326 lines
32 KiB
XML
326 lines
32 KiB
XML
<document ID-DOI="10.3897/zookeys.28.325" ID-GBIF-Dataset="cac72c82-2c83-401f-b6d0-8afb5321fde3" ID-ISSN="1313–2970" ID-Zenodo-Dep="576570" ID-ZooBank="A979ECDE-871F-4AFC-9ABA-63A0FD6DC323" approvalRequired="46" approvalRequired_for_textStreams="46" checkinTime="1587979922717" checkinUser="plazi" docAuthor="Sereno, Paul & Larsson, Hans" docDate="2009" docId="039B2B68FFD39B01A5FAB1BFFD9A50CD" docLanguage="en" docName="ZK_article_2104.pdf" docOrigin="ZooKeys 28 (28)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@fc5bfa5" docStyleName="zookeys.2008.journal_article" docTitle="Laganosuchus maghrebensis Sereno & Larsson 2009, sp. n." docType="treatment" docUuid="E5BCEEAD-DFEF-4EF6-AA62-110EF91FCD0F" docUuidSource="ZooBank" docVersion="6" lastPageNumber="87" masterDocId="FFA25310FF879B57A576B119FF895175" masterDocTitle="Cretaceous Crocodyliforms from the Sahara" masterLastPageNumber="143" masterPageNumber="1" pageNumber="85" updateTime="1643567712823" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cretaceous Crocodyliforms from the Sahara</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sereno, Paul</mods:namePart>
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<mods:affiliation>University of Chicago, Chicago, United States of America</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Larsson, Hans</mods:namePart>
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<mods:affiliation>Redpath Museum, McGill University, Montreal ,, Canada</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-11-19</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>28</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>28</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>143</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.28.325</mods:identifier>
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<mods:identifier type="GBIF-Dataset">cac72c82-2c83-401f-b6d0-8afb5321fde3</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576570</mods:identifier>
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<mods:identifier type="ZooBank">A979ECDE-871F-4AFC-9ABA-63A0FD6DC323</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790381" ID-GBIF-Taxon="163750066" ID-Zenodo-Dep="3790381" LSID="urn:lsid:zoobank.org:act:E5BCEEAD-DFEF-4EF6-AA62-110EF91FCD0F" httpUri="http://treatment.plazi.org/id/039B2B68FFD39B01A5FAB1BFFD9A50CD" lastPageId="86" lastPageNumber="87" pageId="84" pageNumber="85">
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<subSubSection pageId="84" pageNumber="85" type="nomenclature">
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||
<paragraph blockId="84.[140,999,166,298]" box="[140,543,166,193]" pageId="84" pageNumber="85">
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||
<heading box="[140,543,166,193]" fontSize="11" level="2" pageId="84" pageNumber="85" reason="3">
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<emphasis bold="true" box="[140,543,166,193]" pageId="84" pageNumber="85">
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<taxonomicName authority="Sereno & Larsson, 2009" authorityName="Sereno & Larsson" authorityYear="2009" box="[140,471,166,193]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="84" pageNumber="85" phylum="Chordata" rank="species" status="sp. n.">
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<emphasis bold="true" box="[140,471,166,193]" italics="true" pageId="84" pageNumber="85">
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<emphasis box="[140,306,166,193]" inLineHeading="true" pageId="84" pageNumber="85" reason="1">Laganosuchus</emphasis>
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maghrebensis
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[478,543,167,193]" pageId="84" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="84.[140,999,166,298]" box="[140,999,202,229]" pageId="84" pageNumber="85">
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||
<uri box="[140,999,202,229]" pageId="84" pageNumber="85">
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||
urn:lsid:zoobank.org:act:
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<uuid box="[414,999,202,229]" pageId="84" pageNumber="85">E5BCEEAD-DFEF-4EF6-AA62-110EF91FCD0F</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="84" pageNumber="85" type="description">
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<paragraph blockId="84.[140,999,166,298]" box="[140,219,237,263]" pageId="84" pageNumber="85">
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<figureCitation box="[140,219,237,263]" captionStart="Figure 42" captionStartId="84.[140,212,1494,1516]" captionTargetBox="[215,1044,624,1446]" captionTargetId="figure@84.[185,1063,600,1474]" captionTargetPageId="84" captionText="Figure 42. Dentary fragment of the crocodyliform Laganosuchus maghrebensis gen. n. sp. n. Anterior end of the left dentary (UCRC PV2). A Left lateral view. B Medial view (reversed). C Dorsal view. D Ventral view. E Replacement tooth in first dentary alveolus in lingual view. Scale bar in A-D equals 3 cm; scale bar in E equals 1 cm. Abbreviations: ad1, 3, alveolus for dentary tooth 1, 3; afo, alveolar foramina; asp, articular surface for the splenial; d1, 4, dentary tooth 1, 4; fl, fluting; fo, foramen; Mc, Meckel’s canal; rt, replacement tooth; sym, symphysis; tasp, tip of the articular surface for the splenial." figureDoi="http://doi.org/10.5281/zenodo.3768383" httpUri="https://zenodo.org/record/3768383/files/figure.png" pageId="84" pageNumber="85">Fig. 42</figureCitation>
