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<document id="8EF174D86551085082A74E6BA4EFE229" ID-DOI="10.11646/zootaxa.2096.1.27" ID-ISSN="1175-5326" ID-Zenodo-Dep="5321990" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621422523" checkinUser="felipe" docAuthor="Strugnell, Jan, Voight, Janet R., Collins, Patrick C. &amp; Allcock, A. Louise" docDate="2009" docId="03914A02FF9A9C1CFF6693B4FE89FEF7" docLanguage="en" docName="zt02096p459.pdf" docOrigin="Zootaxa 2096 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2096.1.27" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Vulcanoctopus hydrothermalis Gonzalez &amp; Guerra 1998" docType="treatment" docVersion="4" lastPageNumber="450" masterDocId="FFA8327AFF929C15FFF1914AFFD0FFD1" masterDocTitle="Molecular phylogenetic analysis of a known and a new hydrothermal vent octopod: their relationships with the genus Benthoctopus (Cephalopoda: Octopodidae)" masterLastPageNumber="459" masterPageNumber="442" pageNumber="450" updateTime="1699084934570" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="E39A3A43A30C8E445548A76C7B853690" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1851DCFC1DD29A2CC71C43F699E5CEC2">Molecular phylogenetic analysis of a known and a new hydrothermal vent octopod: their relationships with the genus Benthoctopus (Cephalopoda: Octopodidae)</mods:title>
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<mods:name id="3107CDC80D289A717B494CFA6D69A316" type="personal">
<mods:role id="FBF5ACDCD8A8B6F27D4E532C51BA6E69">
<mods:roleTerm id="CEBFC3863BD540ABED8D664BC3978658">Author</mods:roleTerm>
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<mods:namePart id="C43FB7C6F8AE0E2A62C8EBA9EB808BBD">Strugnell, Jan</mods:namePart>
<mods:affiliation id="D1E5DA9A8EDDAF8725741DF71C4ECD3F">Department of Zoology, University of Cambridge, Downing Street, Cambridge CB 2 3 EJ, UK School of Biological Sciences, Queen's University Belfast, 97 Lisburn Road, Belfast BT 9 7 BL, UK Department of Zoology, Field Museum of Natural History, 1400 S. Lake Shore Dr, Chicago, IL 60605 - 2496, USA Duke Marine Laboratory, 135 Duke Marine Lab Road, Beaufort, NC 28516, USA The Martin Ryan Marine Science Institute, National University of Ireland, Galway, University Road, Galway, Ireland Corresponding author: jan. strugnell @ gmail. com</mods:affiliation>
</mods:name>
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<mods:namePart id="D282F3BD74DC16CF286B13A3FE8410E5">Voight, Janet R.</mods:namePart>
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<mods:name id="B443BBA89F518B85D41DD75323996390" type="personal">
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<mods:namePart id="6112E16F68B5157EE9A21321D45CAD3C">Collins, Patrick C.</mods:namePart>
</mods:name>
<mods:name id="A0DACA6D89048882967AFA2ECF4236E8" type="personal">
<mods:role id="23873CDEA01E661A8718532BEE0E1C1D">
<mods:roleTerm id="90F719BDA49F5B48441A9D6CE545968C">Author</mods:roleTerm>
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<mods:namePart id="5D548B40B97677142BAC7A192389F3DF">Allcock, A. Louise</mods:namePart>
</mods:name>
<mods:typeOfResource id="DEDEFDA6D247310548B4BEFABC075B5A">text</mods:typeOfResource>
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<mods:title id="29D89094170B2EAC6A1E7D560E1F6D3F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D3FCE0FE802688F38A157F7598B0474D">
<mods:date id="6A0E7E49B83901BF8346E389DF928EB3">2009</mods:date>
<mods:detail id="8CE34BFB992AA3FA5B7779AD573832B6" type="pubDate">
<mods:number id="EA4B4A2302900950727F806E2E8B1E1A">2009-05-11</mods:number>
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<mods:detail id="3A9A613DFC3E145D6E9A4FF667256AFA" type="volume">
<mods:number id="0A6DBA465124E6B9027F9DA1D02E0FE5">2096</mods:number>
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<mods:detail id="A687A5F8B66262C8359B491A1B54413A" type="issue">
