405 lines
70 KiB
XML
405 lines
70 KiB
XML
<document id="818D5ACD383465889ACA22EB2FC543E6" ID-CLB-Dataset="30873" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-Zenodo-Dep="1163422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. & Yağmur, Ersen A." docDate="2018" docId="03B9B44C56730C2153818143FAAC4AB2" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola Zonstein, 1987: 1014" docType="treatment" docVersion="12" lastPageNumber="7" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="5" updateTime="1698518020641" updateUser="ExternalLinkService">
|
||
<mods:mods id="1CABA03AFB1DC45322CBA00BF9BCB472" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="6273DF5762F8EF00B0CED73D4B98889C">
|
||
<mods:title id="CD1B260EDDB67FBB06FCC0A96DD34DCB">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="93A91FC1EB6E7C9619F23632F48FE6A7" type="personal">
|
||
<mods:role id="68AAE36D607B6FF41BFEE522C6C4B57E">
|
||
<mods:roleTerm id="8FF0C823932E9B5537F0A83DE4C070B4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D8377B8564EB0DE92D0B5DF9A39F4FC7">Zonstein, Sergei</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F8F0A8155C0864A7E4DA44D96C89BFD4" type="personal">
|
||
<mods:role id="EF8A9393B0D63DD7AAF866344FD51697">
|
||
<mods:roleTerm id="3AE53FAB4595F641E60E9DE4E56FCB39">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3EE2C1B4FE531A6B109C5BA405179499">Kunt, Kadir B.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="AD36B84FD2DD12A6374F511BDA506CB6" type="personal">
|
||
<mods:role id="8BE3000B4254F4D571EDDDD5B91EE176">
|
||
<mods:roleTerm id="550D0A7D850F131CA32630B7E1EC4B05">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B7098556F6C9D2F04AD1C49D1434785D">Yağmur, Ersen A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="AEDA516EB85494345D63143668BD22D7">text</mods:typeOfResource>
|
||
<mods:relatedItem id="128023E07554F45FE2A11F556D9901CC" type="host">
|
||
<mods:titleInfo id="23C98D948FAA85CC94E2A23EE727A79C">
|
||
<mods:title id="6F7F8FDDAF9A27BAD963ADB0EAC269A8">European Journal of Taxonomy</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1035570FDCDF879A3179A4AB40FA751B">
|
||
<mods:date id="67072E27D222F8B3B3B2833C1E5FDA8A">2018</mods:date>
|
||
<mods:detail id="800540F63199BB383320B7C24B4BCAC4" type="pubDate">
|
||
<mods:number id="A3274C5D9EC3004E7A66C683C43E775E">2018-01-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="0603EE8D867474B8674D657FB59E09D0" type="volume">
|
||
<mods:number id="2DA3E3688BB6E00DC2E2BCD17704EC39">399</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="53261B5C253372B8697DFE14D3632F17" unit="page">
|
||
<mods:start id="091E4567F427B12BB51D939D7CAD2F80">1</mods:start>
|
||
<mods:end id="72CD1726C30786FB23FEBE7F02C9C1BD">93</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="99AE57FAE76DF0D7E462989B06F30C49">journal article</mods:classification>
|
||
<mods:identifier id="D0F376CEF7E8D88945DF164DADE8EF91" type="CLB-Dataset">30873</mods:identifier>
|
||
<mods:identifier id="A1E63BF263C932750FBBD16684764F03" type="DOI">10.5852/ejt.2018.399</mods:identifier>
|
||
<mods:identifier id="6F52FB178004284D631C1D7F233AD03B" type="GBIF-Dataset">b389e1f3-82be-4258-9b09-b7ef9d839f99</mods:identifier>
|
||
<mods:identifier id="D95B836909C120DE1F1A23E1DFC6AD65" type="Zenodo-Dep">1163422</mods:identifier>
|
||
<mods:identifier id="EE7EEE4AF43EC13EB3FE8B8FC2670827" type="ZooBank">E836E138-D6E2-4F62-B4B3-CE2E073F2B24</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03B9B44C56730C2153818143FAAC4AB2" ID-DOI="http://doi.org/10.5281/zenodo.5980273" ID-GBIF-Taxon="140561333" ID-Zenodo-Dep="5980273" LSID="urn:lsid:plazi:treatment:03B9B44C56730C2153818143FAAC4AB2" httpUri="http://treatment.plazi.org/id/03B9B44C56730C2153818143FAAC4AB2" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
|
||
<subSubSection id="C30A56D156730C2353818143FE824CAE" pageId="4" pageNumber="5" type="nomenclature">
|
||
<paragraph id="8BAF055A56730C2353818143FC354C6F" blockId="4.[189,1007,1104,1131]" box="[189,1007,1104,1131]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C107ED956730C2353818143FEEE4C6F" ID-CoL="77BJ" authorityName="Zonstein, 1987:" authorityYear="1014" box="[189,308,1104,1130]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856730C2353818143FEEE4C6F" box="[189,308,1104,1130]" italics="true" pageId="4" pageNumber="5">Raveniola</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<bibRefCitation id="EF8178AB56730C23526D8143FD5B4C6F" author="Zonstein S. & Marusik Y. M." box="[337,641,1104,1130]" journalOrPublisher="ZooKeys" pageId="4" pageNumber="5" pagination="71 - 99" part="211" publicationUrl="https://doi.org/10.3897/zookeys.211.3060" refString="Zonstein S. & Marusik Y. M. 2012. A review of the genus Raveniola (Araneae, Nemesiidae) in China, with notes on allied genera and description of four new species from Yunnan. ZooKeys 211: 71 - 99. https: // doi. org / 10.3897 / zookeys. 211.3060" title="A review of the genus Raveniola (Araneae, Nemesiidae) in China, with notes on allied genera and description of four new species from Yunnan" type="journal article" year="2012">Zonstein & Marusik 2012</bibRefCitation>
|
||
: 74. —
|
||
<bibRefCitation id="EF8178AB56730C2351DD8143FC104C6E" author="Li S. & Zonstein S." box="[737,970,1104,1131]" journalOrPublisher="ZooKeys" pageId="4" pageNumber="5" pagination="1 - 32" part="519" publicationUrl="https://doi.org/10.3897/zookeys.519.8784" refString="Li S. & Zonstein S. 2015. Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae). ZooKeys 519: 1 - 32. https: // doi. org / 10.3897 / zookeys. 519.8784" title="Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae)" type="journal article" year="2015">Li & Zonstein 2015</bibRefCitation>
|
||
: 3.