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</paragraph>
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<paragraph blockId="84.[140,999,166,298]" box="[140,238,272,298]" pageId="84" pageNumber="85">Table 14</paragraph>
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</subSubSection>
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<subSubSection pageId="84" pageNumber="85" type="etymology">
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<paragraph blockId="84.[140,1108,343,545]" pageId="84" pageNumber="85">
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<emphasis bold="true" box="[140,275,343,369]" pageId="84" pageNumber="85">
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<emphasis box="[140,270,343,369]" inLineHeading="true" pageId="84" pageNumber="85" reason="1">Etymology</emphasis>
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.
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</emphasis>
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<emphasis box="[281,378,343,369]" italics="true" pageId="84" pageNumber="85">Maghreb</emphasis>
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, western (Arabic); -
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<emphasis box="[600,648,343,368]" italics="true" pageId="84" pageNumber="85">ensis</emphasis>
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(Latin), from. Named for the area where the
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<typeStatus box="[183,282,378,404]" pageId="84" pageNumber="85">holotype</typeStatus>
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was discovered in the Kem Kem Beds of southeastern
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<collectingCountry box="[899,1004,378,404]" name="Morocco" pageId="84" pageNumber="85">Morocco</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="materials_examined">
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<paragraph blockId="84.[140,1108,343,545]" pageId="84" pageNumber="85">
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<materialsCitation ID-GBIF-Occurrence="2608868530" collectionCode="UCRC" pageId="84" pageNumber="85" specimenCode="PV2" specimenCount="1" typeStatus="holotype">
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<emphasis bold="true" box="[185,306,413,439]" pageId="84" pageNumber="85">
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<emphasis box="[185,301,413,439]" inLineHeading="true" pageId="84" pageNumber="85" reason="1">
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<typeStatus box="[185,301,413,439]" pageId="84" pageNumber="85">Holotype</typeStatus>
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</emphasis>
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.
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</emphasis>
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<collectionCode box="[312,393,413,439]" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="84" pageNumber="85" type="University or college">UCRC</collectionCode>
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<specimenCode box="[399,453,413,439]" collectionCode="UCRC" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="84" pageNumber="85" type="University or college">PV2</specimenCode>
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; anterior portion of the left dentary preserving four alveoli and one replacement tooth in the anteriormost tooth position (
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<figureCitation box="[854,938,448,474]" captionStart="Figure 42" captionStartId="84.[140,212,1494,1516]" captionTargetBox="[215,1044,624,1446]" captionTargetId="figure@84.[185,1063,600,1474]" captionTargetPageId="84" captionText="Figure 42. Dentary fragment of the crocodyliform Laganosuchus maghrebensis gen. n. sp. n. Anterior end of the left dentary (UCRC PV2). A Left lateral view. B Medial view (reversed). C Dorsal view. D Ventral view. E Replacement tooth in first dentary alveolus in lingual view. Scale bar in A-D equals 3 cm; scale bar in E equals 1 cm. Abbreviations: ad1, 3, alveolus for dentary tooth 1, 3; afo, alveolar foramina; asp, articular surface for the splenial; d1, 4, dentary tooth 1, 4; fl, fluting; fo, foramen; Mc, Meckel’s canal; rt, replacement tooth; sym, symphysis; tasp, tip of the articular surface for the splenial." figureDoi="http://doi.org/10.5281/zenodo.3768383" httpUri="https://zenodo.org/record/3768383/files/figure.png" pageId="84" pageNumber="85">Fig. 42</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="84.[140,1108,343,545]" pageId="84" pageNumber="85">
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<emphasis bold="true" box="[185,404,483,509]" pageId="84" pageNumber="85">
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<emphasis box="[185,287,483,509]" inLineHeading="true" pageId="84" pageNumber="85" reason="1">Referred</emphasis>
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material.
