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<document id="01E6481EC4CCC95C527DB261B247DBD4" ID-DOI="10.5281/zenodo.214372" ID-GBIF-Dataset="2b697d24-e22e-4b28-9f45-51b1c63a3e95" ID-ISSN="1175-5326" ID-Zenodo-Dep="214372" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460503232220" checkinUser="plazi" docAuthor="Packer, Laurence" docDate="2012" docId="03A987CAF45C5257FF59769DFEA7F985" docLanguage="en" docName="zt03408p058.pdf" docOrigin="Zootaxa 3408" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Penapis larraini Packer, new species" docType="treatment" docVersion="9" lastPageNumber="55" masterDocId="FF90FFB2F45D5256FFCE7643FF88FFC8" masterDocTitle="Penapis larraini Packer, a new species of rophitine bee (Hymenoptera: Halictidae) from a fog oasis in Northern Chile" masterLastPageNumber="58" masterPageNumber="54" pageNumber="55" updateTime="1698312281271" updateUser="plazi">
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<mods:title id="4C10CEFA697BE1AA870DD80FFB278D64">Penapis larraini Packer, a new species of rophitine bee (Hymenoptera: Halictidae) from a fog oasis in Northern Chile</mods:title>
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<mods:namePart id="7C8BAC55CC5D8C156B5439685A1EB193">Packer, Laurence</mods:namePart>
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<mods:date id="EABE7617770CCB9E53F9432283B2B95C">2012</mods:date>
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<treatment id="03A987CAF45C5257FF59769DFEA7F985" ID-DOI="http://doi.org/10.5281/zenodo.6165856" ID-GBIF-Taxon="119522233" ID-Zenodo-Dep="6165856" LSID="urn:lsid:plazi:treatment:03A987CAF45C5257FF59769DFEA7F985" httpUri="http://treatment.plazi.org/id/03A987CAF45C5257FF59769DFEA7F985" lastPageNumber="55" pageId="1" pageNumber="55">
<subSubSection id="C31A6557F45C5257FF59769DFEB9FED1" pageId="1" pageNumber="55" type="nomenclature">
<paragraph id="8BBF36DCF45C5257FF59769DFDDEFF30" blockId="1.[151,598,222,281]" box="[151,598,222,248]" pageId="1" pageNumber="55">
<heading id="D0F781B0F45C5257FF59769DFDDEFF30" bold="true" box="[151,598,222,248]" fontSize="11" level="1" pageId="1" pageNumber="55" reason="1">
<emphasis id="B974EACEF45C5257FF59769DFDDEFF30" bold="true" box="[151,598,222,248]" pageId="1" pageNumber="55">
<taxonomicName id="4C004D5FF45C5257FF59769DFE31FF30" authority="Packer" authorityName="Packer" box="[151,441,222,248]" class="Insecta" family="Halictidae" genus="Penapis" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="larraini" status="sp. nov.">
<emphasis id="B974EACEF45C5257FF59769DFED3FF30" bold="true" box="[151,347,222,248]" italics="true" pageId="1" pageNumber="55">Penapis larraini</emphasis>
Packer
</taxonomicName>
,
<taxonomicNameLabel id="A24757B5F45C5257FE08769DFDDEFF30" box="[454,598,222,248]" pageId="1" pageNumber="55" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF36DCF45C5257FF597742FEB9FED1" blockId="1.[151,598,222,281]" box="[151,305,257,281]" pageId="1" pageNumber="55">
(
<figureCitation id="133B2A59F45C5257FF517742FF65FED1" box="[159,237,257,281]" captionStart="FIGURES 1 A K. A J" captionStartId="2.[151,264,1844,1867]" captionTargetBox="[164,1418,228,1771]" captionTargetId="figure@2.[151,1436,195,1802]" captionTargetPageId="2" captionText="FIGURES 1 A K. A J. Penapis larraini Packer, n. sp. A H male, I J female. K. Penapis toroi female. A. head lateral view. B. S 4 ventral view. C. apex of metasoma ventral view. D. S 5 ventral view. E. S 5 lateral view. F. S 6 ventral view. G. S 8 ventral view. H. Genital capsule ventral view. I. Head lateral view. J. T 1 T 3 dorsal view. K. T 1 T 3 dorsal view. Note that the individual shown in J has the most sparse but large punctures of all the specimens available. Scale line = 1 mm." httpUri="https://zenodo.org/record/214373/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
