269 lines
32 KiB
XML
269 lines
32 KiB
XML
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<mods:title id="231B1F010D214ABD55C1E0138BB8B685">Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål</mods:title>
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<treatment id="039E355D643AFFB78F9CFC45FCDCFF4F" ID-DOI="http://doi.org/10.5281/zenodo.6102889" ID-GBIF-Taxon="127671709" ID-Zenodo-Dep="6102889" LSID="urn:lsid:plazi:treatment:039E355D643AFFB78F9CFC45FCDCFF4F" httpUri="http://treatment.plazi.org/id/039E355D643AFFB78F9CFC45FCDCFF4F" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
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<subSubSection id="C32DD7C0643AFFB08F9CFC45FE84F9B6" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8B88844B643AFFB18F9CFC45FC60FC74" blockId="18.[151,910,980,1075]" box="[151,910,980,1006]" pageId="18" pageNumber="19">
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<heading id="D0C03327643AFFB18F9CFC45FC60FC74" bold="true" box="[151,910,980,1006]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<emphasis id="B9435859643AFFB18F9CFC45FC60FC74" bold="true" box="[151,910,980,1006]" pageId="18" pageNumber="19">
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<taxonomicName id="4C37FFC8643AFFB18F9CFC45FE2BFC74" box="[151,453,980,1006]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geographica">
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<emphasis id="B9435859643AFFB18F9CFC45FE2BFC74" bold="true" box="[151,453,980,1006]" italics="true" pageId="18" pageNumber="19">Calligrapha geographica</emphasis>
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</taxonomicName>
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subsp.
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<taxonomicName id="4C37FFC8643AFFB18D29FC45FCF6FC74" authority="Stal, 1860" authorityName="Stal" authorityYear="1860" box="[546,792,980,1006]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="famularis" status="stat. nov.">
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<emphasis id="B9435859643AFFB18D29FC45FD79FC74" bold="true" box="[546,663,980,1006]" italics="true" pageId="18" pageNumber="19">famularis</emphasis>
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Stål, 1860
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</taxonomicName>
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<taxonomicNameLabel id="A270E522643AFFB18C14FC45FC60FC74" box="[799,910,980,1006]" pageId="18" pageNumber="19">stat. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B88844B643AFFB18F9CFC67FE87FB95" blockId="18.[151,910,980,1075]" box="[151,361,1014,1039]" pageId="18" pageNumber="19">
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<bibRef id="529710BD643AFFB18F9CFC67FE87FB95" box="[151,361,1014,1039]" pageId="18" pageNumber="19" type="book chapter">
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<author id="538A6CAE643AFFB18F9CFC67FF04FB95" box="[151,234,1014,1039]" fns="I" npo="LF" pageId="18" pageNumber="19">Stål, C.</author>
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<year id="FF9FC518643AFFB18FFAFC67FEC3FB95" box="[241,301,1014,1039]" pageId="18" pageNumber="19">1860</year>
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:
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<pagination id="B2FE7ABF643AFFB18E3CFC67FE8BFB95" box="[311,357,1014,1039]" pageId="18" pageNumber="19" type="pagination">462</pagination>
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.