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56730C2353818181FE824CAE" blockId="4.[189,872,1169,1242]" box="[189,344,1169,1196]" pageId="4" pageNumber="5">
|
||
<heading id="D0E7B23656730C2353818181FE824CAE" bold="true" box="[189,344,1169,1196]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="B964D94856730C2353818181FE824CAE" bold="true" box="[189,344,1169,1196]" pageId="4" pageNumber="5">Type species</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A56D156730C23538181D3FCB24CDF" box="[189,872,1216,1242]" pageId="4" pageNumber="5" type="synonymic_list">
|
||
<paragraph id="8BAF055A56730C23538181D3FCB24CDF" blockId="4.[189,872,1169,1242]" box="[189,872,1216,1242]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C107ED956730C23538181D3FD9F4CDF" authority="Simon, 1891" authorityName="Simon" authorityYear="1891" box="[189,581,1216,1242]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="virgata">
|
||
<emphasis id="B964D94856730C23538181D3FE7C4CDF" box="[189,422,1216,1242]" italics="true" pageId="4" pageNumber="5">Brachythele virgata</emphasis>
|
||
Simon, 1891
|
||
</taxonomicName>
|
||
, by original designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A56D156730C2253818011FB514B2A" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
|
||
<paragraph id="8BAF055A56730C2353818011FE754D1E" blockId="4.[189,431,1281,1308]" box="[189,431,1281,1308]" pageId="4" pageNumber="5">
|
||
<heading id="D0E7B23656730C2353818011FE754D1E" bold="true" box="[189,431,1281,1308]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="B964D94856730C2353818011FE754D1E" bold="true" box="[189,431,1281,1308]" pageId="4" pageNumber="5">Emended diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56730C2353818023FB664E1A" blockId="4.[189,1399,1328,1567]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C107ED956730C2353818023FEEE4D4F" authorityName="Zonstein, 1987:" authorityYear="1014" box="[189,308,1328,1354]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856730C2353818023FEEE4D4F" box="[189,308,1328,1354]" italics="true" pageId="4" pageNumber="5">Raveniola</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C107ED956730C23524D8022FE014D4E" authorityName="Raven & Schwendinger" authorityYear="1995" box="[369,475,1329,1355]" class="Arachnida" family="Nemesiidae" genus="Sinopesa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856730C23524D8022FE014D4E" box="[369,475,1329,1355]" italics="true" pageId="4" pageNumber="5">Sinopesa</emphasis>
|
||
</taxonomicName>
|
||
share two (sometimes three) retroventral megaspines located sequentially on tibia I in males (the unique position among male nemesiids) and the paired spermathecae in females, each carrying two individual receptacular heads; within these taxa the PMS are reduced in size (lost entirely in some species) and the maxillary serrula and preening combs are absent.
|
||
<taxonomicName id="4C107ED956730C23574B8089FB344DB1" authorityName="Zonstein, 1987:" authorityYear="1014" box="[1143,1262,1434,1460]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856730C23574B8089FB344DB1" box="[1143,1262,1434,1460]" italics="true" pageId="4" pageNumber="5">Raveniola</emphasis>
|
||
</taxonomicName>
|
||
is currently known to differ from
|
||
<taxonomicName id="4C107ED956730C23528A80ADFDFA4DDD" authorityName="Raven & Schwendinger" authorityYear="1995" box="[438,544,1470,1496]" class="Arachnida" family="Nemesiidae" genus="Sinopesa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856730C23528A80ADFDFA4DDD" box="[438,544,1470,1496]" italics="true" pageId="4" pageNumber="5">Sinopesa</emphasis>
|
||
</taxonomicName>
|
||
by a noticeably longer and denser leg scopula (vs a short and rare scopula in
|
||
<taxonomicName id="4C107ED956730C2353E680F2FE9E4DFE" authorityName="Raven & Schwendinger" authorityYear="1995" box="[218,324,1505,1531]" class="Arachnida" family="Nemesiidae" genus="Sinopesa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856730C2353E680F2FE9E4DFE" box="[218,324,1505,1531]" italics="true" pageId="4" pageNumber="5">Sinopesa</emphasis>
|
||
</taxonomicName>
|
||
), and by a developed leg and carapace setation. The male palpal organ is moderate in size in
|
||
<taxonomicName id="4C107ED956730C2353818317FEEE4E1B" authorityName="Zonstein, 1987:" authorityYear="1014" box="[189,308,1540,1566]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856730C2353818317FEEE4E1B" box="[189,308,1540,1566]" italics="true" pageId="4" pageNumber="5">Raveniola</emphasis>
|
||
</taxonomicName>
|
||
but appears to be enlarged (with respect to the entire male palp) in
|
||
<taxonomicName id="4C107ED956730C2357778316FB6F4E1A" authorityName="Raven & Schwendinger" authorityYear="1995" box="[1099,1205,1541,1567]" class="Arachnida" family="Nemesiidae" genus="Sinopesa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856730C2357778316FB6F4E1A" box="[1099,1205,1541,1567]" italics="true" pageId="4" pageNumber="5">Sinopesa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56730C2353818355FCAC4E65" blockId="4.[189,1399,1606,1714]" box="[189,886,1606,1632]" pageId="4" pageNumber="5">
|
||
<heading id="D0E7B23656730C2353818355FCAC4E65" bold="true" box="[189,886,1606,1632]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="B964D94856730C2353818355FCAC4E65" bold="true" box="[189,886,1606,1632]" pageId="4" pageNumber="5">Morphological peculiarities of Western Asian congeners</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56730C2353818366FE7F4EB7" blockId="4.[189,1399,1606,1714]" pageId="4" pageNumber="5">
|
||
HABITUS (
|
||
<figureCitation id="132B19DF56730C2352118367FE6F4E8A" box="[301,437,1652,1679]" captionStart-0="Figs43–51" captionStart-1="Figs 52–58" captionStartId-0="57.[189,241,1838,1864]" captionStartId-1="58.[189,241,1833,1859]" captionTargetBox-0="[190,1398,265,1787]" captionTargetBox-1="[189,1398,265,1770]" captionTargetId-0="figure@57.[189,1398,265,1787]" captionTargetId-1="figure@58.[189,1398,265,1771]" captionTargetPageId-0="57" captionText-0="Figs43–51. Raveniolaspp.,habitusof ♂♂(43,45–46,48,50–51)and ♀♀(44,47,49),dorsalviews.— 43–44. R. adjarica sp. nov. 45. R. dunini sp. nov. 46–47. R. hyrcanica Dunin, 1988. 48–49. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." captionText-1="Figs 52–58. Raveniola spp., habitus of ♂♂ (52–53, 55, 57) and ♀♀ (54, 56, 58), lateral (52) and dorsal (53–58) views. — 52. R. biresekensis sp. nov. (dwarf conspecific male?). 53–54. R. micropa (Ausserer, 1871). 55–56. R. zaizevi (Charitonov, 1948). 57–58. R. niedermeyeri (Brignoli, 1972). Scale bars: 5.0 mm." httpUri-0="https://zenodo.org/record/1163438/files/figure.png" httpUri-1="https://zenodo.org/record/1163440/files/figure.png" pageId="4" pageNumber="5">Figs 43–58</figureCitation>
|
||
). The studied spiders are mostly small or medium-sized nemesiids with a carapace length of 2–10 mm.
