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<mods:title id="6B48E2E37F25B0F7077C513A38C9AB11">Description of two new species of scorpions from China (Tibet) belonging to the genera Mesobuthus Vachon (Buthidae) and Heterometrus Ehrenberg (Scorpionidae)</mods:title>
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<mods:namePart id="25A33B804141B5260291A696722EB346">Lourenço, Wilson R.</mods:namePart>
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<mods:namePart id="208CD2D3D0DB04D027D76E53127427E5">Qi, Jian-Xin</mods:namePart>
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<mods:date id="550CEE4B8A0B2CA3857A3C0C356766FB">2005</mods:date>
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<paragraph id="8BC1DA58FF95FFD9FEA8FAFB0D81FA93" blockId="1.[264,673,1313,1339]" box="[264,673,1313,1339]" pageId="1" pageNumber="2">
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Genus
<taxonomicName id="4C7EA1DBFF95FFD9FEFFFAFB0D81FA93" ID-CoL="5PMS" authority="Vachon, 1950" authorityName="Vachon" authorityYear="1950" box="[351,673,1313,1339]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FEFFFAFB0ED3FA93" bold="true" box="[351,499,1313,1339]" italics="true" pageId="1" pageNumber="2">Mesobuthus</emphasis>
<bibRefCitation id="EFEFA7A9FF95FFD9FE5BFAFB0D81FA93" author="Vachon" box="[507,673,1313,1339]" pageId="1" pageNumber="16" refString="Vachon, M. (1950) Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 28, 152 - 216." type="journal article" year="1950">Vachon, 1950</bibRefCitation>
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<emphasis id="B90A064AFF95FFD9FEA8FAB60EC3FA2D" box="[264,483,1388,1413]" italics="true" pageId="1" pageNumber="2">Historical account</emphasis>
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<paragraph id="8BC1DA58FF95FFDAFE98FA4E0ED0FE06" blockId="1.[264,1323,1388,1894]" lastBlockId="2.[264,1323,284,1150]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
In the early 1940s,
<bibRefCitation id="EFEFA7A9FF95FFD9FDBCFA4E0DEAFA06" author="Vachon" box="[540,714,1428,1454]" pageId="1" pageNumber="16" refString="Vachon, M. (1952) Etudes sur les scorpions. Publications de l'Institut Pasteur d'Algerie, Alger, 482 pp." type="book" year="1952">Vachon (1952)</bibRefCitation>
began a series of studies on the scorpions of North of Africa. One of his main preoccupations was with the definition of the various groups of species within the genus
<taxonomicName id="4C7EA1DBFF95FFD9FD8BFA3E0C2DFA56" authority="Leach. This" authorityName="Leach. This" box="[555,781,1508,1534]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FD8BFA3E0D5CFA55" box="[555,636,1508,1533]" italics="true" pageId="1" pageNumber="2">Buthus</emphasis>
Leach. This
</taxonomicName>
led to the subdivision of what was at that time the genus
<taxonomicName id="4C7EA1DBFF95FFD9FE24F9D60EF6F98D" box="[388,470,1548,1573]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FE24F9D60EF6F98D" box="[388,470,1548,1573]" italics="true" pageId="1" pageNumber="2">Buthus</emphasis>
</taxonomicName>
into about 10 separate genera. Two of the genera proposed by Vachon were
<taxonomicName id="4C7EA1DBFF95FFD9FEECF9EE0D46F9E6" authority="Vachon, 1949" authorityName="Vachon" authorityYear="1949" box="[332,614,1588,1614]" class="Arachnida" family="Buthidae" genus="Buthotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FEECF9EE0E94F9E5" box="[332,436,1588,1613]" italics="true" pageId="1" pageNumber="2">Buthotus</emphasis>
<bibRefCitation id="EFEFA7A9FF95FFD9FE60F9EE0D46F9E6" author="Vachon" box="[448,614,1588,1614]" pageId="1" pageNumber="16" refString="Vachon, M. (1949) Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 27, 134 - 169." type="journal article" year="1949">Vachon, 1949</bibRefCitation>
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and
<taxonomicName id="4C7EA1DBFF95FFD9FD07F9EE0CC9F9E6" authority="Vachon, 1950" authorityName="Vachon" authorityYear="1950" box="[679,1001,1588,1614]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FD07F9EE0C15F9E5" box="[679,821,1588,1613]" italics="true" pageId="1" pageNumber="2">Mesobuthus</emphasis>
<bibRefCitation id="EFEFA7A9FF95FFD9FCE0F9EE0CC9F9E6" author="Vachon" box="[832,1001,1588,1614]" pageId="1" pageNumber="16" refString="Vachon, M. (1950) Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 28, 152 - 216." type="journal article" year="1950">Vachon, 1950</bibRefCitation>
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. The first,
<taxonomicName id="4C7EA1DBFF95FFD9FBD3F9EE0BFBF9E5" box="[1139,1243,1588,1613]" class="Arachnida" family="Buthidae" genus="Buthotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FBD3F9EE0BFBF9E5" box="[1139,1243,1588,1613]" italics="true" pageId="1" pageNumber="2">Buthotus</emphasis>
</taxonomicName>
, comprised the majority of species that had been included in the old subgenus
<taxonomicName id="4C7EA1DBFF95FFD9FBCDF9860E5AF936" authority="Birula, 1908" authorityName="Birula" authorityYear="1908" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FBCDF9860BD2F9DD" box="[1133,1266,1628,1653]" italics="true" pageId="1" pageNumber="2">Hottentotta</emphasis>
<bibRefCitation id="EFEFA7A9FF95FFD9FB5BF9860E5AF936" author="Birula" pageId="1" pageNumber="14" refString="Birula, A. A. (1908) Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werners nach dem Anglo-Aegyptischen Sudan und Nord- Uganda. XIV. Skorpiones und Solifugae. Sitzungsberichte der kaiserlich-koniglichen Akademie der Wissenschaften, Wien, 117, 121 - 152." type="journal article" year="1908">Birula, 1908</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFEFA7A9FF95FFD9FE26F95E0D71F936" author="Kraepelin" box="[390,593,1668,1694]" pageId="1" pageNumber="15" refString="Kraepelin, K. (1891) Revision der Skorpione. I. Die Familie der Androctonidae. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 8, 1 - 144." type="journal article" year="1891">Kraepelin (1891)</bibRefCitation>
was in fact the first author to distinguish a
<taxonomicName id="4C7EA1DBFF95FFD9FBF8F95E0BFEF935" box="[1112,1246,1668,1693]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FBF8F95E0BFEF935" box="[1112,1246,1668,1693]" italics="true" pageId="1" pageNumber="2">Hottentotta</emphasis>
</taxonomicName>
group (species­group) within the genus
<taxonomicName id="4C7EA1DBFF95FFD9FD06F9760DD7F96D" box="[678,759,1708,1733]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FD06F9760DD7F96D" box="[678,759,1708,1733]" italics="true" pageId="1" pageNumber="2">Buthus</emphasis>
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. Most of the species included were allied to
<taxonomicName id="4C7EA1DBFF95FFD9FEA8F90E0E79F945" box="[264,345,1748,1773]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FEA8F90E0E79F945" box="[264,345,1748,1773]" italics="true" pageId="1" pageNumber="2">Buthus</emphasis>
</taxonomicName>
<taxonomicName id="4C7EA1DBFF95FFD9FEC3F90E0D57F946" authority="(Fabricius)" authorityName="Fabricius" baseAuthorityName="Fabricius" box="[355,631,1748,1774]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FEC3F90E0EC9F945" box="[355,489,1748,1773]" italics="true" pageId="1" pageNumber="2">Hottentotta</emphasis>
(Fabricius)
</taxonomicName>
.