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</bibRef>
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</paragraph>
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<paragraph id="8B88844B643AFFB18F9CFB8BFEF9FBA9" blockId="18.[151,910,980,1075]" box="[151,279,1050,1075]" pageId="18" pageNumber="19">
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(
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<figureCitation id="130C98CE643AFFB18F94FB8BFF0BFBA9" box="[159,229,1050,1075]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[291,1305,794,1908]" captionTargetId="figure@9.[278,1309,783,1914]" captionTargetPageId="9" captionText="FIGURE 3. Dorsal and lateral habitus of Holotype of Calligrapha consputa Stål (a), Lectotype of C. geographica ssp. geographica Stål (b), Lectotype of C. geographica ssp. famularis Stål (c), and Lectotype of C. multipustulata Stål (d). Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3539688" httpUri="https://zenodo.org/record/3539688/files/figure.png" pageId="18" pageNumber="19">Figs 3</figureCitation>
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c, 5)
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</paragraph>
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<paragraph id="8B88844B643AFFB18F9CFBF3FCC7FBE2" blockId="18.[151,1287,1122,1328]" box="[151,809,1122,1144]" pageId="18" pageNumber="19">
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<taxonomicName id="4C37FFC8643AFFB18F9CFBF3FE00FBE2" authority="Stal, 1865" authorityName="Stal" authorityYear="1865" box="[151,494,1122,1144]" class="Insecta" family="Chrysomelidae" genus="Chrysomela" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="famularis">
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<emphasis id="B9435859643AFFB18F9CFBF3FE96FBE2" box="[151,376,1122,1144]" italics="true" pageId="18" pageNumber="19">Chrysomela famularis</emphasis>
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: Stål, 1865
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</taxonomicName>
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, Mon. Chrys. Amer., 3, p. 277.
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</paragraph>
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<paragraph id="8B88844B643AFFB18F9CFB10FC98FB2C" blockId="18.[151,1287,1122,1328]" pageId="18" pageNumber="19">
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<taxonomicName id="4C37FFC8643AFFB18F9CFB10FE6AFB0D" box="[151,388,1153,1175]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="formularis">
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<emphasis id="B9435859643AFFB18F9CFB10FE6AFB0D" box="[151,388,1153,1175]" italics="true" pageId="18" pageNumber="19">Calligrapha formularis</emphasis>
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</taxonomicName>
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[lapsus calami]: Gemminger & Harold, 1874, Cat. Col., XI, p. 3433.
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<emphasis id="B9435859643AFFB18F9CFB31FE2DFB2C" box="[151,451,1184,1206]" italics="true" pageId="18" pageNumber="19">
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<taxonomicName id="4C37FFC8643AFFB18F9CFB31FED8FB2C" box="[151,310,1184,1206]" class="Insecta" family="Chrysomelidae" genus="Polyspila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="argus">Polyspila argus</taxonomicName>
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ab.
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<taxonomicName id="4C37FFC8643AFFB18E69FB31FE2DFB2C" authority="Weise, 1916" authorityName="Weise" authorityYear="1916" box="[354,451,1184,1206]" class="Insecta" family="Chrysomelidae" genus="Chrysomela" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="famularis">famularis</taxonomicName>
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</emphasis>
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: Weise, 1916, Col. Cat., pars 68, 12, p. 38.
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</paragraph>
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<paragraph id="8B88844B643AFFB18F9CFB2FFC27FB4E" blockId="18.[151,1287,1122,1328]" box="[151,969,1214,1237]" pageId="18" pageNumber="19">
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<emphasis id="B9435859643AFFB18F9CFB2FFE3CFB4E" box="[151,466,1214,1237]" italics="true" pageId="18" pageNumber="19">
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<taxonomicName id="4C37FFC8643AFFB18F9CFB2FFEBCFB4F" box="[151,338,1214,1237]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="argus">Calligrapha argus</taxonomicName>
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a.
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<taxonomicName id="4C37FFC8643AFFB18E7AFB2FFE3CFB4E" authority="Blackwelder, 1946" authorityName="Blackwelder" authorityYear="1946" box="[369,466,1214,1236]" class="Insecta" family="Chrysomelidae" genus="Chrysomela" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="famularis">famularis</taxonomicName>
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</emphasis>
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: Blackwelder, 1946, Checklist Col., Pt. 4, p. 674.
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</paragraph>
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<paragraph id="8B88844B643AFFB18F9CFB4CFAE9FAAA" blockId="18.[151,1287,1122,1328]" pageId="18" pageNumber="19">
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<taxonomicName id="4C37FFC8643AFFB18F9CFB4CFD2EFB69" authority="Bechyne, 1952" authorityName="Bechyne" authorityYear="1952" box="[151,704,1245,1267]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="variety" species="ramulifera" variety="famularis">
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<emphasis id="B9435859643AFFB18F9CFB4CFDF6FB69" box="[151,536,1245,1267]" italics="true" pageId="18" pageNumber="19">Calligrapha ramulifera var. famularis</emphasis>
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: Bechyné, 1952
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</taxonomicName>
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, Entom.