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56730C23538183C9FE514FEE" blockId="4.[189,1399,1753,2027]" pageId="4" pageNumber="5">
|
||
PROSOMA. Carapace broad-oval and hirsute, with cephalic part slightly to noticeably higher than thoracic part. Eye tubercle predominantly low in males and females (best developed in
|
||
<taxonomicName id="4C107ED956730C23574D83EEFABF4F12" authorityName="Dunin" authorityYear="1988" box="[1137,1381,1789,1815]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hyrcanica">
|
||
<emphasis id="B964D94856730C23574D83EEFABF4F12" box="[1137,1381,1789,1815]" italics="true" pageId="4" pageNumber="5">Raveniola hyrcanica</emphasis>
|
||
</taxonomicName>
|
||
). Cheliceral mound and rastellum absent (see
|
||
<figureCitation id="132B19DF56730C2351838233FCD84F3F" box="[703,770,1824,1850]" captionStart="Figs 1–6" captionStartId="50.[189,241,1734,1760]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@50.[189,1399,265,1680]" captionText="Figs 1–6.Raveniola spp., ♀♀ (1–3, 5–6) and ♂ (4). Chelicerae (1), apical edge of maxilla (2–3), palpal organ (4) and tarsus I (5–6); ventroapical (1), ventral (2–4, 6) and dorsal (5) views. — 1–2. R. micropa (Ausserer, 1871). 3–5. R. pontica (Spassky, 1937). 6. R. hyrcanica Dunin, 1988." httpUri="https://zenodo.org/record/1163424/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
|
||
). In males chelicerae with a small, softened, pale area probably corresponding to a vestigial or poorly-developed intercheliceral male tumescence and located in the well-developed cavity on the ventral cheliceral surface (
|
||
<figureCitation id="132B19DF56730C2350B28274FBCC4F84" box="[910,1046,1894,1921]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="4" pageNumber="5">Figs 59–60</figureCitation>
|
||
). Sternal sigilla medium-sized to small, broad-oval, submarginal (see
|
||
<figureCitation id="132B19DF56730C2351BA8299FCD44FA0" box="[646,782,1930,1957]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="4" pageNumber="5">Figs 62–64</figureCitation>
|
||
). Maxillae with few to numerous maxillary cuspules confined to probasal edge. Serrula absent, but cuticle near the anterior maxillary edge has remnants of scales (
|
||
<figureCitation id="132B19DF56730C23522F82C2FEA44FEE" box="[275,382,2001,2027]" captionStart="Figs 1–6" captionStartId="50.[189,241,1734,1760]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@50.[189,1399,265,1680]" captionText="Figs 1–6.Raveniola spp., ♀♀ (1–3, 5–6) and ♂ (4). Chelicerae (1), apical edge of maxilla (2–3), palpal organ (4) and tarsus I (5–6); ventroapical (1), ventral (2–4, 6) and dorsal (5) views. — 1–2. R. micropa (Ausserer, 1871). 3–5. R. pontica (Spassky, 1937). 6. R. hyrcanica Dunin, 1988." httpUri="https://zenodo.org/record/1163424/files/figure.png" pageId="4" pageNumber="5">Figs 2–3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56720C2253818418FBD849DC" blockId="5.[189,1399,267,473]" pageId="5" pageNumber="6">
|
||
STRUCTURES OF LEGS I–IV. Male tibia and metatarsus I as shown in
|
||
<figureCitation id="132B19DF56720C22508E8418FB8F4920" box="[946,1109,267,293]" captionStart-0="Figs129–132" captionStart-1="Figs 133–136" captionStart-2="Figs 137–140" captionStart-3="Figs 141–144" captionStartId-0="68.[189,241,1870,1896]" captionStartId-1="69.[189,241,1907,1933]" captionStartId-2="70.[189,241,1831,1857]" captionStartId-3="71.[189,241,1858,1884]" captionTargetBox-0="[220,1291,268,1789]" captionTargetBox-1="[255,1325,265,1873]" captionTargetBox-2="[211,1305,274,1777]" captionTargetBox-3="[239,1352,266,1791]" captionTargetId-0="figure@68.[189,1323,265,1791]" captionTargetId-1="figure@69.[227,1361,265,1874]" captionTargetId-2="figure@70.[197,1331,272,1777]" captionTargetId-3="figure@71.[206,1378,265,1793]" captionTargetPageId-0="68" captionTargetPageId-1="69" captionTargetPageId-2="70" captionText-0="Figs129–132.Raveniolaspp.,♂♂.TibiaandmetatarsusI,retrolateralviews.—129.R.adjaricasp.nov. (holotype).130.R.duninisp.nov.(holotype).131.R.hyrcanicaDunin,1988(holotype).132.R.pontica (Spassky, 1937) (Lazarevskoe). Scalebars: 1.0 mm." captionText-1="Figs 133–136. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 133. R. birecikensis sp. nov.(holotype).134. Same speciesas preceding (paratype, Birecik). 135. Samespecies as preceding (dwarf conspecific male?). 136. R. anadolu sp. nov.(holotype). Scale bars: 1.0 mm." captionText-2="Figs 137–140. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 137. R. arthuri Kunt & Yağmur, 2010 (holotype). 138. R. turcica sp. nov. (holotype). 139. R. micropa (Ausserer, 1871) (Uludağ).140. R. mazandaranica Marusik, Zamani &Mirshamsi, 2014 (holotype). Scalebars: 1.0mm." captionText-3="Figs 141–144. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 141. R. zaitzevi (Charitonov, 1948) (Car). 142. R. marusiki sp. nov. (holotype). 143. R. vonwicki Zonstein, 2000 (holotype). 144. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri-0="https://zenodo.org/record/1163460/files/figure.png" httpUri-1="https://zenodo.org/record/1163462/files/figure.png" httpUri-2="https://zenodo.org/record/1163464/files/figure.png" httpUri-3="https://zenodo.org/record/1163466/files/figure.png" pageId="5" pageNumber="6">Figs 129–144</figureCitation>
|
||
. Leg scopula varies from dense and moderately long to thin and short; distal on metatarsi I–II, generally entire on tarsus I, mostly divided on tarsus II; females also with entire scopula on palpal tarsus. Tarsal organ low, domed, with weak to sharp concentric ridges (
|
||
<figureCitation id="132B19DF56720C2251728464FD114994" box="[590,715,375,401]" captionStart="Figs 7–12" captionStartId="51.[189,241,1719,1745]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@51.[189,1399,265,1680]" captionText="Figs 7–12. Raveniola spp., ♀♀. Tarsal organ, legs I (7–8, 10, 12) and IV (9, 11), dorsal views. — 7–9. R. hyrcanica Dunin, 1988. 10. R. pontica (Spassky, 1937). 11. R. micropa (Ausserer, 1871). 12. R. zaitzevi (Charitonov, 1948)." httpUri="https://zenodo.org/record/1163426/files/figure.png" pageId="5" pageNumber="6">Figs 7–12</figureCitation>
|
||
). Trichobothrial bases corrugiform (
|
||
<figureCitation id="132B19DF56720C2257418464FAD34994" box="[1149,1289,375,401]" captionStart="Figs 13–18" captionStartId="52.[189,241,1725,1751]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@52.[189,1399,265,1680]" captionText="Figs 13–18. Raveniola spp., ♀♀. Bases of trichobothria, legs I (13, 15–16, 18) and IV (14, 17), dorsal views.—13–14.R.hyrcanicaDunin,1988.15.R.pontica(Spassky,1937).16–17.R.micropa(Ausserer, 1871). 18. R. zaitzevi (Charitonov, 1948)." httpUri="https://zenodo.org/record/1163428/files/figure.png" pageId="5" pageNumber="6">Figs 13–18</figureCitation>
|
||
). Female palp with tarsal claw dentate on promargin (
|
||