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</emphasis>
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<accessionNumber box="[415,572,483,510]" httpUri="http://www.ncbi.nlm.nih.gov/protein/CMN50838" pageId="84" pageNumber="85">
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<collectionCode box="[415,485,484,510]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="84" pageNumber="85" type="Museum">CMN</collectionCode>
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50838
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</accessionNumber>
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, anterior left dentary fragment preserving the symphyseal end and alveoli 1–3.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3768383" ID-Zenodo-Dep="3768383" httpUri="https://zenodo.org/record/3768383/files/figure.png" pageId="84" pageNumber="85" startId="84.[140,212,1494,1516]" targetBox="[215,1044,624,1446]" targetPageId="84">
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<paragraph blockId="84.[140,1108,1494,1676]" pageId="84" pageNumber="85">
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<emphasis bold="true" box="[140,1049,1494,1516]" pageId="84" pageNumber="85">
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Figure 42. Dentary fragment of the crocodyliform
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<taxonomicName authority="Sereno & Larsson, 2009" authorityName="Sereno & Larsson" authorityYear="2009" box="[646,917,1494,1516]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="84" pageNumber="85" phylum="Chordata" rank="species" status="gen. n.">
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<emphasis bold="true" box="[646,917,1494,1516]" italics="true" pageId="84" pageNumber="85">Laganosuchus maghrebensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[923,990,1494,1515]" pageId="84" pageNumber="85" rank="genus">gen. n.</taxonomicNameLabel>
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sp. n.
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</emphasis>
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Anterior end of the left dentary (UCRC PV2).
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<emphasis bold="true" box="[527,546,1526,1548]" pageId="84" pageNumber="85">A</emphasis>
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Left lateral view.
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<emphasis bold="true" box="[706,723,1526,1548]" pageId="84" pageNumber="85">B</emphasis>
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Medial view (reversed).
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||
<emphasis bold="true" box="[946,965,1526,1548]" pageId="84" pageNumber="85">C</emphasis>
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Dorsal view.
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<emphasis bold="true" box="[1088,1107,1526,1548]" pageId="84" pageNumber="85">D</emphasis>
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Ventral view.
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<emphasis bold="true" box="[264,279,1558,1580]" pageId="84" pageNumber="85">E</emphasis>
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Replacement tooth in first dentary alveolus in lingual view. Scale bar in A-D equals 3 cm; scale bar in E equals 1 cm. Abbreviations:
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<emphasis box="[535,593,1590,1612]" italics="true" pageId="84" pageNumber="85">ad1, 3</emphasis>
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, alveolus for dentary tooth 1, 3;
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<emphasis box="[901,928,1590,1611]" italics="true" pageId="84" pageNumber="85">afo</emphasis>
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, alveolar foramina;
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<emphasis box="[140,168,1623,1644]" italics="true" pageId="84" pageNumber="85">asp</emphasis>
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, articular surface for the splenial;
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<emphasis box="[479,532,1622,1644]" italics="true" pageId="84" pageNumber="85">d1, 4,</emphasis>
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dentary tooth 1, 4;
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<emphasis box="[716,722,1622,1643]" italics="true" pageId="84" pageNumber="85">fl</emphasis>
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, fluting;
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<emphasis box="[813,829,1622,1643]" italics="true" pageId="84" pageNumber="85">fo</emphasis>
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, foramen;
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<emphasis box="[929,958,1623,1644]" italics="true" pageId="84" pageNumber="85">Mc</emphasis>
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, Meckel’s canal;
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<emphasis box="[140,155,1655,1676]" italics="true" pageId="84" pageNumber="85">rt</emphasis>
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, replacement tooth;
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<emphasis box="[348,381,1655,1676]" italics="true" pageId="84" pageNumber="85">sym</emphasis>
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, symphysis;
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<emphasis box="[498,533,1655,1676]" italics="true" pageId="84" pageNumber="85">tasp</emphasis>
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, tip of the articular surface for the splenial.