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<collectingCountry id="F320C4DB643AFFB18C11FB4CFCAAFB69" box="[794,836,1245,1267]" name="Aruba" pageId="18" pageNumber="19">Arb</collectingCountry>
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. Mus. Frey 3, p. 4.
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<emphasis id="B9435859643AFFB18F9CFB6DFDEDFA88" box="[151,515,1276,1298]" italics="true" pageId="18" pageNumber="19">
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<taxonomicName id="4C37FFC8643AFFB18F9CFB6DFE6BFA88" box="[151,389,1276,1298]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ramulifera">Calligrapha ramulifera</taxonomicName>
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s.
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<taxonomicName id="4C37FFC8643AFFB18EA9FB6DFDEDFA88" authority="Blackwelder, 1957" authorityName="Blackwelder" authorityYear="1957" box="[418,515,1276,1298]" class="Insecta" family="Chrysomelidae" genus="Chrysomela" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="famularis">famularis</taxonomicName>
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</emphasis>
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: Blackwelder, 1957, Checklist Col., Pt. 6, p. 1436.
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<taxonomicName id="4C37FFC8643AFFB18F9CFA8BFD34FAAA" authority="Bechyne & Springlova" authorityName="Bechyne & Springlova" box="[151,730,1306,1328]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="variety" species="argus" variety="famularis">
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<emphasis id="B9435859643AFFB18F9CFA8BFE0BFAAA" box="[151,485,1306,1328]" italics="true" pageId="18" pageNumber="19">Calligrapha argus var. famularis</emphasis>
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: Bechyné & Springlová
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</taxonomicName>
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de Bechyné, 1965, Rev. Fac. Agron. Maracay 3, p. 48.
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</paragraph>
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<paragraph id="8B88844B643AFFB18F9CFACCFA7DF8F4" blockId="18.[151,1436,1373,2010]" pageId="18" pageNumber="19">
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Carl Stål (1865) treated this taxon with affinities with his almost indistinguishable
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<taxonomicName id="4C37FFC8643AFFB18B72FACFFA89FAEC" authority="Stal" authorityName="Stal" box="[1145,1383,1373,1398]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geographica">
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<emphasis id="B9435859643AFFB18B72FACFFAC3FAEC" box="[1145,1325,1373,1398]" italics="true" pageId="18" pageNumber="19">C. geographica</emphasis>
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Stål
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</taxonomicName>
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, but highlighting that the latter was always smaller and with differences in their elytral patterns. Judging from the specimen figured in
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<emphasis id="B9435859643AFFB18E8AFA34FD24FA27" box="[385,714,1445,1470]" italics="true" pageId="18" pageNumber="19">Biologia Centrali Americana</emphasis>
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(i.e., fig. 19, Table 10), Jacoby (1882) interpreted
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<taxonomicName id="4C37FFC8643AFFB18A05FA37FA72FA24" box="[1294,1436,1445,1470]" class="Insecta" family="Chrysomelidae" genus="Chrysomela" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="famularis">
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<emphasis id="B9435859643AFFB18A05FA37FA72FA24" box="[1294,1436,1445,1470]" italics="true" pageId="18" pageNumber="19">C. famularis</emphasis>
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</taxonomicName>
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possibly in a different way than the original author, finding understandable similarities with
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<taxonomicName id="4C37FFC8643AFFB18BB2FA5BFAA1FA78" authority="Stal" authorityName="Stal" box="[1209,1359,1481,1506]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="argus">
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<emphasis id="B9435859643AFFB18BB2FA5BFAF5FA7B" box="[1209,1307,1482,1505]" italics="true" pageId="18" pageNumber="19">C. argus</emphasis>
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Stål
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</taxonomicName>
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which compelled him to propose the former as a synonym of the latter (see below). For example, Stål's description remarks the black tint of elytral maculae, while they are reddish in