<figureCitation id="132B19DF56720C2251FE8488FCC349B0" box="[706,793,411,437]" captionStart="Figs 19–24" captionStartId="53.[189,241,1725,1751]" captionTargetBox="[189,1399,265,1678]" captionTargetId="figure@53.[189,1399,265,1680]" captionText="Figs 19–24. Raveniola spp., ♀♀. Palpal (19) and tarsal (20–24) claws; apical (19, 21, 23) and lateral (20, 22, 24) views. — 19–20. R. pontica (Spassky, 1937). 21–22. R. hyrcanica Dunin, 1988, legs I and IV. 23–24. R. micropa (Ausserer, 1871), legs I and IV." httpUri="https://zenodo.org/record/1163430/files/figure.png" pageId="5" pageNumber="6">Fig. 19</figureCitation>
|
||
). Paired tarsal claws of legs I–IV biserially dentate, unpaired claw always present, moderately small and bare (
|
||
<figureCitation id="132B19DF56720C22505084ADFC2C49DC" box="[876,1014,446,473]" captionStart="Figs 19–24" captionStartId="53.[189,241,1725,1751]" captionTargetBox="[189,1399,265,1678]" captionTargetId="figure@53.[189,1399,265,1680]" captionText="Figs 19–24. Raveniola spp., ♀♀. Palpal (19) and tarsal (20–24) claws; apical (19, 21, 23) and lateral (20, 22, 24) views. — 19–20. R. pontica (Spassky, 1937). 21–22. R. hyrcanica Dunin, 1988, legs I and IV. 23–24. R. micropa (Ausserer, 1871), legs I and IV." httpUri="https://zenodo.org/record/1163430/files/figure.png" pageId="5" pageNumber="6">Figs 20–24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56720C2253818712FB964A67" blockId="5.[189,1399,512,611]" pageId="5" pageNumber="6">
|
||
MALE PALP. Tibia of moderate to short length, spinose. Cymbium moderately short, with or without spines (
|
||
<figureCitation id="132B19DF56720C22522B8737FE654A3A" box="[279,447,548,575]" captionStart-0="Figs 149–152" captionStart-1="Figs 153–156" captionStart-2="Figs 157–160" captionStartId-0="73.[189,241,1930,1956]" captionStartId-1="74.[189,241,1894,1920]" captionStartId-2="75.[189,241,1894,1920]" captionTargetBox-0="[194,1398,266,1866]" captionTargetBox-1="[189,1398,265,1868]" captionTargetBox-2="[189,1395,266,1870]" captionTargetId-0="figure@73.[189,1398,265,1875]" captionTargetId-1="figure@74.[189,1398,265,1868]" captionTargetId-2="figure@75.[189,1398,265,1875]" captionTargetPageId-0="73" captionTargetPageId-1="74" captionText-0="Figs 149–152. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 149. R. adjarica sp. nov. (holotype). 150. R. dunini sp. nov. (holotype). 151. R. hyrcanica Dunin, 1988 (holotype). 152. R. pontica (Spassky, 1937) (Lazarevskoe). Scale bars: 0.5 mm." captionText-1="Figs 153–156. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 153. R. birecikensis sp. nov. (holotype). 154. Same species as preceding (paratype, Birecik, revolved bulbshowninventralposition).155.Samespeciesaspreceding(dwarf conspecific♂?). 156.R.anadolu sp. nov. (holotype). Scale bars: 0.5 mm." captionText-2="Figs 157–160. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 157. R. arthuri Kunt & Yağmur, 2010 (holotype). 158. R. turcica sp. nov. (holotype). 159. R. micropa (Ausserer, 1871) (Uludağ). 160. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014 (holotype). Scale bars: 0.5 mm." httpUri-0="https://zenodo.org/record/1163470/files/figure.png" httpUri-1="https://zenodo.org/record/1163472/files/figure.png" httpUri-2="https://zenodo.org/record/1163474/files/figure.png" pageId="5" pageNumber="6">Figs 149–160</figureCitation>
|
||
). Male palpal bulb pyriform, inserted submedially. Embolus short to moderately long, tapering or spiraled, with or without keels and ridges (
|
||
<figureCitation id="132B19DF56720C225047875BFC124A67" box="[891,968,584,610]" captionStart="Figs 1–6" captionStartId="50.[189,241,1734,1760]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@50.[189,1399,265,1680]" captionText="Figs 1–6.Raveniola spp., ♀♀ (1–3, 5–6) and ♂ (4). Chelicerae (1), apical edge of maxilla (2–3), palpal organ (4) and tarsus I (5–6); ventroapical (1), ventral (2–4, 6) and dorsal (5) views. — 1–2. R. micropa (Ausserer, 1871). 3–5. R. pontica (Spassky, 1937). 6. R. hyrcanica Dunin, 1988." httpUri="https://zenodo.org/record/1163424/files/figure.png" pageId="5" pageNumber="6">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="132B19DF56720C2250E8875BFBE54A67" box="[980,1087,584,610]" captionStart-0="Figs 171–179" captionStart-1="Figs 180–188" captionStart-2="Figs 189–197" captionStartId-0="78.[189,241,1846,1872]" captionStartId-1="79.[189,241,1858,1884]" captionStartId-2="80.[189,241,1823,1849]" captionTargetBox-0="[189,1399,265,1803]" captionTargetBox-1="[189,1399,265,1830]" captionTargetBox-2="[200,1386,265,1801]" captionTargetId-0="figure@78.[189,1399,265,1803]" captionTargetId-1="figure@79.[189,1399,265,1830]" captionTargetId-2="figure@80.[200,1386,265,1801]" captionTargetPageId-0="78" captionTargetPageId-1="79" captionText-0="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." captionText-1="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt & Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." captionText-2="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri-0="https://zenodo.org/record/1163480/files/figure.png" httpUri-1="https://zenodo.org/record/1163482/files/figure.png" httpUri-2="https://zenodo.org/record/1163484/files/figure.png" pageId="5" pageNumber="6">171–197</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56720C2253818799FADC4AA1" blockId="5.[189,1286,650,677]" box="[189,1286,650,677]" pageId="5" pageNumber="6">
|
||
SPERMATHECAE. Two wide or narrow spermathecae, each with two receptacles (
|
||
<figureCitation id="132B19DF56720C22576E8799FB234AA1" box="[1106,1273,650,676]" captionStart-0="Figs 198–203" captionStart-1="Figs 204–212" captionStartId-0="81.[189,241,1819,1845]" captionStartId-1="82.[189,241,1656,1682]" captionTargetBox-0="[222,1376,271,1766]" captionTargetBox-1="[198,1407,277,1637]" captionTargetId-0="figure@81.[189,1398,265,1770]" captionTargetId-1="figure@82.[198,1407,265,1652]" captionTargetPageId-0="81" captionText-0="Figs 198–203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." captionText-1="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri-0="https://zenodo.org/record/1163486/files/figure.png" httpUri-1="https://zenodo.org/record/1163488/files/figure.png" pageId="5" pageNumber="6">Figs 198–212</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56720C22538187DEFB514B2A" blockId="5.[189,1398,716,815]" pageId="5" pageNumber="6">
|
||
SPINNERETS. Two pairs of spinnerets (PMS always present). PMS small but with a few fully functional spigots confined to the apex (
|
||
<figureCitation id="132B19DF56720C22511E87E3FD744B0E" box="[546,686,752,779]" captionStart="Figs 25–30" captionStartId="54.[189,241,1739,1765]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@54.[189,1399,265,1680]" captionText="Figs 25–30. Raveniolaspp.,♀♀. Posteriormedian spinnerets (25–27),detailsof spigots (28–30);apical (25–26,28–30)andventral(27)views.—25,29–30.R.hyrcanicaDunin,1988.26.R.pontica(Spassky, 1937). 27–28. R. zaitzevi (Charitonov, 1948)." httpUri="https://zenodo.org/record/1163432/files/figure.png" pageId="5" pageNumber="6">Figs 25–30</figureCitation>