, AJ)
</paragraph>
</subSubSection>
<subSubSection id="C31A6557F45C5257FF59770BFCCBFD01" pageId="1" pageNumber="55" type="diagnosis">
<paragraph id="8BBF36DCF45C5257FF59770BFCCBFD01" blockId="1.[151,1437,328,1613]" pageId="1" pageNumber="55">
<emphasis id="B974EACEF45C5257FF59770BFE99FEA9" bold="true" box="[151,273,328,353]" pageId="1" pageNumber="55">Diagnosis.</emphasis>
The male can be differentiated from other species of the genus by the absence of sublateral processes on S6 and the form of the sublateral processes on S5 (
<figureCitation id="133B2A59F45C5257FD10772EFCE8FE4D" box="[734,864,365,389]" captionStart="FIGURES 1 A K. A J" captionStartId="2.[151,264,1844,1867]" captionTargetBox="[164,1418,228,1771]" captionTargetId="figure@2.[151,1436,195,1802]" captionTargetPageId="2" captionText="FIGURES 1 A K. A J. Penapis larraini Packer, n. sp. A H male, I J female. K. Penapis toroi female. A. head lateral view. B. S 4 ventral view. C. apex of metasoma ventral view. D. S 5 ventral view. E. S 5 lateral view. F. S 6 ventral view. G. S 8 ventral view. H. Genital capsule ventral view. I. Head lateral view. J. T 1 T 3 dorsal view. K. T 1 T 3 dorsal view. Note that the individual shown in J has the most sparse but large punctures of all the specimens available. Scale line = 1 mm." httpUri="https://zenodo.org/record/214373/files/figure.png" pageId="1" pageNumber="55">Figs. 1CF</figureCitation>
). In side view, the latter have a narrow, digitiform apex, which is dorsoventrally flattened (
<figureCitation id="133B2A59F45C5257FD9977D2FD24FE61" box="[599,684,401,425]" captionStart="FIGURES 1 A K. A J" captionStartId="2.[151,264,1844,1867]" captionTargetBox="[164,1418,228,1771]" captionTargetId="figure@2.[151,1436,195,1802]" captionTargetPageId="2" captionText="FIGURES 1 A K. A J. Penapis larraini Packer, n. sp. A H male, I J female. K. Penapis toroi female. A. head lateral view. B. S 4 ventral view. C. apex of metasoma ventral view. D. S 5 ventral view. E. S 5 lateral view. F. S 6 ventral view. G. S 8 ventral view. H. Genital capsule ventral view. I. Head lateral view. J. T 1 T 3 dorsal view. K. T 1 T 3 dorsal view. Note that the individual shown in J has the most sparse but large punctures of all the specimens available. Scale line = 1 mm." httpUri="https://zenodo.org/record/214373/files/figure.png" pageId="1" pageNumber="55">Fig. 1E</figureCitation>
). All other species in the genus have rounded (in
<taxonomicName id="4C004D5FF45C5257FB1A77D1FAA5FE61" box="[1236,1325,402,425]" class="Insecta" family="Halictidae" genus="Penapis" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="B974EACEF45C5257FB1A77D1FAA5FE61" box="[1236,1325,402,425]" italics="true" pageId="1" pageNumber="55">P. penai</emphasis>
</taxonomicName>
angulate) sublateral processes on S6 and approximately triangular processes on S5, which are not dorsoventrally flattened, rather they are uniformly obliquely compressed. The female is most easily distinguished by the punctation of the metasomal terga: T1T4 have large, shallow punctures on the apical half (excepting the apical impressed areas) and T2T4 have minute, denser punctures bearing minute setae that are restricted to the anterior half of the tergum (slightly more extensive medially) (