<mods:number id="AEC1B47C73AEF14ABA2FE65DF88C6DE8">1</mods:number>
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<mods:start id="87548345119E3F5426AF71A0CBEBDE58">442</mods:start>
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<mods:url id="39BC2EB399092D220C7C419ECB0A4C91">https://biotaxa.org/Zootaxa/article/view/zootaxa.2096.1.27</mods:url>
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<mods:classification id="C20B6BFD0C1E5025A685EC825448C8E5">journal article</mods:classification>
<mods:identifier id="38AD03649EE79DF520DD09264FAFB3CA" type="DOI">10.11646/zootaxa.2096.1.27</mods:identifier>
<mods:identifier id="A0BC9F126945AF9052B896987FC5439E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8E7826E2983C3095B8C7C75CB4F12936" type="Zenodo-Dep">5321990</mods:identifier>
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<treatment id="03914A02FF9A9C1CFF6693B4FE89FEF7" ID-DOI="http://doi.org/10.5281/zenodo.5334614" ID-Zenodo-Dep="5334614" LSID="urn:lsid:plazi:treatment:03914A02FF9A9C1CFF6693B4FE89FEF7" httpUri="http://treatment.plazi.org/id/03914A02FF9A9C1CFF6693B4FE89FEF7" lastPageId="9" lastPageNumber="450" pageId="8" pageNumber="450">
<subSubSection id="C322A89FFF9A9C1DFF6693B4FC81FCC9" box="[151,849,766,792]" pageId="8" pageNumber="450" type="nomenclature">
<paragraph id="8B87FB14FF9A9C1DFF6693B4FC81FCC9" blockId="8.[151,1439,731,2026]" box="[151,849,766,792]" pageId="8" pageNumber="450">
<heading id="D0CF4C78FF9A9C1DFF6693B4FC81FCC9" bold="true" box="[151,849,766,792]" fontSize="11" level="1" pageId="8" pageNumber="450" reason="1">
<taxonomicName id="4C388097FF9A9C1DFF6693B4FC81FCC9" ID-CoL="5BSC6" authority="Gonzalez &amp; Guerra, 1998" authorityName="Gonzalez &amp; Guerra" authorityYear="1998" box="[151,849,766,792]" class="Cephalopoda" family="Enteroctopodidae" genus="Vulcanoctopus" kingdom="Animalia" order="Octopoda" pageId="8" pageNumber="450" phylum="Mollusca" rank="species" species="hydrothermalis">
<emphasis id="B94C2706FF9A9C1DFF6693B4FC81FCC9" bold="true" box="[151,849,766,792]" pageId="8" pageNumber="450">
<emphasis id="B94C2706FF9A9C1DFF6693B4FDD9FCC9" bold="true" box="[151,521,766,792]" italics="true" pageId="8" pageNumber="450">Vulcanoctopus hydrothermalis</emphasis>
<bibRefCitation id="EFA986E5FF9A9C1DFDE193B4FC81FCC9" author="Gonzalez &amp; Guerra" box="[528,849,766,792]" firstAuthor="Gonzalez" pageId="8" pageNumber="450" pagination="169 - 184" refId="ref10672" refString="Gonzalez, A. F., Guerra, A., Pascual, S. &amp; Briand, P. (1998) Vulcanoctopus hydrothermalis gen. et sp. nov. (Mollusca, Cephalopoda): an octopod from a deep-sea hydrothermal vent site. Cahiers de Biologie Marine, 39, 169 - 184." type="journal article" year="1998">González &amp; Guerra, 1998</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C322A89FFF9A9C1CFF66926BFE89FEF7" lastPageId="9" lastPageNumber="451" pageId="8" pageNumber="450" type="materials_examined">
<paragraph id="8B87FB14FF9A9C1DFF66926BFDCEFC59" blockId="8.[151,1439,731,2026]" pageId="8" pageNumber="450">
<emphasis id="B94C2706FF9A9C1DFF66926BFE19FCEB" box="[151,457,801,826]" italics="true" pageId="8" pageNumber="450">Type material examined:</emphasis>
<taxonomicName id="4C388097FF9A9C1DFE23926BFB40FCEA" authority="Gonzalez &amp; Guerra, 1998" authorityName="Gonzalez &amp; Guerra" authorityYear="1998" box="[466,1168,801,827]" class="Cephalopoda" family="Enteroctopodidae" genus="Vulcanoctopus" kingdom="Animalia" order="Octopoda" pageId="8" pageNumber="450" phylum="Mollusca" rank="species" species="hydrothermalis">
<emphasis id="B94C2706FF9A9C1DFE23926BFC99FCEB" box="[466,841,801,826]" italics="true" pageId="8" pageNumber="450">Vulcanoctopus hydrothermalis</emphasis>