|
||
). PLS with spigots confined to ventral surface of segments (
|
||
<figureCitation id="132B19DF56720C2253F98607FE964B2A" box="[197,332,788,815]" captionStart="Figs 31–36" captionStartId="55.[189,241,1739,1765]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@55.[189,1399,265,1680]" captionText="Figs 31–36. Raveniola spp., ♀♀. Posterior lateral spinnerets (31–33), with spigots in close-up view (34–36); proventral (31), apical (32, 36) and ventral (33–35) views. — 31, 34. R. zaitzevi (Charitonov, 1948). 32, 36. R. micropa (Ausserer, 1871). 33, 35. R. hyrcanica Dunin, 1988." httpUri="https://zenodo.org/record/1163434/files/figure.png" pageId="5" pageNumber="6">Figs 33–36</figureCitation>
|
||
) and with apical segment triangular to digitiform (
|
||
<figureCitation id="132B19DF56720C2250A18607FBFE4B2B" box="[925,1060,788,814]" captionStart="Figs 31–36" captionStartId="55.[189,241,1739,1765]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@55.[189,1399,265,1680]" captionText="Figs 31–36. Raveniola spp., ♀♀. Posterior lateral spinnerets (31–33), with spigots in close-up view (34–36); proventral (31), apical (32, 36) and ventral (33–35) views. — 31, 34. R. zaitzevi (Charitonov, 1948). 32, 36. R. micropa (Ausserer, 1871). 33, 35. R. hyrcanica Dunin, 1988." httpUri="https://zenodo.org/record/1163434/files/figure.png" pageId="5" pageNumber="6">Figs 31–32</figureCitation>
|
||
,
|
||
<figureCitation id="132B19DF56720C22570C8607FBA44B2A" box="[1072,1150,788,815]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="5" pageNumber="6">65–67</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A56D156720C2253818645FDF94C56" pageId="5" pageNumber="6" type="synonymic_list">
|
||
<paragraph id="8BAF055A56720C2253818645FE564B75" blockId="5.[189,396,854,880]" box="[189,396,854,880]" pageId="5" pageNumber="6">
|
||
<heading id="D0E7B23656720C2253818645FE564B75" bold="true" box="[189,396,854,880]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="B964D94856720C2253818645FE564B75" bold="true" box="[189,396,854,880]" pageId="5" pageNumber="6">Species included</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56720C2253818696FDF94C56" blockId="5.[189,1399,901,1107]" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C107ED956720C2253818696FEEE4B9A" authorityName="Zonstein, 1987:" authorityYear="1014" box="[189,308,901,927]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856720C2253818696FEEE4B9A" box="[189,308,901,927]" italics="true" pageId="5" pageNumber="6">Raveniola</emphasis>
|
||
</taxonomicName>
|
||
currently comprises 37 species, including the new species described here; 16 of them occur in Western Asia:
|
||
<taxonomicName id="4C107ED956720C22525986B9FE324BC6" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[357,488,937,963]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="adjarica" status="sp. nov.">
|
||
<emphasis id="B964D94856720C22525986B9FE324BC6" box="[357,488,937,963]" italics="true" pageId="5" pageNumber="6">R. adjarica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356720C2252D386B9FD914BC1" box="[495,587,938,964]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C107ED956720C22516A86B9FD0C4BC6" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[598,726,937,963]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="anadolu" status="sp. nov.">
|
||
<emphasis id="B964D94856720C22516A86B9FD0C4BC6" box="[598,726,937,963]" italics="true" pageId="5" pageNumber="6">R. anadolu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356720C2251E086B9FCE24BC1" box="[732,824,938,964]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C107ED956720C22507F86B9FB1B4BC6" authority="Kunt & Yagmur, 2010" authorityName="Kunt & Yagmur" authorityYear="2010" box="[835,1217,937,964]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="arthuri">
|
||
<emphasis id="B964D94856720C22507F86B9FC6C4BC6" box="[835,950,937,963]" italics="true" pageId="5" pageNumber="6">R. arthuri</emphasis>
|
||
<bibRefCitation id="EF8178AB56720C22508086B9FB1B4BC6" author="Kunt K. B. & Yagmur E. A." box="[956,1217,937,964]" journalOrPublisher="Turkish Journal of Zoology" pageId="5" pageNumber="6" pagination="305 - 309" part="34" publicationUrl="https://doi.org/10.3906/zoo-0906-40" refString="Kunt K. B. & Yagmur E. A. 2010. A new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae; Nemesiidae) from Turkey. Turkish Journal of Zoology 34: 305 - 309. https: // doi. org / 10.3906 / zoo- 0906 - 40" title="A new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae; Nemesiidae) from Turkey" type="journal article" year="2010">Kunt & Yağmur, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C107ED956720C2257F086B9FAAC4BC6" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[1228,1398,937,963]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="birecikensis" status="sp. nov.">
|
||
<emphasis id="B964D94856720C2257F086B9FAAC4BC6" box="[1228,1398,937,963]" italics="true" pageId="5" pageNumber="6">R. birecikensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356720C22538186DDFEC04BED" box="[189,282,974,1000]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C107ED956720C22521986DDFE4A4BE2" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[293,400,973,999]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dunini" status="sp. nov.">
|
||
<emphasis id="B964D94856720C22521986DDFE4A4BE2" box="[293,400,973,999]" italics="true" pageId="5" pageNumber="6">R. dunini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356720C2252AB86DDFE294BED" box="[407,499,974,1000]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C107ED956720C2252C386DDFCF44BE2" authority="Dunin, 1988" authorityName="Dunin" authorityYear="1988" box="[511,814,973,999]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="hyrcanica">
|
||
<emphasis id="B964D94856720C2252C386DDFD494BE2" box="[511,659,973,999]" italics="true" pageId="5" pageNumber="6">R. hyrcanica</emphasis>
|
||
<bibRefCitation id="EF8178AB56720C2251A686DEFCF44BE2" author="Dunin P. M." box="[666,814,973,999]" journalOrPublisher="Zoologicheskii Zhurnal" pageId="5" pageNumber="6" pagination="1245 - 1248" part="67" refString="Dunin P. M. 1988. Mygalomorphic spiders (Aranei, Mygalomorphae) of Azerbaijan. Zoologicheskii Zhurnal 67 (8): 1245 - 1248. [In Russian with English summary]" title="Mygalomorphic spiders (Aranei, Mygalomorphae) of Azerbaijan" type="journal article" year="1988">Dunin, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C107ED956720C22500686DDFC1B4BE2" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[826,961,973,999]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
|
||
<emphasis id="B964D94856720C22500686DDFC1B4BE2" box="[826,961,973,999]" italics="true" pageId="5" pageNumber="6">R. marusiki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356720C2250F486DDFBFF4BED" box="[968,1061,974,1000]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C107ED956720C22570C86DDFDD24C0E" authority="Marusik, Zamani & Mirshamsi, 2014" authorityName="Marusik, Zamani & Mirshamsi" authorityYear="2014" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mazandaranica">