<bibRefCitation id="EFEFA7A9FF95FFD9FD25F90E0C06F946" author="Birula" box="[645,806,1748,1774]" pageId="1" pageNumber="14" refString="Birula, A. A. (1908) Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werners nach dem Anglo-Aegyptischen Sudan und Nord- Uganda. XIV. Skorpiones und Solifugae. Sitzungsberichte der kaiserlich-koniglichen Akademie der Wissenschaften, Wien, 117, 121 - 152." type="journal article" year="1908">Birula (1908)</bibRefCitation>
created the subgenus
<taxonomicName id="4C7EA1DBFF95FFD9FB93F90E0B98F945" box="[1075,1208,1748,1773]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FB93F90E0B98F945" box="[1075,1208,1748,1773]" italics="true" pageId="1" pageNumber="2">Hottentotta</emphasis>
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, without, however, a clear diagnosis.
<bibRefCitation id="EFEFA7A9FF95FFD9FDEFF9260DDCF8BE" author="Vachon" box="[591,764,1788,1814]" pageId="1" pageNumber="16" refString="Vachon, M. (1949) Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 27, 134 - 169." type="journal article" year="1949">Vachon (1949)</bibRefCitation>
disregarded
<taxonomicName id="4C7EA1DBFF95FFD9FC37F9260B4DF8BE" authority="Birula" authorityName="Birula" box="[919,1133,1788,1814]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FC37F9260B3DF8BD" box="[919,1053,1788,1813]" italics="true" pageId="1" pageNumber="2">Hottentotta</emphasis>
Birula
</taxonomicName>
and established a new name,
<taxonomicName id="4C7EA1DBFF95FFD9FE06F8FE0D2EF895" box="[422,526,1828,1853]" class="Arachnida" family="Buthidae" genus="Buthotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FE06F8FE0D2EF895" box="[422,526,1828,1853]" italics="true" pageId="1" pageNumber="2">Buthotus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C7EA1DBFF95FFD9FDBDF8FE0D83F895" box="[541,675,1828,1853]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FDBDF8FE0D83F895" box="[541,675,1828,1853]" italics="true" pageId="1" pageNumber="2">Hottentotta</emphasis>
</taxonomicName>
is, however, a valid senior synonym for
<taxonomicName id="4C7EA1DBFF95FFD9FB28F8FE0BD0F895" box="[1160,1264,1828,1853]" class="Arachnida" family="Buthidae" genus="Buthotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF95FFD9FB28F8FE0BD0F895" box="[1160,1264,1828,1853]" italics="true" pageId="1" pageNumber="2">Buthotus</emphasis>
</taxonomicName>
, and was re­established by
<bibRefCitation id="EFEFA7A9FF95FFD9FDAAF8960DE1F8CE" author="Francke" box="[522,705,1868,1894]" pageId="1" pageNumber="15" refString="Francke, O. F. (1985) Conspectus genericus scorpionorum 1758 - 1982 (Arachnida: Scorpiones). Occasional Papers of the Museum, Texas Tech University, 98, 1 - 32." type="journal article" year="1985">Francke (1985)</bibRefCitation>
. Other valid subgenera in addition to the nominotypical
<taxonomicName id="4C7EA1DBFF96FFDAFE9DFEC60EE2FE9D" box="[317,450,284,309]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFE9DFEC60EE2FE9D" box="[317,450,284,309]" italics="true" pageId="2" pageNumber="3">Hottentotta</emphasis>
</taxonomicName>
are
<taxonomicName id="4C7EA1DBFF96FFDAFE5BFEC60DB4FE9D" box="[507,660,284,309]" class="Arachnida" family="Buthidae" genus="Balfourianus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFE5BFEC60DB4FE9D" box="[507,660,284,309]" italics="true" pageId="2" pageNumber="3">Balfourianus</emphasis>
</taxonomicName>
which was described by