<materialsCitation id="3B50F149FF9A9C1DFCA3926BFB40FCEA" box="[850,1168,801,827]" collectingDate="1996-02-19" county="Guerra" latitude="12.8" location="Briand" longLatPrecision="1292" longitude="-103.933334" municipality="Pascual" pageId="8" pageNumber="450" specimenCount="1" stateProvince="Monaghan" typeStatus="holotype">
<bibRefCitation id="EFA986E5FF9A9C1DFCA3926BFB40FCEA" author="Gonzalez &amp; Guerra" box="[850,1168,801,827]" firstAuthor="Gonzalez" pageId="8" pageNumber="450" pagination="169 - 184" refId="ref10672" refString="Gonzalez, A. F., Guerra, A., Pascual, S. &amp; Briand, P. (1998) Vulcanoctopus hydrothermalis gen. et sp. nov. (Mollusca, Cephalopoda): an octopod from a deep-sea hydrothermal vent site. Cahiers de Biologie Marine, 39, 169 - 184." type="journal article" year="1998">
González &amp; Guerra,
<quantity id="4CC056F1FF9A9C1DFBA5926BFB40FCEA" box="[1108,1168,801,827]" metricMagnitude="1" metricUnit="m" metricValue="5.07492" pageId="8" pageNumber="450" unit="in" value="1998.0">
<locationDeviation id="9A35E236FF9A9C1DFBA5926BFB40FCEA" box="[1108,1168,801,827]" location="Gonzalez" pageId="8" pageNumber="450">1998</locationDeviation>
</quantity>
</bibRefCitation>
</materialsCitation>
</taxonomicName>
in González,
<collectingCounty id="62E68398FF9A9C1DFACE926BFA47FCEA" box="[1343,1431,801,827]" pageId="8" pageNumber="450">Guerra</collectingCounty>
,
<collectingMunicipality id="6BE3616EFF9A9C1DFF669202FF21FCB3" box="[151,241,840,866]" pageId="8" pageNumber="450">Pascual</collectingMunicipality>
&amp;
<location id="8EE7ADCFFF9A9C1DFEE99202FEBBFCB3" LSID="urn:lsid:plazi:treatment:03914A02FF9A9C1CFF6693B4FE89FEF7:8EE7ADCFFF9A9C1DFEE99202FEBBFCB3" box="[280,363,840,866]" county="Guerra" latitude="12.8" longLatPrecision="1292" longitude="-103.933334" municipality="Pascual" name="Briand" pageId="8" pageNumber="450" stateProvince="Monaghan">Briand</location>
,
<typeStatus id="548345B6FF9A9C1DFE899202FE32FCB3" box="[376,482,840,866]" pageId="8" pageNumber="450" type="holotype">holotype</typeStatus>
,
<collectingRegion id="49FC35F6FF9A9C1DFE1F9202FD98FCB3" box="[494,584,840,866]" country="Ireland" name="Monaghan" pageId="8" pageNumber="450">MNHN</collectingRegion>
2885, HOT 96
<location id="8EE7ADCFFF9A9C1DFCF59202FC00FCB3" LSID="urn:lsid:plazi:treatment:03914A02FF9A9C1CFF6693B4FE89FEF7:8EE7ADCFFF9A9C1DFCF59202FC00FCB3" box="[772,976,840,866]" county="Guerra" latitude="12.8" longLatPrecision="1292" longitude="-103.933334" municipality="Pascual" name="East Pacific Rise" pageId="8" pageNumber="450" stateProvince="Monaghan">East Pacific Rise</location>
,
<location id="8EE7ADCFFF9A9C1DFC2C9202FB7AFCB3" LSID="urn:lsid:plazi:treatment:03914A02FF9A9C1CFF6693B4FE89FEF7:8EE7ADCFFF9A9C1DFC2C9202FB7AFCB3" box="[989,1194,840,866]" county="Guerra" latitude="12.8" longLatPrecision="1292" longitude="-103.933334" municipality="Pascual" name="North of Genesis" pageId="8" pageNumber="450" stateProvince="Monaghan">North of Genesis</location>
,
<geoCoordinate id="EE0C9DD3FF9A9C1DFB469202FAC9FCB3" box="[1207,1305,839,866]" degrees="12" direction="north" minutes="48" orientation="latitude" pageId="8" pageNumber="450" precision="925" value="12.8">12°48'N</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF9A9C1DFAD09202FA4AFCB3" box="[1313,1434,839,866]" degrees="103" direction="west" minutes="56" orientation="longitude" pageId="8" pageNumber="450" precision="925" value="-103.933334">103°56'W</geoCoordinate>
,
<date id="FF86DDD4FF9A9C1DFF669224FEB9FC59" box="[151,361,878,904]" pageId="8" pageNumber="450" value="1996-02-19">
<collectingDate id="EFC2243CFF9A9C1DFF669224FEB9FC59" box="[151,361,878,904]" pageId="8" pageNumber="450" value="1996-02-19">
19 February
<quantity id="4CC056F1FF9A9C1DFEDD9224FEB9FC59" box="[300,361,878,904]" metricMagnitude="3" metricUnit="m" metricValue="1.9962646999999998" pageId="8" pageNumber="450" unit="m" value="1996.2647">1996</quantity>
</collectingDate>
</date>
, 2647 m, male.
</paragraph>
<paragraph id="8B87FB14FF9A9C1DFF3792DFFEEEFA89" blockId="8.[151,1439,731,2026]" pageId="8" pageNumber="450">
<emphasis id="B94C2706FF9A9C1DFF3792DFFDE1FC7F" box="[198,561,917,942]" italics="true" pageId="8" pageNumber="450">Additional material examined:</emphasis>
<taxonomicName id="4C388097FF9A9C1DFDCD92DFFBD9FC7E" authority=": FMNH" authorityName="FMNH" box="[572,1033,917,943]" class="Cephalopoda" family="Enteroctopodidae" genus="Vulcanoctopus" kingdom="Animalia" order="Octopoda" pageId="8" pageNumber="450" phylum="Mollusca" rank="species" species="hydrothermalis">
<emphasis id="B94C2706FF9A9C1DFDCD92DFFC7AFC7F" box="[572,938,917,942]" italics="true" pageId="8" pageNumber="450">Vulcanoctopus hydrothermalis:</emphasis>
FMNH
</taxonomicName>
307183,
<collectionCode id="ED2963D1FF9A9C1DFB8892DFFB4FFC7E" box="[1145,1183,917,943]" country="Italy" name="Collection of Leptospira Strains" pageId="8" pageNumber="450">RV</collectionCode>
<taxonomicName id="4C388097FF9A9C1DFB5892DFFA45FC7F" authority="HOV Alvin" authorityName="HOV Alvin" box="[1193,1429,917,943]" class="Insecta" family="Curculionidae" genus="Atlantis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="450" phylum="Arthropoda" rank="genus">
Atlantis HOV
<emphasis id="B94C2706FF9A9C1DFAA692DFFA45FC7F" box="[1367,1429,917,942]" italics="true" pageId="8" pageNumber="450">Alvin</emphasis>
</taxonomicName>
,
<geoCoordinate id="EE0C9DD3FF9A9C1DFF6692F6FF29FC07" box="[151,249,955,982]" degrees="10" direction="north" minutes="46" orientation="latitude" pageId="8" pageNumber="450" precision="925" value="10.766666">10°46'N</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF9A9C1DFEF392F6FEABFC07" box="[258,379,955,982]" degrees="103" direction="west" minutes="39" orientation="longitude" pageId="8" pageNumber="450" precision="925" value="-103.65">103°39'W</geoCoordinate>
(East Pacific Rise),
<emphasis id="B94C2706FF9A9C1DFD8492F6FD64FC04" box="[629,692,956,981]" italics="true" pageId="8" pageNumber="450">Alvin</emphasis>
Dive 3936,
<date id="FF86DDD4FF9A9C1DFCBA92F6FBE7FC07" box="[843,1079,956,982]" pageId="8" pageNumber="450" value="2003-11-17">
17 November
<quantity id="4CC056F1FF9A9C1DFC0892F6FBE7FC07" box="[1017,1079,956,982]" metricMagnitude="3" metricUnit="m" metricValue="2.0032574" pageId="8" pageNumber="450" unit="m" value="2003.2574">2003</quantity>
</date>
, 2574 m, male, coll &amp; det JR Voight. *
<collectionCode id="ED2963D1FF9A9C1DFF0E92A8FE87FC2D" box="[255,343,994,1020]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="450" type="Museum">FMNH</collectionCode>
307184,
<collectionCode id="ED2963D1FF9A9C1DFE3792A8FE3CFC2D" box="[454,492,994,1020]" country="Italy" name="Collection of Leptospira Strains" pageId="8" pageNumber="450">RV</collectionCode>
<taxonomicName id="4C388097FF9A9C1DFE0592A8FD0DFC2A" authority="HOV Alvin" authorityName="HOV Alvin" box="[500,733,994,1020]" class="Insecta" family="Curculionidae" genus="Atlantis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="B94C2706FF9A9C1DFE0592A8FD9FFC2A" box="[500,591,994,1019]" italics="true" pageId="8" pageNumber="450">Atlantis</emphasis>
HOV
<emphasis id="B94C2706FF9A9C1DFD6E92A8FD0DFC2A" box="[671,733,994,1019]" italics="true" pageId="8" pageNumber="450">Alvin</emphasis>
</taxonomicName>
,
<geoCoordinate id="EE0C9DD3FF9A9C1DFD1D92A8FC9CFC2D" box="[748,844,994,1020]" degrees="08" direction="north" minutes="38" orientation="latitude" pageId="8" pageNumber="450" precision="925" value="8.633333">08°38'N</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF9A9C1DFCA492A8FC1BFC2D" box="[853,971,994,1020]" degrees="104" direction="west" minutes="13" orientation="longitude" pageId="8" pageNumber="450" precision="925" value="-104.21667">104°13'W</geoCoordinate>
(East Pacific Rise),
<emphasis id="B94C2706FF9A9C1DFB4C92A8FB2BFC2A" box="[1213,1275,994,1019]" italics="true" pageId="8" pageNumber="450">Alvin</emphasis>
Dive 3926,
<date id="FF86DDD4FF9A9C1DFA7C92A8FE8AFBF2" pageId="8" pageNumber="450" value="2003-11-07">
7 November
<quantity id="4CC056F1FF9A9C1DFEED9543FE8AFBF2" box="[284,346,1033,1059]" metricMagnitude="3" metricUnit="m" metricValue="2.0032541" pageId="8" pageNumber="450" unit="m" value="2003.2541">2003</quantity>
</date>
, 2541 m, male, coll &amp; det JR Voight.
<collectionCode id="ED2963D1FF9A9C1DFCD29543FCA8FBF2" box="[803,888,1033,1059]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="450" type="Museum">FMNH</collectionCode>
278064,
<collectionCode id="ED2963D1FF9A9C1DFC189543FBC0FBF2" box="[1001,1040,1033,1059]" country="Italy" name="Collection of Leptospira Strains" pageId="8" pageNumber="450">RV</collectionCode>
<taxonomicName id="4C388097FF9A9C1DFBE89543FAD5FBF3" authority="HOV Alvin" authorityName="HOV Alvin" box="[1049,1285,1033,1059]" class="Insecta" family="Curculionidae" genus="Atlantis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="B94C2706FF9A9C1DFBE89543FBA6FBF3" box="[1049,1142,1033,1058]" italics="true" pageId="8" pageNumber="450">Atlantis</emphasis>
HOV
<emphasis id="B94C2706FF9A9C1DFB379543FAD5FBF3" box="[1222,1285,1033,1058]" italics="true" pageId="8" pageNumber="450">Alvin</emphasis>
</taxonomicName>
,
<geoCoordinate id="EE0C9DD3FF9A9C1DFAE49543FA4CFBF2" box="[1301,1436,1032,1059]" degrees="9" direction="north" minutes="50.333" orientation="latitude" pageId="8" pageNumber="450" precision="1" value="9.838883">9°50.333'N</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF9A9C1DFF66957AFE97FB9B" box="[151,327,1071,1098]" degrees="104" direction="west" minutes="17.482" orientation="longitude" pageId="8" pageNumber="450" precision="1" value="-104.29137">104°17.482'W</geoCoordinate>
(East Pacific Rise), Dive
<date id="FF86DDD4FF9A9C1DFD7C957AFCE5FB9B" box="[653,821,1072,1098]" pageId="8" pageNumber="450" value="2846-10-24">2846, 24 Oct.</date>
<quantity id="4CC056F1FF9A9C1DFCCF957AFC3CFB9B" box="[830,1004,1072,1098]" metricMagnitude="3" metricUnit="m" metricValue="1.9942512" pageId="8" pageNumber="450" unit="m" value="1994.2512">1994, 2512 m</quantity>
, male, coll
<collectionCode id="ED2963D1FF9A9C1DFB8F957AFB45FB9B" box="[1150,1173,1072,1098]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="450">R</collectionCode>
. Lutz, det JR Voight.
<collectionCode id="ED2963D1FF9A9C1DFF66951CFF3CFBA1" box="[151,236,1110,1136]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="450" type="Museum">FMNH</collectionCode>
287365,
<collectionCode id="ED2963D1FF9A9C1DFEAD951CFE53FBA1" box="[348,387,1110,1136]" country="Italy" name="Collection of Leptospira Strains" pageId="8" pageNumber="450">RV</collectionCode>
<taxonomicName id="4C388097FF9A9C1DFE7A951DFDA5FBA1" authority="HOV Alvin" authorityName="HOV Alvin" box="[395,629,1110,1136]" class="Insecta" family="Curculionidae" genus="Atlantis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="B94C2706FF9A9C1DFE7A951DFE37FBA1" box="[395,487,1111,1136]" italics="true" pageId="8" pageNumber="450">Atlantis</emphasis>
HOV
<emphasis id="B94C2706FF9A9C1DFDC6951DFDA5FBA1" box="[567,629,1111,1136]" italics="true" pageId="8" pageNumber="450">Alvin</emphasis>
</taxonomicName>
,
<geoCoordinate id="EE0C9DD3FF9A9C1DFD74951CFCDBFBA1" box="[645,779,1110,1136]" degrees="9" direction="north" minutes="49.136" orientation="latitude" pageId="8" pageNumber="450" precision="1" value="9.8189335">9°49.136'N</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF9A9C1DFCE5951CFC6EFBA1" box="[788,958,1110,1136]" degrees="104" direction="west" minutes="17.332" orientation="longitude" pageId="8" pageNumber="450" precision="1" value="-104.288864">104°17.332'W</geoCoordinate>
(East Pacific Rise), Dive 3217,
<date id="FF86DDD4FF9A9C1DFAB1951CFF07FB46" pageId="8" pageNumber="450" value="1998-05-17">
17 May
<quantity id="4CC056F1FF9A9C1DFF669537FF07FB46" box="[151,215,1149,1175]" metricMagnitude="3" metricUnit="m" metricValue="1.9982495" pageId="8" pageNumber="450" unit="m" value="1998.2495">1998</quantity>
</date>
, 2495 m, male, coll C. Fisher, det JR Voight.
<collectionCode id="ED2963D1FF9A9C1DFCE19537FCB6FB46" box="[784,870,1149,1175]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="450" type="Museum">FMNH</collectionCode>
287366,
<collectionCode id="ED2963D1FF9A9C1DFC2D9537FBD3FB46" box="[988,1027,1149,1175]" country="Italy" name="Collection of Leptospira Strains" pageId="8" pageNumber="450">RV</collectionCode>
<taxonomicName id="4C388097FF9A9C1DFBFD9537FB2FFB47" authority="HOV Alvin" authorityName="HOV Alvin" box="[1036,1279,1149,1175]" class="Insecta" family="Curculionidae" genus="Atlantis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="B94C2706FF9A9C1DFBFD9537FBBCFB47" box="[1036,1132,1149,1174]" italics="true" pageId="8" pageNumber="450">Atlantis</emphasis>
HOV
<emphasis id="B94C2706FF9A9C1DFB4E9537FB2FFB47" box="[1215,1279,1149,1174]" italics="true" pageId="8" pageNumber="450">Alvin</emphasis>
</taxonomicName>
,
<geoCoordinate id="EE0C9DD3FF9A9C1DFAE09537FA4CFB46" box="[1297,1436,1148,1175]" degrees="9" direction="north" minutes="50.624" orientation="latitude" pageId="8" pageNumber="450" precision="1" value="9.843734">9°50.624'N</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF9A9C1DFF6695EEFEE6FB6F" box="[151,310,1187,1214]" degrees="104" direction="west" minutes="17.48" orientation="longitude" pageId="8" pageNumber="450" precision="9" value="-104.291336">104°17.48'W</geoCoordinate>
(East Pacific Rise), Dive 3212,
<date id="FF86DDD4FF9A9C1DFD3995EEFCBEFB6F" box="[712,878,1188,1214]" pageId="8" pageNumber="450" value="1998-05-13">
13 May
<quantity id="4CC056F1FF9A9C1DFCDE95EEFCBEFB6F" box="[815,878,1188,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.9982509" pageId="8" pageNumber="450" unit="m" value="1998.2509">1998</quantity>
</date>
, 2509 m, male, coll C. Fisher, det JR Voight.
<collectionCode id="ED2963D1FF9A9C1DFF669580FF3DFB35" box="[151,237,1226,1252]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="450" type="Museum">FMNH</collectionCode>
307185,
<collectionCode id="ED2963D1FF9A9C1DFEAE9580FE56FB35" box="[351,390,1226,1252]" country="Italy" name="Collection of Leptospira Strains" pageId="8" pageNumber="450">RV</collectionCode>
<taxonomicName id="4C388097FF9A9C1DFE7F9581FDACFB35" authority="HOV Alvin" authorityName="HOV Alvin" box="[398,636,1226,1252]" class="Insecta" family="Curculionidae" genus="Atlantis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="B94C2706FF9A9C1DFE7F9581FE3BFB35" box="[398,491,1227,1252]" italics="true" pageId="8" pageNumber="450">Atlantis</emphasis>
HOV
<emphasis id="B94C2706FF9A9C1DFDCC9581FDACFB35" box="[573,636,1227,1252]" italics="true" pageId="8" pageNumber="450">Alvin</emphasis>
</taxonomicName>
,
<geoCoordinate id="EE0C9DD3FF9A9C1DFD7D9580FD3FFB35" box="[652,751,1226,1252]" degrees="12" direction="north" minutes="49" orientation="latitude" pageId="8" pageNumber="450" precision="925" value="12.816667">12°49'N</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF9A9C1DFD069580FCBFFB35" box="[759,879,1226,1252]" degrees="103" direction="west" minutes="56" orientation="longitude" pageId="8" pageNumber="450" precision="925" value="-103.933334">103°56'W</geoCoordinate>
(East Pacific Rise), Dive 3938,
<date id="FF86DDD4FF9A9C1DFB099580FF0AFADA" pageId="8" pageNumber="450" value="2003-11-19">
19 November
<quantity id="4CC056F1FF9A9C1DFF6695BBFF0AFADA" box="[151,218,1265,1291]" metricMagnitude="3" metricUnit="m" metricValue="2.0032619" pageId="8" pageNumber="450" unit="m" value="2003.2619">2003</quantity>
</date>
, 2619 m, male, coll &amp; det JR Voight.
<collectionCode id="ED2963D1FF9A9C1DFD2795BBFCFFFADA" box="[726,815,1265,1291]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="450" type="Museum">FMNH</collectionCode>
306500,
<collectionCode id="ED2963D1FF9A9C1DFC5F95BBFC06FADA" box="[942,982,1265,1291]" country="Italy" name="Collection of Leptospira Strains" pageId="8" pageNumber="450">RV</collectionCode>
<taxonomicName id="4C388097FF9A9C1DFC1095BBFB34FADB" authority="HOV Alvin" authorityName="HOV Alvin" box="[993,1252,1265,1291]" class="Insecta" family="Curculionidae" genus="Atlantis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="B94C2706FF9A9C1DFC1095BBFB97FADB" box="[993,1095,1265,1290]" italics="true" pageId="8" pageNumber="450">Atlantis</emphasis>
HOV
<emphasis id="B94C2706FF9A9C1DFB5195BBFB34FADB" box="[1184,1252,1265,1290]" italics="true" pageId="8" pageNumber="450">Alvin</emphasis>
</taxonomicName>
,
<geoCoordinate id="EE0C9DD3FF9A9C1DFB0995BBFA4CFADA" box="[1272,1436,1264,1291]" degrees="12" direction="north" minutes="48.673" orientation="latitude" pageId="8" pageNumber="450" precision="1" value="12.811216">12°48.673'N</geoCoordinate>
<geoCoordinate id="EE0C9DD3FF9A9C1DFF669452FE91FAE3" box="[151,321,1303,1330]" degrees="103" direction="west" minutes="56.408" orientation="longitude" pageId="8" pageNumber="450" precision="1" value="-103.94013">103°56.408'W</geoCoordinate>
(East Pacific Rise), Dive 3939,
<date id="FF86DDD4FF9A9C1DFD309452FC79FAE3" box="[705,937,1304,1330]" pageId="8" pageNumber="450" value="2003-11-20">
20 November
<quantity id="4CC056F1FF9A9C1DFC9A9452FC79FAE3" box="[875,937,1304,1330]" metricMagnitude="3" metricUnit="m" metricValue="2.003262" pageId="8" pageNumber="450" unit="m" value="2003.262">2003</quantity>
</date>
, 2620 m, male, coll A. Berry &amp; J. Voight, det JR Voight.
</paragraph>
<paragraph id="8B87FB14FF9A9C1DFF37942FFD4AF8F9" blockId="8.[151,1439,731,2026]" pageId="8" pageNumber="450">
<emphasis id="B94C2706FF9A9C1DFF37942FFE94FAAF" box="[198,324,1381,1406]" italics="true" pageId="8" pageNumber="450">Diagnosis.</emphasis>
<materialsCitation id="3B50F149FF9A9C1DFEBD942FFD45F8F9" collectionCode="ML" collectorName="Arm &amp; Arm I &amp; Funnel &amp; Gills &amp; Ligula &amp; Digestive &amp; Maximum &amp; Mature &amp; Rachidian &amp; Integument &amp; These" pageId="8" pageNumber="450" specimenCount="8" typeStatus="holotype">
Incirrate octopod. Biserial Suckers. Ink sac and anal flaps absent. Arms 2–4.5 times
<collectionCode id="ED2963D1FF9A9C1DFADA942FFA8BFAAE" box="[1323,1371,1381,1407]" country="France" name="Musee de Lectoure" pageId="8" pageNumber="450">ML</collectionCode>
.
<collectorName id="26CD9EC2FF9A9C1DFA97942FFA4CFAAE" box="[1382,1436,1381,1407]" pageId="8" pageNumber="450">Arm</collectorName>
formula 1&gt;2&gt;3&gt;4; arm length somewhat variable among
<specimenCount id="9D3E309DFF9A9C1DFC8094C6FBDCFA77" box="[881,1036,1420,1446]" pageId="8" pageNumber="450" type="generic">8 specimens</specimenCount>
with seemingly intact (or fully regenerated arms): arm I at least
<quantity id="4CC056F1FF9A9C1DFDEA94F8FDBEFA1D" box="[539,622,1458,1484]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="450" unit="mm" value="10.0">10 mm</quantity>
longer than II in 5, subequal in 3; arms II at least
<quantity id="4CC056F1FF9A9C1DFB4C94F8FADFFA1D" box="[1213,1295,1458,1484]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="450" unit="mm" value="10.0">10 mm</quantity>
longer than III in 4 of 8; arms III and IV subequal in 5 of 8.
<collectorName id="26CD9EC2FF9A9C1DFD0C9493FC97FA22" box="[765,839,1497,1523]" pageId="8" pageNumber="450">Arm I</collectorName>
with up to 166 suckers, other arms with fewer; hectocotylus carries between 58 and 83 suckers.
<collectorName id="26CD9EC2FF9A9C1DFD14974AFCE8F9CB" box="[741,824,1536,1562]" pageId="8" pageNumber="450">Funnel</collectorName>
organ W-shaped, lateral limbs shorter than medial limbs.
<collectorName id="26CD9EC2FF9A9C1DFF16976CFEF0F991" box="[231,288,1574,1600]" pageId="8" pageNumber="450">Gills</collectorName>
with 8 to 10 lamellae per outer demibranch.
<collectorName id="26CD9EC2FF9A9C1DFCC1976CFCACF991" box="[816,892,1574,1600]" pageId="8" pageNumber="450">Ligula</collectorName>
of males without strong transverse ridges LLI 8–10.
<collectorName id="26CD9EC2FF9A9C1DFF149707FE8BF9B6" box="[229,347,1613,1639]" pageId="8" pageNumber="450">Digestive</collectorName>
gland multilobed.
<collectorName id="26CD9EC2FF9A9C1DFDB89707FD17F9B6" box="[585,711,1613,1639]" pageId="8" pageNumber="450">Maximum</collectorName>
spermatophore length 62, maximum SpLI 125, maximum number of spermatophores 114.
<collectorName id="26CD9EC2FF9A9C1DFDE9973EFDBDF95F" box="[536,621,1652,1678]" pageId="8" pageNumber="450">Mature</collectorName>
males without enlarged suckers.
<collectorName id="26CD9EC2FF9A9C1DFC0B973EFBA2F95F" box="[1018,1138,1652,1678]" pageId="8" pageNumber="450">Rachidian</collectorName>
multicuspid.
<collectorName id="26CD9EC2FF9A9C1DFAE7973EFA4CF95F" box="[1302,1436,1652,1678]" pageId="8" pageNumber="450">Integument</collectorName>
of
<typeStatus id="548345B6FF9A9C1DFF4697D0FECEF965" box="[183,286,1690,1716]" pageId="8" pageNumber="450" type="holotype">holotype</typeStatus>
shows evidence of some sculpture in the form of widely spaced small papillae on the dorsal and ventral mantle, but most conspicuous are the low wrinkles that cover the animal, including the oral web.
<collectorName id="26CD9EC2FF9A9C1DFF6697A2FF32F8D3" box="[151,226,1768,1794]" pageId="8" pageNumber="450">These</collectorName>
are present in both fresh and preserved specimens, not as the description stated “wrinkled in preservation” (
<bibRefCitation id="EFA986E5FF9A9C1DFEB59644FD5DF8F9" author="Gonzalez" box="[324,653,1806,1832]" etAl="et al." firstAuthor="Gonzalez" page="173" pageId="8" pageNumber="450" pagination="169 - 184" refId="ref10672" refString="Gonzalez, A. F., Guerra, A., Pascual, S. &amp; Briand, P. (1998) Vulcanoctopus hydrothermalis gen. et sp. nov. (Mollusca, Cephalopoda): an octopod from a deep-sea hydrothermal vent site. Cahiers de Biologie Marine, 39, 169 - 184." type="journal article" year="1998">
González
<emphasis id="B94C2706FF9A9C1DFE4D9645FE26F8F9" box="[444,502,1807,1832]" italics="true" pageId="8" pageNumber="450">et al.</emphasis>
1998 p. 173
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B87FB14FF9A9C1CFF37967FFE89FEF7" blockId="8.[151,1439,731,2026]" lastBlockId="9.[151,1436,152,294]" lastPageId="9" lastPageNumber="451" pageId="8" pageNumber="450">
<emphasis id="B94C2706FF9A9C1DFF37967FFEE5F89F" box="[198,309,1845,1870]" italics="true" pageId="8" pageNumber="450">Remarks.</emphasis>
A female of this species was recently described (
<bibRefCitation id="EFA986E5FF9A9C1DFC6F967FFB4FF89E" author="Gonzalez" box="[926,1183,1845,1871]" etAl="et al." firstAuthor="Gonzalez" pageId="8" pageNumber="450" pagination="375 - 379" refId="ref10779" refString="Gonzalez, A. F., Guerra, A., Pascual, S. &amp; Segonzac, M. (2008) Female description of the hydrothermal vent cephalopod Vulcanoctopus hydrothermalis. Journal of the Marine Biological Association of the United Kingdom, 88, 375 - 379." type="journal article" year="2008">
González
<emphasis id="B94C2706FF9A9C1DFBE8967FFB88F89F" box="[1049,1112,1845,1870]" italics="true" pageId="8" pageNumber="450">et al.</emphasis>
2008
</bibRefCitation>
) as lacking the dark swelling described in the intestine of males (
<bibRefCitation id="EFA986E5FF9A9C1DFD539616FC49F8A7" author="Gonzalez" box="[674,921,1884,1910]" etAl="et al." firstAuthor="Gonzalez" pageId="8" pageNumber="450" pagination="169 - 184" refId="ref10672" refString="Gonzalez, A. F., Guerra, A., Pascual, S. &amp; Briand, P. (1998) Vulcanoctopus hydrothermalis gen. et sp. nov. (Mollusca, Cephalopoda): an octopod from a deep-sea hydrothermal vent site. Cahiers de Biologie Marine, 39, 169 - 184." type="journal article" year="1998">
González
<emphasis id="B94C2706FF9A9C1DFCE99616FC82F8A4" box="[792,850,1884,1909]" italics="true" pageId="8" pageNumber="450">et al.</emphasis>
1998
</bibRefCitation>
). Neither has the dark swelling been seen in 13 additional specimens, 8 from the type locality, in Field Museum collections. The condition of the optic chiasma is nearly unexplored in other deep-sea octopodids. Information on spermatophores given in diagnoses was extracted from
<bibRefCitation id="EFA986E5FF9A9C1DFE08969AFCD1F83B" author="Gonzalez" box="[505,769,2000,2026]" etAl="et al." firstAuthor="Gonzalez" pageId="8" pageNumber="450" pagination="289 - 298" refId="ref10731" refString="Gonzalez, A. F., Guerra, A., Rocha, F. &amp; Briand, P. (2002) Morphological variation in males of Vulcanoctopus hydrothermalis (Mollusca, Cephalopoda). Bulletin of Marine Science, 71, 289 - 298." type="journal article" year="2002">
González
<emphasis id="B94C2706FF9A9C1DFD9E969AFD79F838" box="[623,681,2000,2025]" italics="true" pageId="8" pageNumber="450">et al.</emphasis>
(2002)
</bibRefCitation>
, which details variation in male specimens from the type locality. The
<typeStatus id="548345B6FF9B9C1CFEC291D2FE4AFF63" box="[307,410,152,178]" pageId="9" pageNumber="451" type="holotype">holotype</typeStatus>
has been heavily dissected; the reproductive tract, for example, is in eight parts. Only 69 spermatophores remain in the
<typeStatus id="548345B6FF9B9C1CFDC091F4FD4FFF09" box="[561,671,190,216]" pageId="9" pageNumber="451" type="holotype">holotype</typeStatus>
; this is similar to the next highest counts we found in
<specimenCount id="9D3E309DFF9B9C1CFACB91F4FA4BFF09" box="[1338,1435,190,216]" pageId="9" pageNumber="451" type="male">30 male</specimenCount>
specimens (70, 72, 74). The funnel organ shape is herein described for the first time based on specimens from the type locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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