|
||
<emphasis id="B964D94856720C22570C86DDFADD4BE2" box="[1072,1287,973,999]" italics="true" pageId="5" pageNumber="6">R. mazandaranica</emphasis>
|
||
<bibRefCitation id="EF8178AB56720C22563186DEFDD24C0E" author="Marusik Y. M. & Zamani A. & Mirshamsi O." journalOrPublisher="ZooKeys" pageId="5" pageNumber="6" pagination="1 - 10" part="463" publicationUrl="https://doi.org/10.3897/zookeys.463.8692" refString="Marusik Y. M., Zamani A. & Mirshamsi O. 2014. Three new species of mygalomorph and filistatid spiders from Iran (Araneae, Cyrtaucheniidae, Nemesiidae and Filistatidae). ZooKeys 463: 1 - 10. https: // doi. org / 10.3897 / zookeys. 463.8692" title="Three new species of mygalomorph and filistatid spiders from Iran (Araneae, Cyrtaucheniidae, Nemesiidae and Filistatidae)" type="journal article" year="2014">Marusik, Zamani & Mirshamsi, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C107ED956720C22512886E1FCBF4C0E" authority="(Ausserer, 1871)" authorityName="in Uludag Mts" authorityYear="1871" baseAuthorityName="Ausserer" baseAuthorityYear="1871" box="[532,869,1009,1035]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="micropa">
|
||
<emphasis id="B964D94856720C22512886E1FD4C4C0E" box="[532,662,1010,1035]" italics="true" pageId="5" pageNumber="6">R. micropa</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8178AB56720C22519A86E2FC864C0E" author="Ausserer A." box="[678,860,1009,1035]" journalOrPublisher="Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien" pageId="5" pageNumber="6" pagination="177 - 224" part="21" refString="Ausserer A. 1871. Beitrage zur Kenntnis der Arachniden-Familie Territelariae Thorell (Mygalidae Autor). Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien 21: 177 - 224." title="Beitrage zur Kenntnis der Arachniden-Familie Territelariae Thorell (Mygalidae Autor)" type="journal article" year="1871">Ausserer, 1871</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C107ED956720C22504E86E1FC144C0E" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[882,974,1010,1035]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nana" status="sp. nov.">
|
||
<emphasis id="B964D94856720C22504E86E1FC144C0E" box="[882,974,1010,1035]" italics="true" pageId="5" pageNumber="6">R. nana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356720C2250EA86E2FBE94C0E" box="[982,1075,1009,1035]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C107ED956720C22577C86E1FEDE4C2A" authority="(Brignoli, 1972)" baseAuthorityName="Brignoli" baseAuthorityYear="1972" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="niedermeyeri">
|
||
<emphasis id="B964D94856720C22577C86E1FB264C0E" box="[1088,1276,1009,1035]" italics="true" pageId="5" pageNumber="6">R. niedermeyeri</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8178AB56720C22563086E2FF264C2A" author="Brignoli P. M." journalOrPublisher="Revue suisse de Zoologie" pageId="5" pageNumber="6" pagination="409 - 413" part="79" refString="Brignoli P. M. 1972. Une nouvelle Brachythele de l'Iran (Arachnida, Araneae, Dipluridae). Revue suisse de Zoologie 79: 409 - 413." title="Une nouvelle Brachythele de l'Iran (Arachnida, Araneae, Dipluridae)" type="journal article" year="1972">Brignoli, 1972</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C107ED956720C2252338105FD904C2A" authority="(Spassky, 1937)" baseAuthorityName="Spassky" baseAuthorityYear="1937" box="[271,586,1045,1071]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pontica">
|
||
<emphasis id="B964D94856720C2252338105FE5D4C2A" box="[271,391,1046,1071]" italics="true" pageId="5" pageNumber="6">R. pontica</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8178AB56720C2252A98106FD9B4C2A" author="Spassky S. A." box="[405,577,1045,1071]" journalOrPublisher="Festschrift Embrik Strand" pageId="5" pageNumber="6" pagination="361 - 368" part="3" refString="Spassky S. A. 1937. Araneae palaearcticae novae. Mygalomorphae. I. Festschrift Embrik Strand 3: 361 - 368." title="Araneae palaearcticae novae. Mygalomorphae. I" type="journal article" year="1937">Spassky, 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C107ED956720C2251698105FD674C2A" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[597,701,1046,1071]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sinani" status="sp. nov.">
|
||
<emphasis id="B964D94856720C2251698105FD674C2A" box="[597,701,1046,1071]" italics="true" pageId="5" pageNumber="6">R. sinani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356720C2251FF8106FCC54C2A" box="[707,799,1045,1071]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C107ED956720C2250168105FC414C2A" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[810,923,1046,1071]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="turcica" status="sp. nov.">
|
||
<emphasis id="B964D94856720C2250168105FC414C2A" box="[810,923,1046,1071]" italics="true" pageId="5" pageNumber="6">R. turcica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356720C22509E8106FC244C2A" box="[930,1022,1045,1071]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C107ED956720C2257348105FA9C4C2A" authority="Zonstein, 2000" authorityName="Zonstein" authorityYear="2000" box="[1032,1350,1045,1071]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vonwicki">
|
||
<emphasis id="B964D94856720C2257348105FB4A4C2A" box="[1032,1168,1045,1071]" italics="true" pageId="5" pageNumber="6">R. vonwicki</emphasis>
|
||
<bibRefCitation id="EF8178AB56720C2257AB8106FA9C4C2A" author="Zonstein S. L." box="[1175,1350,1045,1071]" journalOrPublisher="Tethys Entomological Research" pageId="5" pageNumber="6" pagination="49 - 52" part="2" refString="Zonstein S. L. 2000. Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia. Tethys Entomological Research 2: 49 - 52." title="Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia" type="journal article" year="2000">Zonstein, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C107ED956720C225381812AFDFA4C56" authority="(Charitonov, 1948)" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[189,544,1080,1107]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="zaitzevi">
|
||
<emphasis id="B964D94856720C225381812AFEEC4C57" box="[189,310,1081,1106]" italics="true" pageId="5" pageNumber="6">R. zaitzevi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8178AB56720C225279812BFDCC4C56" author="Charitonov D. E." box="[325,534,1080,1107]" journalOrPublisher="Soobshcheniya Akademii Nauk Gruzinskoi SSR" pageId="5" pageNumber="6" pagination="135 - 139" part="9" refString="Charitonov D. E. 1948. A new species of Brachythele from Georgia. Soobshcheniya Akademii Nauk Gruzinskoi SSR 9: 135 - 139. [In Russian and Latin]" title="A new species of Brachythele from Georgia" type="journal article" year="1948">Charitonov, 1948</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A56D156720C2253818169FE4A4D72" pageId="5" pageNumber="6" type="discussion">