<bibRefCitation id="EFEFA7A9FF96FFDAFC6AFEC60B5AFE9E" author="Vachon" box="[970,1146,284,310]" pageId="2" pageNumber="16" refString="Vachon, M. (1979) Notes on the types of scorpions in the British Museum (Natural History), London. Buthus socotrensis Pocock, 1889 (Family: Buthidae). Bulletin of the British Museum of Natural History (Zoology), 36, 233 - 237." type="journal article" year="1979">Vachon (1979)</bibRefCitation>
to include the species
<taxonomicName id="4C7EA1DBFF96FFDAFEC7FE9E0DCDFEF6" authority="Pocock" authorityName="Pocock" box="[359,749,324,350]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="socotrensis">
<emphasis id="B90A064AFF96FFDAFEC7FE9E0D59FEF5" box="[359,633,324,349]" italics="true" pageId="2" pageNumber="3">Hottentotta socotrensis</emphasis>
(Pocock)
</taxonomicName>
which is endemic to the Island of
<collectingCountry id="F3699AC8FF96FFDAFB36FE9E0BD2FEF6" box="[1174,1266,324,350]" name="Yemen" pageId="2" pageNumber="3">Socotra</collectingCountry>
, and
<taxonomicName id="4C7EA1DBFF96FFDAFEA8FEB60E9FFE2D" box="[264,447,364,389]" class="Arachnida" family="Buthidae" genus="Deccanobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFEA8FEB60E9FFE2D" box="[264,447,364,389]" italics="true" pageId="2" pageNumber="3">Deccanobuthus</emphasis>
</taxonomicName>
described by
<bibRefCitation id="EFEFA7A9FF96FFDAFDC3FEB60C3BFE2E" author="Loureno" box="[611,795,364,390]" pageId="2" pageNumber="15" refString="Lourenco, W. R. (2000) The genus Hottentotta Birula 1908 with the description of a new subgenus and species from India (Scorpiones, Buthidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 13 (162), 191 - 195." type="journal article" year="2000">Loureno (2000)</bibRefCitation>
for the species
<taxonomicName id="4C7EA1DBFF96FFDAFC74FEB60E44FE05" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="geffardi" subGenus="Deccanobuthus">
<emphasis id="B90A064AFF96FFDAFC74FEB60B79FE2D" box="[980,1113,364,389]" italics="true" pageId="2" pageNumber="3">Hottentotta</emphasis>
(
<emphasis id="B90A064AFF96FFDAFBCBFEB60A01FE2D" box="[1131,1313,364,389]" italics="true" pageId="2" pageNumber="3">Deccanobuthus</emphasis>
)
<emphasis id="B90A064AFF96FFDAFEA8FE4E0E44FE05" box="[264,356,404,429]" italics="true" pageId="2" pageNumber="3">geffardi</emphasis>
</taxonomicName>
from
<collectingCountry id="F3699AC8FF96FFDAFE0CFE4E0ECCFE06" box="[428,492,404,430]" name="India" pageId="2" pageNumber="3">India</collectingCountry>
.
</paragraph>
<paragraph id="8BC1DA58FF96FFDAFE98FE660E9EFBD6" blockId="2.[264,1323,284,1150]" pageId="2" pageNumber="3">
In their exhaustive study of the genus
<taxonomicName id="4C7EA1DBFF96FFDAFCAEFE660C56FE7D" box="[782,886,444,469]" class="Arachnida" family="Buthidae" genus="Buthotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFCAEFE660C56FE7D" box="[782,886,444,469]" italics="true" pageId="2" pageNumber="3">Buthotus</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C7EA1DBFF96FFDAFC06FE660B0BFE7D" box="[934,1067,444,469]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFC06FE660B0BFE7D" box="[934,1067,444,469]" italics="true" pageId="2" pageNumber="3">Hottentotta</emphasis>
</taxonomicName>
),
<bibRefCitation id="EFEFA7A9FF96FFDAFBE7FE660E84FE56" author="Vachon" pageId="2" pageNumber="16" refString="Vachon, M. &amp; Stockmann, R. (1968) Contribution a l'etude des scorpions africains appartenant au genre Buthotus Vachon, 1949 et etude de la variabilite. Monitore Zoologico Italiano (N. S.) 2 (Suppl.), 81 - 149." type="journal article" year="1968">Vachon and Stockmann (1968)</bibRefCitation>
recognized and defined several lineages and sub­lineages of
<taxonomicName id="4C7EA1DBFF96FFDAFB3EFE3E0A03FE55" box="[1182,1315,484,509]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFB3EFE3E0A03FE55" box="[1182,1315,484,509]" italics="true" pageId="2" pageNumber="3">Hottentotta</emphasis>
</taxonomicName>
. Both authors, as well as
<bibRefCitation id="EFEFA7A9FF96FFDAFDE8FDD60CEAFD8E" author="Vachon" box="[584,970,524,550]" pageId="2" pageNumber="16" refString="Vachon, M. &amp; Kinzelbach, R. (1987) On the taxonomy and distribution of the scorpions of the Middle East. In: Krupp, F., Schneider, W. &amp; Kinzelbach, R. (Eds.). Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East, Mainz, 1985. Beihefte zum Tubinger Atlas des vorderen Orients, Reihe A (Naturwissenschaften), 28, 91 - 103." type="journal article" year="1987">Vachon and Kinzelbach (1987)</bibRefCitation>
, suggested that the African, Saharo­Sindian and Indian lineages might be different. In fact, several species originally thought to belong to the Indian lineage are currently placed in the genus
<taxonomicName id="4C7EA1DBFF96FFDAFB3DFD860E8DFD36" authority="Vachon, 1950" authorityName="Vachon" authorityYear="1950" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFB3DFD860A0BFDDD" box="[1181,1323,604,629]" italics="true" pageId="2" pageNumber="3">Mesobuthus</emphasis>
<bibRefCitation id="EFEFA7A9FF96FFDAFEA8FD5E0E8DFD36" author="Vachon" box="[264,429,644,670]" pageId="2" pageNumber="16" refString="Vachon, M. (1950) Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie, 28, 152 - 216." type="journal article" year="1950">Vachon, 1950</bibRefCitation>
</taxonomicName>
(see also
<bibRefCitation id="EFEFA7A9FF96FFDAFD8CFD5E0C5BFD36" author="Tikader" box="[556,891,644,670]" pageId="2" pageNumber="16" refString="Tikader, B. K. &amp; Bastawade, D. B. (1983) The Fauna of India. Vol. 3. Scorpions (Scorpionida: Arachnida). Zoological Survey of India, Calcutta, 671 pp." type="book" year="1983">Tikader &amp; Bastawade 1983</bibRefCitation>
). In addition to its association with
<taxonomicName id="4C7EA1DBFF96FFDAFEA8FD760EADFD6D" box="[264,397,684,709]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFEA8FD760EADFD6D" box="[264,397,684,709]" italics="true" pageId="2" pageNumber="3">Hottentotta</emphasis>
</taxonomicName>
, the genus
<taxonomicName id="4C7EA1DBFF96FFDAFDBEFD760DF4FD6D" box="[542,724,684,709]" class="Arachnida" family="Buthidae" genus="Deccanobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFDBEFD760DF4FD6D" box="[542,724,684,709]" italics="true" pageId="2" pageNumber="3">Deccanobuthus</emphasis>
</taxonomicName>
also shows affinities with the genus
<taxonomicName id="4C7EA1DBFF96FFDAFB3DFD760A0BFD6D" box="[1181,1323,684,709]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFB3DFD760A0BFD6D" box="[1181,1323,684,709]" italics="true" pageId="2" pageNumber="3">Mesobuthus</emphasis>
</taxonomicName>
and, because of its very marked carination, the new
<taxonomicName id="4C7EA1DBFF96FFDAFC21FD0E0B2FFD45" box="[897,1039,724,749]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFC21FD0E0B2FFD45" box="[897,1039,724,749]" italics="true" pageId="2" pageNumber="3">Mesobuthus</emphasis>
</taxonomicName>
species described here, could also be accommodated in this last genus. However, according to
<bibRefCitation id="EFEFA7A9FF96FFDAFB20FD260E78FC96" author="Fet" pageId="2" pageNumber="15" refString="Fet, V. &amp; Lowe, G. (2000) Family Buthidae C. L. Koch, 1837. In: Fet, V., Sissom, W. D., Lowe, G. &amp; Braunwalder, M. E. (Eds.). Catalog of the Scorpions of the world (1758 - 1998). The New York Entomological Society, New York, pp. 54 - 286." type="book chapter" year="2000">Fet and Lowe (2000)</bibRefCitation>
, the relationships of
<taxonomicName id="4C7EA1DBFF96FFDAFDF7FCFE0DC5FC95" box="[599,741,804,829]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFDF7FCFE0DC5FC95" box="[599,741,804,829]" italics="true" pageId="2" pageNumber="3">Mesobuthus</emphasis>
</taxonomicName>
to other genera, and in particular to
<taxonomicName id="4C7EA1DBFF96FFDAFB05FCFE0A0AFC95" box="[1189,1322,804,829]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFB05FCFE0A0AFC95" box="[1189,1322,804,829]" italics="true" pageId="2" pageNumber="3">Hottentotta</emphasis>
</taxonomicName>
remain poorly defined. According to V. Fet (personal communication), the clarification of this situation will require a careful analysis of clusters of true
<taxonomicName id="4C7EA1DBFF96FFDAFC48FCAE0B56FC25" box="[1000,1142,884,909]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFC48FCAE0B56FC25" box="[1000,1142,884,909]" italics="true" pageId="2" pageNumber="3">Mesobuthus</emphasis>
</taxonomicName>
versus African and Middle­East
<taxonomicName id="4C7EA1DBFF96FFDAFE71FC460D76FC1D" box="[465,598,924,949]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFE71FC460D76FC1D" box="[465,598,924,949]" italics="true" pageId="2" pageNumber="3">Hottentotta</emphasis>
</taxonomicName>
, with the possible definition of new characters. The DNA tree proposed by
<bibRefCitation id="EFEFA7A9FF96FFDAFE07FC1E0D4FFC76" box="[423,623,964,990]" pageId="2" pageNumber="15" refString="Fet, V., Gantenbein, B., Gromov, A. V, Lowe, G. &amp; Lourenco, W. R. (2003) The first molecular phylogeny of Buthidae (Scorpiones). Euscorpius, 4, 1 - 10." type="journal article">
Fet
<emphasis id="B90A064AFF96FFDAFE78FC1E0D36FC75" box="[472,534,964,989]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
(2003)
</bibRefCitation>
shows that
<taxonomicName id="4C7EA1DBFF96FFDAFCA6FC1E0CB4FC75" box="[774,916,964,989]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFCA6FC1E0CB4FC75" box="[774,916,964,989]" italics="true" pageId="2" pageNumber="3">Mesobuthus</emphasis>
</taxonomicName>
is much closer to
<taxonomicName id="4C7EA1DBFF96FFDAFBDCFC1E0E40FBAE" authority="Vachon" authorityName="Vachon" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFBDCFC1E0A0BFC75" box="[1148,1323,964,989]" italics="true" pageId="2" pageNumber="3">Compsobuthus</emphasis>
Vachon
</taxonomicName>
that it is to
<taxonomicName id="4C7EA1DBFF96FFDAFE4EFC360D53FBAD" box="[494,627,1004,1029]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFE4EFC360D53FBAD" box="[494,627,1004,1029]" italics="true" pageId="2" pageNumber="3">Hottentotta</emphasis>
</taxonomicName>
, but, for this study only few
<taxonomicName id="4C7EA1DBFF96FFDAFC6AFC360B70FBAD" box="[970,1104,1004,1029]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90A064AFF96FFDAFC6AFC360B70FBAD" box="[970,1104,1004,1029]" italics="true" pageId="2" pageNumber="3">Hottentotta</emphasis>
</taxonomicName>
species have been analysed. For this reason, we believe that only after a revision of all the species assigned to these different genera has been made, a definitive conclusion regarding their precise status can be reached.
</paragraph>
</subSubSection>
</treatment>
</document>