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</paragraph>
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</caption>
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<paragraph blockId="85.[140,1108,166,440]" pageId="85" pageNumber="86">
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<emphasis bold="true" box="[185,347,166,193]" pageId="85" pageNumber="86">
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<emphasis box="[185,243,167,193]" inLineHeading="true" pageId="85" pageNumber="86" reason="1">Type</emphasis>
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locality.
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</emphasis>
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Er Rachidia District,
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<collectingCountry box="[607,709,167,193]" name="Morocco" pageId="85" pageNumber="86">Morocco</collectingCountry>
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(exact locality unknown). The referred specimen (
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<accessionNumber box="[334,488,202,229]" httpUri="http://www.ncbi.nlm.nih.gov/protein/CMN50838" pageId="85" pageNumber="86">
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<collectionCode box="[334,405,202,228]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="85" pageNumber="86" type="Museum">CMN</collectionCode>
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50838
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</accessionNumber>
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) probably was found south of Erfoud (Fig. 1A, B).
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</paragraph>
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<paragraph blockId="85.[140,1108,166,440]" pageId="85" pageNumber="86">
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<emphasis bold="true" box="[185,294,237,263]" pageId="85" pageNumber="86">
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<emphasis box="[185,288,237,263]" inLineHeading="true" pageId="85" pageNumber="86" reason="1">Horizon</emphasis>
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.
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</emphasis>
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Kem Kem Beds, upper member; Upper Cretaceous (Cenomanian), ca. 95 Mya (
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<bibRefCitation author="Sereno PC & Dutheil DB & Iarochene M & Larsson HCE & Lyon GH" box="[242,452,272,299]" pageId="85" pageNumber="86" pagination="986 - 991" refId="ref47291" refString="Sereno PC, Dutheil DB, Iarochene M, Larsson HCE, Lyon GH et al. (1996) Predatory dinosaurs from the Sahara and Late Cretaceous faunal differentiation. Science 272: 986 - 991." type="journal article" year="1996">Sereno et al. 1996</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86" type="diagnosis">
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<paragraph blockId="85.[140,1108,166,440]" pageId="85" pageNumber="86">
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<emphasis bold="true" box="[185,312,307,333]" pageId="85" pageNumber="86">
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<emphasis box="[185,307,307,333]" inLineHeading="true" pageId="85" pageNumber="86" reason="1">Diagnosis</emphasis>
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.
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</emphasis>
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Metasuchian with a narrow, well defined groove on the ventral aspect of the anterior dentary immediately lateral to the splenial that arcs to the posterior aspect of the symphysis; shallow, anteriorly tapering trough on the anterior dentary just lateral to the more sharply defined groove.
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</paragraph>
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<paragraph blockId="85.[140,1108,483,1003]" pageId="85" pageNumber="86">
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<emphasis bold="true" box="[140,244,483,509]" pageId="85" pageNumber="86">
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<emphasis box="[140,239,483,509]" inLineHeading="true" pageId="85" pageNumber="86" reason="1">Dentary</emphasis>
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.
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</emphasis>
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The anterior portion of the dentary is preserved in two specimens of
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<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="85" pageNumber="86" phylum="Chordata" rank="species">
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<emphasis italics="true" pageId="85" pageNumber="86">Laganosuchus maghrebensis</emphasis>
|
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</taxonomicName>
|
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. Th e very rugose dentary symphysis suggests that it may have fused with maturity, and that its size, which is somewhat smaller than
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<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="85" pageNumber="86" phylum="Chordata" rank="species">
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<emphasis italics="true" pageId="85" pageNumber="86">Laganosuchus thaumastos</emphasis>
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</taxonomicName>
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, may not be significant.
