treatments-xml/data/03/9E/B8/039EB86FA96E0442FF15CD3192EFFE8B.xml

<document id="4A0B996567797AA0939B4081A13AD6FC" ID-CLB-Dataset="53371" ID-DOI="10.11646/zootaxa.5293.2.9" ID-GBIF-Dataset="f3941643-9de3-4323-be08-ff975d5fcd76" ID-ISSN="1175-5326" ID-Zenodo-Dep="7960306" ID-ZooBank="A0295A9C-7731-4AE6-BC94-C9DC94CE140E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1684798617567" checkinUser="plazi" docAuthor="Martens, Jochen, Maghradze, Eter &amp; Barjadze, Shalva" docDate="2023" docId="039EB86FA96E0442FF15CD3192EFFE8B" docLanguage="en" docName="zootaxa.5293.2.9.pdf" docOrigin="Zootaxa 5293 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.2.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nemaspela kotia Martens &amp; Maghradze &amp; Barjadze 2023, sp. nov." docType="treatment" docVersion="6" lastPageNumber="368" masterDocId="FFA7C017A96F0445FF82CB4F937AFE5E" masterDocTitle="An additional species of the genus Nemaspela Šilhavý, 1966 from a cave in Georgia with a note on Nemaspela melouri (Opiliones: Nemastomatidae)" masterLastPageNumber="370" masterPageNumber="361" pageNumber="362" updateTime="1699508118997" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="51DF7F7BDC30BB109F65335D4846DA3D">An additional species of the genus Nemaspela Šilhavý, 1966 from a cave in Georgia with a note on Nemaspela melouri (Opiliones: Nemastomatidae)</mods:title>
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<mods:namePart id="E1C6C0797F880DA18EB4FE0708B9EE92">Martens, Jochen</mods:namePart>
<mods:affiliation id="F403C3AAF62025CF927240D35ABE5119">Johannes Gutenberg-Universität, Institut für Organismische und Molekulare Evolutionsbiologie (iomE), D- 55099 Mainz, Germany. Senckenberg Research Institute, Arachnology, D- 60325 Frankfurt am Main, Germany.</mods:affiliation>
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<mods:namePart id="3378025F3BFA1CC64476455DDB5C793B">Maghradze, Eter</mods:namePart>
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<mods:affiliation id="EEE810CBBC30ECF019A16273C5392B84">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia &amp; Eter Magradze: eter. magradze. 1 @ iliauni. edu. ge; https: // orcid. org / 0000 - 0002 - 4796 - 9439</mods:affiliation>
<mods:nameIdentifier id="5B5C0EF58F595495F05044F68DC14B03" type="email">eter.magradze.1@iliauni.edu.ge</mods:nameIdentifier>
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<mods:namePart id="CA79107038165B77531D2B2B8A6C7AEF">Barjadze, Shalva</mods:namePart>
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<mods:affiliation id="EF1FEC3815F9DF5676E26596E8E5E0AC">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia &amp; Shalva Bajardze: shalva. barjadze @ yahoo. com; https: // orcid. org / 0000 - 0001 - 8992 - 4987</mods:affiliation>
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<taxonomicName id="4C3772FAA96E0444FF15CD31921AF8C7" ID-CoL="BTD3F" authority="Martens &amp; Maghradze &amp; Barjadze, 2023" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2023" box="[151,352,1662,1689]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="kotia" status="sp. nov.">
<emphasis id="B943D56BA96E0444FF15CD31921AF8C7" bold="true" box="[151,352,1662,1689]" italics="true" pageId="1" pageNumber="362">Nemaspela kotia</emphasis>
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<paragraph id="8B880979A96E0444FF15CDEC92AAF8E3" blockId="1.[151,464,1662,1725]" box="[151,464,1699,1725]" pageId="1" pageNumber="362">
(
<figureCitation id="130C15FCA96E0444FF1DCDEC926BF8E3" box="[159,273,1699,1725]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2-3" captionStart-2="FIGURES 4-11" captionStart-3="FIGURES 12-16" captionStartId-0="2.[151,250,1945,1969]" captionStartId-1="3.[151,264,878,902]" captionStartId-2="4.[151,264,1943,1967]" captionStartId-3="6.[151,264,1398,1422]" captionTargetBox-0="[151,1436,958,1920]" captionTargetBox-1="[151,1436,181,854]" captionTargetBox-2="[185,1411,186,1920]" captionTargetBox-3="[156,1431,314,1370]" captionTargetId-0="figure-363@2.[151,1436,957,1920]" captionTargetId-1="figure-18@3.[151,1436,181,854]" captionTargetId-2="figure-32@4.[175,1412,181,1920]" captionTargetId-3="figure-95@6.[151,1436,309,1375]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionText-0="FIGURE 1. Nemaspela kotia sp. nov. at the type locality, Kotia cave, presumably female in lateral view. Photograph by Dragan Antić." captionText-1="FIGURES 2-3. Nemaspela kotia sp. nov. Preserved specimens, lateral view. 2. Male, 3. Female (bar: 2 mm). Photographs by Jean Severin." captionText-2="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." captionText-3="FIGURES 12-16. Nemaspela kotia sp. nov. Chelicerae in retro-lateral view. 12. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Chelicera in pro-lateral view. 13. Male (bar: 0.3 mm). 15. Female (bar: 0.3 mm). 16. Patellar apophysis of female, retro-lateral views (bar: 0.1 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.7960308" figureDoi-1="http://doi.org/10.5281/zenodo.7960310" figureDoi-2="http://doi.org/10.5281/zenodo.7960312" figureDoi-3="http://doi.org/10.5281/zenodo.7960314" httpUri-0="https://zenodo.org/record/7960308/files/figure.png" httpUri-1="https://zenodo.org/record/7960310/files/figure.png" httpUri-2="https://zenodo.org/record/7960312/files/figure.png" httpUri-3="https://zenodo.org/record/7960314/files/figure.png" pageId="1" pageNumber="362">Figs 1–16</figureCitation>
, 
<figureCitation id="130C15FCA96E0444FE9CCDEC921CF8E3" box="[286,358,1699,1725]" captionStart="MAP 1" captionStartId="5.[152,211,1578,1602]" captionTargetBox="[151,1436,151,1554]" captionTargetId="figure-18@5.[151,1436,151,1554]" captionTargetPageId="5" captionText="MAP 1. Distributional map of the Nemaspela species in the Caucasus. 1. Vorontsovskaya Cave (N. sokolovi); 2. Labirintovaya Cave (N. sokolovi); 3. Dolgaya Cave (N. sokolovi); 4. Fontanka Cave (N. kovali); 5. Omega-15 Cave (N. kovali); 6. Omega-12 Cave (N. kovali); 7. Otte-Shik Cave (N. kovali); 8. Akhunskaya Cave (N. abchasica); 9. Beloskalskaya Cave (N. abchasica); 10. Kolokolnaya Cave (N. abchasica); 11. Kuibishevi Cave (N. abchasica); 12. L 24 Cave (N. abchasica); 13. Kveda Shakurani (=Nizhnyaya Shakuranskaya) Cave (N. abchasica); 14. Zemo Eshera Cave (N. birsteini); 15. Tarkili Cave (N. birsteini); 16. Sarma Cave (N. gagrica); 17. Veryovkina Cave (N. gagrica); 18. Sakishore Cave (N. femorecurvata); 19. Prometheus Cave (N. prometheus); 20. Melouri Cave (N. melouri); 21. Solkota Cave (N. melouri); 22. Kotia Cave (=Kotiasklde Cave) (N. kotia) (for details of localities and collection depositories see Martens et al. 2021)." figureDoi="http://doi.org/10.5281/zenodo.7971988" httpUri="https://zenodo.org/record/7971988/files/figure.png" pageId="1" pageNumber="362">Map 1</figureCitation>
, 
<tableCitation id="C6B53CC2A96E0444FEF6CDEC92BEF8E3" box="[372,452,1699,1725]" captionStart="TABLE 1" captionStartId="2.[152,245,620,644]" captionTargetPageId="2" captionText="TABLE 1. Measurements (mm) of legs and pedipalp in male holotype and female paratype (in parentheses) of Nemaspela kotia sp. nov." httpUri="http://table.plazi.org/id/DF4859F1A96D0447FF1AC9239253FCF7" pageId="1" pageNumber="362" tableUuid="DF4859F1A96D0447FF1AC9239253FCF7">Table 1</tableCitation>
)
</paragraph>
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<emphasis id="B943D56BA96E0444FF15CDA4922AF95A" bold="true" box="[151,336,1770,1797]" pageId="1" pageNumber="362">Type material:</emphasis>
<materialsCitation id="3B5F0324A96E0444FEE2CDA4910DF913" ID-GBIF-Occurrence="4116970301" collectingDate="2021-10-08" country="Georgia" county="Male" elevation="725" latitude="42.213306" location="Kotiasklde Cave" longLatPrecision="1" longitude="43.324417" municipality="Kotia Cave" pageId="1" pageNumber="362" specimenCount="1" stateProvince="Imereti" typeStatus="holotype">
<emphasis id="B943D56BA96E0444FEE2CDA492ACF95B" bold="true" box="[352,470,1771,1797]" pageId="1" pageNumber="362">
<typeStatus id="548CB7DBA96E0444FEE2CDA492ABF95B" box="[352,465,1771,1797]" pageId="1" pageNumber="362" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCounty id="62E971F5A96E0444FE64CDA49158F95B" box="[486,546,1771,1797]" pageId="1" pageNumber="362">Male</collectingCounty>
, ISIZU-TO-T-00001, leg. 
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<collectingDate id="EFCDD651A96E0444FCE6CDA490B3F95B" box="[868,969,1771,1797]" pageId="1" pageNumber="362" value="2021-10-08">8.x.2021</collectingDate>
</date>
, 
<collectingCountry id="F32049E9A96E0444FC5FCDA49724F95B" box="[989,1118,1771,1797]" name="Georgia" pageId="1" pageNumber="362">GEORGIA</collectingCountry>
, 
<collectingMunicipality id="6BEC9303A96E0444FBF0CDA49780F95B" box="[1138,1274,1771,1797]" pageId="1" pageNumber="362">Kotia Cave</collectingMunicipality>
(= 
<location id="8EE85FA2A96E0444FAA2CDA493AEF977" LSID="urn:lsid:plazi:treatment:039EB86FA96E0442FF15CD3192EFFE8B:8EE85FA2A96E0444FAA2CDA493AEF977" country="Georgia" county="Male" latitude="42.213306" longLatPrecision="1" longitude="43.324417" municipality="Kotia Cave" name="Kotiasklde Cave" pageId="1" pageNumber="362" stateProvince="Imereti">Kotiasklde Cave</location>
), close to the village 
<location id="8EE85FA2A96E0444FE4FCC409171F977" LSID="urn:lsid:plazi:treatment:039EB86FA96E0442FF15CD3192EFFE8B:8EE85FA2A96E0444FE4FCC409171F977" box="[461,523,1807,1833]" country="Georgia" county="Male" latitude="42.213306" longLatPrecision="1" longitude="43.324417" municipality="Kotia Cave" name="Sveri" pageId="1" pageNumber="362" stateProvince="Imereti">Sveri</location>
, 
<location id="8EE85FA2A96E0444FD9ACC40906EF977" LSID="urn:lsid:plazi:treatment:039EB86FA96E0442FF15CD3192EFFE8B:8EE85FA2A96E0444FD9ACC40906EF977" box="[536,788,1807,1833]" country="Georgia" county="Male" latitude="42.213306" longLatPrecision="1" longitude="43.324417" municipality="Kotia Cave" name="Chiatura Municipality" pageId="1" pageNumber="362" stateProvince="Imereti">Chiatura Municipality</location>
, 
<location id="8EE85FA2A96E0444FCA3CC409018F977" LSID="urn:lsid:plazi:treatment:039EB86FA96E0442FF15CD3192EFFE8B:8EE85FA2A96E0444FCA3CC409018F977" box="[801,866,1807,1833]" country="Georgia" county="Male" latitude="42.213306" longLatPrecision="1" longitude="43.324417" municipality="Kotia Cave" name="Zemo" pageId="1" pageNumber="362" stateProvince="Imereti">Zemo</location>
Imereti plateau, 
<collectingRegion id="49F3C79BA96E0444FBA5CC4097AFF977" box="[1063,1237,1807,1833]" country="Georgia" name="Imereti" pageId="1" pageNumber="362">Imereti Region</collectingRegion>
, 
<location id="8EE85FA2A96E0444FB60CC4096E7F977" LSID="urn:lsid:plazi:treatment:039EB86FA96E0442FF15CD3192EFFE8B:8EE85FA2A96E0444FB60CC4096E7F977" box="[1250,1437,1807,1833]" country="Georgia" county="Male" latitude="42.213306" longLatPrecision="1" longitude="43.324417" municipality="Kotia Cave" name="western Georgia" pageId="1" pageNumber="362" stateProvince="Imereti">western Georgia</location>
(
<geoCoordinate id="EE036FBEA96E0444FF1DCC7C9243F913" box="[159,313,1843,1869]" degrees="42" direction="north" minutes="12" orientation="latitude" pageId="1" pageNumber="362" precision="1" seconds="47.9" value="42.213306">42°12'47.9&quot;N</geoCoordinate>
<geoCoordinate id="EE036FBEA96E0444FEC2CC7C92A3F913" box="[320,473,1843,1869]" degrees="43" direction="east" minutes="19" orientation="longitude" pageId="1" pageNumber="362" precision="1" seconds="27.9" value="43.324417">43°19'27.9&quot;E</geoCoordinate>
, 
<elevation id="001AEE4AA96E0444FE66CC7D911DF913" box="[484,615,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="1" pageNumber="362" unit="m" value="725.0">
<quantity id="4CCFA49CA96E0444FE66CC7D9151F913" box="[484,555,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="1" pageNumber="362" unit="m" value="725.0">725 m</quantity>
a.s.l.
</elevation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B880979A96E0444FF45CC189098F9CB" blockId="1.[151,1437,1770,2013]" pageId="1" pageNumber="362">
<materialsCitation id="3B5F0324A96E0444FF45CC189098F9CB" ID-GBIF-Occurrence="4116970302" collectingDate="2021-10-08" collectorName="All &amp; E. Maghradze &amp; Sh. Barjadze &amp; L. Shavadze." country="Georgia" county="Male" elevation="725" latitude="42.213306" location="Kotiasklde Cave" longLatPrecision="1" longitude="43.324417" municipality="Kotia Cave" pageId="1" pageNumber="362" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Imereti" typeStatus="paratype">
<emphasis id="B943D56BA96E0444FF45CC189232F92F" bold="true" box="[199,328,1879,1905]" pageId="1" pageNumber="362">
<typeStatus id="548CB7DBA96E0444FF45CC189239F92F" box="[199,323,1879,1905]" pageId="1" pageNumber="362" type="paratype">Paratypes</typeStatus>
:
</emphasis>
The same locality as holotype. 
<specimenCount id="9D31C2F0A96E0444FD28CC189074F92F" box="[682,782,1879,1905]" pageId="1" pageNumber="362" type="female">1 female</specimenCount>
, 
<specimenCount id="9D31C2F0A96E0444FC9BCC18901EF92F" box="[793,868,1879,1905]" pageId="1" pageNumber="362" type="male">1 male</specimenCount>
CJM; 
<specimenCount id="9D31C2F0A96E0444FC32CC18976EF92F" box="[944,1044,1879,1905]" pageId="1" pageNumber="362" type="female">1 female</specimenCount>
, ISIZU-TO-T-00002; 
<date id="FF892FB9A96E0444FA88CC189616F92F" box="[1290,1388,1879,1905]" pageId="1" pageNumber="362" value="2021-10-08">
<collectingDate id="EFCDD651A96E0444FA88CC189616F92F" box="[1290,1388,1879,1905]" pageId="1" pageNumber="362" value="2021-10-08">8.x.2021</collectingDate>
</date>
. 
<collectorName id="26C26CAFA96E0444FAFACC1896E6F92F" box="[1400,1436,1879,1905]" pageId="1" pageNumber="362">All</collectorName>
specimens were collected by 
<collectorName id="26C26CAFA96E0444FE5DCC349105F9CB" box="[479,639,1915,1941]" pageId="1" pageNumber="362">E. Maghradze</collectorName>
, 
<collectorName id="26C26CAFA96E0444FD08CC349060F9CB" box="[650,794,1915,1941]" pageId="1" pageNumber="362">Sh. Barjadze</collectorName>
and 
<collectorName id="26C26CAFA96E0444FCD2CC349098F9CB" box="[848,994,1915,1941]" pageId="1" pageNumber="362">L. Shavadze.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32D5AF2A96E0447FF45CCD190E7FF43" lastPageId="2" lastPageNumber="363" pageId="1" pageNumber="362" type="diagnosis">
<paragraph id="8B880979A96E0447FF45CCD190E7FF43" blockId="1.[151,1437,1770,2013]" lastBlockId="2.[151,1437,151,573]" lastPageId="2" lastPageNumber="363" pageId="1" pageNumber="362">
<emphasis id="B943D56BA96E0444FF45CCD19238F9E6" bold="true" box="[199,322,1950,1976]" pageId="1" pageNumber="362">Diagnosis.</emphasis>
A minute, long-legged species with pale soft cuticle, generally similar in size and habitus to three other closely related 
<taxonomicName id="4C3772FAA96E0444FE00CC8C912DF983" box="[386,599,1987,2013]" pageId="1" pageNumber="362">
<emphasis id="B943D56BA96E0444FE00CC8C9284F983" box="[386,510,1987,2013]" italics="true" pageId="1" pageNumber="362">Nemaspela</emphasis>
species
</taxonomicName>
from the Georgian caves, i.e. 
<taxonomicName id="4C3772FAA96E0444FC28CC8C9663F983" authority="Martens, 2006" authorityName="Martens" authorityYear="2006" box="[938,1305,1987,2013]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="femorecurvata">
<emphasis id="B943D56BA96E0444FC28CC8C9716F983" box="[938,1132,1987,2013]" italics="true" pageId="1" pageNumber="362">N. femorecurvata</emphasis>
<bibRefCitation id="EFA67488A96E0444FBF6CC8C9663F983" author="Martens, J." box="[1140,1305,1987,2013]" pageId="1" pageNumber="362" pagination="145 - 210" refId="ref5792" refString="Martens, J. (2006) Weberknechte aus dem Kaukasus (Arachnida, Opiliones, Nemastomatidae). Senckenbergiana biologica, 86, 145 - 210." type="journal article" year="2006">Martens, 2006</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3772FAA96E0447FAA6CC8C912DFEEF" authority="Martens, Maghradze &amp; Barjadze, 2021" authorityName="Martens, Maghradze &amp; Sh. Barjadze" authorityYear="2021" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" lastPageId="2" lastPageNumber="363" order="Opiliones" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="melouri">
<emphasis id="B943D56BA96E0444FAA6CC8C96E6F983" box="[1316,1436,1987,2013]" italics="true" pageId="1" pageNumber="362">N. melouri</emphasis>
<bibRefCitation id="EFA67488A96D0447FF15CBD8912DFEEF" author="Martens, J. &amp; Maghradze, E. &amp; Barjadze, S." box="[151,599,151,177]" pageId="2" pageNumber="363" pagination="541 - 558" refId="ref5820" refString="Martens, J., Maghradze, E. &amp; Barjadze, S. (2021) Two new species of the genus Nemaspela Silhavy from caves in Georgia (Opiliones: Nemastomatidae). Zootaxa, 4951 (3), 541 - 558. https: // doi. org / 10.11646 / zootaxa. 4951.3.7" type="journal article" year="2021">Martens, Maghradze &amp; Barjadze, 2021</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C3772FAA96D0447FD17CBD89784FEEF" authority="Martens, Maghradze &amp; Barjadze, 2021" authorityName="Martens, Maghradze &amp; Barjadze" authorityYear="2021" box="[661,1278,151,177]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="363" phylum="Arthropoda" rank="species" species="prometheus">
<emphasis id="B943D56BA96D0447FD17CBD89043FEEF" box="[661,825,151,177]" italics="true" pageId="2" pageNumber="363">N. prometheus</emphasis>
<bibRefCitation id="EFA67488A96D0447FCC0CBD89784FEEF" author="Martens, J. &amp; Maghradze, E. &amp; Barjadze, S." box="[834,1278,151,177]" pageId="2" pageNumber="363" pagination="541 - 558" refId="ref5820" refString="Martens, J., Maghradze, E. &amp; Barjadze, S. (2021) Two new species of the genus Nemaspela Silhavy from caves in Georgia (Opiliones: Nemastomatidae). Zootaxa, 4951 (3), 541 - 558. https: // doi. org / 10.11646 / zootaxa. 4951.3.7" type="journal article" year="2021">Martens, Maghradze &amp; Barjadze, 2021</bibRefCitation>
</taxonomicName>
characterized by specific genital morphology, i.e. extremely slender truncus and small size of basal inflated part of truncus, glans penis in dorsal/ventral view and lateral view markedly slenderer than in 
<taxonomicName id="4C3772FAA96D0447FC5CCB909722FEA7" authorityName="Martens, Maghradze &amp; Sh. Barjadze" authorityYear="2021" box="[990,1112,223,249]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="363" phylum="Arthropoda" rank="species" species="melouri">
<emphasis id="B943D56BA96D0447FC5CCB909722FEA7" box="[990,1112,223,249]" italics="true" pageId="2" pageNumber="363">N. melouri</emphasis>
</taxonomicName>
and 
<emphasis id="B943D56BA96D0447FB16CB90963AFEA7" box="[1172,1344,223,249]" italics="true" pageId="2" pageNumber="363">
<taxonomicName id="4C3772FAA96D0447FB16CB909647FEA7" authorityName="Martens, Maghradze &amp; Barjadze" authorityYear="2021" box="[1172,1341,223,249]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="363" phylum="Arthropoda" rank="species" species="prometheus">N. prometheus</taxonomicName>
,
</emphasis>
truncus below glans slightly inflated, glans distinctly separated from truncus.
</paragraph>
</subSubSection>
<subSubSection id="C32D5AF2A96D0443FF45CA699153F9F4" lastPageId="6" lastPageNumber="367" pageId="2" pageNumber="363" type="description">
<paragraph id="8B880979A96D0447FF45CA69977BFF8E" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FF45CA699223FF1E" bold="true" box="[199,345,294,320]" pageId="2" pageNumber="363">Description.</emphasis>
Male (n 
<emphasis id="B943D56BA96D0447FE3DCA6892B5FF1F" bold="true" box="[447,463,295,321]" pageId="2" pageNumber="363">=</emphasis>
2). Body length: 1.8, 2.0. 
<emphasis id="B943D56BA96D0447FC82CA6890B3FF1F" box="[768,969,295,321]" italics="true" pageId="2" pageNumber="363">Body, dorsal side</emphasis>
(
<figureCitation id="130C15FCA96D0447FC59CA68975CFF1F" box="[987,1062,295,321]" captionStart="FIGURE 1" captionStartId="2.[151,250,1945,1969]" captionTargetBox="[151,1436,958,1920]" captionTargetId="figure-363@2.[151,1436,957,1920]" captionTargetPageId="2" captionText="FIGURE 1. Nemaspela kotia sp. nov. at the type locality, Kotia cave, presumably female in lateral view. Photograph by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.7960308" httpUri="https://zenodo.org/record/7960308/files/figure.png" pageId="2" pageNumber="363">Figs 1</figureCitation>
, 
<figureCitation id="130C15FCA96D0447FBB5CA68972DFF1F" box="[1079,1111,295,321]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="2" pageNumber="363">10</figureCitation>
): Prosoma and opisthosoma forming two well discernible body parts because of a depression between them, opisthosoma strongly globular, less so the prosoma, eyes and Tu oc absent; dorsal scutal parts of prosoma and opisthosoma uniformly light yellowish, smooth, difficult to distinguish from neighbouring exoskeletal parts, without any distinct spots or markings, no setae present on dorsal side except for few minute ones on rear end of opisthosoma.
</paragraph>
<paragraph id="8B880979A96D0447FF45CA94962DFC63" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FF45CA9492ECFFAB" box="[199,406,475,501]" italics="true" pageId="2" pageNumber="363">Body, ventral side</emphasis>
(
<figureCitation id="130C15FCA96D0447FE2BCA94928BFFAB" box="[425,497,475,501]" captionStart="FIGURES 2-3" captionStartId="3.[151,264,878,902]" captionTargetBox="[151,1436,181,854]" captionTargetId="figure-18@3.[151,1436,181,854]" captionTargetPageId="3" captionText="FIGURES 2-3. Nemaspela kotia sp. nov. Preserved specimens, lateral view. 2. Male, 3. Female (bar: 2 mm). Photographs by Jean Severin." figureDoi="http://doi.org/10.5281/zenodo.7960310" httpUri="https://zenodo.org/record/7960310/files/figure.png" pageId="2" pageNumber="363">Fig. 2</figureCitation>
): Op gen and the Cx I–IV brownish contrasting with the much lighter remaining ventral side, irregularly dark brown to blackish spots on Op gen and Cx I–IV depending on interval from final moult, all sternites with a single row of black setae contrasting with light cuticle, Op gen with a few scattered setae.
</paragraph>
<caption id="DF4859F1A96D0447FF1AC9239253FCF7" ID-Table-UUID="DF4859F1A96D0447FF1AC9239253FCF7" httpUri="http://table.plazi.org/id/DF4859F1A96D0447FF1AC9239253FCF7" pageId="2" pageNumber="363" startId="2.[152,245,620,644]" targetBox="[163,1377,696,932]" targetIsTable="true" targetPageId="2" targetType="table">
<paragraph id="8B880979A96D0447FF1AC9239253FCF7" blockId="2.[151,1437,620,681]" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FF1AC9239277FCDA" bold="true" box="[152,269,620,644]" pageId="2" pageNumber="363">TABLE 1.</emphasis>
Measurements (mm) of legs and pedipalp in male holotype and female paratype (in parentheses) of 
<taxonomicName id="4C3772FAA96D0447FAA4C92393B6FCF6" authority="Martens &amp; Maghradze &amp; Barjadze, 2023" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2023" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="363" phylum="Arthropoda" rank="species" species="kotia" status="sp. nov.">
<emphasis id="B943D56BA96D0447FAA4C92393B6FCF6" italics="true" pageId="2" pageNumber="363">Nemaspela kotia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2706810A96D0447FF51C9DE9253FCF7" box="[211,297,657,681]" pageId="2" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B880979A96D0447FF21C9F7963DFDFA" pageId="2" pageNumber="363">
<table id="F937FBD9A96DFBBAFF21C9F7961BFDFA" box="[163,1377,696,932]" gridcols="8" gridrows="6" pageId="2" pageNumber="363">
<tr id="35070B3BA96DFBBAFF21C9F7961BFC8E" box="[163,1377,696,720]" gridrow="0" pageId="2" pageNumber="363">
<th id="76D66247A96DFBBAFF21C9F79279FC8E" box="[163,259,696,720]" gridcol="0" gridrow="0" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FF21C9F793B6FC8E" bold="true" box="[163,204,696,720]" pageId="2" pageNumber="363">Leg</emphasis>
</th>
<th id="76D66247A96DFBBAFEC3C9F792E2FC8E" box="[321,408,696,720]" gridcol="1" gridrow="0" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FEC3C9F79221FC8E" bold="true" box="[321,347,696,720]" pageId="2" pageNumber="363">Tr</emphasis>
</th>
<th id="76D66247A96DFBBAFE62C9F7914DFC8E" box="[480,567,696,720]" gridcol="2" gridrow="0" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FE62C9F79281FC8E" bold="true" box="[480,507,696,720]" pageId="2" pageNumber="363">Fe</emphasis>
</th>
<th id="76D66247A96DFBBAFDFCC9F791AFFC8E" box="[638,725,696,720]" gridcol="3" gridrow="0" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FDFCC9F791ECFC8E" bold="true" box="[638,662,696,720]" pageId="2" pageNumber="363">Pt</emphasis>
</th>
<th id="76D66247A96DFBBAFC9EC9F79009FC8E" box="[796,883,696,720]" gridcol="4" gridrow="0" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FC9EC9F79049FC8E" bold="true" box="[796,819,696,720]" pageId="2" pageNumber="363">Ti</emphasis>
</th>
<th id="76D66247A96DFBBAFC38C9F7976BFC8E" box="[954,1041,696,720]" gridcol="5" gridrow="0" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FC38C9F790A0FC8E" bold="true" box="[954,986,696,720]" pageId="2" pageNumber="363">Mt</emphasis>
</th>
<th id="76D66247A96DFBBAFBDAC9F797D5FC8E" box="[1112,1199,696,720]" gridcol="6" gridrow="0" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FBDAC9F79709FC8E" bold="true" box="[1112,1139,696,720]" pageId="2" pageNumber="363">Ta</emphasis>
</th>
<th id="76D66247A96DFBBAFB73C9F7961BFC8E" box="[1265,1377,696,720]" gridcol="7" gridrow="0" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FB73C9F79652FC8E" bold="true" box="[1265,1320,696,720]" pageId="2" pageNumber="363">Total</emphasis>
</th>
</tr>
<tr id="35070B3BA96DFBBAFF21C9A4961BFD5D" box="[163,1377,747,771]" gridrow="1" pageId="2" pageNumber="363">
<th id="76D66247A96DFBBAFF21C9A49279FD5D" box="[163,259,747,771]" gridcol="0" gridrow="1" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FF21C9A493D7FD5D" bold="true" box="[163,173,747,771]" pageId="2" pageNumber="363">I</emphasis>
</th>
<td id="76D66247A96DFBBAFEC3C9A492E2FD5D" box="[321,408,747,771]" gridcol="1" gridrow="1" pageId="2" pageNumber="363">0.2 (0.2)</td>
<td id="76D66247A96DFBBAFE62C9A4914DFD5D" box="[480,567,747,771]" gridcol="2" gridrow="1" pageId="2" pageNumber="363">3.2 (3.2)</td>
<td id="76D66247A96DFBBAFDFCC9A491AFFD5D" box="[638,725,747,771]" gridcol="3" gridrow="1" pageId="2" pageNumber="363">0.6 (0.6)</td>
<td id="76D66247A96DFBBAFC9EC9A49009FD5D" box="[796,883,747,771]" gridcol="4" gridrow="1" pageId="2" pageNumber="363">2.0 (2.1)</td>
<td id="76D66247A96DFBBAFC38C9A4976BFD5D" box="[954,1041,747,771]" gridcol="5" gridrow="1" pageId="2" pageNumber="363">4.3 (4.2)</td>
<td id="76D66247A96DFBBAFBDAC9A497D5FD5D" box="[1112,1199,747,771]" gridcol="6" gridrow="1" pageId="2" pageNumber="363">2.0 (1.9)</td>
<td id="76D66247A96DFBBAFB73C9A4961BFD5D" box="[1265,1377,747,771]" gridcol="7" gridrow="1" pageId="2" pageNumber="363">12.3 (12.2)</td>
</tr>
<tr id="35070B3BA96DFBBAFF21C85C961BFD75" box="[163,1377,787,811]" gridrow="2" pageId="2" pageNumber="363">
<th id="76D66247A96DFBBAFF21C85C9279FD75" box="[163,259,787,811]" gridcol="0" gridrow="2" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FF21C85C93CDFD75" bold="true" box="[163,183,787,811]" pageId="2" pageNumber="363">II</emphasis>
</th>
<td id="76D66247A96DFBBAFEC3C85C92E2FD75" box="[321,408,787,811]" gridcol="1" gridrow="2" pageId="2" pageNumber="363">0.3 (0.3)</td>
<td id="76D66247A96DFBBAFE62C85C914DFD75" box="[480,567,787,811]" gridcol="2" gridrow="2" pageId="2" pageNumber="363">5.2 (5.7)</td>
<td id="76D66247A96DFBBAFDFCC85C91AFFD75" box="[638,725,787,811]" gridcol="3" gridrow="2" pageId="2" pageNumber="363">0.7 (0.7)</td>
<td id="76D66247A96DFBBAFC9EC85C9009FD75" box="[796,883,787,811]" gridcol="4" gridrow="2" pageId="2" pageNumber="363">4.9 (4.8)</td>
<td id="76D66247A96DFBBAFC38C85C976BFD75" box="[954,1041,787,811]" gridcol="5" gridrow="2" pageId="2" pageNumber="363">8.2 (7.8)</td>
<td id="76D66247A96DFBBAFBDAC85C97D5FD75" box="[1112,1199,787,811]" gridcol="6" gridrow="2" pageId="2" pageNumber="363">4.3 (3.9)</td>
<td id="76D66247A96DFBBAFB73C85C961BFD75" box="[1265,1377,787,811]" gridcol="7" gridrow="2" pageId="2" pageNumber="363">23.6 (23.2)</td>
</tr>
<tr id="35070B3BA96DFBBAFF21C874961BFD0A" box="[163,1377,827,852]" gridrow="3" pageId="2" pageNumber="363">
<th id="76D66247A96DFBBAFF21C8749279FD0A" box="[163,259,827,852]" gridcol="0" gridrow="3" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FF21C87493BBFD0D" bold="true" box="[163,193,827,851]" pageId="2" pageNumber="363">III</emphasis>
</th>
<td id="76D66247A96DFBBAFEC3C87492E2FD0A" box="[321,408,827,852]" gridcol="1" gridrow="3" pageId="2" pageNumber="363">0.2 (0.2)</td>
<td id="76D66247A96DFBBAFE62C874914DFD0A" box="[480,567,827,852]" gridcol="2" gridrow="3" pageId="2" pageNumber="363">3.5 (3.4)</td>
<td id="76D66247A96DFBBAFDFCC87491AFFD0A" box="[638,725,827,852]" gridcol="3" gridrow="3" pageId="2" pageNumber="363">0.6 (0.6)</td>
<td id="76D66247A96DFBBAFC9EC8749009FD0A" box="[796,883,827,852]" gridcol="4" gridrow="3" pageId="2" pageNumber="363">2.1 (2.0)</td>
<td id="76D66247A96DFBBAFC38C874976BFD0A" box="[954,1041,827,852]" gridcol="5" gridrow="3" pageId="2" pageNumber="363">4.7 (4.5)</td>
<td id="76D66247A96DFBBAFBDAC87497D5FD0A" box="[1112,1199,827,852]" gridcol="6" gridrow="3" pageId="2" pageNumber="363">2.1 (2.9)</td>
<td id="76D66247A96DFBBAFB73C874961BFD0A" box="[1265,1377,827,852]" gridcol="7" gridrow="3" pageId="2" pageNumber="363">13.2 (13.6)</td>
</tr>
<tr id="35070B3BA96DFBBAFF21C82C961BFD22" box="[163,1377,867,892]" gridrow="4" pageId="2" pageNumber="363">
<th id="76D66247A96DFBBAFF21C82C9279FD22" box="[163,259,867,892]" gridcol="0" gridrow="4" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FF21C82C93C5FD25" bold="true" box="[163,191,867,891]" pageId="2" pageNumber="363">IV</emphasis>
</th>
<td id="76D66247A96DFBBAFEC3C82C92E2FD22" box="[321,408,867,892]" gridcol="1" gridrow="4" pageId="2" pageNumber="363">0.2 (0.2)</td>
<td id="76D66247A96DFBBAFE62C82C914DFD22" box="[480,567,867,892]" gridcol="2" gridrow="4" pageId="2" pageNumber="363">4.8 (4.9)</td>
<td id="76D66247A96DFBBAFDFCC82C91AFFD22" box="[638,725,867,892]" gridcol="3" gridrow="4" pageId="2" pageNumber="363">0.6 (0.6)</td>
<td id="76D66247A96DFBBAFC9EC82C9009FD22" box="[796,883,867,892]" gridcol="4" gridrow="4" pageId="2" pageNumber="363">2.8 (3.0)</td>
<td id="76D66247A96DFBBAFC38C82C976BFD22" box="[954,1041,867,892]" gridcol="5" gridrow="4" pageId="2" pageNumber="363">6.5 (6.5)</td>
<td id="76D66247A96DFBBAFBDAC82C97D5FD22" box="[1112,1199,867,892]" gridcol="6" gridrow="4" pageId="2" pageNumber="363">2.7 (2.2)</td>
<td id="76D66247A96DFBBAFB73C82C961BFD22" box="[1265,1377,867,892]" gridcol="7" gridrow="4" pageId="2" pageNumber="363">17.6 (17.4)</td>
</tr>
<tr id="35070B3BA96DFBBAFF21C8C4961BFDFA" box="[163,1377,907,932]" gridrow="5" pageId="2" pageNumber="363">
<th id="76D66247A96DFBBAFF21C8C49279FDFA" box="[163,259,907,932]" gridcol="0" gridrow="5" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FF21C8C49279FDFD" bold="true" box="[163,259,907,931]" pageId="2" pageNumber="363">Pedipalp</emphasis>
</th>
<td id="76D66247A96DFBBAFEC3C8C492E2FDFA" box="[321,408,907,932]" gridcol="1" gridrow="5" pageId="2" pageNumber="363">0.7 (0.6)</td>
<td id="76D66247A96DFBBAFE62C8C4914DFDFA" box="[480,567,907,932]" gridcol="2" gridrow="5" pageId="2" pageNumber="363">1.7 (1.7)</td>
<td id="76D66247A96DFBBAFDFCC8C491AFFDFA" box="[638,725,907,932]" gridcol="3" gridrow="5" pageId="2" pageNumber="363">1.7 (1.6)</td>
<td id="76D66247A96DFBBAFC9EC8C49009FDFA" box="[796,883,907,932]" gridcol="4" gridrow="5" pageId="2" pageNumber="363">1.4 (1.3)</td>
<td id="76D66247A96DFBBAFC38C8C4976BFDFA" box="[954,1041,907,932]" gridcol="5" gridrow="5" pageId="2" pageNumber="363">-- (--)</td>
<td id="76D66247A96DFBBAFBDAC8C497D5FDFA" box="[1112,1199,907,932]" gridcol="6" gridrow="5" pageId="2" pageNumber="363">0.9 (0.8)</td>
<td id="76D66247A96DFBBAFB73C8C4961BFDFA" box="[1265,1377,907,932]" gridcol="7" gridrow="5" pageId="2" pageNumber="363">6.4 (6.0)</td>
</tr>
</table>
</paragraph>
<caption id="DF4859F1A96D0447FF15CCD693ADF98B" ID-DOI="http://doi.org/10.5281/zenodo.7960308" ID-Zenodo-Dep="7960308" httpUri="https://zenodo.org/record/7960308/files/figure.png" pageId="2" pageNumber="363" startId="2.[151,250,1945,1969]" targetBox="[151,1436,958,1920]" targetPageId="2" targetType="figure">
<paragraph id="8B880979A96D0447FF15CCD693ADF98B" blockId="2.[151,1437,1945,2005]" pageId="2" pageNumber="363">
<emphasis id="B943D56BA96D0447FF15CCD69268F9EF" bold="true" box="[151,274,1945,1969]" pageId="2" pageNumber="363">FIGURE 1.</emphasis>
<taxonomicName id="4C3772FAA96D0447FE9ACCD692BBF9EF" authority="Martens &amp; Maghradze &amp; Barjadze, 2023" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2023" box="[280,449,1945,1969]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="363" phylum="Arthropoda" rank="species" species="kotia" status="sp. nov.">
<emphasis id="B943D56BA96D0447FE9ACCD692BBF9EF" box="[280,449,1945,1969]" italics="true" pageId="2" pageNumber="363">Nemaspela kotia</emphasis>
</taxonomicName>
<emphasis id="B943D56BA96D0447FE44CCD6916CF9EF" bold="true" box="[454,534,1945,1969]" pageId="2" pageNumber="363">
<taxonomicNameLabel id="A2706810A96D0447FE44CCD6916CF9EF" box="[454,534,1945,1969]" pageId="2" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at the type locality, Kotia cave, presumably female in lateral view. Photograph by Dragan Antić.
</paragraph>
</caption>
<caption id="DF4859F1A96C0446FF15C8219265FDF4" ID-DOI="http://doi.org/10.5281/zenodo.7960310" ID-Zenodo-Dep="7960310" httpUri="https://zenodo.org/record/7960310/files/figure.png" pageId="3" pageNumber="364" startId="3.[151,264,878,902]" targetBox="[151,1436,181,854]" targetPageId="3" targetType="figure">
<paragraph id="8B880979A96C0446FF15C8219265FDF4" blockId="3.[151,1436,878,938]" pageId="3" pageNumber="364">
<emphasis id="B943D56BA96C0446FF15C821924DFDD8" bold="true" box="[151,311,878,902]" pageId="3" pageNumber="364">FIGURES 2-3.</emphasis>
<taxonomicName id="4C3772FAA96C0446FEBCC8219293FDD8" authority="Martens &amp; Maghradze &amp; Barjadze, 2023" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2023" box="[318,489,878,902]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="kotia" status="sp. nov.">
<emphasis id="B943D56BA96C0446FEBCC8219293FDD8" box="[318,489,878,902]" italics="true" pageId="3" pageNumber="364">Nemaspela kotia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2706810A96C0446FE6DC8219138FDD8" box="[495,578,878,902]" pageId="3" pageNumber="364" rank="species">
<emphasis id="B943D56BA96C0446FE6DC8219146FDD8" bold="true" box="[495,572,878,902]" pageId="3" pageNumber="364">sp. nov</emphasis>
.
</taxonomicNameLabel>
Preserved specimens, lateral view. 
<emphasis id="B943D56BA96C0446FC2CC82190BBFDD8" bold="true" box="[942,961,878,902]" pageId="3" pageNumber="364">2.</emphasis>
Male, 
<emphasis id="B943D56BA96C0446FB88C8219767FDD8" bold="true" box="[1034,1053,878,902]" pageId="3" pageNumber="364">3.</emphasis>
Female (bar: 2 mm). Photographs by Jean Severin.
</paragraph>
</caption>
<paragraph id="8B880979A96C0446FF45C89793ADFA95" blockId="3.[151,1437,984,1983]" pageId="3" pageNumber="364">
<emphasis id="B943D56BA96C0446FF45C897924FFDAC" box="[199,309,984,1010]" italics="true" pageId="3" pageNumber="364">Chelicera</emphasis>
(
<figureCitation id="130C15FCA96C0446FECAC89792B5FDAC" box="[328,463,984,1010]" captionStart="FIGURES 12-16" captionStartId="6.[151,264,1398,1422]" captionTargetBox="[156,1431,314,1370]" captionTargetId="figure-95@6.[151,1436,309,1375]" captionTargetPageId="6" captionText="FIGURES 12-16. Nemaspela kotia sp. nov. Chelicerae in retro-lateral view. 12. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Chelicera in pro-lateral view. 13. Male (bar: 0.3 mm). 15. Female (bar: 0.3 mm). 16. Patellar apophysis of female, retro-lateral views (bar: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960314" httpUri="https://zenodo.org/record/7960314/files/figure.png" pageId="3" pageNumber="364">Figs 12–13</figureCitation>
): Very slender and elongated; basal article slightly constricted close to its proximal end, from there slightly and continuously widened distally, distal third slightly bent on ventral side, several setae on dorsal side at proximal portion, additionally two long setae dorsal sub-distally, no dorsal Apo on basal article present; 2 
<superScript id="7C42A431A96C0446FE86CF0B926EFA0C" attach="left" box="[260,276,1092,1106]" fontSize="6" pageId="3" pageNumber="364">nd</superScript>
cheliceral article slender and longish, slightly inflated and nearly parallel-sided (in pro-lateral view, 
<figureCitation id="130C15FCA96C0446FAF2CF0B93C9FADC" captionStart="FIGURES 12-16" captionStartId="6.[151,264,1398,1422]" captionTargetBox="[156,1431,314,1370]" captionTargetId="figure-95@6.[151,1436,309,1375]" captionTargetPageId="6" captionText="FIGURES 12-16. Nemaspela kotia sp. nov. Chelicerae in retro-lateral view. 12. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Chelicera in pro-lateral view. 13. Male (bar: 0.3 mm). 15. Female (bar: 0.3 mm). 16. Patellar apophysis of female, retro-lateral views (bar: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960314" httpUri="https://zenodo.org/record/7960314/files/figure.png" pageId="3" pageNumber="364">Fig. 13</figureCitation>
) but slightly enlarged toward movable third article, with a loose tuft of long and strong setae all over the frontal surface, several shorter ones densely packed ventro-distally. On second article pro-laterally a group of few shorter setae.
</paragraph>
<paragraph id="8B880979A96C0446FF45CF9B92C2FBB4" blockId="3.[151,1437,984,1983]" pageId="3" pageNumber="364">
<emphasis id="B943D56BA96C0446FF45CF9B9256FAB0" box="[199,300,1236,1262]" italics="true" pageId="3" pageNumber="364">Pedipalp</emphasis>
(
<figureCitation id="130C15FCA96C0446FEB9CF9B92FFFAB0" box="[315,389,1236,1262]" captionStart="FIGURES 2-3" captionStartId="3.[151,264,878,902]" captionTargetBox="[151,1436,181,854]" captionTargetId="figure-18@3.[151,1436,181,854]" captionTargetPageId="3" captionText="FIGURES 2-3. Nemaspela kotia sp. nov. Preserved specimens, lateral view. 2. Male, 3. Female (bar: 2 mm). Photographs by Jean Severin." figureDoi="http://doi.org/10.5281/zenodo.7960310" httpUri="https://zenodo.org/record/7960310/files/figure.png" pageId="3" pageNumber="364">Figs 2</figureCitation>
, 
<figureCitation id="130C15FCA96C0446FE13CF9B92D8FAB0" box="[401,418,1236,1262]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="3" pageNumber="364">4</figureCitation>
): Very slender and extremely long (in terms of 
<taxonomicName id="4C3772FAA96C0446FC34CF9B9748FAB0" authorityName="Silhavy" authorityYear="1966" box="[950,1074,1236,1262]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="B943D56BA96C0446FC34CF9B9748FAB0" box="[950,1074,1236,1262]" italics="true" pageId="3" pageNumber="364">Nemaspela</emphasis>
</taxonomicName>
morphology), all articles nearly parallel-sided, Fe slightly widened towards distal end, distal end slightly more enlarged and bent downwards, Pt slender, slightly tapering towards distal end, distal fifth slightly bent to ventral side, Ti slightly shorter than Fe, tapering towards distal end, distal half slightly curved upwards, Ta slightly curved to ventral side, slightly enlarged toward distal part, all articles except Tr bearing scattered clavate setae, these most conspicuous on Fe ventrally, few on dorsal side, on Pt, Ti and Ta on all sides and densely packed. Setae on Fe longer than breadth of article at the position, generally slightly shorter on dorsal side; on Ti and Ta of the same length. No article with special armament in males (but see female).
</paragraph>
<paragraph id="8B880979A96C0446FF45CEBA915CF89C" blockId="3.[151,1437,984,1983]" pageId="3" pageNumber="364">
<emphasis id="B943D56BA96C0446FF45CEBA9386F850" box="[199,252,1525,1550]" italics="true" pageId="3" pageNumber="364">Legs</emphasis>
(
<figureCitation id="130C15FCA96C0446FE8FCEBB920FF850" box="[269,373,1524,1550]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2-3" captionStartId-0="2.[151,250,1945,1969]" captionStartId-1="3.[151,264,878,902]" captionTargetBox-0="[151,1436,958,1920]" captionTargetBox-1="[151,1436,181,854]" captionTargetId-0="figure-363@2.[151,1436,957,1920]" captionTargetId-1="figure-18@3.[151,1436,181,854]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Nemaspela kotia sp. nov. at the type locality, Kotia cave, presumably female in lateral view. Photograph by Dragan Antić." captionText-1="FIGURES 2-3. Nemaspela kotia sp. nov. Preserved specimens, lateral view. 2. Male, 3. Female (bar: 2 mm). Photographs by Jean Severin." figureDoi-0="http://doi.org/10.5281/zenodo.7960308" figureDoi-1="http://doi.org/10.5281/zenodo.7960310" httpUri-0="https://zenodo.org/record/7960308/files/figure.png" httpUri-1="https://zenodo.org/record/7960310/files/figure.png" pageId="3" pageNumber="364">Figs 1–2</figureCitation>
): Light brownish tending to yellowish brownish and yellowish toward distal end, Tr mottled dark brownish, articles (in comparison to body size) extremely long and slender, thread-like, round in crosssection, armed with some scattered short black setae. Many pseudo-articulations on Fe I–IV (
<figureCitation id="130C15FCA96C0446FB4ACD739655F808" box="[1224,1327,1596,1622]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2-3" captionStartId-0="2.[151,250,1945,1969]" captionStartId-1="3.[151,264,878,902]" captionTargetBox-0="[151,1436,958,1920]" captionTargetBox-1="[151,1436,181,854]" captionTargetId-0="figure-363@2.[151,1436,957,1920]" captionTargetId-1="figure-18@3.[151,1436,181,854]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Nemaspela kotia sp. nov. at the type locality, Kotia cave, presumably female in lateral view. Photograph by Dragan Antić." captionText-1="FIGURES 2-3. Nemaspela kotia sp. nov. Preserved specimens, lateral view. 2. Male, 3. Female (bar: 2 mm). Photographs by Jean Severin." figureDoi-0="http://doi.org/10.5281/zenodo.7960308" figureDoi-1="http://doi.org/10.5281/zenodo.7960310" httpUri-0="https://zenodo.org/record/7960308/files/figure.png" httpUri-1="https://zenodo.org/record/7960310/files/figure.png" pageId="3" pageNumber="364">Figs 1–2</figureCitation>
), densely packed near basis of articles, towards distal end of article distances between them consecutively more extended; no pseudo-articulations near distal end. Number of femoral pseudo-articulations: Fe I 15, Fe II 19, Fe III 13, Fe IV 18. Measurements are given in 
<tableCitation id="C6B53CC2A96C0446FE4ECDE7915AF89C" box="[460,544,1704,1730]" captionStart="TABLE 1" captionStartId="2.[152,245,620,644]" captionTargetPageId="2" captionText="TABLE 1. Measurements (mm) of legs and pedipalp in male holotype and female paratype (in parentheses) of Nemaspela kotia sp. nov." httpUri="http://table.plazi.org/id/DF4859F1A96D0447FF1AC9239253FCF7" pageId="3" pageNumber="364" tableUuid="DF4859F1A96D0447FF1AC9239253FCF7">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8B880979A96C0446FF45CD8392A7F9E0" blockId="3.[151,1437,984,1983]" pageId="3" pageNumber="364">
<emphasis id="B943D56BA96C0446FF45CD83929BF8B8" box="[199,481,1740,1766]" italics="true" pageId="3" pageNumber="364">Male genital morphology</emphasis>
(
<figureCitation id="130C15FCA96C0446FE6DCD83912FF8B8" box="[495,597,1740,1766]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="3" pageNumber="364">Figs 6–9</figureCitation>
): Truncus penis (
<figureCitation id="130C15FCA96C0446FC92CD839009F8B8" box="[784,883,1740,1766]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="3" pageNumber="364">Figs 6–7</figureCitation>
) extremely slender; basis forming a small inflated part (occupying about one fifth of whole penis length) and markedly separated from the rest of the truncus; inflated part slender, evenly broad, laterally rounded and deeply split medially, completely filled by two penial muscles, their tendons spanning the hole truncus length up to the basis of the glans. Truncus nearly parallel-sided in dorsal/ ventral view nearly all over its length (
<figureCitation id="130C15FCA96C0446FDC8CC1391F5F928" box="[586,655,1884,1910]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="3" pageNumber="364">Fig. 6</figureCitation>
), but slightly narrower subterminally, than broadening up close to the glans from where slightly narrowed to glans basis; in lateral view (
<figureCitation id="130C15FCA96C0446FC0ACCCF90B6F9C4" box="[904,972,1920,1946]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="3" pageNumber="364">Fig. 7</figureCitation>
) truncus narrow as well, strictly parallel-sided over most of its length.
</paragraph>
<caption id="DF4859F1A96B0441FF15CCD8974AF9A6" ID-DOI="http://doi.org/10.5281/zenodo.7960312" ID-Zenodo-Dep="7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="4" pageNumber="365" startId="4.[151,264,1943,1967]" targetBox="[185,1411,186,1920]" targetPageId="4" targetType="figure">
<paragraph id="8B880979A96B0441FF15CCD8974AF9A6" blockId="4.[151,1437,1943,2040]" pageId="4" pageNumber="365">
<emphasis id="B943D56BA96B0441FF15CCD8923EF9F1" bold="true" box="[151,324,1943,1967]" pageId="4" pageNumber="365">FIGURES 4-11.</emphasis>
<taxonomicName id="4C3772FAA96B0441FECECCD79282F9EE" authority="Martens &amp; Maghradze &amp; Barjadze, 2023" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2023" box="[332,504,1944,1968]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="kotia" status="sp. nov.">
<emphasis id="B943D56BA96B0441FECECCD79282F9EE" box="[332,504,1944,1968]" italics="true" pageId="4" pageNumber="365">Nemaspela kotia</emphasis>
</taxonomicName>
<emphasis id="B943D56BA96B0441FD82CCD79128F9EE" bold="true" box="[512,594,1944,1968]" pageId="4" pageNumber="365">
<taxonomicNameLabel id="A2706810A96B0441FD82CCD79128F9EE" box="[512,594,1944,1968]" pageId="4" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Pedipalps in retro-lateral view. 
<emphasis id="B943D56BA96B0441FC22CCD890C9F9F1" bold="true" box="[928,947,1943,1967]" pageId="4" pageNumber="365">4.</emphasis>
Male (bar: 0.5 mm); 
<emphasis id="B943D56BA96B0441FB1ACCD797D0F9EE" bold="true" box="[1176,1194,1944,1968]" pageId="4" pageNumber="365">5.</emphasis>
Female (bar: 0.5 mm). Penis. 
<emphasis id="B943D56BA96B0441FF58CCF49397F98D" bold="true" box="[218,237,1979,2003]" pageId="4" pageNumber="365">6.</emphasis>
Ventral view (bar: 0.2 mm); 
<emphasis id="B943D56BA96B0441FD8DCCF39158F98A" bold="true" box="[527,546,1980,2004]" pageId="4" pageNumber="365">7.</emphasis>
Lateral view (bar: 0.2 mm). Glans penis. 
<emphasis id="B943D56BA96B0441FC46CCF490ADF98D" bold="true" box="[964,983,1979,2003]" pageId="4" pageNumber="365">8.</emphasis>
Ventral view (bar: 0.15 mm); 
<emphasis id="B943D56BA96B0441FA87CCF49662F98D" bold="true" box="[1285,1304,1979,2003]" pageId="4" pageNumber="365">9.</emphasis>
Lateral view (bar: 0.15 mm). Habitus in lateral view. 
<emphasis id="B943D56BA96B0441FDACCC909137F9A9" bold="true" box="[558,589,2015,2039]" pageId="4" pageNumber="365">10.</emphasis>
Female (bar: 1.0 mm); 
<emphasis id="B943D56BA96B0441FCBCCC909026F9A9" bold="true" box="[830,860,2015,2039]" pageId="4" pageNumber="365">11.</emphasis>
Male (bar: 1.0 mm).
</paragraph>
</caption>
<caption id="DF4859F1A96A0440FF1ACD659014F960" ID-DOI="http://doi.org/10.5281/zenodo.7971988" ID-Zenodo-Dep="7971988" httpUri="https://zenodo.org/record/7971988/files/figure.png" pageId="5" pageNumber="366" startId="5.[152,211,1578,1602]" targetBox="[151,1436,151,1554]" targetPageId="5">
<paragraph id="8B880979A96A0440FF1ACD659014F960" blockId="5.[151,1437,1578,1854]" pageId="5" pageNumber="366">
<emphasis id="B943D56BA96A0440FF1ACD659396F81C" bold="true" box="[152,236,1578,1602]" pageId="5" pageNumber="366">MAP 1.</emphasis>
Distributional map of the 
<taxonomicName id="4C3772FAA96A0440FE7ACD6591CDF81C" box="[504,695,1578,1602]" pageId="5" pageNumber="366">
<emphasis id="B943D56BA96A0440FE7ACD659113F81C" box="[504,617,1578,1602]" italics="true" pageId="5" pageNumber="366">Nemaspela</emphasis>
species
</taxonomicName>
in the Caucasus. 1. Vorontsovskaya Cave (
<taxonomicName id="4C3772FAA96A0440FBEFCD6497A5F81C" baseAuthorityName="Ljovuschkin &amp; Starobogatov" baseAuthorityYear="1963" box="[1133,1247,1578,1602]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="sokolovi">
<emphasis id="B943D56BA96A0440FBEFCD6497A5F81C" box="[1133,1247,1578,1602]" italics="true" pageId="5" pageNumber="366">N. sokolovi</emphasis>
</taxonomicName>
); 2. Labirintovaya Cave (
<taxonomicName id="4C3772FAA96A0440FF58CD009236F838" baseAuthorityName="Ljovuschkin &amp; Starobogatov" baseAuthorityYear="1963" box="[218,332,1614,1638]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="sokolovi">
<emphasis id="B943D56BA96A0440FF58CD009236F838" box="[218,332,1614,1638]" italics="true" pageId="5" pageNumber="366">N. sokolovi</emphasis>
</taxonomicName>
); 3. Dolgaya Cave (
<taxonomicName id="4C3772FAA96A0440FD9ECD0091F4F838" baseAuthorityName="Ljovuschkin &amp; Starobogatov" baseAuthorityYear="1963" box="[540,654,1614,1638]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="sokolovi">
<emphasis id="B943D56BA96A0440FD9ECD0091F4F838" box="[540,654,1614,1638]" italics="true" pageId="5" pageNumber="366">N. sokolovi</emphasis>
</taxonomicName>
); 4. Fontanka Cave (
<taxonomicName id="4C3772FAA96A0440FCE4CD0090B8F838" authorityName="Chemeris" authorityYear="2009" box="[870,962,1614,1638]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="kovali">
<emphasis id="B943D56BA96A0440FCE4CD0090B8F838" box="[870,962,1614,1638]" italics="true" pageId="5" pageNumber="366">N. kovali</emphasis>
</taxonomicName>
); 5. Omega-15 Cave (
<taxonomicName id="4C3772FAA96A0440FB24CD009678F838" authorityName="Chemeris" authorityYear="2009" box="[1190,1282,1614,1638]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="kovali">
<emphasis id="B943D56BA96A0440FB24CD009678F838" box="[1190,1282,1614,1638]" italics="true" pageId="5" pageNumber="366">N. kovali</emphasis>
</taxonomicName>
); 6. Omega-12 Cave (
<taxonomicName id="4C3772FAA96A0440FF59CD3C9242F8D4" authorityName="Chemeris" authorityYear="2009" box="[219,312,1650,1674]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="kovali">
<emphasis id="B943D56BA96A0440FF59CD3C9242F8D4" box="[219,312,1650,1674]" italics="true" pageId="5" pageNumber="366">N. kovali</emphasis>
</taxonomicName>
); 7. Otte-Shik Cave (
<taxonomicName id="4C3772FAA96A0440FD9BCD3C910CF8D4" authorityName="Chemeris" authorityYear="2009" box="[537,630,1650,1674]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="kovali">
<emphasis id="B943D56BA96A0440FD9BCD3C910CF8D4" box="[537,630,1650,1674]" italics="true" pageId="5" pageNumber="366">N. kovali</emphasis>
</taxonomicName>
); 8. Akhunskaya Cave (
<taxonomicName id="4C3772FAA96A0440FCF3CD3C908DF8D4" baseAuthorityName="Ljovushkin &amp; Starobogatov" baseAuthorityYear="1963" box="[881,1015,1650,1674]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="abchasica">
<emphasis id="B943D56BA96A0440FCF3CD3C908DF8D4" box="[881,1015,1650,1674]" italics="true" pageId="5" pageNumber="366">N. abchasica</emphasis>
</taxonomicName>
); 9. Beloskalskaya Cave (
<taxonomicName id="4C3772FAA96A0440FA85CD3C96F7F8D4" baseAuthorityName="Ljovushkin &amp; Starobogatov" baseAuthorityYear="1963" box="[1287,1421,1650,1674]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="abchasica">
<emphasis id="B943D56BA96A0440FA85CD3C96F7F8D4" box="[1287,1421,1650,1674]" italics="true" pageId="5" pageNumber="366">N. abchasica</emphasis>
</taxonomicName>
); 10. Kolokolnaya Cave (
<taxonomicName id="4C3772FAA96A0440FE05CDD89171F8F0" baseAuthorityName="Ljovushkin &amp; Starobogatov" baseAuthorityYear="1963" box="[391,523,1686,1710]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="abchasica">
<emphasis id="B943D56BA96A0440FE05CDD89171F8F0" box="[391,523,1686,1710]" italics="true" pageId="5" pageNumber="366">N. abchasica</emphasis>
</taxonomicName>
); 11. Kuibishevi Cave (
<taxonomicName id="4C3772FAA96A0440FD7ECDD890FAF8F0" baseAuthorityName="Ljovushkin &amp; Starobogatov" baseAuthorityYear="1963" box="[764,896,1686,1710]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="abchasica">
<emphasis id="B943D56BA96A0440FD7ECDD890FAF8F0" box="[764,896,1686,1710]" italics="true" pageId="5" pageNumber="366">N. abchasica</emphasis>
</taxonomicName>
); 12. L 24 Cave (
<taxonomicName id="4C3772FAA96A0440FBADCDD897C9F8F0" baseAuthorityName="Ljovushkin &amp; Starobogatov" baseAuthorityYear="1963" box="[1071,1203,1686,1710]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="abchasica">
<emphasis id="B943D56BA96A0440FBADCDD897C9F8F0" box="[1071,1203,1686,1710]" italics="true" pageId="5" pageNumber="366">N. abchasica</emphasis>
</taxonomicName>
); 13. Kveda Shakurani (=Nizhnyaya Shakuranskaya) Cave (
<taxonomicName id="4C3772FAA96A0440FD91CDF491E3F88C" baseAuthorityName="Ljovushkin &amp; Starobogatov" baseAuthorityYear="1963" box="[531,665,1722,1746]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="abchasica">
<emphasis id="B943D56BA96A0440FD91CDF491E3F88C" box="[531,665,1722,1746]" italics="true" pageId="5" pageNumber="366">N. abchasica</emphasis>
</taxonomicName>
); 14. Zemo Eshera Cave (
<taxonomicName id="4C3772FAA96A0440FC2CCDF4975BF88C" baseAuthorityName="Ljovushkin" baseAuthorityYear="1972" box="[942,1057,1722,1746]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="birsteini">
<emphasis id="B943D56BA96A0440FC2CCDF4975BF88C" box="[942,1057,1722,1746]" italics="true" pageId="5" pageNumber="366">N. birsteini</emphasis>
</taxonomicName>
); 15. Tarkili Cave (
<taxonomicName id="4C3772FAA96A0440FB73CDF4961EF88C" baseAuthorityName="Ljovushkin" baseAuthorityYear="1972" box="[1265,1380,1722,1746]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="birsteini">
<emphasis id="B943D56BA96A0440FB73CDF4961EF88C" box="[1265,1380,1722,1746]" italics="true" pageId="5" pageNumber="366">N. birsteini</emphasis>
</taxonomicName>
); 16. Sarma Cave (
<taxonomicName id="4C3772FAA96A0440FEA3CD9092F7F8A8" authorityName="Tchemeris" authorityYear="2013" box="[289,397,1759,1782]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="gagrica">
<emphasis id="B943D56BA96A0440FEA3CD9092F7F8A8" box="[289,397,1759,1782]" italics="true" pageId="5" pageNumber="366">N. gagrica</emphasis>
</taxonomicName>
); 17. Veryovkina Cave (
<taxonomicName id="4C3772FAA96A0440FD07CD90918BF8A8" authorityName="Tchemeris" authorityYear="2013" box="[645,753,1759,1782]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="gagrica">
<emphasis id="B943D56BA96A0440FD07CD90918BF8A8" box="[645,753,1759,1782]" italics="true" pageId="5" pageNumber="366">N. gagrica</emphasis>
</taxonomicName>
); 18. Sakishore Cave (
<taxonomicName id="4C3772FAA96A0440FC5BCD9097F0F8A8" authorityName="Martens" authorityYear="2006" box="[985,1162,1758,1782]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="femorecurvata">
<emphasis id="B943D56BA96A0440FC5BCD9097F0F8A8" box="[985,1162,1758,1782]" italics="true" pageId="5" pageNumber="366">N. femorecurvata</emphasis>
</taxonomicName>
); 19. 
<taxonomicName id="4C3772FAA96A0440FB47CD91960CF8A8" authority="Cave" authorityName="Cave" box="[1221,1398,1758,1782]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="prometheus">Prometheus Cave</taxonomicName>
(
<taxonomicName id="4C3772FAA96A0440FA07CD909276F944" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="prometheus">
<emphasis id="B943D56BA96A0440FA07CD909276F944" italics="true" pageId="5" pageNumber="366">N. prometheus</emphasis>
</taxonomicName>
); 20. Melouri Cave (
<taxonomicName id="4C3772FAA96A0440FE61CC4C9135F944" authorityName="Martens, Maghradze &amp; Sh. Barjadze" authorityYear="2021" box="[483,591,1794,1818]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="melouri">
<emphasis id="B943D56BA96A0440FE61CC4C9135F944" box="[483,591,1794,1818]" italics="true" pageId="5" pageNumber="366">N. melouri</emphasis>
</taxonomicName>
); 21. Solkota Cave (
<taxonomicName id="4C3772FAA96A0440FCA3CC4C90F7F944" authorityName="Martens, Maghradze &amp; Sh. Barjadze" authorityYear="2021" box="[801,909,1794,1818]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="melouri">
<emphasis id="B943D56BA96A0440FCA3CC4C90F7F944" box="[801,909,1794,1818]" italics="true" pageId="5" pageNumber="366">N. melouri</emphasis>
</taxonomicName>
); 22. Kotia Cave (=Kotiasklde Cave) (
<taxonomicName id="4C3772FAA96A0440FA95CC4C9612F944" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2023" box="[1303,1384,1794,1818]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="kotia">
<emphasis id="B943D56BA96A0440FA95CC4C9612F944" box="[1303,1384,1794,1818]" italics="true" pageId="5" pageNumber="366">N. kotia</emphasis>
</taxonomicName>
) (for details of localities and collection depositories see 
<bibRefCitation id="EFA67488A96A0440FD18CC689019F960" author="Martens, J. &amp; Maghradze, E. &amp; Barjadze, S." box="[666,867,1830,1854]" pageId="5" pageNumber="366" pagination="541 - 558" refId="ref5820" refString="Martens, J., Maghradze, E. &amp; Barjadze, S. (2021) Two new species of the genus Nemaspela Silhavy from caves in Georgia (Opiliones: Nemastomatidae). Zootaxa, 4951 (3), 541 - 558. https: // doi. org / 10.11646 / zootaxa. 4951.3.7" type="journal article" year="2021">
Martens 
<emphasis id="B943D56BA96A0440FD71CC68905CF960" box="[755,806,1830,1854]" italics="true" pageId="5" pageNumber="366">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="8B880979A96A0443FF45CC2391C5FF43" blockId="5.[151,1436,1900,2035]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="367" pageId="5" pageNumber="366">
Glans (
<figureCitation id="130C15FCA96A0440FE9ACC239207F9D8" box="[280,381,1900,1926]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="5" pageNumber="366">Figs 8–9</figureCitation>
) conspicuously separated from truncus (dorsal/ventral view), distinctly fusiform, just slightly broader than truncus at mid-part (
<figureCitation id="130C15FCA96A0440FDA3CCDF9113F9F4" box="[545,617,1936,1962]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="5" pageNumber="366">Fig. 9</figureCitation>
), short, starting where two tendons are attached to inner truncus wall, lateral outline only evenly rounded, continuously tapering toward stylus, stylus short, a continuation of the glans, tapering to a distinctly pointed distal tip. Glans in lateral view (
<figureCitation id="130C15FCA96A0440FCE7CC9790D3F9AC" box="[869,937,2008,2034]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="5" pageNumber="366">Fig. 8</figureCitation>
) a continuation of the distal part of truncus, but slightly enlarged dorsally for most of its length, short stylus distinctly bent to dorsal side. A few inconspicuous short unspecialized spicules forming the armament of glans, their arrangement symmetrical in ventral, dorsal and lateral views; one double row each on dorsal, ventral and lateral sides, distally extended to the basis of stylus, there slightly closer to each other than on glans proper.
</paragraph>
<caption id="DF4859F1A9690443FF15CE399216FB88" ID-DOI="http://doi.org/10.5281/zenodo.7960314" ID-Zenodo-Dep="7960314" httpUri="https://zenodo.org/record/7960314/files/figure.png" pageId="6" pageNumber="367" startId="6.[151,264,1398,1422]" targetBox="[156,1431,314,1370]" targetPageId="6" targetType="figure">
<paragraph id="8B880979A9690443FF15CE399216FB88" blockId="6.[151,1437,1398,1494]" pageId="6" pageNumber="367">
<emphasis id="B943D56BA9690443FF15CE399235FBD0" bold="true" box="[151,335,1398,1422]" pageId="6" pageNumber="367">FIGURES 12-16.</emphasis>
<taxonomicName id="4C3772FAA9690443FED7CE399284FBD0" authority="Martens &amp; Maghradze &amp; Barjadze, 2023" authorityName="Martens &amp; Maghradze &amp; Barjadze" authorityYear="2023" box="[341,510,1398,1422]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="367" phylum="Arthropoda" rank="species" species="kotia" status="sp. nov.">
<emphasis id="B943D56BA9690443FED7CE399284FBD0" box="[341,510,1398,1422]" italics="true" pageId="6" pageNumber="367">Nemaspela kotia</emphasis>
</taxonomicName>
<emphasis id="B943D56BA9690443FD86CE399129FBD0" bold="true" box="[516,595,1398,1422]" pageId="6" pageNumber="367">
<taxonomicNameLabel id="A2706810A9690443FD86CE399129FBD0" box="[516,595,1398,1422]" pageId="6" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Chelicerae in retro-lateral view. 
<emphasis id="B943D56BA9690443FC1CCE3990C4FBD0" bold="true" box="[926,958,1398,1422]" pageId="6" pageNumber="367">12.</emphasis>
Male (bar: 0.3 mm); 
<emphasis id="B943D56BA9690443FB17CE3997CFFBD0" bold="true" box="[1173,1205,1398,1422]" pageId="6" pageNumber="367">14.</emphasis>
Female (bar: 0.3 mm). Chelicera in pro-lateral view. 
<emphasis id="B943D56BA9690443FE44CED5929FFBEC" bold="true" box="[454,485,1434,1458]" pageId="6" pageNumber="367">13.</emphasis>
Male (bar: 0.3 mm). 
<emphasis id="B943D56BA9690443FD42CED5919AFBEC" bold="true" box="[704,736,1434,1458]" pageId="6" pageNumber="367">15.</emphasis>
Female (bar: 0.3 mm). 
<emphasis id="B943D56BA9690443FC53CED5908BFBEC" bold="true" box="[977,1009,1434,1458]" pageId="6" pageNumber="367">16.</emphasis>
Patellar apophysis of female, retro-lateral views (bar: 0.1 mm).
</paragraph>
</caption>
<paragraph id="8B880979A9690443FF45CD4B91BBF945" blockId="6.[151,1437,1540,2035]" pageId="6" pageNumber="367">
<emphasis id="B943D56BA9690443FF45CD4B9170F840" bold="true" box="[199,522,1540,1567]" pageId="6" pageNumber="367">Description. Female (n=2).</emphasis>
Body length: 1.9, 2.0. External morphology of body generally similar to male, opisthosoma more elevated than in male (
<figureCitation id="130C15FCA9690443FDFCCD6791B3F81D" box="[638,713,1576,1603]" captionStart="FIGURE 1" captionStartId="2.[151,250,1945,1969]" captionTargetBox="[151,1436,958,1920]" captionTargetId="figure-363@2.[151,1436,957,1920]" captionTargetPageId="2" captionText="FIGURE 1. Nemaspela kotia sp. nov. at the type locality, Kotia cave, presumably female in lateral view. Photograph by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.7960308" httpUri="https://zenodo.org/record/7960308/files/figure.png" pageId="6" pageNumber="367">Figs 1</figureCitation>
, 
<figureCitation id="130C15FCA9690443FD58CD669193F81D" box="[730,745,1577,1603]" captionStart="FIGURES 2-3" captionStartId="3.[151,264,878,902]" captionTargetBox="[151,1436,181,854]" captionTargetId="figure-18@3.[151,1436,181,854]" captionTargetPageId="3" captionText="FIGURES 2-3. Nemaspela kotia sp. nov. Preserved specimens, lateral view. 2. Male, 3. Female (bar: 2 mm). Photographs by Jean Severin." figureDoi="http://doi.org/10.5281/zenodo.7960310" httpUri="https://zenodo.org/record/7960310/files/figure.png" pageId="6" pageNumber="367">3</figureCitation>
, 
<figureCitation id="130C15FCA9690443FD7BCD66906CF81D" box="[761,790,1577,1603]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="6" pageNumber="367">11</figureCitation>
). Ventral side similar to male, light yellowish, only Op gen more sclerotized and thus darker, irregular blackish spots on Op gen, apparently dependent on age after final moult; spots lacking in younger specimens. 
<emphasis id="B943D56BA9690443FD1ECD3F9070F8D4" box="[668,778,1648,1674]" italics="true" pageId="6" pageNumber="367">Chelicera</emphasis>
(
<figureCitation id="130C15FCA9690443FC9CCD3F90D8F8D5" box="[798,930,1648,1675]" captionStart="FIGURES 12-16" captionStartId="6.[151,264,1398,1422]" captionTargetBox="[156,1431,314,1370]" captionTargetId="figure-95@6.[151,1436,309,1375]" captionTargetPageId="6" captionText="FIGURES 12-16. Nemaspela kotia sp. nov. Chelicerae in retro-lateral view. 12. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Chelicera in pro-lateral view. 13. Male (bar: 0.3 mm). 15. Female (bar: 0.3 mm). 16. Patellar apophysis of female, retro-lateral views (bar: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960314" httpUri="https://zenodo.org/record/7960314/files/figure.png" pageId="6" pageNumber="367">Figs 14–15</figureCitation>
) similar to male, proportions of 1 
<superScript id="7C42A431A9690443FAB6CD3F9645F820" attach="left" box="[1332,1343,1648,1662]" fontSize="6" pageId="6" pageNumber="367">st</superScript>
and 2 
<superScript id="7C42A431A9690443FA0ECD3F96E6F820" attach="none" box="[1420,1436,1648,1662]" fontSize="6" pageId="6" pageNumber="367">nd</superScript>
articles and their setation similar to male (in retro-lateral view, see 
<figureCitation id="130C15FCA9690443FC16CDDB9092F8F1" box="[916,1000,1684,1711]" captionStart="FIGURES 12-16" captionStartId="6.[151,264,1398,1422]" captionTargetBox="[156,1431,314,1370]" captionTargetId="figure-95@6.[151,1436,309,1375]" captionTargetPageId="6" captionText="FIGURES 12-16. Nemaspela kotia sp. nov. Chelicerae in retro-lateral view. 12. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Chelicera in pro-lateral view. 13. Male (bar: 0.3 mm). 15. Female (bar: 0.3 mm). 16. Patellar apophysis of female, retro-lateral views (bar: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960314" httpUri="https://zenodo.org/record/7960314/files/figure.png" pageId="6" pageNumber="367">Fig. 14</figureCitation>
). 
<emphasis id="B943D56BA9690443FC7FCDDB9718F8F0" box="[1021,1122,1684,1710]" italics="true" pageId="6" pageNumber="367">Pedipalp</emphasis>
(
<figureCitation id="130C15FCA9690443FBF1CDDB97C1F8F1" box="[1139,1211,1684,1711]" captionStart="FIGURE 1" captionStartId="2.[151,250,1945,1969]" captionTargetBox="[151,1436,958,1920]" captionTargetId="figure-363@2.[151,1436,957,1920]" captionTargetPageId="2" captionText="FIGURE 1. Nemaspela kotia sp. nov. at the type locality, Kotia cave, presumably female in lateral view. Photograph by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.7960308" httpUri="https://zenodo.org/record/7960308/files/figure.png" pageId="6" pageNumber="367">Figs 1</figureCitation>
, 
<figureCitation id="130C15FCA9690443FB48CDDA97A3F8F1" box="[1226,1241,1685,1711]" captionStart="FIGURES 2-3" captionStartId="3.[151,264,878,902]" captionTargetBox="[151,1436,181,854]" captionTargetId="figure-18@3.[151,1436,181,854]" captionTargetPageId="3" captionText="FIGURES 2-3. Nemaspela kotia sp. nov. Preserved specimens, lateral view. 2. Male, 3. Female (bar: 2 mm). Photographs by Jean Severin." figureDoi="http://doi.org/10.5281/zenodo.7960310" httpUri="https://zenodo.org/record/7960310/files/figure.png" pageId="6" pageNumber="367">3</figureCitation>
, 
<figureCitation id="130C15FCA9690443FB6ACDDA9679F8F1" box="[1256,1283,1685,1711]" captionStart="FIGURES 4-11" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1411,186,1920]" captionTargetId="figure-32@4.[175,1412,181,1920]" captionTargetPageId="4" captionText="FIGURES 4-11. Nemaspela kotia sp. nov. Pedipalps in retro-lateral view. 4. Male (bar: 0.5 mm); 5. Female (bar: 0.5 mm). Penis. 6. Ventral view (bar: 0.2 mm); 7. Lateral view (bar: 0.2 mm). Glans penis. 8. Ventral view (bar: 0.15 mm); 9. Lateral view (bar: 0.15 mm). Habitus in lateral view. 10. Female (bar: 1.0 mm); 11. Male (bar: 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960312" httpUri="https://zenodo.org/record/7960312/files/figure.png" pageId="6" pageNumber="367">11</figureCitation>
) very similar to male including setation, all articles of equal size (see 
<tableCitation id="C6B53CC2A9690443FC88CDF79026F88D" box="[778,860,1720,1747]" captionStart="TABLE 1" captionStartId="2.[152,245,620,644]" captionTargetPageId="2" captionText="TABLE 1. Measurements (mm) of legs and pedipalp in male holotype and female paratype (in parentheses) of Nemaspela kotia sp. nov." httpUri="http://table.plazi.org/id/DF4859F1A96D0447FF1AC9239253FCF7" pageId="6" pageNumber="367" tableUuid="DF4859F1A96D0447FF1AC9239253FCF7">Table 1</tableCitation>
). On Ti retro-lateral at distal end a small apophysis surrounded by few clavate setae (
<figureCitation id="130C15FCA9690443FD93CD93911FF8A8" box="[529,613,1756,1782]" captionStart="FIGURES 12-16" captionStartId="6.[151,264,1398,1422]" captionTargetBox="[156,1431,314,1370]" captionTargetId="figure-95@6.[151,1436,309,1375]" captionTargetPageId="6" captionText="FIGURES 12-16. Nemaspela kotia sp. nov. Chelicerae in retro-lateral view. 12. Male (bar: 0.3 mm); 14. Female (bar: 0.3 mm). Chelicera in pro-lateral view. 13. Male (bar: 0.3 mm). 15. Female (bar: 0.3 mm). 16. Patellar apophysis of female, retro-lateral views (bar: 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7960314" httpUri="https://zenodo.org/record/7960314/files/figure.png" pageId="6" pageNumber="367">Fig. 16</figureCitation>
). Number of femoral pseudo-articulations (n=1): Fe I 19, Fe II 24, Fe III 17, Fe IV 22. Measurements are given in 
<tableCitation id="C6B53CC2A9690443FDE5CC4F91C1F945" box="[615,699,1792,1819]" captionStart="TABLE 1" captionStartId="2.[152,245,620,644]" captionTargetPageId="2" captionText="TABLE 1. Measurements (mm) of legs and pedipalp in male holotype and female paratype (in parentheses) of Nemaspela kotia sp. nov." httpUri="http://table.plazi.org/id/DF4859F1A96D0447FF1AC9239253FCF7" pageId="6" pageNumber="367" tableUuid="DF4859F1A96D0447FF1AC9239253FCF7">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8B880979A9690443FF45CC6B91FBF9D8" blockId="6.[151,1437,1540,2035]" pageId="6" pageNumber="367">
<emphasis id="B943D56BA9690443FF45CC6B9245F960" bold="true" box="[199,319,1828,1854]" pageId="6" pageNumber="367">Variation.</emphasis>
The available specimens are rather homogeneous in size including appendages and coloration. Coloration of appendages and body depends on age after final moult; older specimens appear to be darker. For minor differences between sexes see above.
</paragraph>
<paragraph id="8B880979A9690443FF45CCDF9153F9F4" blockId="6.[151,1437,1540,2035]" box="[199,553,1936,1963]" pageId="6" pageNumber="367">
<emphasis id="B943D56BA9690443FF45CCDF920AF9F4" bold="true" box="[199,368,1936,1962]" pageId="6" pageNumber="367">Relationships.</emphasis>
See Discussion.
</paragraph>
</subSubSection>
<subSubSection id="C32D5AF2A9690443FF45CCFA92AEF9AC" pageId="6" pageNumber="367" type="etymology">
<paragraph id="8B880979A9690443FF45CCFA92AEF9AC" blockId="6.[151,1437,1540,2035]" pageId="6" pageNumber="367">
<emphasis id="B943D56BA9690443FF45CCFA9234F991" bold="true" box="[199,334,1973,1999]" pageId="6" pageNumber="367">Etymology.</emphasis>
The name is given according to the 
<typeStatus id="548CB7DBA9690443FD7BCCFA9053F991" box="[761,809,1973,1999]" pageId="6" pageNumber="367">type</typeStatus>
locality, Kotia Cave, the single known locality of this species. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32D5AF2A9680442FF45CBD992EFFE8B" pageId="7" pageNumber="368" type="distribution">
<paragraph id="8B880979A9680442FF45CBD992EFFE8B" blockId="7.[151,1436,150,213]" pageId="7" pageNumber="368">
<emphasis id="B943D56BA9680442FF45CBD99223FEEE" bold="true" box="[199,345,150,176]" pageId="7" pageNumber="368">Distribution</emphasis>
(
<figureCitation id="130C15FCA9680442FEE8CBD892CCFEEF" box="[362,438,151,177]" captionStart="MAP 1" captionStartId="5.[152,211,1578,1602]" captionTargetBox="[151,1436,151,1554]" captionTargetId="figure-18@5.[151,1436,151,1554]" captionTargetPageId="5" captionText="MAP 1. Distributional map of the Nemaspela species in the Caucasus. 1. Vorontsovskaya Cave (N. sokolovi); 2. Labirintovaya Cave (N. sokolovi); 3. Dolgaya Cave (N. sokolovi); 4. Fontanka Cave (N. kovali); 5. Omega-15 Cave (N. kovali); 6. Omega-12 Cave (N. kovali); 7. Otte-Shik Cave (N. kovali); 8. Akhunskaya Cave (N. abchasica); 9. Beloskalskaya Cave (N. abchasica); 10. Kolokolnaya Cave (N. abchasica); 11. Kuibishevi Cave (N. abchasica); 12. L 24 Cave (N. abchasica); 13. Kveda Shakurani (=Nizhnyaya Shakuranskaya) Cave (N. abchasica); 14. Zemo Eshera Cave (N. birsteini); 15. Tarkili Cave (N. birsteini); 16. Sarma Cave (N. gagrica); 17. Veryovkina Cave (N. gagrica); 18. Sakishore Cave (N. femorecurvata); 19. Prometheus Cave (N. prometheus); 20. Melouri Cave (N. melouri); 21. Solkota Cave (N. melouri); 22. Kotia Cave (=Kotiasklde Cave) (N. kotia) (for details of localities and collection depositories see Martens et al. 2021)." figureDoi="http://doi.org/10.5281/zenodo.7971988" httpUri="https://zenodo.org/record/7971988/files/figure.png" pageId="7" pageNumber="368">Map 1</figureCitation>
). Known only from the 
<typeStatus id="548CB7DBA9680442FD50CBD89078FEEF" box="[722,770,151,177]" pageId="7" pageNumber="368">type</typeStatus>
locality, the Kotia Cave in Chiatura Municipality, 
<collectingRegion id="49F3C79BA9680442FAC9CBD896E6FEEF" box="[1355,1436,151,177]" country="Georgia" name="Imereti" pageId="7" pageNumber="368">Imereti</collectingRegion>
Region, West 
<collectingCountry id="F32049E9A9680442FEB6CBF492EBFE8B" box="[308,401,187,213]" name="Georgia" pageId="7" pageNumber="368">Georgia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>