId="562" pageNumber="563">
<th id="768C5DE3ED63C5CAFCEFFD8BFABFFDA1" box="[810,1402,581,608]" colspan="5" colspanRight="4" gridcol="0" gridrow="5" pageId="562" pageNumber="563">Legs 5 and 6 each represented by pair of small setae.</th>
</tr>
</table>
</paragraph>
<paragraph id="8BD236DDED633807FF78FE11FEE3FAC9" blockId="562.[136,778,173,2070]" pageId="562" pageNumber="563">
Labrum (
<figureCitation id="13562A58ED633807FEE3FE11FE50FE3B" box="[294,405,479,506]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="562" pageNumber="563">Fig. 375F</figureCitation>
) shortand broad, unornamented, with convex posterior margin. Mandible (
<figureCitation id="13562A58ED633807FD92FDCFFD0FFDDD" box="[599,714,513,540]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="562" pageNumber="563">Fig. 375G</figureCitation>
) with broad coxalgnathobase bearing along medial margin, strong distal tooth, uneven serrate middle region, and pointed proximal corner: palp (fused basis, exopod and endopod) bilobed distally; short outer lobe (exopod) with 4 naked setae; longer inner lobe (distal part of basis + unsegmented endopod) with 6 setae, proximal seta (representing medial seta on basis), second proximal seta (originally medial seta of first endopodal segment), and 4 setae from original second endopodal segment; first 2 proximal setae bearing few spinules on margins, other setae naked. Maxillule (
<figureCitation id="13562A58ED633807FE4BFCB9FDC5FC53" box="[398,512,887,914]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="562" pageNumber="563">Fig. 375H</figureCitation>
) not articulated, distally bilobed; outer lobe (palp, comprising fused basis, exopod and endopod) bearing 1 outer proximal and 4 distal setae, 1 distal seta weakly pinnate; inner lobe (precoxal arthrite) larger than outer lobe, armed with 5 pinnate setae on medial margin. Maxilla (
<figureCitation id="13562A58ED633807FE5EFBEFFDC6FBFD" box="[411,515,1057,1084]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="562" pageNumber="563">Fig. 375I</figureCitation>
) 3-segmented; syncoxa with 2 endites bearing 2 setae on first endite and 1 large seta on second; basis with robust claw ornamented with serrate membrane in distal third; unsegmented endopod with 2 largeand 2 small setae, all naked. Maxilliped (
<figureCitation id="13562A58ED633807FD19FB67FF7BFB27" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="562" pageNumber="563">Fig. 375J</figureCitation>
) as distally narrowing lobe bearing 2 weakly pinnate setaeon apex.
</paragraph>
<paragraph id="8BD236DDED633807FF78FADEFD69F99A" blockId="562.[136,778,173,2070]" pageId="562" pageNumber="563">
Legs 1–4 (
<figureCitation id="13562A58ED633807FE84FAC1FE28FAEB" box="[321,493,1295,1322]" captionStart="FIGURE 375" captionStartId="560.[136,245,1907,1933]" captionTargetBox="[233,1333,210,1848]" captionTargetId="figure-19@560.[195,1392,185,1870]" captionTargetPageId="560" captionText="FIGURE 375. Scoliosoma haplomerosum gen. et sp. nov., female.A, habitus, right; B, caudal rami, dorsal; C, rostrum; D, antennule; E, antenna; F, labrum; G, mandible; H, maxillule; I, maxilla; J, maxilliped; K, leg 1; L, leg 2. Scale bars: A, 0.2 mm; B, C, E–L, 0.02 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592064" httpUri="https://zenodo.org/record/4592064/files/figure.png" pageId="562" pageNumber="563">Figs. 375K, L</figureCitation>
,
<figureCitation id="13562A58ED633807FE3AFAC1FDA0FAEB" box="[511,613,1295,1322]" captionStart="FIGURE 376" captionStartId="561.[136,245,1907,1933]" captionTargetBox="[228,1334,212,1845]" captionTargetId="figure-17@561.[195,1393,185,1870]" captionTargetPageId="561" captionText="FIGURE 376. Scoliosoma haplomerosum gen. et sp. nov., female: A, leg 3; B, leg 4. Copepodid V female: C, habitus, dorsal; D, habitus, left; E, urosome, dorsal; F, antennule; G, leg 1; H, leg 2; I, leg 3; J, leg 4. Scale bars: A, B, F–J, 0.02 mm; C, D, 0.1 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592066" httpUri="https://zenodo.org/record/4592066/files/figure.png" pageId="562" pageNumber="563">376A, B</figureCitation>
) rudimentary, lacking defined protopod. Legs 1–3 bilobed, unsegmented; both rami covered with minute spinules. Inner lobe (endopod) as large as outer lobe (exopod) in leg 1, but much smaller than outer lobe in legs 2 and 3. Leg 4 represented by single lobe (exopod). Legs 1 and 2 with 1 outer proximal seta on outer lobe; this seta absent in legs 3 and 4. Other setae 1, 3, 5, and 7 on exopods of legs 1–4, respectively, and 0, 2, and 3 on endopods of legs 1–3, respectively; all setae rudimentary. Leg 5 absent.
</paragraph>
<paragraph id="8BD236DDED633807FF78F9AAFEBBF9BC" blockId="562.[136,778,173,2070]" box="[189,382,1635,1662]" pageId="562" pageNumber="563">
<emphasis id="B919EACFED633807FF78F9AAFF3FF9BF" bold="true" box="[189,250,1636,1662]" pageId="562" pageNumber="563">Male</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BD236DDED633807FF78F94BFA47FEB0" blockId="562.[136,778,173,2070]" lastBlockId="562.[810,1452,173,369]" pageId="562" pageNumber="563">
<emphasis id="B919EACFED633807FF78F94BFDB2F95E" bold="true" box="[189,631,1669,1696]" pageId="562" pageNumber="563">Description of Copepodid V female</emphasis>
. Body (
<figureCitation id="13562A58ED633807FD19F94BFF29F903" captionStart="FIGURE 376" captionStartId="561.[136,245,1907,1933]" captionTargetBox="[228,1334,212,1845]" captionTargetId="figure-17@561.[195,1393,185,1870]" captionTargetPageId="561" captionText="FIGURE 376. Scoliosoma haplomerosum gen. et sp. nov., female: A, leg 3; B, leg 4. Copepodid V female: C, habitus, dorsal; D, habitus, left; E, urosome, dorsal; F, antennule; G, leg 1; H, leg 2; I, leg 3; J, leg 4. Scale bars: A, B, F–J, 0.02 mm; C, D, 0.1 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592066" httpUri="https://zenodo.org/record/4592066/files/figure.png" pageId="562" pageNumber="563">Fig. 376C, D</figureCitation>
) not swollen, slightlycurvedventrally, poorly segmented, much smaller than adult (body length of figured specimen 533 μm). Cephalic region widest. Prosome-urosome division obscure. Fifth pedigerous and genital somites (
<figureCitation id="13562A58ED633807FEB1F8FEFE22F88B" box="[372,487,1839,1866]" captionStart="FIGURE 376" captionStartId="561.[136,245,1907,1933]" captionTargetBox="[228,1334,212,1845]" captionTargetId="figure-17@561.[195,1393,185,1870]" captionTargetPageId="561" captionText="FIGURE 376. Scoliosoma haplomerosum gen. et sp. nov., female: A, leg 3; B, leg 4. Copepodid V female: C, habitus, dorsal; D, habitus, left; E, urosome, dorsal; F, antennule; G, leg 1; H, leg 2; I, leg 3; J, leg 4. Scale bars: A, B, F–J, 0.02 mm; C, D, 0.1 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592066" httpUri="https://zenodo.org/record/4592066/files/figure.png" pageId="562" pageNumber="563">Fig. 376E</figureCitation>
) not articulated from one another.Abdomen 2-segmented in dorsal view, but second abdominal somite with additional suture line on ventral surface. Caudal rami 2.3 times longer than wide (37×16 μm); armedwith 3 smallsetae. Antennule (
<figureCitation id="13562A58ED633807FD4BF876FCC7F812" box="[654,770,1976,2003]" captionStart="FIGURE 376" captionStartId="561.[136,245,1907,1933]" captionTargetBox="[228,1334,212,1845]" captionTargetId="figure-17@561.[195,1393,185,1870]" captionTargetPageId="561" captionText="FIGURE 376. Scoliosoma haplomerosum gen. et sp. nov., female: A, leg 3; B, leg 4. Copepodid V female: C, habitus, dorsal; D, habitus, left; E, urosome, dorsal; F, antennule; G, leg 1; H, leg 2; I, leg 3; J, leg 4. Scale bars: A, B, F–J, 0.02 mm; C, D, 0.1 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592066" httpUri="https://zenodo.org/record/4592066/files/figure.png" pageId="562" pageNumber="563">Fig. 376F</figureCitation>
) much narrower than in adult, 4-segmented; armature formula 13, 11, 3, and 14; setae distinct.Antenna, labrum, mandible, maxillule, maxilla, and maxilliped as in adult female. Legs 1–4 (
<figureCitation id="13562A58ED633807FBD7FF01FBBFFF2B" box="[1042,1146,207,234]" captionStart="FIGURE 376" captionStartId="561.[136,245,1907,1933]" captionTargetBox="[228,1334,212,1845]" captionTargetId="figure-17@561.[195,1393,185,1870]" captionTargetPageId="561" captionText="FIGURE 376. Scoliosoma haplomerosum gen. et sp. nov., female: A, leg 3; B, leg 4. Copepodid V female: C, habitus, dorsal; D, habitus, left; E, urosome, dorsal; F, antennule; G, leg 1; H, leg 2; I, leg 3; J, leg 4. Scale bars: A, B, F–J, 0.02 mm; C, D, 0.1 mm; E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4592066" httpUri="https://zenodo.org/record/4592066/files/figure.png" pageId="562" pageNumber="563">Fig. 376</figureCitation>
G-J) biramous, each with unsegmented or partially segmented protopod, and welldefined exopod and endopod; exopods incompletely 2- segmented, and endopods unsegmented; setae distinct and naked. Armature formula for legs 1–4 as follows:
</paragraph>
</subSubSection>
<subSubSection id="C3776556ED633807FC9AFD47FAE2F8C7" pageId="562" pageNumber="563" type="discussion">
<paragraph id="8BD236DDED633807FC9AFD47FAA6FC8C" blockId="562.[810,1452,649,1798]" pageId="562" pageNumber="563">
<emphasis id="B919EACFED633807FC9AFD47FC09FD62" bold="true" box="[863,972,649,675]" pageId="562" pageNumber="563">Remarks</emphasis>
. The galls inhabited by the adult females were flattened and about 1.2 mm in diameter, with irregular margins. Each gall contained a single female and one was found with its anal somite protruding out of the gall. Six copepodids discovered were among fibrillary network on the external surface of one of the galls.
</paragraph>
<paragraph id="8BD236DDED633807FC9AFC9BFA6EFAAC" blockId="562.[810,1452,649,1798]" pageId="562" pageNumber="563">
<taxonomicName id="4C6D4D5EED633807FC9AFC9BFBB2FCAE" authorityName="Sluiter" authorityYear="1885" box="[863,1143,853,879]" class="Ascidiacea" family="Styelidae" genus="Polycarpa" kingdom="Animalia" order="Pleurogona" pageId="562" pageNumber="563" phylum="Chordata" rank="species" species="cryptocarpa">
<emphasis id="B919EACFED633807FC9AFC9BFBB2FCAE" box="[863,1143,853,879]" italics="true" pageId="562" pageNumber="563">Polycarpa cryptocarpa</emphasis>
</taxonomicName>
, the ascidian host of
<taxonomicName id="4C6D4D5EED633807FCEFFCB9FB9DFC50" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[810,1112,887,913]" class="Maxillopoda" family="Notodelphyidae" genus="Scoliosoma" kingdom="Animalia" order="Cyclopoida" pageId="562" pageNumber="563" phylum="Arthropoda" rank="species" species="haplomerosum" status="gen. et sp. nov.">
<emphasis id="B919EACFED633807FCEFFCB9FB9DFC50" box="[810,1112,887,913]" italics="true" pageId="562" pageNumber="563">Scoliosoma haplomerosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4ED633807FBA4FCB9FAD0FC53" box="[1121,1301,887,914]" pageId="562" pageNumber="563" rank="species">
<emphasis id="B919EACFED633807FBA4FCB9FACAFC50" bold="true" box="[1121,1295,887,913]" pageId="562" pageNumber="563">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, is known to harbour several other species of ascidicolous copepods. One of them,
<taxonomicName id="4C6D4D5EED633807FC0CFC75FA62FC14" authority="Sluiter, 1884" authorityName="Sluiter" authorityYear="1884" box="[969,1447,955,981]" class="Maxillopoda" family="Notodelphyidae" genus="Sphaerothylacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="562" pageNumber="563" phylum="Arthropoda" rank="species" species="polycarpae">
<emphasis id="B919EACFED633807FC0CFC75FACFFC14" box="[969,1290,955,981]" italics="true" pageId="562" pageNumber="563">Sphaerothylacus polycarpae</emphasis>
<bibRefCitation id="EFFC4B2CED633807FAD6FC75FA62FC14" author="Sluiter, C. P." box="[1299,1447,955,981]" journalOrPublisher="Natuurkundig Tijdschrift voor Nederlandsch- Indie" pageId="562" pageNumber="563" pagination="201 - 224" part="43" refId="ref330965" refString="Sluiter, C. P. (1884) Uber einen in Ascidien schmarotzenden Wurzelkrebs. Natuurkundig Tijdschrift voor Nederlandsch- Indie, 43 (4), 201 - 224, pl. 1." title="Uber einen in Ascidien schmarotzenden Wurzelkrebs" type="journal article" year="1884">Sluiter, 1884</bibRefCitation>
</taxonomicName>
, is remarkable as it was first recognized as a cirripede by its original describer (
<bibRefCitation id="EFFC4B2CED633807FBE3FC31FB78FBD8" author="Sluiter, C. P." box="[1062,1213,1023,1049]" journalOrPublisher="Natuurkundig Tijdschrift voor Nederlandsch- Indie" pageId="562" pageNumber="563" pagination="201 - 224" part="43" refId="ref330965" refString="Sluiter, C. P. (1884) Uber einen in Ascidien schmarotzenden Wurzelkrebs. Natuurkundig Tijdschrift voor Nederlandsch- Indie, 43 (4), 201 - 224, pl. 1." title="Uber einen in Ascidien schmarotzenden Wurzelkrebs" type="journal article" year="1884">Sluiter, 1884</bibRefCitation>
), but
<bibRefCitation id="EFFC4B2CED633807FAC5FC31FA6EFBD8" author="Grygier, M. J." box="[1280,1451,1023,1049]" journalOrPublisher="Monoculus Copepod Newsletter, Oldenburg" pageId="562" pageNumber="563" pagination="23 - 25" part="25" refId="ref328298" refString="Grygier, M. J. (1993) Sphaerothylacus is a notodelphyid copepod, not a parasitic cirripede. Monoculus Copepod Newsletter, Oldenburg, 25, 23 - 25." title="Sphaerothylacus is a notodelphyid copepod, not a parasitic cirripede" type="journal article" year="1993">Grygier (1993)</bibRefCitation>
later concluded that it was a copepod. The absence of frontolateral horns in the nauplii of
<taxonomicName id="4C6D4D5EED633807FACFFB8AFA6EFB9C" authorityName="Sluiter" authorityYear="1884" box="[1290,1451,1091,1118]" class="Maxillopoda" family="Notodelphyidae" genus="Sphaerothylacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="562" pageNumber="563" phylum="Arthropoda" rank="species" species="polycarpae">
<emphasis id="B919EACFED633807FACFFB8AFADDFB9F" box="[1290,1304,1092,1118]" italics="true" pageId="562" pageNumber="563">S</emphasis>
.
<emphasis id="B919EACFED633807FAE8FB8DFA6EFB9C" box="[1325,1451,1091,1117]" italics="true" pageId="562" pageNumber="563">polycarpae</emphasis>
</taxonomicName>
illustrated by
<bibRefCitation id="EFFC4B2CED633807FC14FBABFBB0FBBE" author="Sluiter, C. P." box="[977,1141,1125,1151]" journalOrPublisher="Natuurkundig Tijdschrift voor Nederlandsch- Indie" pageId="562" pageNumber="563" pagination="201 - 224" part="43" refId="ref330965" refString="Sluiter, C. P. (1884) Uber einen in Ascidien schmarotzenden Wurzelkrebs. Natuurkundig Tijdschrift voor Nederlandsch- Indie, 43 (4), 201 - 224, pl. 1." title="Uber einen in Ascidien schmarotzenden Wurzelkrebs" type="journal article" year="1884">Sluiter (1884)</bibRefCitation>
indicates that his species cannot be classified as a cirripede, but the presence of transverse sculpturing appearing to represent articulations on the hind body of the nauplii is unusual for copepods.
<taxonomicName id="4C6D4D5EED633807FCEFFB23FBB3FAC6" authorityName="Sluiter" authorityYear="1884" box="[810,1142,1261,1287]" class="Maxillopoda" family="Notodelphyidae" genus="Sphaerothylacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="562" pageNumber="563" phylum="Arthropoda" rank="species" species="polycarpae">
<emphasis id="B919EACFED633807FCEFFB23FBB3FAC6" box="[810,1142,1261,1287]" italics="true" pageId="562" pageNumber="563">Sphaerothylacus polycarpae</emphasis>
</taxonomicName>
is not confusable with
<taxonomicName id="4C6D4D5EED633807FCEFFAC1FB9DFAE8" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[810,1112,1295,1321]" class="Maxillopoda" family="Notodelphyidae" genus="Scoliosoma" kingdom="Animalia" order="Cyclopoida" pageId="562" pageNumber="563" phylum="Arthropoda" rank="species" species="haplomerosum" status="gen. et sp. nov.">
<emphasis id="B919EACFED633807FCEFFAC1FB9DFAE8" box="[810,1112,1295,1321]" italics="true" pageId="562" pageNumber="563">Scoliosoma haplomerosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4ED633807FBA5FAC1FAD4FAEB" box="[1120,1297,1295,1322]" pageId="562" pageNumber="563" rank="species">
<emphasis id="B919EACFED633807FBA5FAC1FACEFAE8" bold="true" box="[1120,1291,1295,1321]" pageId="562" pageNumber="563">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, because it is an almost spherical parasite with a root-like holdfast and a pore on the opposite side of the body from the holdfast.
</paragraph>
<paragraph id="8BD236DDED633807FC9AFABBFAE2F8C7" blockId="562.[810,1452,649,1798]" pageId="562" pageNumber="563">
Another remarkably transformed copepod associate of
<taxonomicName id="4C6D4D5EED633807FCEFFA56FC15FA70" authorityName="Sluiter" authorityYear="1885" box="[810,976,1432,1458]" class="Ascidiacea" family="Styelidae" genus="Polycarpa" kingdom="Animalia" order="Pleurogona" pageId="562" pageNumber="563" phylum="Chordata" rank="species" species="cryptocarpa">
<emphasis id="B919EACFED633807FCEFFA56FCFEFA70" box="[810,827,1432,1457]" italics="true" pageId="562" pageNumber="563">P</emphasis>
.
<emphasis id="B919EACFED633807FC8FFA56FC15FA70" box="[842,976,1432,1457]" italics="true" pageId="562" pageNumber="563">cryptocarpa</emphasis>
</taxonomicName>
is
<taxonomicName id="4C6D4D5EED633807FC30FA59FA63FA73" authority="Gotto, 1981" authorityName="Gotto" authorityYear="1981" box="[1013,1446,1431,1458]" class="Maxillopoda" family="Notodelphyidae" genus="Paralobodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="562" pageNumber="563" phylum="Arthropoda" rank="species" species="setipoda">
<emphasis id="B919EACFED633807FC30FA59FAD3FA70" box="[1013,1302,1431,1457]" italics="true" pageId="562" pageNumber="563">Paralobodelphys setipoda</emphasis>
<bibRefCitation id="EFFC4B2CED633807FADAFA56FA63FA73" author="Gotto, R. V." box="[1311,1446,1432,1458]" journalOrPublisher="Crustaceana" pageId="562" pageNumber="563" pagination="173 - 177" part="40" refId="ref328154" refString="Gotto, R. V. (1981) Paralobodelphys setipoda n. gen. n. sp. (Cyclopoida, Notodelphyidae): an ascidicolous copepod from the Great Barrier Reef, Australia. Crustaceana, 40 (2), 173 - 177. https: // doi. org / 10.1163 / 156854081 X 00570" title="Paralobodelphys setipoda n. gen. n. sp. (Cyclopoida, Notodelphyidae): an ascidicolous copepod from the Great Barrier Reef, Australia" type="journal article" year="1981">Gotto, 1981</bibRefCitation>
</taxonomicName>
, which was described as a new genus and species from the Great Barrier Reef,
<collectingCountry id="F37A764DED633807FB80FA15FB68FA34" box="[1093,1197,1499,1525]" name="Australia" pageId="562" pageNumber="563">Australia</collectingCountry>
. The specimens were “contained in a circular area” (Gotto, 1881), suggesting that the species is a gall-inhabitant as is
<taxonomicName id="4C6D4D5EED633807FB29F9EEFA6EF9F8" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1260,1451,1567,1594]" class="Maxillopoda" family="Notodelphyidae" genus="Scoliosoma" kingdom="Animalia" order="Cyclopoida" pageId="562" pageNumber="563" phylum="Arthropoda" rank="species" species="haplomerosum" status="gen. et sp. nov.">
<emphasis id="B919EACFED633807FB29F9EEFB3FF9FB" box="[1260,1274,1568,1594]" italics="true" pageId="562" pageNumber="563">S</emphasis>
.
<emphasis id="B919EACFED633807FACCF9D1FA6EF9F8" box="[1289,1451,1567,1593]" italics="true" pageId="562" pageNumber="563">haplomerosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4ED633807FCEFF98CFC19F99D" box="[810,988,1602,1628]" pageId="562" pageNumber="563" rank="species">
<emphasis id="B919EACFED633807FCEFF98CFC13F99D" bold="true" box="[810,982,1602,1628]" pageId="562" pageNumber="563">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
Although this species has a body similar to that of
<taxonomicName id="4C6D4D5EED633807FC5BF9AAFB9AF9BC" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[926,1119,1635,1662]" class="Maxillopoda" family="Notodelphyidae" genus="Scoliosoma" kingdom="Animalia" order="Cyclopoida" pageId="562" pageNumber="563" phylum="Arthropoda" rank="species" species="haplomerosum" status="gen. et sp. nov">
<emphasis id="B919EACFED633807FC5BF9AAFC69F9BF" box="[926,940,1636,1662]" italics="true" pageId="562" pageNumber="563">S</emphasis>
.
<emphasis id="B919EACFED633807FC78F9ADFB9AF9BC" box="[957,1119,1635,1661]" italics="true" pageId="562" pageNumber="563">haplomerosum</emphasis>
</taxonomicName>
<emphasis id="B919EACFED633807FBAFF9AAFADEF9BF" bold="true" box="[1130,1307,1636,1662]" pageId="562" pageNumber="563">
<taxonomicNameLabel id="A22A57B4ED633807FBAFF9AAFADEF9BF" box="[1130,1307,1636,1662]" pageId="562" pageNumber="563" rank="species">gen. et sp. nov</taxonomicNameLabel>
</emphasis>
, its cephalic appendages and legs are quite different from those of
<taxonomicName id="4C6D4D5EED633807FA52F948FC09F900" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" class="Maxillopoda" family="Notodelphyidae" genus="Scoliosoma" kingdom="Animalia" order="Cyclopoida" pageId="562" pageNumber="563" phylum="Arthropoda" rank="species" species="haplomerosum" status="gen. et sp. nov.">
<emphasis id="B919EACFED633807FA52F948FA60F961" box="[1431,1445,1670,1696]" italics="true" pageId="562" pageNumber="563">S</emphasis>
.
<emphasis id="B919EACFED633807FCEFF969FC09F900" box="[810,972,1703,1729]" italics="true" pageId="562" pageNumber="563">haplomerosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4ED633807FC16F966FBBAF903" box="[979,1151,1704,1730]" pageId="562" pageNumber="563" rank="species">
<emphasis id="B919EACFED633807FC16F966FBBDF903" bold="true" box="[979,1144,1704,1730]" pageId="562" pageNumber="563">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
For example, the mandible lacks a coxal gnathobase, the maxilla is only 2-segmented, and the legs have elongate rami in
<taxonomicName id="4C6D4D5EED633807FB60F922FAE4F8C4" authorityName="Gotto" authorityYear="1981" box="[1189,1313,1771,1798]" class="Maxillopoda" family="Notodelphyidae" genus="Paralobodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="562" pageNumber="563" phylum="Arthropoda" rank="species" species="setipoda">
<emphasis id="B919EACFED633807FB60F922FB73F8C4" box="[1189,1206,1772,1797]" italics="true" pageId="562" pageNumber="563">P</emphasis>
.
<emphasis id="B919EACFED633807FB01F925FAE4F8C4" box="[1220,1313,1771,1797]" italics="true" pageId="562" pageNumber="563">setipoda</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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