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<taxonomicName id="4C37FFC8643AFFB18C94F983FBEFF9B3" box="[927,1025,1554,1577]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="argus">
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<emphasis id="B9435859643AFFB18C94F983FBEFF9B3" box="[927,1025,1554,1577]" italics="true" pageId="18" pageNumber="19">C. argus</emphasis>
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</taxonomicName>
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and also in the specimen named
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<taxonomicName id="4C37FFC8643AFFB18A89F983FEEDF9D4" class="Insecta" family="Chrysomelidae" genus="Chrysomela" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="famularis">
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<emphasis id="B9435859643AFFB18A89F983FEEDF9D4" italics="true" pageId="18" pageNumber="19">C. famularis</emphasis>
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</taxonomicName>
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by Jacoby. Nonetheless, Jacoby's taxonomic decision was perpetuated in subsequent catalogues (Weise 1916; Blackwelder 1946), and formalized by Bechyné (1952), considering this taxon a variety with confluent spots (possibly interpreting this taxon in the same way as Jacoby did seven decades earlier). However, after examining the
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<typeStatus id="548C3AE9643AFFB18FCAF933FF1FF920" box="[193,241,1698,1722]" pageId="18" pageNumber="19">type</typeStatus>
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material for this species, I found Stål's judgement sound in discussing this taxon close to
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<taxonomicName id="4C37FFC8643AFFB18BEDF933FA7AF920" box="[1254,1428,1697,1722]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geographica">
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<emphasis id="B9435859643AFFB18BEDF933FA7AF920" box="[1254,1428,1697,1722]" italics="true" pageId="18" pageNumber="19">C. geographica</emphasis>
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</taxonomicName>
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. Indeed, I can only separate both taxa on the basis of their midlateral spots, free and very narrow in the case of
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<taxonomicName id="4C37FFC8643AFFB18A89F957FECAF898" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geographica">
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<emphasis id="B9435859643AFFB18A89F957FECAF898" italics="true" pageId="18" pageNumber="19">C. geographica</emphasis>
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</taxonomicName>
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and wide, connected to dark elytral margin by dark suffusions in
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<taxonomicName id="4C37FFC8643AFFB18B09F97BFB60F898" box="[1026,1166,1769,1794]" class="Insecta" family="Chrysomelidae" genus="Chrysomela" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="famularis">
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<emphasis id="B9435859643AFFB18B09F97BFB60F898" box="[1026,1166,1769,1794]" italics="true" pageId="18" pageNumber="19">C. famularis</emphasis>
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</taxonomicName>
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(besides its larger size). With the limited distribution data available to me, both forms seem to be parapatric, whereby
|
||
<taxonomicName id="4C37FFC8643AFFB18BE1F89FFA75F8BC" box="[1258,1435,1805,1830]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geographica">
|
||
<emphasis id="B9435859643AFFB18BE1F89FFA75F8BC" box="[1258,1435,1805,1830]" italics="true" pageId="18" pageNumber="19">C. geographica</emphasis>
|
||
</taxonomicName>
|
||
occupies a northwestern range and
|
||
<taxonomicName id="4C37FFC8643AFFB18D21F8A3FD56F8D0" box="[554,696,1841,1866]" class="Insecta" family="Chrysomelidae" genus="Chrysomela" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="famularis">
|
||
<emphasis id="B9435859643AFFB18D21F8A3FD56F8D0" box="[554,696,1841,1866]" italics="true" pageId="18" pageNumber="19">C. famularis</emphasis>
|
||
</taxonomicName>
|
||
a southeastern range in their combined distribution (
|
||
<figureCitation id="130C98CE643AFFB18A1DF8A0FAB5F8D0" box="[1302,1371,1841,1866]" captionStart="FIGURE 5" captionStartId="14.[151,250,1907,1929]" captionTargetBox="[165,1427,1142,1879]" captionTargetId="figure@14.[150,1437,1132,1886]" captionTargetPageId="14" captionText="FIGURE 5. Geographic distribution of species in the Calligrapha multipustulata - group." httpUri="https://zenodo.org/record/287934/files/figure.png" pageId="18" pageNumber="19">Fig. 5</figureCitation>
|
||
), and until more data becomes available to resolve their taxonomic status, they are proposed here with subspecific rank.
