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<document id="402D7D8FE20C41F9936784AC1C8E7BB0" ID-DOI="10.5281/zenodo.174106" ID-GBIF-Dataset="2239677a-be03-489b-a288-2112c6d383c8" ID-ISSN="1175­5326" ID-Zenodo-Dep="174106" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459790819789" checkinUser="plazi" docAuthor="Tan, Jingjing, Ren, Dong &amp; Shih, Chungkun" docDate="2006" docId="038D87F9FFD998107120F91DD6EFF884" docLanguage="en" docName="zt01326p068.pdf" docOrigin="Zootaxa 1326" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Priacma tuberculosa Tan, Ren &amp; Shih, 2006, sp. nov." docType="treatment" docVersion="8" lastPageNumber="61" masterDocId="FFB4FF81FFDD98167028FFC0D71BFFCA" masterDocTitle="First record of fossil Priacma (Coleoptera: Archostemata: Cupedidae) from the Jehol Biota of western Liaoning, China" masterLastPageNumber="68" masterPageNumber="55" pageNumber="59" updateTime="1698219209044" updateUser="plazi">
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<mods:title id="A2E61C6BC84EF629A1261A487AE9E801">First record of fossil Priacma (Coleoptera: Archostemata: Cupedidae) from the Jehol Biota of western Liaoning, China</mods:title>
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<mods:namePart id="18E9BB318C5620624F5F1D1622F30C84">Tan, Jingjing</mods:namePart>
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<mods:namePart id="92F028703B06DE6C1532F1C2B8B85C22">Ren, Dong</mods:namePart>
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<mods:namePart id="CBF8B88614C5C0F049D232A90975CBBD">Shih, Chungkun</mods:namePart>
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<mods:date id="D91E3E7C3CC3BFAA1E4787D60D866416">2006</mods:date>
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<treatment id="038D87F9FFD998107120F91DD6EFF884" ID-DOI="http://doi.org/10.5281/zenodo.6259433" ID-GBIF-Taxon="119350375" ID-Zenodo-Dep="6259433" LSID="urn:lsid:plazi:treatment:038D87F9FFD998107120F91DD6EFF884" httpUri="http://treatment.plazi.org/id/038D87F9FFD998107120F91DD6EFF884" lastPageId="6" lastPageNumber="61" pageId="4" pageNumber="59">
<subSubSection id="C33E6564FFD998127120F91DD6DFF8D3" pageId="4" pageNumber="59" type="nomenclature">
<paragraph id="8B9B36EFFFD998127120F91DD573F93C" blockId="4.[264,616,1756,1817]" box="[264,616,1756,1783]" pageId="4" pageNumber="59">
<heading id="D0D38183FFD998127120F91DD573F93C" bold="true" box="[264,616,1756,1783]" fontSize="11" level="1" pageId="4" pageNumber="59" reason="1">
<emphasis id="B950EAFDFFD998127120F91DD573F93C" bold="true" box="[264,616,1756,1783]" pageId="4" pageNumber="59">
<taxonomicName id="4C244D6CFFD998127120F91DD51FF93D" box="[264,516,1757,1783]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="tuberculosa" status="sp. nov.">
<emphasis id="B950EAFDFFD998127120F91DD51FF93D" bold="true" box="[264,516,1757,1783]" italics="true" pageId="4" pageNumber="59">Priacma tuberculosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2635786FFD998127223F91CD573F93C" box="[523,616,1756,1782]" pageId="4" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B36EFFFD998127120F93FD6DFF8D3" blockId="4.[264,616,1756,1817]" box="[264,452,1791,1817]" pageId="4" pageNumber="59">
(
<figureCitation id="131F2A6AFFD998127139F93FD67FF8D3" box="[273,356,1791,1817]" captionStart="FIGURES 1 4" captionStartId="3.[264,382,1795,1819]" captionTargetBox="[314,1277,326,1768]" captionTargetId="figure@3.[314,1277,326,1769]" captionTargetPageId="3" captionText="FIGURES 1 4. Habiti of new Priacma species, photographs of rock impressions. 1 — Priacma latidentata sp. nov., 2 — Priacma tuberculosa sp. nov., 3 — Priacma clavata sp. nov., 4 — Priacma renaria sp. nov." httpUri="https://zenodo.org/record/174107/files/figure.png" pageId="4" pageNumber="59">Figs. 2</figureCitation>
,
<figureCitation id="131F2A6AFFD998127158F93FD6A0F8D3" box="[368,443,1791,1817]" captionStart-0="FIGURES 10 12" captionStart-1="FIGURES 13 14" captionStartId-0="7.[264,382,1017,1041]" captionStartId-1="7.[264,382,1546,1570]" captionTargetBox-0="[271,1322,321,974]" captionTargetBox-1="[264,1323,1129,1522]" captionTargetId-0="figure@7.[264,1323,312,992]" captionTargetId-1="figure@7.[264,1323,1129,1522]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="FIGURES 10 12. Priacma tuberculosa sp. nov., holotype. 10 — dorsal view, 11 — ventral view, 12 — outline of elytral cells." captionText-1="FIGURES 13 14. Priacma tuberculosa sp. nov., holotype. 13 — head, 14 — elytral cells." httpUri-0="https://zenodo.org/record/174110/files/figure.png" httpUri-1="https://zenodo.org/record/174111/files/figure.png" pageId="4" pageNumber="59">1014</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E6564FFD998107120F88FD466FD84" lastPageId="6" lastPageNumber="61" pageId="4" pageNumber="59" type="diagnosis">
<paragraph id="8B9B36EFFFD998107120F88FD466FD84" blockId="4.[264,1314,1871,1897]" lastBlockId="6.[264,1325,284,1870]" lastPageId="6" lastPageNumber="61" pageId="4" pageNumber="59">
<emphasis id="B950EAFDFFD998127120F88FD693F8A3" bold="true" box="[264,392,1871,1897]" pageId="4" pageNumber="59">Diagnosis.</emphasis>
<taxonomicName id="4C244D6CFFD9981271A7F890D52EF8A3" box="[399,565,1872,1897]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="tuberculosa" status="sp. nov.">
<emphasis id="B950EAFDFFD9981271A7F890D52EF8A3" box="[399,565,1872,1897]" italics="true" pageId="4" pageNumber="59">P. tuberculosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2635786FFD998127214F88FD58CF8A3" box="[572,663,1871,1897]" pageId="4" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="4C244D6CFFD9981273CCF890D34FF8A3" box="[996,1108,1872,1897]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="serrata">
<emphasis id="B950EAFDFFD9981273CCF890D34FF8A3" box="[996,1108,1872,1897]" italics="true" pageId="4" pageNumber="59">P. serrata</emphasis>
</taxonomicName>
in the absence of spines on edge of elytra, from
<taxonomicName id="4C244D6CFFDB981072ADFEDCD417FEFF" box="[645,780,284,309]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="corrupta">
<emphasis id="B950EAFDFFDB981072ADFEDCD417FEFF" box="[645,780,284,309]" italics="true" pageId="6" pageNumber="61">P. corrupta</emphasis>
</taxonomicName>
in the mesosternum of new species without cross suture, from
<taxonomicName id="4C244D6CFFDB981071C3FE84D58DFE97" box="[491,662,324,349]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="longicapitis">
<emphasis id="B950EAFDFFDB981071C3FE84D58DFE97" box="[491,662,324,349]" italics="true" pageId="6" pageNumber="61">P. longicapitis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C244D6CFFDB981072FCFE84D456FE97" box="[724,845,324,349]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="B950EAFDFFDB981072FCFE84D456FE97" box="[724,845,324,349]" italics="true" pageId="6" pageNumber="61">P. oculata</emphasis>
</taxonomicName>
in the ventral head surface of new one without grooves for inserting antennae, from
<taxonomicName id="4C244D6CFFDB9810730EFEACD48BFE4F" box="[806,912,364,389]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="striata">
<emphasis id="B950EAFDFFDB9810730EFEACD48BFE4F" box="[806,912,364,389]" italics="true" pageId="6" pageNumber="61">P. striata</emphasis>
</taxonomicName>
in the elytron of new one without paler flecks, from
<taxonomicName id="4C244D6CFFDB981071CCFE54D55CFE67" box="[484,583,404,429]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="sanzii">
<emphasis id="B950EAFDFFDB981071CCFE54D55CFE67" box="[484,583,404,429]" italics="true" pageId="6" pageNumber="61">P. sanzii</emphasis>
</taxonomicName>
in new ones antennal second segment shorter than third one. New species differs from
<taxonomicName id="4C244D6CFFDB9810727AFE7CD5EFFE1F" box="[594,756,444,469]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="latidentata" status="sp. nov.">
<emphasis id="B950EAFDFFDB9810727AFE7CD5EFFE1F" box="[594,756,444,469]" italics="true" pageId="6" pageNumber="61">P. latidentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2635786FFDB9810732AFE7BD47CFE1F" box="[770,871,443,469]" pageId="6" pageNumber="61" rank="species">
<emphasis id="B950EAFDFFDB9810732AFE7BD47CFE1F" bold="true" box="[770,871,443,469]" pageId="6" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
in mandibles not flattened at apex, posterior pair of dorsal head tubercles separated and elytral cells with 3 black macula on their margins, from both
<taxonomicName id="4C244D6CFFDB98107211FDCCD5ABFDEF" box="[569,688,524,549]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="renaria" status="sp. nov.">
<emphasis id="B950EAFDFFDB98107211FDCCD5ABFDEF" box="[569,688,524,549]" italics="true" pageId="6" pageNumber="61">P. renaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2635786FFDB98107292FDCBD400FDEF" box="[698,795,523,549]" pageId="6" pageNumber="61" rank="species">
<emphasis id="B950EAFDFFDB98107292FDCBD400FDEF" bold="true" box="[698,795,523,549]" pageId="6" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C244D6CFFDB98107371FDCCD4D4FDEF" box="[857,975,524,549]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="clavata" status="sp. nov.">
<emphasis id="B950EAFDFFDB98107371FDCCD4D4FDEF" box="[857,975,524,549]" italics="true" pageId="6" pageNumber="61">P. clavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2635786FFDB981073F2FDCBD321FDEF" box="[986,1082,523,549]" pageId="6" pageNumber="61" rank="species">
<emphasis id="B950EAFDFFDB981073F2FDCBD321FDEF" bold="true" box="[986,1082,523,549]" pageId="6" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
in the pedicel of
<taxonomicName id="4C244D6CFFDB9810753DFDCCD68AFD87" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="tuberculosa">
<emphasis id="B950EAFDFFDB9810753DFDCCD68AFD87" italics="true" pageId="6" pageNumber="61">P. tuberculosa</emphasis>
</taxonomicName>
distinctly shorter than third antennomere.
</paragraph>
</subSubSection>
<caption id="DF5B6667FFD898137120F88DD39CF8AF" box="[264,1159,1869,1893]" httpUri="https://zenodo.org/record/174109/files/figure.png" pageId="5" pageNumber="60" targetBox="[284,1309,316,1852]" targetPageId="5">
<paragraph id="8B9B36EFFFD898137120F88DD39CF8AF" blockId="5.[264,1159,1869,1893]" box="[264,1159,1869,1893]" pageId="5" pageNumber="60">
<emphasis id="B950EAFDFFD898137120F88DD6AFF8AF" bold="true" box="[264,436,1869,1893]" pageId="5" pageNumber="60">FIGURES 89.</emphasis>
<taxonomicName id="4C244D6CFFD898137193F88ED58BF8AF" box="[443,656,1870,1893]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="latidentata" status="sp. nov.">
<emphasis id="B950EAFDFFD898137193F88ED58BF8AF" box="[443,656,1870,1893]" italics="true" pageId="5" pageNumber="60">Priacma latidentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2635786FFD8981372BFF88DD5F6F8AF" box="[663,749,1869,1893]" pageId="5" pageNumber="60" rank="species">
<emphasis id="B950EAFDFFD8981372BFF88DD5F6F8AF" bold="true" box="[663,749,1869,1893]" pageId="5" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype. 8— head, 9—metaventrite.
</paragraph>
</caption>
<subSubSection id="C33E6564FFDB98107110FD9BD36FFA2C" pageId="6" pageNumber="61" type="description">
<paragraph id="8B9B36EFFFDB98107110FD9BD340FD54" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="61">
<emphasis id="B950EAFDFFDB98107110FD9BD6D4FDBF" bold="true" box="[312,463,603,629]" pageId="6" pageNumber="61">Description.</emphasis>
Body length
<quantity id="4CDC9B0AFFDB981072A8FD9CD5EAFDBC" box="[640,753,604,630]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="61" unit="mm" value="10.5">10.5 mm</quantity>
, body width
<quantity id="4CDC9B0AFFDB981073B2FD9CD4E6FDBC" box="[922,1021,604,630]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="61" unit="mm" value="2.5">2.5 mm</quantity>
, elytron length
<quantity id="4CDC9B0AFFDB981074ECFD9CD233FDBC" box="[1220,1320,604,630]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="61" unit="mm" value="7.5">7.5 mm</quantity>
. Medium­sized and subcylindrical beetle, covered with tubercles (
<figureCitation id="131F2A6AFFDB9810742DFD44D356FD54" box="[1029,1101,644,670]" captionStart="FIGURES 1 4" captionStartId="3.[264,382,1795,1819]" captionTargetBox="[314,1277,326,1768]" captionTargetId="figure@3.[314,1277,326,1769]" captionTargetPageId="3" captionText="FIGURES 1 4. Habiti of new Priacma species, photographs of rock impressions. 1 — Priacma latidentata sp. nov., 2 — Priacma tuberculosa sp. nov., 3 — Priacma clavata sp. nov., 4 — Priacma renaria sp. nov." httpUri="https://zenodo.org/record/174107/files/figure.png" pageId="6" pageNumber="61">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8B9B36EFFFDB98107110FD6CD364FCF4" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="61">
Head (
<figureCitation id="131F2A6AFFDB981071AEFD6CD6C7FD0C" box="[390,476,684,710]" captionStart="FIGURES 13 14" captionStartId="7.[264,382,1546,1570]" captionTargetBox="[264,1323,1129,1522]" captionTargetId="figure@7.[264,1323,1129,1522]" captionTargetPageId="7" captionText="FIGURES 13 14. Priacma tuberculosa sp. nov., holotype. 13 — head, 14 — elytral cells." httpUri="https://zenodo.org/record/174111/files/figure.png" pageId="6" pageNumber="61">Fig. 13</figureCitation>
) a little wider than long, bearing two pairs of tubercles, anterior pair of tubercles at base of antennae, conical, small, posterior ones circular, separated from each other, larger than anterior pair; eyes medium­sized; apex of mandibles not flattened, bidentate in horizontal cutting surface, cervical constriction unconspicuous.
</paragraph>
<paragraph id="8B9B36EFFFDB98107110FC8CD458FC7C" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="61">
Antennae filiform, incomplete (with 10 visible segments), less than half as long as entire insect, scape slightly oblong, pedicel 0.5 times as long as third antennomere, following segments homonomous (
<figureCitation id="131F2A6AFFDB9810728EFC5CD412FC7C" box="[678,777,924,950]" captionStart="FIGURES 10 12" captionStartId="7.[264,382,1017,1041]" captionTargetBox="[271,1322,321,974]" captionTargetId="figure@7.[264,1323,312,992]" captionTargetPageId="7" captionText="FIGURES 10 12. Priacma tuberculosa sp. nov., holotype. 10 — dorsal view, 11 — ventral view, 12 — outline of elytral cells." httpUri="https://zenodo.org/record/174110/files/figure.png" pageId="6" pageNumber="61">Figs. 10</figureCitation>
,
<figureCitation id="131F2A6AFFDB9810733DFC5CD42EFC7C" box="[789,821,924,950]" captionStart="FIGURES 13 14" captionStartId="7.[264,382,1546,1570]" captionTargetBox="[264,1323,1129,1522]" captionTargetId="figure@7.[264,1323,1129,1522]" captionTargetPageId="7" captionText="FIGURES 13 14. Priacma tuberculosa sp. nov., holotype. 13 — head, 14 — elytral cells." httpUri="https://zenodo.org/record/174111/files/figure.png" pageId="6" pageNumber="61">13</figureCitation>
).
</paragraph>
<paragraph id="8B9B36EFFFDB98107110FC04D30EFBE4" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="61">Pronotum transverse, wider than head, narrowed towards base, 0.8 times as wide as long at posterior edge, anterior angles sharp, extending, without propleuron, disc of pronotum bearing two circular elevations; scutellum subtriangular.</paragraph>
<paragraph id="8B9B36EFFFDB98107110FBFCD369FB6C" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="61">
Elytra about 1.5 times as wide as prothorax, 4 times as long as wide, with 9 rows of cells, elytral cells polygonal, with 3 black macula on their margins (
<figureCitation id="131F2A6AFFDB98107455FBA4D3F3FBB4" box="[1149,1256,1124,1150]" captionStart="FIGURES 10 12" captionStartId="7.[264,382,1017,1041]" captionTargetBox="[271,1322,321,974]" captionTargetId="figure@7.[264,1323,312,992]" captionTargetPageId="7" captionText="FIGURES 10 12. Priacma tuberculosa sp. nov., holotype. 10 — dorsal view, 11 — ventral view, 12 — outline of elytral cells." httpUri="https://zenodo.org/record/174110/files/figure.png" pageId="6" pageNumber="61">Figs. 12</figureCitation>
,
<figureCitation id="131F2A6AFFDB981074D4FBA4D207FBB4" box="[1276,1308,1124,1150]" captionStart="FIGURES 13 14" captionStartId="7.[264,382,1546,1570]" captionTargetBox="[264,1323,1129,1522]" captionTargetId="figure@7.[264,1323,1129,1522]" captionTargetPageId="7" captionText="FIGURES 13 14. Priacma tuberculosa sp. nov., holotype. 13 — head, 14 — elytral cells." httpUri="https://zenodo.org/record/174111/files/figure.png" pageId="6" pageNumber="61">14</figureCitation>
), elongated in distal part of elytron, approximately 34 cells formed in a row.
</paragraph>
<paragraph id="8B9B36EFFFDB98107110FB74D3F1FA74" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="61">
Ventral surface (
<figureCitation id="131F2A6AFFDB981071D5FB74D54FFB04" box="[509,596,1204,1230]" captionStart="FIGURES 10 12" captionStartId="7.[264,382,1017,1041]" captionTargetBox="[271,1322,321,974]" captionTargetId="figure@7.[264,1323,312,992]" captionTargetPageId="7" captionText="FIGURES 10 12. Priacma tuberculosa sp. nov., holotype. 10 — dorsal view, 11 — ventral view, 12 — outline of elytral cells." httpUri="https://zenodo.org/record/174110/files/figure.png" pageId="6" pageNumber="61">Fig. 11</figureCitation>
) with gula rectangular, reaching posterior ridge of the head, widening posteriorly, genae widely separated ventrally. Prosternal process narrower than fore coxa, extending beyond coxae. Metaventrite oblong, 1.5 times as wide as long (at posterior margin); without longitudinal suture on metaventrite. Abdomen with 5 visible ventrites superimposing each other, narrowed from the base of fourth visible ventrite, first visible abdominal ventrite as long as last one, both of them longer than other visible ventrites, last visible ventrite 2.1 times as long as the previous one, its apex rounded.
</paragraph>
<paragraph id="8B9B36EFFFDB98107110FA0CD36FFA2C" blockId="6.[264,1325,284,1870]" box="[312,1140,1484,1510]" pageId="6" pageNumber="61">Legs with procoxae circular, small, protrochanter large, sub­triangular.</paragraph>
</subSubSection>
<subSubSection id="C33E6564FFDB98107110FA34D6A6F91C" pageId="6" pageNumber="61" type="materials_examined">
<paragraph id="8B9B36EFFFDB98107110FA34D540F994" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="61">
<typeStatus id="549F884DFFDB98107110FA34D6B1F9C4" box="[312,426,1524,1550]" pageId="6" pageNumber="61" type="holotype">
<emphasis id="B950EAFDFFDB98107110FA34D6B1F9C4" bold="true" box="[312,426,1524,1550]" pageId="6" pageNumber="61">Holotype</emphasis>
</typeStatus>
. Nearly complete adult, No.
<collectionCode id="ED35AE2AFFDB98107329FA34D45BF9C4" box="[769,832,1524,1550]" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Chonbuk National University" pageId="6" pageNumber="61">CNU</collectionCode>
CLB2005011 deposited in the Key Lab of Insect Evolution &amp; Environmental Changes, College of Life Science, Capital Normal University in Beijing,
<collectingCountry id="F333767FFFDB98107226F984D543F994" box="[526,600,1604,1630]" name="China" pageId="6" pageNumber="61">China</collectingCountry>
.
</paragraph>
<paragraph id="8B9B36EFFFDB98107110F9ACD6A6F91C" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="61">
<emphasis id="B950EAFDFFDB98107110F9ACD557F94C" bold="true" box="[312,588,1644,1670]" pageId="6" pageNumber="61">Locality and horizon.</emphasis>
Collected near Chaomidian Village, Beipiao City, Liaoning Province,
<collectingCountry id="F333767FFFDB98107157F954D6D2F964" box="[383,457,1684,1710]" name="China" pageId="6" pageNumber="61">China</collectingCountry>
; the Yixian Formation, Late Jurassic­Early Cretaceous (Late Tithonian to the Berriasian).
</paragraph>
</subSubSection>
<subSubSection id="C33E6564FFDB98107110F924D6EFF884" pageId="6" pageNumber="61" type="etymology">
<paragraph id="8B9B36EFFFDB98107110F924D6EFF884" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="61">
<emphasis id="B950EAFDFFDB98107110F924D6DFF934" bold="true" box="[312,452,1764,1790]" pageId="6" pageNumber="61">Etymology.</emphasis>
The specific epithet derives from the Latin noun
<emphasis id="B950EAFDFFDB9810742DF924D392F937" box="[1029,1161,1764,1789]" italics="true" pageId="6" pageNumber="61">tuberculum</emphasis>
, ­
<emphasis id="B950EAFDFFDB98107489F924D3B2F937" box="[1185,1193,1764,1789]" italics="true" pageId="6" pageNumber="61">i</emphasis>
, n (a small swelling, bump, or protuberance). It refers to tubercles densely covering the body and, especially, the head.
</paragraph>
</subSubSection>
</treatment>
</document>