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</paragraph>
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<paragraph blockId="85.[140,1108,483,1003]" pageId="85" pageNumber="86">
|
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The more complete specimen (
|
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<figureCitation box="[539,621,624,650]" captionStart="Figure 42" captionStartId="84.[140,212,1494,1516]" captionTargetBox="[215,1044,624,1446]" captionTargetId="figure@84.[185,1063,600,1474]" captionTargetPageId="84" captionText="Figure 42. Dentary fragment of the crocodyliform Laganosuchus maghrebensis gen. n. sp. n. Anterior end of the left dentary (UCRC PV2). A Left lateral view. B Medial view (reversed). C Dorsal view. D Ventral view. E Replacement tooth in first dentary alveolus in lingual view. Scale bar in A-D equals 3 cm; scale bar in E equals 1 cm. Abbreviations: ad1, 3, alveolus for dentary tooth 1, 3; afo, alveolar foramina; asp, articular surface for the splenial; d1, 4, dentary tooth 1, 4; fl, fluting; fo, foramen; Mc, Meckel’s canal; rt, replacement tooth; sym, symphysis; tasp, tip of the articular surface for the splenial." figureDoi="http://doi.org/10.5281/zenodo.3768383" httpUri="https://zenodo.org/record/3768383/files/figure.png" pageId="85" pageNumber="86">Fig. 42</figureCitation>
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A-D) shows a remarkable similarity to
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<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="85" pageNumber="86" phylum="Chordata" rank="species">
|
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<emphasis italics="true" pageId="85" pageNumber="86">Laganosuchus thaumastos</emphasis>
|
||
</taxonomicName>
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||
. Both have slender U-shaped lower jaws with the symphysis restricted to the dentary, festooned alveoli with enlarged first and second teeth, and a sharply incised Meckel’s canal developed as a narrow groove. Th e teeth are also spikeshaped without recurvature or marginal ornamentation.
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</paragraph>
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<paragraph blockId="85.[140,1108,483,1003]" lastBlockId="86.[140,1108,167,440]" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86">
|
||
Several differences, however, establish
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[631,807,800,826]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="85" pageNumber="86" phylum="Chordata" rank="species">
|
||
<emphasis box="[631,807,800,826]" italics="true" pageId="85" pageNumber="86">L. maghrebensis</emphasis>
|
||
</taxonomicName>
|
||
as a distinct species. The dentary is narrower near the symphysis (
|
||
<figureCitation box="[587,690,835,862]" captionStart="Figure 42" captionStartId="84.[140,212,1494,1516]" captionTargetBox="[215,1044,624,1446]" captionTargetId="figure@84.[185,1063,600,1474]" captionTargetPageId="84" captionText="Figure 42. Dentary fragment of the crocodyliform Laganosuchus maghrebensis gen. n. sp. n. Anterior end of the left dentary (UCRC PV2). A Left lateral view. B Medial view (reversed). C Dorsal view. D Ventral view. E Replacement tooth in first dentary alveolus in lingual view. Scale bar in A-D equals 3 cm; scale bar in E equals 1 cm. Abbreviations: ad1, 3, alveolus for dentary tooth 1, 3; afo, alveolar foramina; asp, articular surface for the splenial; d1, 4, dentary tooth 1, 4; fl, fluting; fo, foramen; Mc, Meckel’s canal; rt, replacement tooth; sym, symphysis; tasp, tip of the articular surface for the splenial." figureDoi="http://doi.org/10.5281/zenodo.3768383" httpUri="https://zenodo.org/record/3768383/files/figure.png" pageId="85" pageNumber="86">Fig. 42C</figureCitation>
|
||
), lacking the internal crest that thickens the dentary in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[350,501,871,897]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="85" pageNumber="86" phylum="Chordata" rank="species">
|
||
<emphasis box="[350,501,871,897]" italics="true" pageId="85" pageNumber="86">L. thaumastos</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[516,619,871,897]" captionStart="Figure 38" captionStartId="79.[140,212,1526,1548]" captionTargetBox="[251,1005,635,1499]" captionTargetId="figure@79.[227,1021,623,1512]" captionTargetPageId="79" captionText="Figure 38. Lower jaws of the crocodyliform Laganosuchus thaumastos gen. n. sp. n. Cast (UCRC PVC9) of the anterior portion of the left dentary and splenial (MNN IGU13). A Left lateral view. B Medial view (reversed). C Dorsal view. D Ventral view. Scale bar equals 5 cm. Abbreviations: ad1–8, alveolus for dentary tooth 1–8; asp, articular surface for the splenial; cr, crest; d, dentary; dpr, dorsal process; gr, groove (for upper teeth); Mc, Meckel’s canal; sp, splenial; vpr, ventral process." figureDoi="http://doi.org/10.5281/zenodo.3768377" httpUri="https://zenodo.org/record/3768377/files/figure.png" pageId="85" pageNumber="86">Fig. 38B</figureCitation>
|
||
). Th e anterior alveoli in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[895,1067,871,897]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="85" pageNumber="86" phylum="Chordata" rank="species">
|
||
<emphasis box="[895,1067,871,897]" italics="true" pageId="85" pageNumber="86">L. maghrebensis</emphasis>
|
||
</taxonomicName>
|
||
are not procumbent or exposed in lateral view as in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[696,848,906,932]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="85" pageNumber="86" phylum="Chordata" rank="species">
|
||
<emphasis box="[696,848,906,932]" italics="true" pageId="85" pageNumber="86">L. thaumastos</emphasis>
|
||
</taxonomicName>
|
||
. Likewise, an articular scar for the splenial in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[391,561,941,967]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="85" pageNumber="86" phylum="Chordata" rank="species">
|
||
<emphasis box="[391,561,941,967]" italics="true" pageId="85" pageNumber="86">L. maghrebensis</emphasis>
|
||
</taxonomicName>
|
||
shows that its anterior end tapers to a narrow tip along the ventral margin (
|
||
<figureCitation box="[430,529,976,1002]" captionStart="Figure 42" captionStartId="84.[140,212,1494,1516]" captionTargetBox="[215,1044,624,1446]" captionTargetId="figure@84.[185,1063,600,1474]" captionTargetPageId="84" captionText="Figure 42. Dentary fragment of the crocodyliform Laganosuchus maghrebensis gen. n. sp. n. Anterior end of the left dentary (UCRC PV2). A Left lateral view. B Medial view (reversed). C Dorsal view. D Ventral view. E Replacement tooth in first dentary alveolus in lingual view. Scale bar in A-D equals 3 cm; scale bar in E equals 1 cm. Abbreviations: ad1, 3, alveolus for dentary tooth 1, 3; afo, alveolar foramina; asp, articular surface for the splenial; d1, 4, dentary tooth 1, 4; fl, fluting; fo, foramen; Mc, Meckel’s canal; rt, replacement tooth; sym, symphysis; tasp, tip of the articular surface for the splenial." figureDoi="http://doi.org/10.5281/zenodo.3768383" httpUri="https://zenodo.org/record/3768383/files/figure.png" pageId="85" pageNumber="86">Fig. 42B</figureCitation>
|
||
) in contrast to the bifurcated flanges in
|
||
<emphasis italics="true" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86">L. thaumas- tos</emphasis>
|
||
(
|
||
<figureCitation box="[183,285,167,193]" captionStart="Figure 38" captionStartId="79.[140,212,1526,1548]" captionTargetBox="[251,1005,635,1499]" captionTargetId="figure@79.[227,1021,623,1512]" captionTargetPageId="79" captionText="Figure 38. Lower jaws of the crocodyliform Laganosuchus thaumastos gen. n. sp. n. Cast (UCRC PVC9) of the anterior portion of the left dentary and splenial (MNN IGU13). A Left lateral view. B Medial view (reversed). C Dorsal view. D Ventral view. Scale bar equals 5 cm. Abbreviations: ad1–8, alveolus for dentary tooth 1–8; asp, articular surface for the splenial; cr, crest; d, dentary; dpr, dorsal process; gr, groove (for upper teeth); Mc, Meckel’s canal; sp, splenial; vpr, ventral process." figureDoi="http://doi.org/10.5281/zenodo.3768377" httpUri="https://zenodo.org/record/3768377/files/figure.png" pageId="86" pageNumber="87">Fig. 38B</figureCitation>
|
||
). Th e incised groove representing Meckel’s canal is located on the ventral, rather than lingual, aspect of the dentary (
|
||
<figureCitation box="[615,723,202,228]" captionStart="Figure 42" captionStartId="84.[140,212,1494,1516]" captionTargetBox="[215,1044,624,1446]" captionTargetId="figure@84.[185,1063,600,1474]" captionTargetPageId="84" captionText="Figure 42. Dentary fragment of the crocodyliform Laganosuchus maghrebensis gen. n. sp. n. Anterior end of the left dentary (UCRC PV2). A Left lateral view. B Medial view (reversed). C Dorsal view. D Ventral view. E Replacement tooth in first dentary alveolus in lingual view. Scale bar in A-D equals 3 cm; scale bar in E equals 1 cm. Abbreviations: ad1, 3, alveolus for dentary tooth 1, 3; afo, alveolar foramina; asp, articular surface for the splenial; d1, 4, dentary tooth 1, 4; fl, fluting; fo, foramen; Mc, Meckel’s canal; rt, replacement tooth; sym, symphysis; tasp, tip of the articular surface for the splenial." figureDoi="http://doi.org/10.5281/zenodo.3768383" httpUri="https://zenodo.org/record/3768383/files/figure.png" pageId="86" pageNumber="87">Fig. 42D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="85.[140,1107,1077,1131]" pageId="85" pageNumber="86">
|
||
<tableNote pageId="85" pageNumber="86" targetBox="[147,1099,1160,1667]" targetPageId="85">
|
||
<emphasis bold="true" box="[140,244,1077,1099]" pageId="85" pageNumber="86">Table |4.</emphasis>
|
||
Dimensions (mm) of the anterior end of the right dentary of
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[861,1107,1077,1098]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="85" pageNumber="86" phylum="Chordata" rank="species">
|
||
<emphasis box="[861,1107,1077,1098]" italics="true" pageId="85" pageNumber="86">Laganosuchus maghrebensis</emphasis>
|
||
</taxonomicName>
|
||
(UCRC PV2).
|
||
</tableNote>
|
||
</paragraph>
|
||
<paragraph pageId="85" pageNumber="86">
|
||
<table box="[147,1099,1160,1667]" gridcols="3" gridrows="6" pageId="85" pageNumber="86">
|
||
<tr box="[147,1099,1160,1323]" gridrow="0" pageId="85" pageNumber="86">
|
||
<th box="[147,269,1160,1323]" gridcol="0" gridrow="0" pageId="85" pageNumber="86">
|
||
<emphasis bold="true" box="[168,269,1160,1184]" pageId="85" pageNumber="86">Structure</emphasis>
|
||
Dentary
|
||
</th>
|
||
<th box="[315,945,1160,1323]" gridcol="1" gridrow="0" pageId="85" pageNumber="86">
|
||
<emphasis bold="true" box="[569,716,1161,1185]" pageId="85" pageNumber="86">Measurement</emphasis>
|
||
Symphysis, dorsoventral height Symphysis, maximum anteroposterior width Dentary ramus between alveolus 2 and 3, dorsoventral height Dentary ramus between alveolus 2 and 3,transverse width
|
||
</th>
|
||
<th box="[1008,1099,1160,1323]" gridcol="2" gridrow="0" pageId="85" pageNumber="86">
|
||
<emphasis bold="true" box="[1008,1085,1161,1185]" pageId="85" pageNumber="86">Length</emphasis>
|
||
25.5 14.6 19.1 11.4
|
||
</th>
|
||
</tr>
|
||
<tr box="[147,1099,1333,1391]" gridrow="1" pageId="85" pageNumber="86">
|
||
<th box="[147,269,1333,1391]" gridcol="0" gridrow="1" pageId="85" pageNumber="86">Alveolus 1</th>
|
||
<td box="[315,945,1333,1391]" gridcol="1" gridrow="1" pageId="85" pageNumber="86">Maximum mesiodistal length Maximum labiolingual width</td>
|
||
<td box="[1008,1099,1333,1391]" gridcol="2" gridrow="1" pageId="85" pageNumber="86">10.1 6.4</td>
|
||
</tr>
|
||
<tr box="[147,1099,1402,1460]" gridrow="2" pageId="85" pageNumber="86">
|
||
<th box="[147,269,1402,1460]" gridcol="0" gridrow="2" pageId="85" pageNumber="86">Alveolus 2</th>
|
||
<td box="[315,945,1402,1460]" gridcol="1" gridrow="2" pageId="85" pageNumber="86">Maximum mesiodistal length Maximum labiolingual width</td>