<figureCitation id="133B2A59F45C5257FDE67406FDFFFD95" box="[552,631,581,605]" captionStart="FIGURES 1 A K. A J" captionStartId="2.[151,264,1844,1867]" captionTargetBox="[164,1418,228,1771]" captionTargetId="figure@2.[151,1436,195,1802]" captionTargetPageId="2" captionText="FIGURES 1 A K. A J. Penapis larraini Packer, n. sp. A H male, I J female. K. Penapis toroi female. A. head lateral view. B. S 4 ventral view. C. apex of metasoma ventral view. D. S 5 ventral view. E. S 5 lateral view. F. S 6 ventral view. G. S 8 ventral view. H. Genital capsule ventral view. I. Head lateral view. J. T 1 T 3 dorsal view. K. T 1 T 3 dorsal view. Note that the individual shown in J has the most sparse but large punctures of all the specimens available. Scale line = 1 mm." httpUri="https://zenodo.org/record/214373/files/figure.png" pageId="1" pageNumber="55">Fig. 1J</figureCitation>
). Other species have the horizontal portion of T1 impunctate, or almost so, and the minute punctures of T2T4 more extensive, at least on T3 approaching the apical impressed areas medially to a distance much less than the length of the impressed area (
<figureCitation id="133B2A59F45C5257FC4474CEFC6CFD6D" box="[906,996,653,677]" captionStart="FIGURES 1 A K. A J" captionStartId="2.[151,264,1844,1867]" captionTargetBox="[164,1418,228,1771]" captionTargetId="figure@2.[151,1436,195,1802]" captionTargetPageId="2" captionText="FIGURES 1 A K. A J. Penapis larraini Packer, n. sp. A H male, I J female. K. Penapis toroi female. A. head lateral view. B. S 4 ventral view. C. apex of metasoma ventral view. D. S 5 ventral view. E. S 5 lateral view. F. S 6 ventral view. G. S 8 ventral view. H. Genital capsule ventral view. I. Head lateral view. J. T 1 T 3 dorsal view. K. T 1 T 3 dorsal view. Note that the individual shown in J has the most sparse but large punctures of all the specimens available. Scale line = 1 mm." httpUri="https://zenodo.org/record/214373/files/figure.png" pageId="1" pageNumber="55">Fig. 1K</figureCitation>
). The male has similar sculpture but is more readily separated by the sternal characters noted above.
</paragraph>
</subSubSection>
<subSubSection id="C31A6557F45C5257FF077496FEA6FB2D" pageId="1" pageNumber="55" type="description">
<paragraph id="8BBF36DCF45C5257FF077496FE20FBC5" blockId="1.[151,1437,328,1613]" pageId="1" pageNumber="55">
<emphasis id="B974EACEF45C5257FF077496FE25FD25" bold="true" box="[201,429,725,750]" pageId="1" pageNumber="55">Description. Male:</emphasis>
Head width
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, ITW
<quantity id="4CF89B39F45C5257FC8C7496FC5BFD25" box="[834,979,725,749]" metricMagnitude="-3" metricUnit="m" metricValue="1.4249999999999998" metricValueMax="1.45" metricValueMin="1.4" pageId="1" pageNumber="55" unit="mm" value="1.4249999999999998" valueMax="1.45" valueMin="1.4">1.41.45mm</quantity>
, wing length 4.85.0mm, body length
<quantity id="4CF89B39F45C5257FF5974BAFE93FCD9" box="[151,283,761,785]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="7.6" metricValueMin="7.4" pageId="1" pageNumber="55" unit="mm" value="7.5" valueMax="7.6" valueMin="7.4">7.47.6mm</quantity>
. Lower part of face moderately projecting (
<figureCitation id="133B2A59F45C5257FCDA74BAFCF8FCD9" box="[788,880,761,785]" captionStart="FIGURES 1 A K. A J" captionStartId="2.[151,264,1844,1867]" captionTargetBox="[164,1418,228,1771]" captionTargetId="figure@2.[151,1436,195,1802]" captionTargetPageId="2" captionText="FIGURES 1 A K. A J. Penapis larraini Packer, n. sp. A H male, I J female. K. Penapis toroi female. A. head lateral view. B. S 4 ventral view. C. apex of metasoma ventral view. D. S 5 ventral view. E. S 5 lateral view. F. S 6 ventral view. G. S 8 ventral view. H. Genital capsule ventral view. I. Head lateral view. J. T 1 T 3 dorsal view. K. T 1 T 3 dorsal view. Note that the individual shown in J has the most sparse but large punctures of all the specimens available. Scale line = 1 mm." httpUri="https://zenodo.org/record/214373/files/figure.png" pageId="1" pageNumber="55">Fig. 1A</figureCitation>
). Pterostigma with pale disc bordered by brown marginal veins. Midtibial spur gently curved almost to base. S4 with more than 10 outwardly recurved plumose setae on each side; apical comb pale amber, more than 20 straight setae on each side, narrowly interrupted medially (
<figureCitation id="133B2A59F45C5257FF517526FF7DFCB5" box="[159,245,869,893]" captionStart="FIGURES 1 A K. A J" captionStartId="2.[151,264,1844,1867]" captionTargetBox="[164,1418,228,1771]" captionTargetId="figure@2.[151,1436,195,1802]" captionTargetPageId="2" captionText="FIGURES 1 A K. A J. Penapis larraini Packer, n. sp. A H male, I J female. K. Penapis toroi female. A. head lateral view. B. S 4 ventral view. C. apex of metasoma ventral view. D. S 5 ventral view. E. S 5 lateral view. F. S 6 ventral view. G. S 8 ventral view. H. Genital capsule ventral view. I. Head lateral view. J. T 1 T 3 dorsal view. K. T 1 T 3 dorsal view. Note that the individual shown in J has the most sparse but large punctures of all the specimens available. Scale line = 1 mm." httpUri="https://zenodo.org/record/214373/files/figure.png" pageId="1" pageNumber="55">Fig. 1B</figureCitation>
), comb setae finer and longer than in other species. S5 with median apical process shorter than sublateral process, laterally compressed and somewhat downcurved apically; sublateral process in lateral view with deep basal portion ending in acute angulation ventrobasad of dorsoventrally flattened, narrow apical projection (
<figureCitation id="133B2A59F45C5257FAAA75EEFF52FC21" captionStart="FIGURES 1 A K. A J" captionStartId="2.[151,264,1844,1867]" captionTargetBox="[164,1418,228,1771]" captionTargetId="figure@2.[151,1436,195,1802]" captionTargetPageId="2" captionText="FIGURES 1 A K. A J. Penapis larraini Packer, n. sp. A H male, I J female. K. Penapis toroi female. A. head lateral view. B. S 4 ventral view. C. apex of metasoma ventral view. D. S 5 ventral view. E. S 5 lateral view. F. S 6 ventral view. G. S 8 ventral view. H. Genital capsule ventral view. I. Head lateral view. J. T 1 T 3 dorsal view. K. T 1 T 3 dorsal view. Note that the individual shown in J has the most sparse but large punctures of all the specimens available. Scale line = 1 mm." httpUri="https://zenodo.org/record/214373/files/figure.png" pageId="1" pageNumber="55">Figs. 1CE</figureCitation>
). S6 lacking sublateral process, surface flat, apical concavity broad, truncate (
<figureCitation id="133B2A59F45C5257FBF57592FB06FC21" box="[1083,1166,977,1001]" captionStart="FIGURES 1 A K. A J" captionStartId="2.[151,264,1844,1867]" captionTargetBox="[164,1418,228,1771]" captionTargetId="figure@2.[151,1436,195,1802]" captionTargetPageId="2" captionText="FIGURES 1 A K. A J. Penapis larraini Packer, n. sp. A H male, I J female. K. Penapis toroi female. A. head lateral view. B. S 4 ventral view. C. apex of metasoma ventral view. D. S 5 ventral view. E. S 5 lateral view. F. S 6 ventral view. G. S 8 ventral view. H. Genital capsule ventral view. I. Head lateral view. J. T 1 T 3 dorsal view. K. T 1 T 3 dorsal view. Note that the individual shown in J has the most sparse but large punctures of all the specimens available. Scale line = 1 mm." httpUri="https://zenodo.org/record/214373/files/figure.png" pageId="1" pageNumber="55">Fig. 1F</figureCitation>
). S8 and genital capsule as in figures 1G and 1H.
</paragraph>
<paragraph id="8BBF36DCF45C5257FF077259FEA6FB2D" blockId="1.[151,1437,328,1613]" pageId="1" pageNumber="55">
<emphasis id="B974EACEF45C5257FF077259FEA1FBFA" bold="true" box="[201,297,1050,1074]" pageId="1" pageNumber="55">Female:</emphasis>
Head width
<quantity id="4CF89B39F45C5257FE74725AFDB4FBF9" box="[442,572,1049,1073]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" pageId="1" pageNumber="55" unit="mm" value="5.0" valueMax="5.2" valueMin="4.8">4.85.2mm</quantity>
, ITW
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, wing length
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, body length
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. Midfemur flat over most of anterior surface, somewhat concave towards apex because of apicodorsal swelling. Scopa pale straw. Metasomal terga with sparse, shallow, large punctures on apical half (apical impressed areas excepted); punctures smaller, denser and more distinct on T5 and T6; anterior half of T2T4 with minute punctures separated by&gt; their diameters; minute punctures extending slightly past midlength of tergum medially, less extensive laterally (
<figureCitation id="133B2A59F45C5257FF1E728EFE97FB2D" box="[208,287,1229,1253]" captionStart="FIGURES 1 A K. A J" captionStartId="2.[151,264,1844,1867]" captionTargetBox="[164,1418,228,1771]" captionTargetId="figure@2.[151,1436,195,1802]" captionTargetPageId="2" captionText="FIGURES 1 A K. A J. Penapis larraini Packer, n. sp. A H male, I J female. K. Penapis toroi female. A. head lateral view. B. S 4 ventral view. C. apex of metasoma ventral view. D. S 5 ventral view. E. S 5 lateral view. F. S 6 ventral view. G. S 8 ventral view. H. Genital capsule ventral view. I. Head lateral view. J. T 1 T 3 dorsal view. K. T 1 T 3 dorsal view. Note that the individual shown in J has the most sparse but large punctures of all the specimens available. Scale line = 1 mm." httpUri="https://zenodo.org/record/214373/files/figure.png" pageId="1" pageNumber="55">Fig. 1J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A6557F45C5257FF0772B2FE8BFA51" pageId="1" pageNumber="55" type="materials_examined">
<paragraph id="8BBF36DCF45C5257FF0772B2FE8BFA51" blockId="1.[151,1437,328,1613]" pageId="1" pageNumber="55">
<emphasis id="B974EACEF45C5257FF0772B2FE1FFAC2" bold="true" box="[201,407,1265,1290]" pageId="1" pageNumber="55">Material studied.</emphasis>
<typeStatus id="54BB887EF45C5257FE5072B2FD8EFAC1" box="[414,518,1265,1289]" pageId="1" pageNumber="55" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F317764CF45C5257FD9A72B2FD2EFAC1" box="[596,678,1265,1289]" name="Chile" pageId="1" pageNumber="55">CHILE</collectingCountry>
Region I, Alto Patache,
<date id="FFBE101CF45C5257FC7072B2FB9EFAC1" box="[958,1046,1265,1289]" pageId="1" pageNumber="55" value="1997-11">xi.1997</date>
, W. Sielfeld. Allotype female, one male
<typeStatus id="54BB887EF45C5257FF197355FEBFFAE6" box="[215,311,1302,1326]" pageId="1" pageNumber="55" type="paratype">paratype</typeStatus>
and two female
<typeStatus id="54BB887EF45C5257FE317355FDE3FAE6" box="[511,619,1302,1326]" pageId="1" pageNumber="55" type="paratype">paratypes</typeStatus>
with identical label data,
<collectionCode id="ED11AE19F45C5257FC557356FC6EFAE5" box="[923,998,1301,1325]" pageId="1" pageNumber="55">PUCV</collectionCode>
with one pair of
<typeStatus id="54BB887EF45C5257FB787355FAA9FAE6" box="[1206,1313,1302,1326]" pageId="1" pageNumber="55" type="paratype">paratypes</typeStatus>
at
<collectionCode id="ED11AE19F45C5257FA847356FA10FAE5" box="[1354,1432,1301,1325]" httpUri="http://grbio.org/cool/66ra-d50z" name="The Packer Collection at York University" pageId="1" pageNumber="55">PCYU</collectionCode>
. Three female
<typeStatus id="54BB887EF45C5257FEF97379FE2AFA9A" box="[311,418,1338,1362]" pageId="1" pageNumber="55" type="paratype">paratypes</typeStatus>
(one missing the head) same locality,
<quantity id="4CF89B39F45C5257FCAC737AFC2DFA99" box="[866,933,1337,1361]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="55" unit="m" value="820.0">820m</quantity>
, one each from
<date id="FFBE101CF45C5257FBAC737AFB54FA99" box="[1122,1244,1337,1361]" pageId="1" pageNumber="55" value="1997-11-30">30.xi.1997</date>
,
<date id="FFBE101CF45C5257FB24737AFAE2FA99" box="[1258,1386,1337,1361]" pageId="1" pageNumber="55" value="1997-12-02">02.xii.1997</date>
and
<date id="FFBE101CF45C5257FF59731EFE91FABD" box="[151,281,1373,1397]" pageId="1" pageNumber="55" value="1997-12-07">07.xii.1997</date>
, H. Larrain, two at
<collectionCode id="ED11AE19F45C5257FE32731EFDC2FABD" box="[508,586,1373,1397]" httpUri="http://grbio.org/cool/66ra-d50z" name="The Packer Collection at York University" pageId="1" pageNumber="55">PCYU</collectionCode>
, one at
<collectionCode id="ED11AE19F45C5257FD6F731EFD64FABD" box="[673,748,1373,1397]" pageId="1" pageNumber="55">PUCV</collectionCode>
. Although not stated on any labels, the locality is at
<geoCoordinate id="EE34501BF45C5257FAF6731EFA1FFABD" box="[1336,1431,1373,1397]" direction="south" orientation="latitude" pageId="1" pageNumber="55" precision="925" value="-20.816668">20°49S</geoCoordinate>
;
<geoCoordinate id="EE34501BF45C5257FF5973C2FF77FA51" box="[151,255,1409,1433]" direction="west" orientation="longitude" pageId="1" pageNumber="55" precision="925" value="-70.15">70°09W</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A6557F45C5257FF0773E6FEA7F985" pageId="1" pageNumber="55" type="etymology">
<paragraph id="8BBF36DCF45C5257FF0773E6FEA7F985" blockId="1.[151,1437,328,1613]" pageId="1" pageNumber="55">
<emphasis id="B974EACEF45C5257FF0773E6FED8FA75" bold="true" box="[201,336,1445,1469]" pageId="1" pageNumber="55">Etymology.</emphasis>
The species is named after Horacio Larrain, eco-anthropologist and archaeologist of the Universidad Boliviariana in Iquique, northern
<collectingCountry id="F317764CF45C5257FDF2738AFDF4FA29" box="[572,636,1481,1505]" name="Chile" pageId="1" pageNumber="55">Chile</collectingCountry>
. The name is appropriate as his wife,
<collectingCountry id="F317764CF45C5257FBEF7389FBEFFA29" box="[1057,1127,1482,1505]" name="Malta" pageId="1" pageNumber="55">Marta</collectingCountry>
, is a sister of the renowned, late Chilean entomologist Luis Peña, after whom the genus and its
<typeStatus id="54BB887EF45C5257FC4573ADFC33F9CE" box="[907,955,1518,1542]" pageId="1" pageNumber="55">type</typeStatus>
species were named. Dr. Larrain collected the first specimens of the species seen by the author and has been involved with numerous research projects on the
<emphasis id="B974EACEF45C5257FF597076FEA0F985" box="[151,296,1589,1613]" italics="true" pageId="1" pageNumber="55">locus typicus</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>