|
||
</paragraph>
|
||
<paragraph id="8B88844B643AFFB08FCCF8E8FAD1FF4F" blockId="18.[151,1436,1373,2010]" lastBlockId="19.[151,1437,151,1725]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
|
||
There is no indication in the original description about this taxon being described based on a
|
||
<typeStatus id="548C3AE9643AFFB18BFEF8EBFAA2F808" box="[1269,1356,1914,1938]" pageId="18" pageNumber="19" type="syntype">syntype</typeStatus>
|
||
series, however there were two specimens identified as
|
||
<typeStatus id="548C3AE9643AFFB18DC9F80FFD13F82C" box="[706,765,1950,1974]" pageId="18" pageNumber="19">types</typeStatus>
|
||
in the entomological collections at NHM in London (from Deyrolle’s collection) and NRM in Stockholm (Stål 1865). The one from NRM explicitly includes locality information (Teapa) matching the one included in the original description, and the size of the specimen is compatible with the measures given in this same description. Thus, this specimen is selected as
|
||
<typeStatus id="548C3AE9643BFFB08BCCFF2CFAD5FF4F" box="[1223,1339,189,213]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B88844B643BFFB08FCCFF71FE0DFE86" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="20">
|
||
<typeStatus id="548C3AE9643BFFB08FCCFF71FED9FF62" box="[199,311,224,248]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
|
||
by present designation: Teapa / Stål /
|
||
<typeStatus id="548C3AE9643BFFB08C0DFF71FCD0FF62" box="[774,830,224,248]" pageId="19" pageNumber="20">Type</typeStatus>
|
||
/
|
||
<typeStatus id="548C3AE9643BFFB08C56FF71FC4FFF62" box="[861,929,224,248]" pageId="19" pageNumber="20">Typus</typeStatus>
|
||
[red] (NRM). Specimen lacking ten apical antennomeres of left antenna.
|
||
</paragraph>
|
||
<paragraph id="8B88844B643BFFB08FCCFEB6FE8BFEFE" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="20">
|
||
<typeStatus id="548C3AE9643BFFB08FCCFEB6FE8AFEDA" box="[199,356,295,320]" pageId="19" pageNumber="20" type="paralectotype">Paralectotype</typeStatus>
|
||
:
|
||
<typeStatus id="548C3AE9643BFFB08E7AFEB9FE47FEDA" box="[369,425,296,320]" pageId="19" pageNumber="20">Type</typeStatus>
|
||
Stål, Coll. Deyrolle / Baly Coll. /
|
||
<taxonomicName id="4C37FFC8643BFFB08C4FFEB6FC0DFEDA" authority="Stal" authorityName="Stal" box="[836,995,295,320]" class="Insecta" family="Chrysomelidae" genus="Chrysomela" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="famularis">famularis Stål</taxonomicName>
|
||
<collectingCountry id="F320C4DB643BFFB08CE5FEB6FBADFEDA" box="[1006,1091,295,320]" name="Mexico" pageId="19" pageNumber="20">Mexico</collectingCountry>
|
||
[underneath:
|
||
<typeStatus id="548C3AE9643BFFB08BEDFEB9FAF3FEDA" box="[1254,1309,296,320]" pageId="19" pageNumber="20">Type</typeStatus>
|
||
Stål, Coll: Deyrolle] (NHM).
|
||
</paragraph>
|
||
<paragraph id="8B88844B643BFFB08FCCFEFEFD94FD82" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="20">
|
||
<emphasis id="B9435859643BFFB08FCCFEFEFEC8FE12" bold="true" box="[199,294,367,392]" pageId="19" pageNumber="20">Habitus</emphasis>
|
||
(
|
||
<figureCitation id="130C98CE643BFFB08E3CFEFEFE92FE12" box="[311,380,367,392]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[291,1305,794,1908]" captionTargetId="figure@9.[278,1309,783,1914]" captionTargetPageId="9" captionText="FIGURE 3. Dorsal and lateral habitus of Holotype of Calligrapha consputa Stål (a), Lectotype of C. geographica ssp. geographica Stål (b), Lectotype of C. geographica ssp. famularis Stål (c), and Lectotype of C. multipustulata Stål (d). Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3539688" httpUri="https://zenodo.org/record/3539688/files/figure.png" pageId="19" pageNumber="20">Fig. 3</figureCitation>
|
||
c). Length:
|
||
<quantity id="4CCF29AE643BFFB08D09FEE1FD96FE1D" box="[514,632,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.029" pageId="19" pageNumber="20" unit="mm" value="10.29">10.29 mm</quantity>
|
||
, width:
|
||
<quantity id="4CCF29AE643BFFB08DDDFEFEFCD1FE1D" box="[726,831,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="6.57" pageId="19" pageNumber="20" unit="mm" value="6.57">6.57 mm</quantity>
|
||
. Body oval, moderately convex. Head and pronotum dark metallic green. Scutellum, ventral parts and appendices dark metallic green with reddish hue, more evident in ventral parts of abdomen. Four basal antennomeres mostly reddish, darkened dorsally. Labrum, knees, apex of onychium and claws red. Elytra dark yellow, almost orange. Epipleura and heavy elytral markings with a hint of green metallic shine, more so on epipleura.
|
||
</paragraph>
|
||
<paragraph id="8B88844B643BFFB08FCCFDB5FE84F9B6" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="20">Head broad, deeply inserted in pronotum; more or less evenly punctured except on supraantennal calli; frontal suture impressed, perpendicularly linked to broadly U-shaped clypeal suture; supraocular furrow short, not surpassing eye edge behind. Eyes dorsoventrally elongated and finely faceted. Antennae relatively thin, short, reaching humeri; first antennomere long, thick, posteriorly curved; all antennomeres rather elongated, 2–6 thinner, relatively smooth and nearly glabrous, second nearly 2/3 as long as first, third longer than first; antennomeres 3–6 shortening progressively, sixth and seventh subequal and slightly longer than second; antennomeres 7–11 thicker, rugose and pubescent, lengthening progressively; eleventh antennomere as long as third. Labrum feebly emarginate anteriorly. Mandibles large, strong, notably convex at exterior margin, largely protruding beyond labrum. Last segment of maxillary palpi broad and truncated at apex, with sides regularly curved. Pronotum transverse (W/L: 2.33) finely margined anteriorly and laterally; surface microreticulate with irregularly sparse punctures on all surface, denser at disc and laterally, with larger and sometimes confluent punctures; seta at anterior angle bent backwards; seta at posterior angle bent forwards. Hypomeral suture impressed along one third of thorax length, obsolete at apex, lacking at base; hypomera transversally wrinkled at base, microreticulate, without punctures. Prosternum convex; process with sparse golden pubescence, narrow between coxae, slightly enlarged and hollowed at apex, cut straight at posterior border. Metanepisterna broadly margined all around, coarsely punctured. Metaventrite finely sparsely punctured. Scutellum relatively small, flat, shiny, unpunctured, longer than wide at base (W/L=0.91). Elytra very finely punctured; punctures surrounding markings and premarginal lateral row slightly larger; scutellar row of punctures present; elytra with fine dark margin. Elytral markings: (i) sutural stripe entire from base of elytra, completely surrounding scutellum, to gradual narrowing at apex; (ii) subsutural stripe contiguous to sutural stripe in its entire length, except in scutellar area, abruptly cut preapically on elytra at level with premarginal row of punctures; (iii) arcuate band complete, fused to subsutural stripe, apically obliquely divergent; (iv) spot of apical declivity large, laterally confluent with subsutural stripe; (v) humeral spot large, elongated, confluent for most of its length to (vi) humeral lunule, reaching well beyond basis of arcuate band; (vii) spot enclosed by humeral lunule large and ovoid; (viii) apical spot large, confluent with surrounding spots; (ix) midlateral spot broadly elliptical, not entirely confluent with elytral margin except for a narrow zone of suffusion; (x) pattern of some ten spots with different confluence among them or with major markings. Legs slender, finely, sparsely punctured; with very fine short golden pubescence; apex of tibiae rugose with dense longer golden pubescence; broad longitudinal furrow externally on apical 2/3 of tibiae, with 1–2 longitudinal internal ridges, and expanding at apex.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD7C0643BFFB08FCCF9A6FE71F902" pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph id="8B88844B643BFFB08FCCF9A6FE71F902" blockId="19.[151,1437,151,1725]" pageId="19" pageNumber="20">
|
||
<emphasis id="B9435859643BFFB08FCCF9A6FE8EF9CA" bold="true" box="[199,352,1591,1616]" pageId="19" pageNumber="20">Distribution.</emphasis>
|
||
This subspecies of
|
||
<taxonomicName id="4C37FFC8643BFFB08D59F9A9FCEBF9CA" box="[594,773,1591,1616]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="geographica">
|
||
<emphasis id="B9435859643BFFB08D59F9A9FCEBF9CA" box="[594,773,1591,1616]" italics="true" pageId="19" pageNumber="20">C. geographica</emphasis>
|
||
</taxonomicName>
|
||
occupies the southern parts of the species range in the southern states of Chiapas and Tabasco in
|
||
<collectingCountry id="F320C4DB643BFFB08D8AF9CDFD38F9EF" box="[641,726,1628,1653]" name="Mexico" pageId="19" pageNumber="20">Mexico</collectingCountry>
|
||
(
|
||
<figureCitation id="130C98CE643BFFB08DE3F9CDFCC3F9EF" box="[744,813,1628,1653]" captionStart="FIGURE 5" captionStartId="14.[151,250,1907,1929]" captionTargetBox="[165,1427,1142,1879]" captionTargetId="figure@14.[150,1437,1132,1886]" captionTargetPageId="14" captionText="FIGURE 5. Geographic distribution of species in the Calligrapha multipustulata - group." httpUri="https://zenodo.org/record/287934/files/figure.png" pageId="19" pageNumber="20">Fig. 5</figureCitation>
|
||
, black squares). Subspecies endemic of the Caribbean Mesoamerican domain.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD7C0643BFFB08FCCF935FE24F823" pageId="19" pageNumber="20" type="materials_examined">
|
||
<paragraph id="8B88844B643BFFB08FCCF935FD68F927" blockId="19.[151,1437,151,1725]" box="[199,646,1700,1725]" pageId="19" pageNumber="20">
|
||
<emphasis id="B9435859643BFFB08FCCF935FE47F927" bold="true" box="[199,425,1700,1725]" pageId="19" pageNumber="20">Material examined</emphasis>
|
||
(seven specimens).
|
||
</paragraph>
|
||
<paragraph id="8B88844B643BFFB08F9CF97DFEE8F89E" blockId="19.[151,1436,1772,1977]" box="[151,262,1772,1796]" pageId="19" pageNumber="20">
|
||
<collectingCountry id="F320C4DB643BFFB08F9CF97DFEE8F89E" box="[151,262,1772,1796]" name="Mexico" pageId="19" pageNumber="20">
|
||
<collectionCode id="ED261C8E643BFFB08F9CF97DFEE8F89E" box="[151,262,1772,1796]" pageId="19" pageNumber="20">MEXICO</collectionCode>
|
||
</collectingCountry>
|
||
</paragraph>
|
||
<paragraph id="8B88844B643BFFB08FCCF89EFE24F823" blockId="19.[151,1436,1772,1977]" pageId="19" pageNumber="20">
|
||
<emphasis id="B9435859643BFFB08FCCF89EFECFF8B2" bold="true" box="[199,289,1807,1832]" pageId="19" pageNumber="20">
|
||
<collectionCode id="ED261C8E643BFFB08FCCF89EFEF2F8B2" box="[199,284,1807,1832]" pageId="19" pageNumber="20">EGRC</collectionCode>
|
||
:
|
||
</emphasis>
|
||
(1) one specimen:
|
||
<collectingCountry id="F320C4DB643BFFB08D0EF89EFDB3F8B2" box="[517,605,1807,1832]" name="Mexico" pageId="19" pageNumber="20">Mexico</collectingCountry>
|
||
, Chiapas, Cuxtepeques area,
|
||
<date id="FF89A28B643BFFB08CB9F89EFBDBF8B2" box="[946,1077,1807,1832]" pageId="19" pageNumber="20" value="1987-06-15">15/06/1987</date>
|
||
, D.B. Thomas & A. Mendoza,
|
||
<taxonomicName id="4C37FFC8643BFFB08F9CF8A5FD61F8D7" authority="Stal J. Gomez-Zurita" authorityName="Stal J. Gomez-Zurita" box="[151,655,1844,1869]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="famularis">
|
||
<emphasis id="B9435859643BFFB08F9CF8A5FE7BF8D7" box="[151,405,1844,1869]" italics="true" pageId="19" pageNumber="20">Calligrapha famularis</emphasis>
|
||
Stål J. Gómez-Zurita
|
||
</taxonomicName>
|
||
det. 2011.
|
||
<emphasis id="B9435859643BFFB08C1EF8A5FC8CF8D7" bold="true" box="[789,866,1844,1869]" pageId="19" pageNumber="20">NHM:</emphasis>
|
||
(1) one specimen: Teapa, Tabasco, Feb., H.H.S.,
|
||
<emphasis id="B9435859643BFFB08F9CF8C6FE77F8EA" box="[151,409,1879,1904]" italics="true" pageId="19" pageNumber="20">
|
||
Calligr.
|
||
<taxonomicName id="4C37FFC8643BFFB08FFCF8C6FE77F8EA" box="[247,409,1879,1904]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="multipustulata">multipustulata</taxonomicName>
|
||
</emphasis>
|
||
var., Godman-Salvin Coll., Biol. Centr.-Amer.; (2) one specimen: Teapa, Tabasco, Jan., H.H.S., Godman-Salvin Coll., Biol. Centr.-Amer.; (3) one specimen: Teapa, Pilate, Baly Coll.
|
||
<emphasis id="B9435859643BFFB08BE4F8EDFAD5F80F" bold="true" box="[1263,1339,1916,1941]" pageId="19" pageNumber="20">
|
||
<collectionCode id="ED261C8E643BFFB08BE4F8EDFAD8F80F" box="[1263,1334,1916,1941]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="19" pageNumber="20">NRM</collectionCode>
|
||
:
|
||
</emphasis>
|
||
(1) one specimen:
|
||
<collectingCountry id="F320C4DB643BFFB08E04F831FE89F823" box="[271,359,1952,1977]" name="Mexico" pageId="19" pageNumber="20">Mexico</collectingCountry>
|
||
, Tarnier.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD7C0643CFFB78FCCFF06FCDCFF4F" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="8B88844B643CFFB78FCCFF06FCDCFF4F" blockId="20.[151,1436,151,213]" pageId="20" pageNumber="21">
|
||
<emphasis id="B9435859643CFFB78FCCFF06FED5FF2A" bold="true" box="[199,315,151,176]" pageId="20" pageNumber="21">Variation.</emphasis>
|
||
Apex of arcuate band subsquare; markings showing a lesser degree of confluence (e.g., spot of apical declivity and apical spot free from subsutural stripe).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |