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<document id="E3193D63EB4399A6EA498CF301BCA35F" ID-DOI="10.5281/zenodo.170669" ID-GBIF-Dataset="cc432084-63d9-4ab2-a3aa-8e2efc7b740a" ID-ISSN="1175­5326" ID-Zenodo-Dep="170669" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459714546813" checkinUser="plazi" docAuthor="Braby, Michael F." docDate="2005" docId="03813712D22BFF9DD062FB03FDC4FE0D" docLanguage="en" docName="zt00832.pdf" docOrigin="Zootaxa 832" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pierinae" docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="FFB84F6AD222FF97D16AFF85FFEFFF8B" masterDocTitle="Provisional checklist of genera of the Pieridae (Lepidoptera: Papilionoidea)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="10" updateTime="1698207631183" updateUser="plazi">
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<mods:title id="1531AB3E30EA5F7A1CBA67DB9AA40482">Provisional checklist of genera of the Pieridae (Lepidoptera: Papilionoidea)</mods:title>
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<treatment id="03813712D22BFF9DD062FB03FDC4FE0D" ID-DOI="http://doi.org/10.5281/zenodo.5688532" ID-GBIF-Taxon="119334307" ID-Zenodo-Dep="5688532" LSID="urn:lsid:plazi:treatment:03813712D22BFF9DD062FB03FDC4FE0D" httpUri="http://treatment.plazi.org/id/03813712D22BFF9DD062FB03FDC4FE0D" lastPageId="10" lastPageNumber="10" pageId="9" pageNumber="10">
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<paragraph id="8B978604D22BFF9ED062FB03FDC9FB2B" blockId="9.[264,1324,1158,1899]" box="[264,550,1158,1184]" pageId="9" pageNumber="10">
<heading id="D0DF3168D22BFF9ED062FB03FDC9FB2B" bold="true" box="[264,550,1158,1184]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B95C5A16D22BFF9ED062FB03FDC9FB2B" bold="true" box="[264,550,1158,1184]" pageId="9" pageNumber="10">
<taxonomicName id="4C28FD87D22BFF9ED062FB03FE9AFB2B" box="[264,373,1158,1184]" class="Insecta" family="Pieridae" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subFamily" subFamily="Pierinae">Pierinae</taxonomicName>
:
<taxonomicName id="4C28FD87D22BFF9ED0EBFB03FE3BFB2B" box="[385,468,1158,1184]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED0EBFB03FE3BFB2B" bold="true" box="[385,468,1158,1184]" italics="true" pageId="9" pageNumber="10">Appias</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="8B978604D22BFF9ED062FB2CFD48F978" blockId="9.[264,1324,1158,1899]" pageId="9" pageNumber="10">
<bibRefCitation id="EFB9FBF5D22BFF9ED062FB2CFE76FB48" author="Yata" box="[264,409,1193,1219]" pageId="9" pageNumber="16" refString="Yata, O. (1985) Part 1: Pieridae. In: Tsukada, E. (Ed), Butterflies of the South East Asian islands. II: Pieridae, Danaidae. Plapac, Tokyo, pp. 33 - 120, 205 - 438." type="book chapter" year="1985">Yata (1985)</bibRefCitation>
, Vane­Wright and de
<bibRefCitation id="EFB9FBF5D22BFF9ED3C3FB2CFCD5FB48" author="Jong" box="[681,826,1193,1219]" pageId="9" pageNumber="16" refString="Vane-Wright, R. I. &amp; de Jong, R. (2003) The butterflies of Sulawesi: annotated checklist for a critical island fauna. Zoologische Verhandelingen, Leiden (343), 3 - 267." type="journal article" year="2003">Jong (2003)</bibRefCitation>
and Yata
<emphasis id="B95C5A16D22BFF9ED2D7FB2CFC1AFB49" box="[957,1013,1193,1218]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. (unpublished data) have revised the higher systematics of
<taxonomicName id="4C28FD87D22BFF9ED3F4FB54FCBDFB60" authority="Hubner" authorityName="Hubner" box="[670,850,1233,1259]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED3F4FB54FD1FFB61" box="[670,752,1233,1258]" italics="true" pageId="9" pageNumber="10">Appias</emphasis>
Hübner
</taxonomicName>
from the Oriental Region.
<bibRefCitation id="EFB9FBF5D22BFF9ED5F5FB54FAC3FB60" author="Yata" box="[1183,1324,1233,1259]" pageId="9" pageNumber="16" refString="Yata, O. (1985) Part 1: Pieridae. In: Tsukada, E. (Ed), Butterflies of the South East Asian islands. II: Pieridae, Danaidae. Plapac, Tokyo, pp. 33 - 120, 205 - 438." type="book chapter" year="1985">Yata (1985)</bibRefCitation>
provisionally recognised seven groups within
<taxonomicName id="4C28FD87D22BFF9ED242FB7CFC96FA99" box="[808,889,1273,1298]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED242FB7CFC96FA99" box="[808,889,1273,1298]" italics="true" pageId="9" pageNumber="10">Appias</emphasis>
</taxonomicName>
, five of which were awarded subgeneric status and two without formal names (the
<taxonomicName id="4C28FD87D22BFF9ED22AFAA4FC6AFAB1" box="[832,901,1313,1338]" class="Insecta" family="Pieridae" genus="Glutophrissa" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sylvia">
<emphasis id="B95C5A16D22BFF9ED22AFAA4FC6AFAB1" box="[832,901,1313,1338]" italics="true" pageId="9" pageNumber="10">sylvia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C28FD87D22BFF9ED2D5FAA4FBFAFAB1" box="[959,1045,1313,1338]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lyncida">
<emphasis id="B95C5A16D22BFF9ED2D5FAA4FBFAFAB1" box="[959,1045,1313,1338]" italics="true" pageId="9" pageNumber="10">lyncida</emphasis>
</taxonomicName>
groups). The subgenus
<taxonomicName id="4C28FD87D22BFF9ED062FACCFE1EFAE8" authority="Butler" authorityName="Butler" box="[264,497,1353,1379]" class="Insecta" family="Pieridae" genus="Glutophrissa" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED062FACCFE4EFAE9" box="[264,417,1353,1378]" italics="true" pageId="9" pageNumber="10">Glutophrissa</emphasis>
Butler
</taxonomicName>
was restricted to include two Neotropical species (
<taxonomicName id="4C28FD87D22BFF9ED52CFACCFB4EFAE9" box="[1094,1185,1353,1378]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="drusilla">
<emphasis id="B95C5A16D22BFF9ED52CFACCFB4EFAE9" box="[1094,1185,1353,1378]" italics="true" pageId="9" pageNumber="10">drusilla</emphasis>
</taxonomicName>
group), but
<bibRefCitation id="EFB9FBF5D22BFF9ED062FAF4FE17FA00" box="[264,504,1393,1419]" pageId="9" pageNumber="14" refString="Ackery, P. R., Smith, C. R. &amp; Vane-Wright, R. I. (1995) Carcasson's African Butterflies. An Annotated Catalogue of the Papilionoidea and Hesperioidea of the Afrotropical Region. CSIRO Publishing, Melbourne, xi + 803 pp." type="book">
Ackery
<emphasis id="B95C5A16D22BFF9ED00CFAF4FE74FA01" box="[358,411,1393,1418]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. (1995)
</bibRefCitation>
expanded the concept of
<taxonomicName id="4C28FD87D22BFF9ED242FAF4FBC0FA01" authority="(Glutophrissa)" baseAuthorityName="Glutophrissa" box="[808,1071,1393,1418]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED242FAF4FBC0FA01" box="[808,1071,1393,1418]" italics="true" pageId="9" pageNumber="10">Appias (Glutophrissa)</emphasis>
</taxonomicName>
to include five Afrotropical species, formerly treated by
<bibRefCitation id="EFB9FBF5D22BFF9ED3AEFA1CFCBEFA38" author="Yata" box="[708,849,1433,1459]" pageId="9" pageNumber="16" refString="Yata, O. (1985) Part 1: Pieridae. In: Tsukada, E. (Ed), Butterflies of the South East Asian islands. II: Pieridae, Danaidae. Plapac, Tokyo, pp. 33 - 120, 205 - 438." type="book chapter" year="1985">Yata (1985)</bibRefCitation>
in the
<taxonomicName id="4C28FD87D22BFF9ED2C3FA1CFC01FA39" box="[937,1006,1433,1458]" class="Insecta" family="Pieridae" genus="Glutophrissa" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sylvia">
<emphasis id="B95C5A16D22BFF9ED2C3FA1CFC01FA39" box="[937,1006,1433,1458]" italics="true" pageId="9" pageNumber="10">sylvia</emphasis>
</taxonomicName>
group. The status of
<taxonomicName id="4C28FD87D22BFF9ED59FFA1CFE9AFA51" class="Insecta" family="Pieridae" genus="Glutophrissa" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED59FFA1CFE9AFA51" italics="true" pageId="9" pageNumber="10">Glutophrissa</emphasis>
</taxonomicName>
, however, is currently uncertain.
<bibRefCitation id="EFB9FBF5D22BFF9ED391FA44FC71FA50" author="Lamas" box="[763,926,1473,1499]" pageId="9" pageNumber="15" refString="Lamas, G. (2005) Pieridae. In: Lamas, G. (Ed), Checklist: Part 4 A. Hesperioidea-Papilionoidea. In: Heppner, J. B. (Ed), Atlas of Neotropical Lepidoptera. Volume 5 A. Association for Tropical Lepidoptera / Scientific Publishers, Gainesville (in press)." type="book" year="2005">Lamas (2005)</bibRefCitation>
provisionally treated it as distinct genus restricted to the
<taxonomicName id="4C28FD87D22BFF9ED37CFA6CFD9EF989" box="[534,625,1513,1538]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="drusilla">
<emphasis id="B95C5A16D22BFF9ED37CFA6CFD9EF989" box="[534,625,1513,1538]" italics="true" pageId="9" pageNumber="10">drusilla</emphasis>
</taxonomicName>
group, and
<collectingCountry id="F33FC694D22BFF9ED26BFA6CFCF6F988" box="[769,793,1513,1539]" name="Iceland" pageId="9" pageNumber="10">O</collectingCountry>
. Yata (pers. comm. 2004) has suggested that phylogenetically it may be outside
<taxonomicName id="4C28FD87D22BFF9ED3C6F994FD12F9A1" box="[684,765,1553,1578]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED3C6F994FD12F9A1" box="[684,765,1553,1578]" italics="true" pageId="9" pageNumber="10">Appias</emphasis>
</taxonomicName>
and
<taxonomicName id="4C28FD87D22BFF9ED251F994FC11F9A0" authority="Distant" authorityName="Distant" box="[827,1022,1553,1579]" class="Insecta" family="Pieridae" genus="Saletara" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED251F994FC71F9A1" box="[827,926,1553,1578]" italics="true" pageId="9" pageNumber="10">Saletara</emphasis>
Distant
</taxonomicName>
, both of which appear to comprise a monophyletic group. Vane­Wright and de
<bibRefCitation id="EFB9FBF5D22BFF9ED2FAF9BCFBF0F9D8" author="Jong" box="[912,1055,1593,1619]" pageId="9" pageNumber="16" refString="Vane-Wright, R. I. &amp; de Jong, R. (2003) The butterflies of Sulawesi: annotated checklist for a critical island fauna. Zoologische Verhandelingen, Leiden (343), 3 - 267." type="journal article" year="2003">Jong (2003)</bibRefCitation>
placed Yatas
<taxonomicName id="4C28FD87D22BFF9ED5BEF9BCFAC5F9D9" box="[1236,1322,1593,1618]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lyncida">
<emphasis id="B95C5A16D22BFF9ED5BEF9BCFAC5F9D9" box="[1236,1322,1593,1618]" italics="true" pageId="9" pageNumber="10">lyncida</emphasis>
</taxonomicName>
group (five species) within the nominate subspecies
<taxonomicName id="4C28FD87D22BFF9ED2E2F9E4FBA5F9F1" box="[904,1098,1633,1658]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED2E2F9E4FBA5F9F1" box="[904,1098,1633,1658]" italics="true" pageId="9" pageNumber="10">Appias (Appias)</emphasis>
</taxonomicName>
, which previously included only two Oriental species. In the Oriental Region, four subgenera are thus currently recognised (
<taxonomicName id="4C28FD87D22BFF9ED08DF934FDD7F941" box="[487,568,1713,1738]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED08DF934FDD7F941" box="[487,568,1713,1738]" italics="true" pageId="9" pageNumber="10">Appias</emphasis>
</taxonomicName>
,
<taxonomicName id="4C28FD87D22BFF9ED323F934FCFEF940" authority="Butler" authorityName="Butler" box="[585,785,1713,1739]" class="Insecta" family="Pieridae" genus="Phrissura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED323F934FD52F941" box="[585,701,1713,1738]" italics="true" pageId="9" pageNumber="10">Phrissura</emphasis>
Butler
</taxonomicName>
,
<taxonomicName id="4C28FD87D22BFF9ED274F934FC1FF940" authority="Geyer" authorityName="Geyer" box="[798,1008,1713,1739]" class="Insecta" family="Pieridae" genus="Hiposcritia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED274F934FC4BF941" box="[798,932,1713,1738]" italics="true" pageId="9" pageNumber="10">Hiposcritia</emphasis>
Geyer
</taxonomicName>
,
<taxonomicName id="4C28FD87D22BFF9ED56BF934FB05F940" authority="Hubner" authorityName="Hubner" box="[1025,1258,1713,1739]" class="Insecta" family="Pieridae" genus="Catophaga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED56BF934FB6CF941" box="[1025,1155,1713,1738]" italics="true" pageId="9" pageNumber="10">Catophaga</emphasis>
Hübner
</taxonomicName>
), two of which are endemic to the region.
</paragraph>
</subSubSection>
<subSubSection id="C332D58FD22BFF9DD052F884FDC4FE0D" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="8B978604D22BFF9DD052F884FDC4FE0D" blockId="9.[264,1324,1158,1899]" lastBlockId="10.[264,1324,284,390]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<bibRefCitation id="EFB9FBF5D22BFF9ED052F884FE20F890" author="Klots" box="[312,463,1793,1819]" pageId="9" pageNumber="15" refString="Klots, A. B. (1933) A generic classification of the Pieridae (Lepidoptera) together with a study of the male genitalia. Entomologica america, 12, 139 - 242." type="journal article" year="1933">Klots (1933)</bibRefCitation>
treated
<taxonomicName id="4C28FD87D22BFF9ED35BF884FD19F890" authority="Distant" authorityName="Distant" box="[561,758,1793,1819]" class="Insecta" family="Pieridae" genus="Udaiana" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED35BF884FD78F891" box="[561,663,1793,1818]" italics="true" pageId="9" pageNumber="10">Udaiana</emphasis>
Distant
</taxonomicName>
as a distinct monotypic genus closely allied to
<taxonomicName id="4C28FD87D22BFF9ED062F8ACFEB6F8C9" box="[264,345,1833,1858]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED062F8ACFEB6F8C9" box="[264,345,1833,1858]" italics="true" pageId="9" pageNumber="10">Appias</emphasis>
</taxonomicName>
, and regarded
<taxonomicName id="4C28FD87D22BFF9ED366F8ACFD90F8C9" box="[524,639,1833,1858]" class="Insecta" family="Pieridae" genus="Phrissura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED366F8ACFD90F8C9" box="[524,639,1833,1858]" italics="true" pageId="9" pageNumber="10">Phrissura</emphasis>
</taxonomicName>
, also monotypic, as a subgenus of
<taxonomicName id="4C28FD87D22BFF9ED549F8ACFB9BF8C9" box="[1059,1140,1833,1858]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D22BFF9ED549F8ACFB9BF8C9" box="[1059,1140,1833,1858]" italics="true" pageId="9" pageNumber="10">Appias</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFB9FBF5D22BFF9ED5EEF8ACFAC5F8C8" author="Corbet" box="[1156,1322,1833,1859]" pageId="9" pageNumber="14" refString="Corbet, A. S. (1946) The conspecificity of Phrissura aegis (C. &amp; R. Feld) and Udaiana cynis (Hew.) (Lepidoptera: Pieridae). The Entomologist, 79, 145 - 146." type="journal article" year="1946">Corbet (1946)</bibRefCitation>
synonymised
<taxonomicName id="4C28FD87D22BFF9ED0C6F8D4FD0AF8E0" authority="Hewitson" authorityName="Hewitson" box="[428,741,1873,1899]" class="Insecta" family="Pieridae" genus="Udaiana" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cynis">
<emphasis id="B95C5A16D22BFF9ED0C6F8D4FDB7F8E1" box="[428,600,1873,1898]" italics="true" pageId="9" pageNumber="10">Udaiana cynis</emphasis>
(Hewitson)
</taxonomicName>
with
<taxonomicName id="4C28FD87D22BFF9ED246F8D4FC08F8E1" box="[812,999,1873,1898]" class="Insecta" family="Pieridae" genus="Phrissura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aegis">
<emphasis id="B95C5A16D22BFF9ED246F8D4FC08F8E1" box="[812,999,1873,1898]" italics="true" pageId="9" pageNumber="10">Phrissura aegis</emphasis>
</taxonomicName>
(C. &amp; R. Felder), but
<bibRefCitation id="EFB9FBF5D22BFF9DD59DF8D4FEB8FEBD" author="Yata" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="16" refString="Yata, O. (1985) Part 1: Pieridae. In: Tsukada, E. (Ed), Butterflies of the South East Asian islands. II: Pieridae, Danaidae. Plapac, Tokyo, pp. 33 - 120, 205 - 438." type="book chapter" year="1985">Yata (1985)</bibRefCitation>
considered the two taxa to be specifically distinct and to belong to separate genera,
<taxonomicName id="4C28FD87D228FF9DD062FEC1FE80FED6" box="[264,367,324,349]" class="Insecta" family="Pieridae" genus="Udaiana" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D228FF9DD062FEC1FE80FED6" box="[264,367,324,349]" italics="true" pageId="10" pageNumber="11">Udaiana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C28FD87D228FF9DD0CDFEC1FD68FED6" authority="(Phrissura)" baseAuthorityName="Phrissura" box="[423,647,324,349]" class="Insecta" family="Pieridae" genus="Appias" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B95C5A16D228FF9DD0CDFEC1FD68FED6" box="[423,647,324,349]" italics="true" pageId="10" pageNumber="11">Appias (Phrissura)</emphasis>
</taxonomicName>
, respectively, based on study of the male and female genitalia and wing venation.
</paragraph>
</subSubSection>
</treatment>
</document>