|
||
<td box="[1008,1099,1402,1460]" gridcol="2" gridrow="2" pageId="85" pageNumber="86">7.3 5.0</td>
|
||
</tr>
|
||
<tr box="[147,1099,1471,1529]" gridrow="3" pageId="85" pageNumber="86">
|
||
<th box="[147,269,1471,1529]" gridcol="0" gridrow="3" pageId="85" pageNumber="86">Alveolus 3</th>
|
||
<td box="[315,945,1471,1529]" gridcol="1" gridrow="3" pageId="85" pageNumber="86">Maximum mesiodistal length Maximum labiolingual width</td>
|
||
<td box="[1008,1099,1471,1529]" gridcol="2" gridrow="3" pageId="85" pageNumber="86">6.4 4.0</td>
|
||
</tr>
|
||
<tr box="[147,1099,1540,1598]" gridrow="4" pageId="85" pageNumber="86">
|
||
<th box="[147,269,1540,1598]" gridcol="0" gridrow="4" pageId="85" pageNumber="86">Alveolus 4</th>
|
||
<td box="[315,945,1540,1598]" gridcol="1" gridrow="4" pageId="85" pageNumber="86">Maximum mesiodistal length Maximum labiolingual width</td>
|
||
<td box="[1008,1099,1540,1598]" gridcol="2" gridrow="4" pageId="85" pageNumber="86">11.9 5.7</td>
|
||
</tr>
|
||
<tr box="[147,1099,1609,1667]" gridrow="5" pageId="85" pageNumber="86">
|
||
<th box="[147,269,1609,1667]" gridcol="0" gridrow="5" pageId="85" pageNumber="86">Crown d1</th>
|
||
<td box="[315,945,1609,1667]" gridcol="1" gridrow="5" pageId="85" pageNumber="86">Replacement crown, height Replacement crown, mesiodistal width of base</td>
|
||
<td box="[1008,1099,1609,1667]" gridcol="2" gridrow="5" pageId="85" pageNumber="86">12.2 5.3</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="86.[140,1108,167,440]" pageId="86" pageNumber="87">
|
||
Although crown size in the two species is very similar, the alveolus of the caniniform tooth (d4) is slightly larger than comparable measurements for d1 (Table 14), the reverse of the condition in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[502,779,307,333]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="86" pageNumber="87" phylum="Chordata" rank="species">
|
||
<emphasis box="[502,779,307,333]" italics="true" pageId="86" pageNumber="87">Laganosuchus thaumastos</emphasis>
|
||
</taxonomicName>
|
||
(Table 13).
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[926,1100,307,333]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="86" pageNumber="87" phylum="Chordata" rank="species">
|
||
<emphasis box="[926,1100,307,333]" italics="true" pageId="86" pageNumber="87">L. maghrebensis</emphasis>
|
||
</taxonomicName>
|
||
, in addition, shows low fluting on the lingual aspect of the crown of the first dentary tooth (
|
||
<figureCitation box="[217,320,378,404]" captionStart="Figure 42" captionStartId="84.[140,212,1494,1516]" captionTargetBox="[215,1044,624,1446]" captionTargetId="figure@84.[185,1063,600,1474]" captionTargetPageId="84" captionText="Figure 42. Dentary fragment of the crocodyliform Laganosuchus maghrebensis gen. n. sp. n. Anterior end of the left dentary (UCRC PV2). A Left lateral view. B Medial view (reversed). C Dorsal view. D Ventral view. E Replacement tooth in first dentary alveolus in lingual view. Scale bar in A-D equals 3 cm; scale bar in E equals 1 cm. Abbreviations: ad1, 3, alveolus for dentary tooth 1, 3; afo, alveolar foramina; asp, articular surface for the splenial; d1, 4, dentary tooth 1, 4; fl, fluting; fo, foramen; Mc, Meckel’s canal; rt, replacement tooth; sym, symphysis; tasp, tip of the articular surface for the splenial." figureDoi="http://doi.org/10.5281/zenodo.3768383" httpUri="https://zenodo.org/record/3768383/files/figure.png" pageId="86" pageNumber="87">Fig. 42E</figureCitation>
|
||
). A comparable crown, however is not available at the anterior end of the dentary series in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[373,524,413,439]" class="Reptilia" family="Stomatosuchidae" genus="Laganosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="86" pageNumber="87" phylum="Chordata" rank="species">
|
||
<emphasis box="[373,524,413,439]" italics="true" pageId="86" pageNumber="87">L. thaumastos</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |