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<mods:title id="DAD24D41AA807F27122C9E34E0FC08EF">Notes on Indian wolf spiders: I. Genus Evippa Simon, 1882 (Araneae: Lycosidae Evippinae)</mods:title>
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<mods:affiliation id="1DB336C1E56498EBC594148F62D23B86">Zoological Survey of India, Prani Vigyan Bhawan, M-Block, New Alipore, Kolkata, West Bengal 700 053, India. caleb 87 woodgate @ gmail. com; https: // orcid. org / 0000 - 0002 - 9471 - 9467</mods:affiliation>
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<paragraph id="8BD236D2FFEF2E3BFF3AF9F2FD405187" blockId="10.[151,735,1657,1719]" box="[151,735,1657,1684]" pageId="10" pageNumber="169">
<heading id="D09A81BEFFEF2E3BFF3AF9F2FD405187" bold="true" box="[151,735,1657,1684]" fontSize="11" level="1" pageId="10" pageNumber="169" reason="1">
<taxonomicName id="4C6D4D51FFEF2E3BFF3AF9F2FD405187" ID-CoL="5BSRJ" authority="(O. Pickard-Cambridge, 1872)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[151,735,1657,1684]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="169" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B919EAC0FFEF2E3BFF3AF9F2FD405187" bold="true" box="[151,735,1657,1684]" pageId="10" pageNumber="169">
<emphasis id="B919EAC0FFEF2E3BFF3AF9F2FEC45180" bold="true" box="[151,347,1657,1684]" italics="true" pageId="10" pageNumber="169">Wadicosa fidelis</emphasis>
(O.
<bibRefCitation id="EFFC4B23FFEF2E3BFE3FF9F2FD485187" author="Pickard-Cambridge, O." box="[402,727,1657,1683]" pageId="10" pageNumber="169" pagination="212 - 354" refId="ref6463" refString="Pickard-Cambridge, O. (1872) General list of the spiders of Palestine and Syria, with descriptions of numerous new species, and characters of two new genera. Proceedings of the Zoological Society of London, 40, 212 - 354, pls. 13 - 16." type="journal article" year="1872">Pickard-Cambridge, 1872</bibRefCitation>
)
</emphasis>
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<figureCitation id="13562A57FFEF2E3BFF3AF916FF7851A3" box="[151,231,1693,1719]" captionStart="FIGURE 10" captionStartId="12.[151,250,1673,1697]" captionTargetBox="[155,1429,186,1642]" captionTargetId="figure-21@12.[151,1436,181,1649]" captionTargetPageId="12" captionText="FIGURE 10. Wadicosa fidelis (O. Pickard-Cambridge, 1872), holotype of Evippa mandlaensis Gajbe, 2004. A female habitus (Ar-1621), dorsal. BC genitalia: B epigynum, ventral; C vulva, dorsal. D label from type tube. Scale bars: A, 2 mm; BC, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4805805" httpUri="https://zenodo.org/record/4805805/files/figure.png" pageId="10" pageNumber="169">Fig. 10</figureCitation>
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<treatmentCitationGroup id="AB7D11FCFFEF2E3BFF3AF968FD1751EE" box="[151,648,1762,1786]" pageId="10" pageNumber="169">
<treatmentCitation id="0ACC10C3FFEF2E3BFF3AF968FD1B51EE" author="Pickard-Cambridge, O." box="[151,644,1762,1786]" page="319" pageId="10" pageNumber="169" year="1872">
<taxonomicName id="4C6D4D51FFEF2E3BFF3AF968FD1B51EE" authority="O. Pickard-Cambridge, 1872: 319" authorityName="O. Pickard-Cambridge" authorityPageNumber="319" authorityYear="1872" box="[151,644,1762,1786]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="169" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B919EAC0FFEF2E3BFF3AF968FEBC51EE" box="[151,291,1762,1786]" italics="true" pageId="10" pageNumber="169">Lycosa fidelis</emphasis>
O.
<bibRefCitation id="EFFC4B23FFEF2E3BFEE5F969FD1B51EE" author="Pickard-Cambridge, O." box="[328,644,1762,1786]" pageId="10" pageNumber="169" pagination="212 - 354" refId="ref6463" refString="Pickard-Cambridge, O. (1872) General list of the spiders of Palestine and Syria, with descriptions of numerous new species, and characters of two new genera. Proceedings of the Zoological Society of London, 40, 212 - 354, pls. 13 - 16." type="journal article" year="1872">Pickard-Cambridge, 1872: 319</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BD236D2FFEF2E3BFF3AF88BFC8F5023" blockId="10.[151,1417,1762,1847]" pageId="10" pageNumber="169">
<treatmentCitationGroup id="AB7D11FCFFEF2E3BFF3AF88BFC8F5023" pageId="10" pageNumber="169">
<treatmentCitation id="0ACC10C3FFEF2E3BFF3AF88BFD16500D" author="Kronestedt, T. &amp; Zyuzin, A. A." box="[151,649,1792,1817]" page="818" pageId="10" pageNumber="169" year="2009">
<taxonomicName id="4C6D4D51FFEF2E3BFF3AF88BFD16500D" ID-CoL="5BSRJ" authority="Kronestedt &amp; Zyuzin 2009: 818" authorityName="Kronestedt &amp; Zyuzin" authorityPageNumber="818" authorityYear="2009" box="[151,649,1792,1817]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="169" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B919EAC0FFEF2E3BFF3AF88BFEA2500C" box="[151,317,1792,1816]" italics="true" pageId="10" pageNumber="169">Wadicosa fidelis</emphasis>
<bibRefCitation id="EFFC4B23FFEF2E3BFEEEF88BFD16500D" author="Kronestedt, T. &amp; Zyuzin, A. A." box="[323,649,1792,1817]" pageId="10" pageNumber="169" pagination="813 - 828" refId="ref6175" refString="Kronestedt, T. &amp; Zyuzin, A. A. (2009) Fixation of Lycosa fidelis O. Pickard-Cambridge, 1872 as the type species for the genus Wadicosa Zyuzin, 1985 (Araneae: Lycosidae), with a redescripton of the species. Contributions to Natural History, 12, 813 - 828. [incl. corrigendum: Vol. 13 / 2010: 89]" type="journal article" year="2009">Kronestedt &amp; Zyuzin 2009: 818</bibRefCitation>
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, figs 120 (for complete list of references, see
<bibRefCitation id="EFFC4B23FFEF2E3BFBC9F88BFAE1500D" author="World Spider Catalog" box="[1124,1406,1792,1817]" pageId="10" pageNumber="169" refId="ref6906" refString="World Spider Catalog (2021) World Spider Catalog. Version 22. Natural History Museum Bern, Bern. https: // doi. org / 10.24436 / 2" type="book" year="2021">World Spider Catalog 2021</bibRefCitation>
).
<treatmentCitation id="0ACC10C3FFEF2E3BFF3AF8ABFD9E5023" author="Gajbe, U. A." box="[151,513,1823,1847]" page="8" pageId="10" pageNumber="169" year="2004">
<taxonomicName id="4C6D4D51FFEF2E3BFF3AF8ABFD9E5023" authority="Gajbe, 2004: 8" authorityName="Gajbe" authorityPageNumber="8" authorityYear="2004" box="[151,513,1823,1847]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="169" phylum="Arthropoda" rank="species" species="mandlaensis">
<emphasis id="B919EAC0FFEF2E3BFF3AF8ABFEFB5023" box="[151,356,1823,1847]" italics="true" pageId="10" pageNumber="169">Evippa mandlaensis</emphasis>
<bibRefCitation id="EFFC4B23FFEF2E3BFEC7F894FD9E5023" author="Gajbe, U. A." box="[362,513,1823,1847]" pageId="10" pageNumber="169" pagination="1 - 40" refId="ref6099" refString="Gajbe, U. A. (2004) Studies on some spiders of the family Lycosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India, Occasional Paper, 221, 1 - 40." type="journal article" year="2004">Gajbe, 2004: 8</bibRefCitation>
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, figs 69.
<emphasis id="B919EAC0FFEF2E3BFDC1F894FC8F5023" bold="true" box="[620,784,1823,1847]" pageId="10" pageNumber="169">New synonymy</emphasis>
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</paragraph>
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<emphasis id="B919EAC0FFEF2E3BFF3AF8EEFD98506B" bold="true" box="[151,519,1893,1920]" pageId="10" pageNumber="169">
Type material.
<taxonomicName id="4C6D4D51FFEF2E3BFEE0F8EDFD9F5094" authorityName="Gajbe" authorityYear="2004" box="[333,512,1893,1920]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="169" phylum="Arthropoda" rank="species" species="mandlaensis">
<emphasis id="B919EAC0FFEF2E3BFEE0F8EDFD9F5094" bold="true" box="[333,512,1893,1920]" italics="true" pageId="10" pageNumber="169">E. mandlaensis</emphasis>
</taxonomicName>
.
</emphasis>
Female
<typeStatus id="54D68870FFEF2E3BFDC5F8EEFD55506B" box="[616,714,1893,1919]" pageId="10" pageNumber="169" type="holotype">holotype</typeStatus>
from
<collectingCountry id="F37A7642FFEF2E3BFCBDF8EDFCFB5094" box="[784,868,1894,1920]" name="India" pageId="10" pageNumber="169">INDIA</collectingCountry>
:
<collectingRegion id="49A9F830FFEF2E3BFCC2F8EEFBAF506B" box="[879,1072,1893,1919]" country="India" name="Madhya Pradesh" pageId="10" pageNumber="169">Madhya Pradesh</collectingRegion>
: Mandla (= Mandala): Narayanganj (
<geoCoordinate id="EE595015FFEF2E3BFF75F802FE1350B7" box="[216,396,1929,1955]" degrees="22" direction="north" minutes="49" orientation="latitude" pageId="10" pageNumber="169" precision="1" seconds="38.41" value="22.827337">22°4938.41N</geoCoordinate>
,
<geoCoordinate id="EE595015FFEF2E3BFE34F802FDD550B7" box="[409,586,1929,1955]" degrees="80" direction="east" minutes="14" orientation="longitude" pageId="10" pageNumber="169" precision="1" seconds="10.73" value="80.23631">80°1410.73E</geoCoordinate>
),
<quantity id="4C959B37FFEF2E3BFDF2F802FD3750B0" box="[607,680,1929,1956]" metricMagnitude="2" metricUnit="m" metricValue="4.31" pageId="10" pageNumber="169" unit="m" value="431.0">431 m</quantity>
alt.; U.A. Gajbe leg.;
<date id="FFD31012FFEF2E3BFC0AF802FBF850B0" box="[935,1127,1929,1956]" pageId="10" pageNumber="169" value="1985-01-23">23 January 1985</date>
; repository NZC-ZSI, Kolkata (no register number specified), examined.
<emphasis id="B919EAC0FFEF2E3BFD1DF825FCBC50D3" bold="true" box="[688,803,1965,1992]" pageId="10" pageNumber="169">
<taxonomicName id="4C6D4D51FFEF2E3BFD1DF825FC8350DC" authorityName="Kronestedt &amp; Zyuzin" authorityYear="2009" box="[688,796,1965,1992]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="169" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B919EAC0FFEF2E3BFD1DF825FC8350DC" bold="true" box="[688,796,1965,1992]" italics="true" pageId="10" pageNumber="169">W. fidelis</emphasis>
</taxonomicName>
.
</emphasis>
<typeStatus id="54D68870FFEF2E3BFC81F825FC0350DC" box="[812,924,1966,1992]" pageId="10" pageNumber="169" type="lectotype">Lectotype</typeStatus>
male from Palestine (
<collectionCode id="ED7CAE17FFEF2E3BFB05F826FA9D50D3" box="[1192,1282,1965,1991]" pageId="10" pageNumber="169">OXUM</collectionCode>
, collection of O. Pickard-Cambridge: bottle 1572, tube 39), not examined.
<bibRefCitation id="EFFC4B23FFEF2E3BFCE5F85AFB1450FF" author="Kronestedt, T. &amp; Zyuzin, A. A." box="[840,1163,2001,2027]" pageId="10" pageNumber="169" pagination="813 - 828" refId="ref6175" refString="Kronestedt, T. &amp; Zyuzin, A. A. (2009) Fixation of Lycosa fidelis O. Pickard-Cambridge, 1872 as the type species for the genus Wadicosa Zyuzin, 1985 (Araneae: Lycosidae), with a redescripton of the species. Contributions to Natural History, 12, 813 - 828. [incl. corrigendum: Vol. 13 / 2010: 89]" type="journal article" year="2009">Kronestedt &amp; Zyuzin (2009)</bibRefCitation>
illustrated the male and female genitalia of this species and their female genitalic illustrations (
<bibRefCitation id="EFFC4B23FFEE2E3AFC0BFF1CFB4857A5" author="Kronestedt, T. &amp; Zyuzin, A. A." box="[934,1239,151,177]" pageId="11" pageNumber="170" pagination="813 - 828" refId="ref6175" refString="Kronestedt, T. &amp; Zyuzin, A. A. (2009) Fixation of Lycosa fidelis O. Pickard-Cambridge, 1872 as the type species for the genus Wadicosa Zyuzin, 1985 (Araneae: Lycosidae), with a redescripton of the species. Contributions to Natural History, 12, 813 - 828. [incl. corrigendum: Vol. 13 / 2010: 89]" type="journal article" year="2009">Kronestedt &amp; Zyuzin 2009</bibRefCitation>
: figs 15, 17) were used for comparative purpose.
</paragraph>
<paragraph id="8BD236D2FFEE2E3AFF6AFF55FE41569D" blockId="11.[151,1437,150,501]" pageId="11" pageNumber="170">
<emphasis id="B919EAC0FFEE2E3AFF6AFF55FDB357ED" bold="true" box="[199,556,222,249]" pageId="11" pageNumber="170">Justification of the synonymy.</emphasis>
<bibRefCitation id="EFFC4B23FFEE2E3AFD99FF54FD5757ED" author="Gajbe, U. A." box="[564,712,223,249]" pageId="11" pageNumber="170" pagination="1 - 40" refId="ref6099" refString="Gajbe, U. A. (2004) Studies on some spiders of the family Lycosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India, Occasional Paper, 221, 1 - 40." type="journal article" year="2004">Gajbe (2004)</bibRefCitation>
described
<taxonomicName id="4C6D4D51FFEE2E3AFCEFFF54FC7257ED" authorityName="Gajbe" authorityYear="2004" box="[834,1005,223,249]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="170" phylum="Arthropoda" rank="species" species="mandlaensis">
<emphasis id="B919EAC0FFEE2E3AFCEFFF54FCCC57EC" box="[834,851,223,248]" italics="true" pageId="11" pageNumber="170">E</emphasis>
.
<emphasis id="B919EAC0FFEE2E3AFCCCFF54FC7257ED" box="[865,1005,223,249]" italics="true" pageId="11" pageNumber="170">mandlaensis</emphasis>
</taxonomicName>
based on a female specimen collected in
<collectingRegion id="49A9F830FFEE2E3AFF1BFE88FEE95609" box="[182,374,259,285]" country="India" name="Madhya Pradesh" pageId="11" pageNumber="170">Madhya Pradesh</collectingRegion>
. Detailed examination of the
<typeStatus id="54D68870FFEE2E3AFD65FE88FCB55609" box="[712,810,259,285]" pageId="11" pageNumber="170" type="holotype">holotype</typeStatus>
showed no differences between this species and
<taxonomicName id="4C6D4D51FFEE2E3AFAF1FE88FF535655" authorityName="Kronestedt &amp; Zyuzin" authorityYear="2009" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="170" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B919EAC0FFEE2E3AFAF1FE88FAEC5608" box="[1372,1395,259,284]" italics="true" pageId="11" pageNumber="170">W</emphasis>
.
<emphasis id="B919EAC0FFEE2E3AFA2EFE88FF535655" italics="true" pageId="11" pageNumber="170">fidelis</emphasis>
</taxonomicName>
, a widely distributed species, which was previously recorded from
<collectingCountry id="F37A7642FFEE2E3AFC6EFEACFC625655" box="[963,1021,295,321]" name="India" pageId="11" pageNumber="170">India</collectingCountry>
(
<bibRefCitation id="EFFC4B23FFEE2E3AFBA1FEACFAAE5655" author="Tikader, B. K. &amp; Malhotra, M. S." box="[1036,1329,295,321]" pageId="11" pageNumber="170" pagination="248 - 447" refId="ref6874" refString="Tikader, B. K. &amp; Malhotra, M. S. (1980) Lycosidae (Wolf-spiders). Fauna India, Araneae, 1, 248 - 447." type="journal article" year="1980">Tikader &amp; Malhotra 1980</bibRefCitation>
;
<bibRefCitation id="EFFC4B23FFEE2E3AFA90FEACFF4E5672" author="Lu, T. &amp; Wang, L. Y. &amp; Hadole, P. &amp; Zhang, Z. S." pageId="11" pageNumber="170" pagination="130 - 142" refId="ref6246" refString="Lu, T., Wang, L. Y., Hadole, P. &amp; Zhang, Z. S. (2016) Redescription of four wolf-spiders (Araneae: Lycosidae) from India. Indian Journal of Arachnology, 5, 130 - 142." type="journal article" year="2016">
Lu
<emphasis id="B919EAC0FFEE2E3AFAC9FEACFA025655" box="[1380,1437,295,321]" italics="true" pageId="11" pageNumber="170">et al.</emphasis>
2016
</bibRefCitation>
) (compare
<figureCitation id="13562A57FFEE2E3AFEFFFEC0FE495671" box="[338,470,331,357]" captionStart="FIGURE 10" captionStartId="12.[151,250,1673,1697]" captionTargetBox="[155,1429,186,1642]" captionTargetId="figure-21@12.[151,1436,181,1649]" captionTargetPageId="12" captionText="FIGURE 10. Wadicosa fidelis (O. Pickard-Cambridge, 1872), holotype of Evippa mandlaensis Gajbe, 2004. A female habitus (Ar-1621), dorsal. BC genitalia: B epigynum, ventral; C vulva, dorsal. D label from type tube. Scale bars: A, 2 mm; BC, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4805805" httpUri="https://zenodo.org/record/4805805/files/figure.png" pageId="11" pageNumber="170">Fig. 10BC</figureCitation>
with
<bibRefCitation id="EFFC4B23FFEE2E3AFDBAFEC0FCD45671" author="Kronestedt, T. &amp; Zyuzin, A. A." box="[535,843,331,357]" pageId="11" pageNumber="170" pagination="813 - 828" refId="ref6175" refString="Kronestedt, T. &amp; Zyuzin, A. A. (2009) Fixation of Lycosa fidelis O. Pickard-Cambridge, 1872 as the type species for the genus Wadicosa Zyuzin, 1985 (Araneae: Lycosidae), with a redescripton of the species. Contributions to Natural History, 12, 813 - 828. [incl. corrigendum: Vol. 13 / 2010: 89]" type="journal article" year="2009">Kronestedt &amp; Zyuzin 2009</bibRefCitation>
: figs 15, 17). Thus, we consider
<taxonomicName id="4C6D4D51FFEE2E3AFB10FEC0FAF65671" authorityName="Gajbe" authorityYear="2004" box="[1213,1385,331,357]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="170" phylum="Arthropoda" rank="species" species="mandlaensis">
<emphasis id="B919EAC0FFEE2E3AFB10FEC0FB515670" box="[1213,1230,331,356]" italics="true" pageId="11" pageNumber="170">E</emphasis>
.
<emphasis id="B919EAC0FFEE2E3AFB70FEC0FAF65671" box="[1245,1385,331,357]" italics="true" pageId="11" pageNumber="170">mandlaensis</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C6D4D51FFEE2E3AFEC0FEE4FE49569D" authorityName="Kronestedt &amp; Zyuzin" authorityYear="2009" box="[365,470,367,393]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="170" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B919EAC0FFEE2E3AFEC0FEE4FE1B569C" box="[365,388,367,392]" italics="true" pageId="11" pageNumber="170">W</emphasis>
.
<emphasis id="B919EAC0FFEE2E3AFE3FFEE4FE49569D" box="[402,470,367,393]" italics="true" pageId="11" pageNumber="170">fidelis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3776559FFEE2E3AFF6AFE18FD8C56E1" pageId="11" pageNumber="170" type="discussion">
<paragraph id="8BD236D2FFEE2E3AFF6AFE18FD8C56E1" blockId="11.[151,1437,150,501]" pageId="11" pageNumber="170">
<emphasis id="B919EAC0FFEE2E3AFF6AFE18FEA456B9" bold="true" box="[199,315,403,429]" pageId="11" pageNumber="170">Remarks.</emphasis>
The NZC-ZSI collection has one glass tube for
<taxonomicName id="4C6D4D51FFEE2E3AFCF9FE18FC6056B9" authorityName="Gajbe" authorityYear="2004" box="[852,1023,403,429]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="170" phylum="Arthropoda" rank="species" species="mandlaensis">
<emphasis id="B919EAC0FFEE2E3AFCF9FE18FC6056B9" box="[852,1023,403,429]" italics="true" pageId="11" pageNumber="170">E. mandlaensis</emphasis>
</taxonomicName>
, labeled as holotype (Ar-1621) and containing
<specimenCount id="9D6BFD5BFFEE2E3AFEBBFE3CFE0C56C5" box="[278,403,439,465]" pageId="11" pageNumber="170" type="female">one female</specimenCount>
specimen in fairly good condition, with broken legs. The same tube has a small glass vial containing the dissected genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>