|
||
<paragraph id="8BAF055A56720C2253818169FE484C91" blockId="5.[189,402,1146,1172]" box="[189,402,1146,1172]" pageId="5" pageNumber="6">
|
||
<heading id="D0E7B23656720C2253818169FE484C91" bold="true" box="[189,402,1146,1172]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="B964D94856720C2253818169FE484C91" bold="true" box="[189,402,1146,1172]" pageId="5" pageNumber="6">Species grouping</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56720C22538181B9FE4A4D72" blockId="5.[189,1399,1194,1399]" pageId="5" pageNumber="6">
|
||
To enhance identifications, the species treated here are assigned to three species groups.These assignments are preliminary, because males and females in some species are unknown, and they are not based on a phylogenetic grouping, though some of the groups may indeed reflect phylogenetic relationships. The similarity in external morphology in all females of Western Asian
|
||
<taxonomicName id="4C107ED956720C2257148006FB454D2A" authorityName="Zonstein, 1987:" authorityYear="1014" box="[1064,1183,1301,1327]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856720C2257148006FB454D2A" box="[1064,1183,1301,1327]" italics="true" pageId="5" pageNumber="6">Raveniola</emphasis>
|
||
</taxonomicName>
|
||
necessitates their assignment both throughout the species and the species groups being entirely based on the structure of the spermathecae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A56D156720C225381808CFB454EE1" pageId="5" pageNumber="6" type="distribution">
|
||
<paragraph id="8BAF055A56720C225381808CFE2B4DBC" blockId="5.[189,497,1439,1465]" box="[189,497,1439,1465]" pageId="5" pageNumber="6">
|
||
<heading id="D0E7B23656720C225381808CFE2B4DBC" bold="true" box="[189,497,1439,1465]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="B964D94856720C225381808CFE2B4DBC" bold="true" box="[189,497,1439,1465]" pageId="5" pageNumber="6">Distribution and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56720C22538180DDFB454EE1" blockId="5.[189,1398,1486,1764]" pageId="5" pageNumber="6">
|
||
Within Western Asia, representatives of
|
||
<taxonomicName id="4C107ED956720C22519A80DDFCC74DED" authorityName="Zonstein, 1987:" authorityYear="1014" box="[678,797,1486,1512]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964D94856720C22519A80DDFCC74DED" box="[678,797,1486,1512]" italics="true" pageId="5" pageNumber="6">Raveniola</emphasis>
|
||
</taxonomicName>
|
||
are distributed from the western part of Anatolia (Turkey: Bursa) through the Caucasus mountain region to the Alborz and Khorasan Mts, Iran. All species with known habitats were found in different types of piedmont and mountainous deciduous broad-leaved forests, and at least some of them may inhabit the higher coniferous forest belt (
|
||
<taxonomicName id="4C107ED956720C2257C88328FAAC4E51" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[1268,1398,1594,1620]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="adjarica" status="sp. nov.">
|
||
<emphasis id="B964D94856720C2257C88328FAAC4E51" box="[1268,1398,1594,1620]" italics="true" pageId="5" pageNumber="6">R. adjarica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356720C225381834DFEC24E7D" box="[189,280,1630,1656]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) or even subalpine grasslands (
|
||
<taxonomicName id="4C107ED956720C2251BB834DFD254E72" baseAuthorityName="Spassky" baseAuthorityYear="1937" box="[647,767,1630,1655]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pontica">
|
||
<emphasis id="B964D94856720C2251BB834DFD254E72" box="[647,767,1630,1655]" italics="true" pageId="5" pageNumber="6">R. pontica</emphasis>
|
||
</taxonomicName>
|
||
); some species (like
|
||
<taxonomicName id="4C107ED956720C2250C9834DFBBA4E7D" authority="Zonstein & Kunt & Yağmur, 2018" authorityName="Zonstein & Kunt & Yağmur" authorityYear="2018" box="[1013,1120,1630,1656]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dunini" status="sp. nov.">
|
||
<emphasis id="B964D94856720C2250C9834DFBBA4E7D" box="[1013,1120,1630,1656]" italics="true" pageId="5" pageNumber="6">R. dunini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A257643356720C22575B834DFB184E7D" box="[1127,1218,1630,1656]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) are also found in the open forest biotopes at low altitudes. In most species, females use cavities under rocks to build primitive burrow-like dwellings lacking any silk-lining. However, females of
|
||
<taxonomicName id="4C107ED956720C22577483B5FB064EBA" authorityName="Dunin" authorityYear="1988" box="[1096,1244,1701,1727]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="hyrcanica">
|
||
<emphasis id="B964D94856720C22577483B5FB064EBA" box="[1096,1244,1701,1727]" italics="true" pageId="5" pageNumber="6">R. hyrcanica</emphasis>
|
||
</taxonomicName>
|
||
, unlike other congeners, dig very simple open burrows, also without silk-lining, in the forest floor.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A56D156720C215381821EFAAC4AB2" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="key">
|
||
<paragraph id="8BAF055A56720C225381821EFC5A4F2F" blockId="5.[189,896,1804,1834]" box="[189,896,1804,1834]" pageId="5" pageNumber="6">
|
||
<emphasis id="B964D94856720C225381821EFC5A4F2F" bold="true" box="[189,896,1804,1834]" pageId="5" pageNumber="6">
|
||
<emphasis id="B964D94856720C225381821EFD354F2C" bold="true" box="[189,751,1804,1833]" italics="true" pageId="5" pageNumber="6">Key to the Western Asian species groups of</emphasis>
|
||
<taxonomicName id="4C107ED956720C2251CB821EFC5A4F2F" ID-CoL="77BJ" authorityName="Zonstein, 1987:" authorityYear="1014" box="[759,896,1805,1834]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="93" phylum="Arthropoda" rank="genus">Raveniola</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56720C2253D68251FEEF4F59" blockId="5.[189,1398,1858,2027]" box="[234,309,1858,1884]" pageId="5" pageNumber="6">
|
||
<emphasis id="B964D94856720C2253D68251FEEF4F59" bold="true" box="[234,309,1858,1884]" pageId="5" pageNumber="6">Males</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56720C2253818276FAAC4FA6" blockId="5.[189,1398,1858,2027]" pageId="5" pageNumber="6">
|
||
1. Palpal tibia subcylindrical, without bare preapical area, low mound and sensilla (as in
|
||
<figureCitation id="132B19DF56720C2257C38276FECD4FA6" captionStart-0="Figs 171–179" captionStart-1="Figs 180–188" captionStartId-0="78.[189,241,1846,1872]" captionStartId-1="79.[189,241,1858,1884]" captionTargetBox-0="[189,1399,265,1803]" captionTargetBox-1="[189,1399,265,1830]" captionTargetId-0="figure@78.[189,1399,265,1803]" captionTargetId-1="figure@79.[189,1399,265,1830]" captionTargetPageId-0="78" captionText-0="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." captionText-1="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt & Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri-0="https://zenodo.org/record/1163480/files/figure.png" httpUri-1="https://zenodo.org/record/1163482/files/figure.png" pageId="5" pageNumber="6">Figs 171– 188</figureCitation>
