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<document id="B3AD72F58713CEC630FC6282ADE2EAE5" ID-DOI="10.5281/zenodo.277392" ID-GBIF-Dataset="8244ec23-791f-4ef0-b5ff-051bd2aaddee" ID-ISSN="1175-5326" ID-Zenodo-Dep="277392" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460298423927" checkinUser="plazi" docAuthor="Montreuil, Olivier &amp; Théry, Thomas" docDate="2011" docId="03C487AA9F4DD64C71C9D94CFC94FA52" docLanguage="en" docName="zt02863p062.pdf" docOrigin="Zootaxa 2863" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pseudonthobium humboldtense Montreuil &amp; Thery, new species" docType="treatment" docVersion="8" lastPageNumber="56" masterDocId="FFFDFFD29F4BD64B715EDF30FFAFFFF6" masterDocTitle="Revision of the New Caledonian genus Pseudonthobium Paulian, 1984 (Coleoptera: Scarabaeidae: Epilissini)" masterLastPageNumber="62" masterPageNumber="49" pageNumber="55" updateTime="1698252329856" updateUser="plazi">
<mods:mods id="70A47FF013DD172859A075314423574A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="25A208449D2691648F8252B0DEECA6FA">
<mods:title id="5AF6720E2EFD5669529B7A5D51F8497C">Revision of the New Caledonian genus Pseudonthobium Paulian, 1984 (Coleoptera: Scarabaeidae: Epilissini)</mods:title>
</mods:titleInfo>
<mods:name id="79C94E092205793F0A985F1DF996BA0D" type="personal">
<mods:role id="47C6C13FFE41F011DA92A0C07F8099FE">
<mods:roleTerm id="A7BE710D3B962FEAD1CF7FD3896D4D5A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2A9722C68BFFE9C8206A40669205907C">Montreuil, Olivier</mods:namePart>
</mods:name>
<mods:name id="4D2958972E8DC9E21C338C5AF3EED3A9" type="personal">
<mods:role id="78D7156EC5A294416F358A459E0007EE">
<mods:roleTerm id="FE0E99FD4E147C1EBE984A0AFEEFEEB7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="48F45D2A20211D9175E85CA775F3EBF8">Théry, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="81ECBA7F565A138427F1E8DAFD34D0CD">text</mods:typeOfResource>
<mods:relatedItem id="A4C6D16B91F3BDFE23A252A0D20612F2" type="host">
<mods:titleInfo id="DAB2D863D7216F3B70C897B596446B78">
<mods:title id="8C52B73E276FE01E6584D788C54AE590">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3289145393387EEC6AC119585E65FC62">
<mods:date id="65D82032B6A6E56E5325477155B68532">2011</mods:date>
<mods:detail id="0D6F5406F9EE85DDDD3482AAB3D8C959" type="volume">
<mods:number id="D83D06FA77AA683A969865AC4BD861D4">2863</mods:number>
</mods:detail>
<mods:extent id="5BAA37241F79A3F55BDB8BA0963E1C6A" unit="page">
<mods:start id="4CC422DB86551BFB1CFBB2488F3970A6">49</mods:start>
<mods:end id="4A0A5776D94BE5A464951BA36D3AD132">62</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="951EAADA9273AD075CAD679FA4777446">journal article</mods:classification>
<mods:identifier id="ED052D55E3625BACB13AFA7AF3410442" type="DOI">10.5281/zenodo.277392</mods:identifier>
<mods:identifier id="72CB81AD437040B4BE0628DB220E079C" type="GBIF-Dataset">8244ec23-791f-4ef0-b5ff-051bd2aaddee</mods:identifier>
<mods:identifier id="23A13160338AFBD5EA85CEB5B9A5B5B6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FCD48375147076D9F3B0AA8A30DB5C5D" type="Zenodo-Dep">277392</mods:identifier>
</mods:mods>
<treatment id="03C487AA9F4DD64C71C9D94CFC94FA52" ID-DOI="http://doi.org/10.5281/zenodo.6191700" ID-GBIF-Taxon="119399414" ID-Zenodo-Dep="6191700" LSID="urn:lsid:plazi:treatment:03C487AA9F4DD64C71C9D94CFC94FA52" httpUri="http://treatment.plazi.org/id/03C487AA9F4DD64C71C9D94CFC94FA52" lastPageId="7" lastPageNumber="56" pageId="6" pageNumber="55">
<subSubSection id="C37765379F4DD64D71C9D94CFEE9F941" pageId="6" pageNumber="55" type="nomenclature">
<paragraph id="8BD236BC9F4DD64D71C9D94CFC05F960" blockId="6.[151,938,1660,1719]" box="[151,938,1660,1686]" pageId="6" pageNumber="55">
<heading id="D09A81D09F4DD64D71C9D94CFC05F960" bold="true" box="[151,938,1660,1686]" fontSize="11" level="1" pageId="6" pageNumber="55" reason="1">
<emphasis id="B919EAAE9F4DD64D71C9D94CFC05F960" bold="true" box="[151,938,1660,1686]" pageId="6" pageNumber="55">
<taxonomicName id="4C6D4D3F9F4DD64D71C9D94CFCA1F960" ID-CoL="78FM3" authority="Montreuil &amp; Thery" authorityName="Montreuil &amp; Thery" box="[151,782,1660,1686]" class="Insecta" family="Scarabaeidae" genus="Pseudonthobium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="55" phylum="Arthropoda" rank="species" species="humboldtense" status="sp. nov.">
<emphasis id="B919EAAE9F4DD64D71C9D94CFDBBF960" bold="true" box="[151,532,1660,1686]" italics="true" pageId="6" pageNumber="55">Pseudonthobium humboldtense</emphasis>
Montreuil &amp; Théry
</taxonomicName>
,
<taxonomicNameLabel id="A22A57D59F4DD64D7245D94CFC05F960" box="[795,938,1660,1686]" pageId="6" pageNumber="55" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD236BC9F4DD64D71C9D9AFFEE9F941" blockId="6.[151,938,1660,1719]" box="[151,326,1695,1719]" pageId="6" pageNumber="55">
Habitus:
<figureCitation id="13562A399F4DD64D71A3D9AFFEEEF941" box="[253,321,1695,1719]" captionStart="FIGURES 1 – 9" captionStartId="3.[151,264,1841,1864]" captionTargetBox="[162,1431,213,1792]" captionTargetId="figure@3.[151,1436,195,1818]" captionTargetPageId="3" captionText="FIGURES 1 – 9. Habitus of Pseudonthobium. 1, P. fracticolloides Paulian, male; 2, P. doense n. sp., male; 3, P. genieri n. sp., male; 4, P. sinuatotibiale n. sp., male; 5, P. humboldtense n. sp., male; 6, P. paniense n. sp., male; 7, P. monteithianum n. sp., male; 8, P. foveolatum n. sp., female; 9, P. fracticolle (Fauvel), male. Scale bar: 1 mm." httpUri="https://zenodo.org/record/277393/files/figure.png" pageId="6" pageNumber="55">Fig. 5</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37765379F4DD64C71C9D9D8FB72FE39" lastPageId="7" lastPageNumber="56" pageId="6" pageNumber="55" type="materials_examined">
<paragraph id="8BD236BC9F4DD64C71C9D9D8FB72FE39" blockId="6.[151,1436,1767,2007]" lastBlockId="7.[151,1437,151,1445]" lastPageId="7" lastPageNumber="56" pageId="6" pageNumber="55">
<emphasis id="B919EAAE9F4DD64D71C9D9D8FEF7F8F6" bold="true" box="[151,344,1767,1792]" pageId="6" pageNumber="55">
<typeStatus id="54D6881E9F4DD64D71C9D9D8FF7DF8F6" box="[151,210,1768,1792]" pageId="6" pageNumber="55">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="54D6881E9F4DD64D7001D9D7FE68F909" box="[351,455,1767,1791]" pageId="6" pageNumber="55" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F37A762C9F4DD64D734DD9D8FD50F909" box="[531,767,1767,1791]" name="New Caledonia" pageId="6" pageNumber="55">
<collectionCode id="ED7CAE799F4DD64D734DD9D8FDFCF909" box="[531,595,1768,1791]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="6" pageNumber="55">NEW</collectionCode>
<collectionCode id="ED7CAE799F4DD64D7307D9D7FD50F909" box="[601,767,1767,1791]" pageId="6" pageNumber="55">CALEDONIA</collectionCode>
</collectingCountry>
,
<geoCoordinate id="EE59507B9F4DD64D7254D9D7FCC8F909" box="[778,871,1767,1791]" direction="south" orientation="latitude" pageId="6" pageNumber="55" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4DD64D7233D9D7FC74F909" box="[877,987,1767,1791]" direction="east" orientation="longitude" pageId="6" pageNumber="55" precision="925" value="166.41667">166°25’E</geoCoordinate>
,
<quantity id="4C959B599F4DD64D72B8D9D7FB98F909" box="[998,1079,1767,1791]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="55" unit="m" value="1350.0">1350m</quantity>
, rainfor., Mt Humboldt, summit track base,
<date id="FFD3107C9F4DD64D7048D83BFE15F8D5" box="[278,442,1803,1827]" pageId="6" pageNumber="55" value="2002-11-05" valueMax="2002-11-07" valueMin="2002-11-05">5–7 Nov 2002</date>
, dung trap, Monteith leg. [QM].
<typeStatus id="54D6881E9F4DD64D7274D83CFC38F8D2" box="[810,919,1804,1828]" pageId="6" pageNumber="55" type="paratype">Paratypes</typeStatus>
(166):
<specimenCount id="9D6BFD359F4DD64D72B4D83BFBE1F8D5" box="[1002,1102,1803,1827]" pageId="6" pageNumber="55" type="male">22 males</specimenCount>
and
<specimenCount id="9D6BFD359F4DD64D75DBD83BFAADF8D5" box="[1157,1282,1803,1827]" pageId="6" pageNumber="55" type="female">14 females</specimenCount>
, same data as
<typeStatus id="54D6881E9F4DD64D71C9D81FFF56F8B1" box="[151,249,1839,1863]" pageId="6" pageNumber="55" type="holotype">holotype</typeStatus>
[QM] —
<specimenCount id="9D6BFD359F4DD64D702FD81FFE78F8B1" box="[369,471,1839,1863]" pageId="6" pageNumber="55" type="male">14 males</specimenCount>
and
<specimenCount id="9D6BFD359F4DD64D734CD81FFD21F8B1" box="[530,654,1839,1863]" pageId="6" pageNumber="55" type="female">22 females</specimenCount>
—
<collectingCountry id="F37A762C9F4DD64D73E3D800FC03F8B1" box="[701,940,1839,1863]" name="New Caledonia" pageId="6" pageNumber="55">
<collectionCode id="ED7CAE799F4DD64D73E3D800FD52F8B1" box="[701,765,1840,1863]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="6" pageNumber="55">NEW</collectionCode>
<collectionCode id="ED7CAE799F4DD64D7258D81FFC03F8B1" box="[774,940,1839,1863]" pageId="6" pageNumber="55">CALEDONIA</collectionCode>
</collectingCountry>
,
<geoCoordinate id="EE59507B9F4DD64D72E4D81FFBB9F8B1" box="[954,1046,1839,1863]" direction="south" orientation="latitude" pageId="6" pageNumber="55" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4DD64D7541D81FFB22F8B1" box="[1055,1165,1839,1863]" direction="east" orientation="longitude" pageId="6" pageNumber="55" precision="925" value="166.4">166°24’E</geoCoordinate>
,
<quantity id="4C959B599F4DD64D75C5D81FFB43F8B1" box="[1179,1260,1839,1863]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="55" unit="m" value="1400.0">1400m</quantity>
, Mt Humboldt, moss forest,
<date id="FFD3107C9F4DD64D7078D863FE01F89D" box="[294,430,1875,1899]" pageId="6" pageNumber="55" value="2002-11-06">6 Nov 2002</date>
, night coll., Burwell &amp; Monteith leg. &amp; Wright [QM];
<specimenCount id="9D6BFD359F4DD64D7544D863FBD0F89D" box="[1050,1151,1875,1899]" pageId="6" pageNumber="55" type="male">17 males</specimenCount>
and
<specimenCount id="9D6BFD359F4DD64D75E9D863FA9EF89D" box="[1207,1329,1875,1899]" pageId="6" pageNumber="55" type="female">10 females</specimenCount>
—
<collectingCountry id="F37A762C9F4DD64D7402D864FE91F879" name="New Caledonia" pageId="6" pageNumber="55">
<collectionCode id="ED7CAE799F4DD64D7402D864FA33F89D" box="[1372,1436,1876,1899]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="6" pageNumber="55">NEW</collectionCode>
<collectionCode id="ED7CAE799F4DD64D71C9D847FE91F879" box="[151,318,1911,1935]" pageId="6" pageNumber="55">CALEDONIA</collectionCode>
</collectingCountry>
,
<geoCoordinate id="EE59507B9F4DD64D7015D847FE07F879" box="[331,424,1911,1935]" direction="south" orientation="latitude" pageId="6" pageNumber="55" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4DD64D70EFD847FDB0F879" box="[433,543,1911,1935]" direction="east" orientation="longitude" pageId="6" pageNumber="55" precision="925" value="166.4">166°24’E</geoCoordinate>
,
<quantity id="4C959B599F4DD64D7370D847FDD1F879" box="[558,638,1911,1935]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="55" unit="m" value="1400.0">1400m</quantity>
, Mt Humboldt, moss forest,
<date id="FFD3107C9F4DD64D7290D847FBD9F879" box="[974,1142,1911,1935]" pageId="6" pageNumber="55" value="2002-11-06" valueMax="2002-11-07" valueMin="2002-11-06">6–7 Nov 2002</date>
, Monteith leg., dung trap [QM];
<specimenCount id="9D6BFD359F4DD64D71B6D8ABFE99F845" box="[232,310,1947,1971]" pageId="6" pageNumber="55" type="male">1 male</specimenCount>
and
<specimenCount id="9D6BFD359F4DD64D702CD8ABFE4FF845" box="[370,480,1947,1971]" pageId="6" pageNumber="55" type="female">2 females</specimenCount>
—
<collectingCountry id="F37A762C9F4DD64D734ED8ACFD50F845" box="[528,767,1947,1971]" name="New Caledonia" pageId="6" pageNumber="55">
<collectionCode id="ED7CAE799F4DD64D734ED8ACFDFFF845" box="[528,592,1948,1971]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="6" pageNumber="55">NEW</collectionCode>
<collectionCode id="ED7CAE799F4DD64D7307D8ABFD50F845" box="[601,767,1947,1971]" pageId="6" pageNumber="55">CALEDONIA</collectionCode>
</collectingCountry>
,
<geoCoordinate id="EE59507B9F4DD64D7253D8ABFCC5F845" box="[781,874,1947,1971]" direction="south" orientation="latitude" pageId="6" pageNumber="55" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4DD64D722DD8ABFC4FF845" box="[883,992,1947,1971]" direction="east" orientation="longitude" pageId="6" pageNumber="55" precision="925" value="166.4">166°24’E</geoCoordinate>
,
<quantity id="4C959B599F4DD64D72B1D8ABFBEFF845" box="[1007,1088,1947,1971]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="55" unit="m" value="1400.0">1400m</quantity>
, Mt Humboldt, moss forest,
<date id="FFD3107C9F4DD64D74D0D8ABFEA5F821" pageId="6" pageNumber="55" value="2002-11-06">6 Nov 2002</date>
, Burwell &amp; Monteith leg. &amp; Wright [QM];
<specimenCount id="9D6BFD359F4DD64D73A9D88FFCECF821" box="[759,835,1983,2007]" pageId="6" pageNumber="55" type="male">1 male</specimenCount>
—
<collectingCountry id="F37A762C9F4DD64D7231D8F0FBF1F821" box="[879,1118,1983,2007]" name="New Caledonia" pageId="6" pageNumber="55">
<collectionCode id="ED7CAE799F4DD64D7231D8F0FC00F821" box="[879,943,1984,2007]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="6" pageNumber="55">NEW</collectionCode>
<collectionCode id="ED7CAE799F4DD64D72E9D88FFBF1F821" box="[951,1118,1983,2007]" pageId="6" pageNumber="55">CALEDONIA</collectionCode>
</collectingCountry>
,
<geoCoordinate id="EE59507B9F4DD64D7537D88FFB6AF821" box="[1129,1221,1983,2007]" direction="south" orientation="latitude" pageId="6" pageNumber="55" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4DD64D7593D88FFA94F821" box="[1229,1339,1983,2007]" direction="east" orientation="longitude" pageId="6" pageNumber="55" precision="925" value="166.41667">166°25’E</geoCoordinate>
,
<quantity id="4C959B599F4DD64D7419D88FFA38F821" box="[1351,1431,1983,2007]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="55" unit="m" value="1350.0">1350m</quantity>
, Mt Humboldt, refuge,
<date id="FFD3107C9F4CD64C70C1DFA7FDE6FF59" box="[415,585,151,175]" pageId="7" pageNumber="56" value="2002-11-05" valueMax="2002-11-08" valueMin="2002-11-05">5–8 Nov 2002</date>
, hand coll., Burwell &amp; Monteith leg. &amp; Wright [QM];
<specimenCount id="9D6BFD359F4CD64C758FDFA7FA85FF59" box="[1233,1322,151,175]" pageId="7" pageNumber="56" type="male">2 males</specimenCount>
—
<collectingCountry id="F37A762C9F4CD64C7402DFA8FE91FF22" name="New Caledonia" pageId="7" pageNumber="56">
<collectionCode id="ED7CAE799F4CD64C7402DFA8FA33FF59" box="[1372,1436,152,175]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="7" pageNumber="56">NEW</collectionCode>
<collectionCode id="ED7CAE799F4CD64C71C9DF8CFE91FF22" box="[151,318,188,212]" pageId="7" pageNumber="56">CALEDONIA</collectionCode>
</collectingCountry>
,
<geoCoordinate id="EE59507B9F4CD64C7016DF8CFE0BFF22" box="[328,420,188,212]" direction="south" orientation="latitude" pageId="7" pageNumber="56" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4CD64C70F4DF8CFDB6FF22" box="[426,537,188,212]" direction="east" orientation="longitude" pageId="7" pageNumber="56" precision="925" value="166.41667">166°25’E</geoCoordinate>
,
<quantity id="4C959B599F4CD64C737DDF8CFDDCFF22" box="[547,627,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="7" pageNumber="56" unit="m" value="1350.0">1350m</quantity>
, Mt Humboldt, refuge,
<date id="FFD3107C9F4CD64C7227DF8CFBB4FF22" box="[889,1051,188,212]" pageId="7" pageNumber="56" value="2002-11-05" valueMax="2002-11-08" valueMin="2002-11-05">5–8 Nov 2002</date>
, night collecting, Burwell &amp; Monteith leg. &amp; Wright [QM];
<specimenCount id="9D6BFD359F4CD64C7097DFD0FD8EFF01" box="[457,545,223,247]" pageId="7" pageNumber="56" type="male">7 males</specimenCount>
and
<specimenCount id="9D6BFD359F4CD64C7302DFEFFD66FF01" box="[604,713,223,247]" pageId="7" pageNumber="56" type="female">5 females</specimenCount>
—
<collectingCountry id="F37A762C9F4CD64C73A9DFD0FC49FF01" box="[759,998,223,247]" name="New Caledonia" pageId="7" pageNumber="56">
<collectionCode id="ED7CAE799F4CD64C73A9DFD0FC98FF01" box="[759,823,224,247]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="7" pageNumber="56">NEW</collectionCode>
<collectionCode id="ED7CAE799F4CD64C721EDFEFFC49FF01" box="[832,998,223,247]" pageId="7" pageNumber="56">CALEDONIA</collectionCode>
</collectingCountry>
,
<geoCoordinate id="EE59507B9F4CD64C72ADDFEFFBE0FF01" box="[1011,1103,223,247]" direction="south" orientation="latitude" pageId="7" pageNumber="56" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4CD64C7506DFEFFB69FF01" box="[1112,1222,223,247]" direction="east" orientation="longitude" pageId="7" pageNumber="56" precision="925" value="166.41667">166°25’E</geoCoordinate>
,
<quantity id="4C959B599F4CD64C758ADFEFFA8BFF01" box="[1236,1316,223,247]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="7" pageNumber="56" unit="m" value="1350.0">1350m</quantity>
, Mt Humboldt, refuge,
<date id="FFD3107C9F4CD64C706BDE34FE76FEEA" box="[309,473,260,284]" pageId="7" pageNumber="56" value="2002-11-05" valueMax="2002-11-08" valueMin="2002-11-05">5–8 Nov 2002</date>
, dung trap, Burwell, Monteith &amp; Wright leg. [QM];
<specimenCount id="9D6BFD359F4CD64C7576DE34FB21FEEA" box="[1064,1166,260,284]" pageId="7" pageNumber="56" type="male">17 males</specimenCount>
and
<specimenCount id="9D6BFD359F4CD64C759ADE34FA9EFEEA" box="[1220,1329,260,284]" pageId="7" pageNumber="56" type="female">6 females</specimenCount>
—
<collectingCountry id="F37A762C9F4CD64C7402DE35FE91FEC9" name="New Caledonia" pageId="7" pageNumber="56">
<collectionCode id="ED7CAE799F4CD64C7402DE35FA33FEEA" box="[1372,1436,261,284]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="7" pageNumber="56">NEW</collectionCode>
<collectionCode id="ED7CAE799F4CD64C71C9DE17FE91FEC9" box="[151,318,295,319]" pageId="7" pageNumber="56">CALEDONIA</collectionCode>
</collectingCountry>
,
<geoCoordinate id="EE59507B9F4CD64C7017DE17FE0AFEC9" box="[329,421,295,319]" direction="south" orientation="latitude" pageId="7" pageNumber="56" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4CD64C70F3DE17FDB4FEC9" box="[429,539,295,319]" direction="east" orientation="longitude" pageId="7" pageNumber="56" precision="925" value="166.41667">166°25’E</geoCoordinate>
,
<quantity id="4C959B599F4CD64C7379DE17FDD8FEC9" box="[551,631,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="56" unit="m" value="1600.0">1600m</quantity>
, Mt Humboldt, summit,
<date id="FFD3107C9F4CD64C72D1DE17FB9CFEC9" box="[911,1075,295,319]" pageId="7" pageNumber="56" value="2002-11-05" valueMax="2002-11-07" valueMin="2002-11-05">5–7 Nov 2002</date>
, Monteith &amp; Burwell leg., dung trap [QM];
<specimenCount id="9D6BFD359F4CD64C7049DE7CFEC2FE92" box="[279,365,332,356]" pageId="7" pageNumber="56" type="male">4 males</specimenCount>
and
<specimenCount id="9D6BFD359F4CD64C70FADE7CFDBFFE92" box="[420,528,332,356]" pageId="7" pageNumber="56" type="female">3 females</specimenCount>
—
<collectingCountry id="F37A762C9F4CD64C7367DE7DFC88FE92" box="[569,807,332,356]" name="New Caledonia" pageId="7" pageNumber="56">
<collectionCode id="ED7CAE799F4CD64C7367DE7DFDD6FE92" box="[569,633,333,356]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="7" pageNumber="56">NEW</collectionCode>
<collectionCode id="ED7CAE799F4CD64C73DEDE7CFC88FE92" box="[640,807,332,356]" pageId="7" pageNumber="56">CALEDONIA</collectionCode>
</collectingCountry>
,
<geoCoordinate id="EE59507B9F4CD64C726FDE7CFC21FE92" box="[817,910,332,356]" direction="south" orientation="latitude" pageId="7" pageNumber="56" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4CD64C72CADE7CFBACFE92" box="[916,1027,332,356]" direction="east" orientation="longitude" pageId="7" pageNumber="56" precision="925" value="166.41667">166°25’E</geoCoordinate>
,
<quantity id="4C959B599F4CD64C7550DE7CFBF1FE92" box="[1038,1118,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="56" unit="m" value="1600.0">1600m</quantity>
, Mt Humboldt, summit,
<date id="FFD3107C9F4CD64C742CDE7CFEA2FE71" pageId="7" pageNumber="56" value="2002-11-05" valueMax="2002-11-07" valueMin="2002-11-05">5–7 Nov 2002</date>
, Burwell, Monteith &amp; Wright leg. [QM];
<specimenCount id="9D6BFD359F4CD64C73A6DE5FFCCFFE71" box="[760,864,367,391]" pageId="7" pageNumber="56" type="male">10 males</specimenCount>
and
<specimenCount id="9D6BFD359F4CD64C72C0DE5FFBA1FE71" box="[926,1038,367,391]" pageId="7" pageNumber="56" type="female">8 females</specimenCount>
—
<collectingCountry id="F37A762C9F4CD64C751EDE40FA9EFE71" box="[1088,1329,367,391]" name="New Caledonia" pageId="7" pageNumber="56">
<collectionCode id="ED7CAE799F4CD64C751EDE40FB2FFE71" box="[1088,1152,368,391]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="7" pageNumber="56">NEW</collectionCode>
<collectionCode id="ED7CAE799F4CD64C75D5DE5FFA9EFE71" box="[1163,1329,367,391]" pageId="7" pageNumber="56">CALEDONIA</collectionCode>
</collectingCountry>
,
<geoCoordinate id="EE59507B9F4CD64C741EDE5FFA33FE71" box="[1344,1436,367,391]" direction="south" orientation="latitude" pageId="7" pageNumber="56" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4CD64C71C9DEA4FEAAFE5A" box="[151,261,404,428]" direction="east" orientation="longitude" pageId="7" pageNumber="56" precision="925" value="166.41667">166°25’E</geoCoordinate>
,
<quantity id="4C959B599F4CD64C704EDEA4FECFFE5A" box="[272,352,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="56" unit="m" value="1300.0">1300m</quantity>
, Mt Humboldt, source,
<date id="FFD3107C9F4CD64C7337DEA4FCA3FE5A" box="[617,780,404,428]" pageId="7" pageNumber="56" value="2002-11-06" valueMax="2002-11-08" valueMin="2002-11-06">6–8 Nov 2002</date>
, Monteith leg., dung trap [QM];
<specimenCount id="9D6BFD359F4CD64C7527DEA4FB75FE5A" box="[1145,1242,404,428]" pageId="7" pageNumber="56" type="female">1 female</specimenCount>
—
<collectionCode id="ED7CAE799F4CD64C745ADEA5FAEBFE5A" box="[1284,1348,405,428]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="7" pageNumber="56">NEW</collectionCode>
<collectionCode id="ED7CAE799F4CD64C7414DEA4FF5BFE39" pageId="7" pageNumber="56">CALE- DONIA</collectionCode>
,
<geoCoordinate id="EE59507B9F4CD64C71A1DE87FEF3FE39" box="[255,348,439,463]" direction="south" orientation="latitude" pageId="7" pageNumber="56" precision="925" value="-21.883333">21°53’S</geoCoordinate>
<geoCoordinate id="EE59507B9F4CD64C703CDE87FE7EFE39" box="[354,465,439,463]" direction="east" orientation="longitude" pageId="7" pageNumber="56" precision="925" value="166.41667">166°25’E</geoCoordinate>
, Mt Humboldt, refuge,
<date id="FFD3107C9F4CD64C7387DE87FCD3FE39" box="[729,892,439,463]" pageId="7" pageNumber="56" value="2002-11-07" valueMax="2002-11-08" valueMin="2002-11-07">7–8 Nov 2002</date>
, Monteith &amp; Wright leg. [QM].
</paragraph>
</subSubSection>
<subSubSection id="C37765379F4CD64C719BDEEBFB37FAAA" pageId="7" pageNumber="56" type="description">
<paragraph id="8BD236BC9F4CD64C719BDEEBFA85FE02" blockId="7.[151,1437,151,1445]" box="[197,1322,475,500]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDEEBFE06FE02" bold="true" box="[197,425,475,500]" pageId="7" pageNumber="56">Description (male).</emphasis>
Length 6.0–8.0 mm. Dark reddish-brown, feebly shiny. Setae yellowish-brown.
</paragraph>
<paragraph id="8BD236BC9F4CD64C719BDD30FA8BFDE1" blockId="7.[151,1437,151,1445]" box="[197,1316,511,536]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDD30FEAEFDEE" box="[197,257,512,536]" italics="true" pageId="7" pageNumber="56">Head</emphasis>
. Frons flat, clypeus feebly depressed at middle. Clypeus bidentate, lateral margins subconvex.
</paragraph>
<paragraph id="8BD236BC9F4CD64C719BDD15FEEDFD72" blockId="7.[151,1437,151,1445]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDD15FE9BFDCA" box="[197,308,549,572]" italics="true" pageId="7" pageNumber="56">Pronotum</emphasis>
. Base strongly bilobate, distinctly concave in the middle, obliquely divergent and concave on each side. Posterior edge obtuse. Lateral margin straight, anteriorly feebly convergent to strongly convergent. Anterior edge not sharp.
</paragraph>
<paragraph id="8BD236BC9F4CD64C719BDDA0FF74FCCC" blockId="7.[151,1437,151,1445]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDDA0FEA4FD5E" box="[197,267,656,680]" italics="true" pageId="7" pageNumber="56">Elytra</emphasis>
. Interstriae flat, strongly microreticulate, with fine punctation; punctures evenly dispersed, much smaller than those on pronotum, separated by about 1–2 times their diameter, bearing thin and very short setae. Anterior margin straight then strongly and abruptly projected obliquely from base of 5th stria to fit the basal margin of pronotum. Elytra of normal shape with evenly curved sides. Setae numerous, short, with even distribution on all interstriae.
</paragraph>
<paragraph id="8BD236BC9F4CD64C719BDC79FC93FC73" blockId="7.[151,1437,151,1445]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDC79FE99FC97" box="[197,310,838,865]" italics="true" pageId="7" pageNumber="56">8th tergite.</emphasis>
Smooth, shiny, with fine and even punctation. Deep furrow along the inner side of margins. Apical margin strongly enlarged, notched in middle of inner border.
</paragraph>
<paragraph id="8BD236BC9F4CD64C719BDCA2FAD0FC3B" blockId="7.[151,1437,151,1445]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDCA2FE89FC5F" box="[197,294,914,937]" italics="true" pageId="7" pageNumber="56">Sternites</emphasis>
. Last sternite strongly enlarged; tegument feebly microreticulate, shiny, completely covered with strong punctation. Other sternites normally developed, feebly microreticulate, less shiny, punctate only on sides.
</paragraph>
<paragraph id="8BD236BC9F4CD64C719BDCEAFCA4FBAB" blockId="7.[151,1437,151,1445]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDCEAFF55FC07" box="[197,250,986,1009]" italics="true" pageId="7" pageNumber="56">Legs</emphasis>
. Outer margin of protibia tridentate in apical third, median tooth closer to apical tooth, basal tooth obsolete. Outer margin indistinctly crenulate at base and between teeth; crenulations bearing very short, white setae. Inner margin sinuate before apex and strongly enlarged from middle, enlargement extended by a lamella (
<figureCitation id="13562A399F4CD64C7465DB11FA3FFBCF" box="[1339,1424,1057,1081]" captionStart="FIGURES 18 – 25" captionStartId="5.[151,264,1208,1231]" captionTargetBox="[415,1172,199,1168]" captionTargetId="figure@5.[412,1174,193,1187]" captionTargetPageId="5" captionText="FIGURES 18 – 25. Right protibia of male Pseudonthobium, dorsal view. 18, P. fracticolloides Paulian; 19, P. doense n. sp.; 20, P. g e n i e r i n. sp.; 21, P. sinuatotibiale n. sp.; 22, P. humboldtense n. sp.; 23, P. paniense n. sp.; 24, P. monteithianum n. sp.; 25, P. fracticolle (Fauvel). Scale bar: 1 mm." httpUri="https://zenodo.org/record/277395/files/figure.png" pageId="7" pageNumber="56">Fig. 22</figureCitation>
). Base of posterior margin of metafemora feebly notched.
</paragraph>
<paragraph id="8BD236BC9F4CD64C719BDB59FDC2FB3F" blockId="7.[151,1437,151,1445]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDB59FE9DFB77" box="[197,306,1129,1153]" italics="true" pageId="7" pageNumber="56">Aedeagus</emphasis>
(
<figureCitation id="13562A399F4CD64C701FDB59FE3AFB77" box="[321,405,1129,1153]" captionStart="FIGURES 10 – 17" captionStartId="4.[151,264,1498,1521]" captionTargetBox="[158,1433,238,1450]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURES 10 – 17. Parameres of Pseudonthobium, right, dorsal and left views. 10, P. fracticolloides Paulian; 11, P. doense n. sp.; 12, P. g e n i e r i n. sp.; 13, P. sinuatotibiale n. sp.; 14, P. humboldtense n. sp.; 15, P. paniense n. sp.; 16, P. monteithianum n. sp.; 17, P. fracticolle (Fauvel). Scale bar: 1 mm." httpUri="https://zenodo.org/record/277394/files/figure.png" pageId="7" pageNumber="56">Fig. 14</figureCitation>
). Left paramere strongly projected laterally in apical third, outer margin more projected forwards, external apical angle spiny. Right paramere strongly curved outwards in dorsal view, feebly spiny at apex, narrow and curved at apex in lateral view.
</paragraph>
<paragraph id="8BD236BC9F4CD64C719BDBE9FA9EFB06" blockId="7.[151,1437,151,1445]" box="[197,1329,1237,1265]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDBE9FEA0FB07" bold="true" box="[197,271,1241,1265]" pageId="7" pageNumber="56">Notes.</emphasis>
Female similar to male, with simple inner margin of protibia; 8th tergite short, very transverse.
</paragraph>
<paragraph id="8BD236BC9F4CD64C719BDBCCFB37FAAA" blockId="7.[151,1437,151,1445]" pageId="7" pageNumber="56">
This species is similar to
<taxonomicName id="4C6D4D3F9F4CD64C70BDDBCDFD36FAE2" box="[483,665,1276,1300]" class="Insecta" family="Scarabaeidae" genus="Pseudonthobium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="sinuatotibiale">
<emphasis id="B919EAAE9F4CD64C70BDDBCDFD36FAE2" box="[483,665,1276,1300]" italics="true" pageId="7" pageNumber="56">P. sinuatotibiale</emphasis>
</taxonomicName>
in its general habitus, but smaller in size, the shape of elytra normal and not sinuate after the middle on the side, metafemora not notched at base and the elytra not elongate. The shape of the aedeagus is similar to
<taxonomicName id="4C6D4D3F9F4CD64C7089DA75FD22FAAA" box="[471,653,1348,1372]" class="Insecta" family="Scarabaeidae" genus="Pseudonthobium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="sinuatotibiale">
<emphasis id="B919EAAE9F4CD64C7089DA75FD22FAAA" box="[471,653,1348,1372]" italics="true" pageId="7" pageNumber="56">P. sinuatotibiale</emphasis>
</taxonomicName>
, except for some differences in the parameres.
</paragraph>
</subSubSection>
<subSubSection id="C37765379F4CD64C719BDA58FAD5FA76" box="[197,1402,1384,1409]" pageId="7" pageNumber="56" type="distribution">
<paragraph id="8BD236BC9F4CD64C719BDA58FAD5FA76" blockId="7.[151,1437,151,1445]" box="[197,1402,1384,1409]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDA58FEF1FA77" bold="true" box="[197,350,1384,1409]" pageId="7" pageNumber="56">Distribution.</emphasis>
This species is known only from Mt Humboldt, its
<typeStatus id="54D6881E9F4CD64C72FFDA59FC7EFA77" box="[929,977,1385,1409]" pageId="7" pageNumber="56">type</typeStatus>
locality, in southeast
<collectingCountry id="F37A762C9F4CD64C759BDA59FAD9FA76" box="[1221,1398,1384,1408]" name="New Caledonia" pageId="7" pageNumber="56">New Caledonia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37765379F4CD64C719BDABDFC94FA52" box="[197,827,1420,1445]" pageId="7" pageNumber="56" type="etymology">
<paragraph id="8BD236BC9F4CD64C719BDABDFC94FA52" blockId="7.[151,1437,151,1445]" box="[197,827,1420,1445]" pageId="7" pageNumber="56">
<emphasis id="B919EAAE9F4CD64C719BDABDFEE3FA53" bold="true" box="[197,332,1421,1445]" pageId="7" pageNumber="56">Etymology.</emphasis>
This species is named after its
<typeStatus id="54D6881E9F4CD64C73F4DABDFD75FA53" box="[682,730,1421,1445]" pageId="7" pageNumber="56">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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