|
||
) …………………………………………………………………………………………………2
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56720C2253FD82BEFAAC4FEE" blockId="5.[189,1398,1858,2027]" pageId="5" pageNumber="6">
|
||
– Palpal tibia dilated subapically, bearing retrolateral hump and bare retroventral-distal area with low mound and sensilla (as in
|
||
<figureCitation id="132B19DF56720C22516582C2FD274FEE" box="[601,765,2001,2027]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="5" pageNumber="6">Figs 189–197</figureCitation>
|
||
) …………………………………
|
||
<taxonomicName id="4C107ED956720C2257B782C3FAF34FEE" ID-CoL="6WT99" baseAuthorityName="Brignoli" baseAuthorityYear="1972" box="[1163,1321,2000,2027]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="niedermeyeri">
|
||
<emphasis id="B964D94856720C2257B782C3FAF34FEE" bold="true" box="[1163,1321,2000,2027]" italics="true" pageId="5" pageNumber="6">niedermeyeri</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56710C2153818418FAAC4923" blockId="6.[189,1399,267,370]" box="[189,1398,267,294]" pageId="6" pageNumber="7">
|
||
2. Embolus always keeled; embolic keel laterally directed (
|
||
<figureCitation id="132B19DF56710C2150A08418FB994920" box="[924,1091,267,293]" captionStart="Figs 171–179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="6" pageNumber="7">Figs 171–179</figureCitation>
|
||
) ………
|
||
<taxonomicName id="4C107ED956710C2157928418FAFD4923" ID-CoL="4RMTW" authorityName="Dunin" authorityYear="1988" box="[1198,1319,267,294]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hyrcanica">
|
||
<emphasis id="B964D94856710C2157928418FAFD4923" bold="true" box="[1198,1319,267,294]" italics="true" pageId="6" pageNumber="7">hyrcanica</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56710C2153FD8421FAAC4977" blockId="6.[189,1399,267,370]" pageId="6" pageNumber="7">
|
||
– Embolus with or without keel; embolic keel, if present, running parallel to embolus (
|
||
<figureCitation id="132B19DF56710C2157C18422FECD4974" captionStart="Figs 180–188" captionStartId="79.[189,241,1858,1884]" captionTargetBox="[189,1399,265,1830]" captionTargetId="figure@79.[189,1399,265,1830]" captionText="Figs 180–188. Raveniola micropa species group, holotypic (180–181, 183–186) and conspecific (182, 187–188) ♂♂. Details of palpal organ; retrolateral (180, 182, 185, 187), ventral (181, 186, 188), retroventral (183) and prolateral (184) views. — 180–181. R. birecikensis sp. nov. 182. Same species as preceding (dwarf specimen?). 183. R. anadolu sp. nov. 184. R. arthuri Kunt & Yağmur, 2010. 185– 186. R. turcica sp. nov. 187–188. R. micropa (Ausserer, 1871). Scalebars: 0.25 mm." httpUri="https://zenodo.org/record/1163482/files/figure.png" pageId="6" pageNumber="7">Figs 180– 188</figureCitation>
|
||
) …………………………………………………………………………………
|
||
<taxonomicName id="4C107ED956710C2157828444FAFB4974" ID-CoL="6WSX9" authorityName="in Uludag Mts" baseAuthorityName="Ausserer" baseAuthorityYear="1871" box="[1214,1313,343,369]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="micropa">
|
||
<emphasis id="B964D94856710C2157828444FAFB4974" bold="true" box="[1214,1313,343,369]" italics="true" pageId="6" pageNumber="7">micropa</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56710C2153D68488FE8B49B0" blockId="6.[189,1398,411,514]" box="[234,337,411,437]" pageId="6" pageNumber="7">
|
||
<emphasis id="B964D94856710C2153D68488FE8B49B0" bold="true" box="[234,337,411,437]" pageId="6" pageNumber="7">Females</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56710C21538184D2FAAC49DE" blockId="6.[189,1398,411,514]" box="[189,1398,449,476]" pageId="6" pageNumber="7">
|
||
1. Bases of spermathecae relatively narrow or moderately wide (
|
||
<figureCitation id="132B19DF56710C2150DC84D2FB5249DE" box="[992,1160,449,475]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="6" pageNumber="7">Figs 204–212</figureCitation>
|
||
) …………………2
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56710C2153FD84FBFAAC4A07" blockId="6.[189,1398,411,514]" box="[193,1398,487,514]" pageId="6" pageNumber="7">
|
||
– Bases of spermathecae very wide (
|
||
<figureCitation id="132B19DF56710C2151A884F4FCE14A04" box="[660,827,487,513]" captionStart="Figs 198–203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198–203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="6" pageNumber="7">Figs 198–203</figureCitation>
|
||
) ………………………………
|
||
<taxonomicName id="4C107ED956710C21579284F4FAFD4A07" ID-CoL="4RMTW" authorityName="Dunin" authorityYear="1988" box="[1198,1319,487,514]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hyrcanica">
|
||
<emphasis id="B964D94856710C21579284F4FAFD4A07" bold="true" box="[1198,1319,487,514]" italics="true" pageId="6" pageNumber="7">hyrcanica</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56710C2153818738FAAC4A69" blockId="6.[189,1399,555,695]" pageId="6" pageNumber="7">
|
||
2. Bases of spermathecae always moderately wide; median and lateral receptacles subequal in shape and size (
|
||
<figureCitation id="132B19DF56710C21525F8742FDD64A6E" box="[355,524,593,619]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="6" pageNumber="7">Figs 204–207</figureCitation>
|
||
) ……………………………………………………………
|
||
<taxonomicName id="4C107ED956710C2157FE8742FAFF4A6E" ID-CoL="6WSX9" authorityName="in Uludag Mts" baseAuthorityName="Ausserer" baseAuthorityYear="1871" box="[1218,1317,593,619]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="micropa">
|
||
<emphasis id="B964D94856710C2157FE8742FAFF4A6E" bold="true" box="[1218,1317,593,619]" italics="true" pageId="6" pageNumber="7">micropa</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
</paragraph>
|
||
<paragraph id="8BAF055A56710C2153FD8764FAAC4AB2" blockId="6.[189,1399,555,695]" pageId="6" pageNumber="7">
|
||
– Bases of spermathecae generally relatively narrow (
|
||
<figureCitation id="132B19DF56710C21506E8764FC214A94" box="[850,1019,631,657]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="6" pageNumber="7">Figs 210–212</figureCitation>
|
||
); median and lateral receptacles different in shape and size (
|
||
<figureCitation id="132B19DF56710C215177878EFD2F4AB2" box="[587,757,669,695]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="6" pageNumber="7">Figs 208–209</figureCitation>
|
||
) …………………………………
|
||
<taxonomicName id="4C107ED956710C2157BB878FFAFF4AB2" ID-CoL="6WT99" baseAuthorityName="Brignoli" baseAuthorityYear="1972" box="[1159,1317,668,695]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="niedermeyeri">
|
||
<emphasis id="B964D94856710C2157BB878FFAFF4AB2" bold="true" box="[1159,1317,668,695]" italics="true" pageId="6" pageNumber="7">niedermeyeri</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |