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<document id="C154423D9DB2352F08CDCBAC8260F836" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE3B9FF2FFF75FD81BA78F85E" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madagaster barbata Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="191" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="191" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:title id="F0A4CF4ECF49BCAC821EE65BFC59D6CC">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name id="10FC2EEC6452823F5D9FE806BB510F7F" type="personal">
<mods:role id="76F7418DA029E5D6FB2B848166415105">
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<mods:namePart id="D139297AA7B6A0027144AE781DC34EB5">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="1ACF11D2520F3C08EDC6012D4775DBC7">text</mods:typeOfResource>
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<mods:title id="21A120E30416D874EF1195DB6E7BDDE3">Zootaxa</mods:title>
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<mods:part id="200C6084AF114C8545179B7F2ECEA745">
<mods:date id="9D27F517E543D7C46F807E543505A062">2017</mods:date>
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<mods:number id="4B7D29FBC36D33F722510BE93BFF1E4C">2017-11-03</mods:number>
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<mods:number id="11608B31EDA271989F794BC76D524000">4342</mods:number>
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<mods:classification id="A57F152E48E3C046D738DD06159C7B29">journal article</mods:classification>
<mods:identifier id="BCAB33F2D1CD8C7CE456BDDA4E110870" type="CLB-Dataset">31639</mods:identifier>
<mods:identifier id="9FFE270D05B3EA0CEBA3A3966DFB6138" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<treatment id="038587BBE3B9FF2FFF75FD81BA78F85E" ID-DOI="http://doi.org/10.5281/zenodo.5323738" ID-GBIF-Taxon="137233383" ID-Zenodo-Dep="5323738" LSID="urn:lsid:plazi:treatment:038587BBE3B9FF2FFF75FD81BA78F85E" httpUri="http://treatment.plazi.org/id/038587BBE3B9FF2FFF75FD81BA78F85E" lastPageNumber="191" pageId="190" pageNumber="191">
<subSubSection id="C3366526E3B9FF2FFF75FD81BB98FDDA" pageId="190" pageNumber="191" type="nomenclature">
<paragraph id="8B9336ADE3B9FF2FFF75FD81BB14FDBB" blockId="190.[151,676,548,607]" box="[151,552,548,574]" pageId="190" pageNumber="191">
<heading id="D0DB81C1E3B9FF2FFF75FD81BB14FDBB" bold="true" box="[151,552,548,574]" fontSize="11" level="1" pageId="190" pageNumber="191" reason="1">
<emphasis id="B958EABFE3B9FF2FFF75FD81BB14FDBB" bold="true" box="[151,552,548,574]" pageId="190" pageNumber="191">
<emphasis id="B958EABFE3B9FF2FFF75FD81B8ADFDBB" bold="true" box="[151,401,548,574]" italics="true" pageId="190" pageNumber="191">
<taxonomicName id="4C2C4D2EE3B9FF2FFF75FD81B8B6FDBB" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,394,548,574]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="barbata" status="sp. nov.">Madagaster barbata</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A26B57C4E3B9FF2FFE7AFD81BB14FDBB" box="[408,552,548,574]" pageId="190" pageNumber="191" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF75FDE3BB98FDDA" blockId="190.[151,676,548,607]" box="[151,676,582,607]" pageId="190" pageNumber="191">
<figureCitation id="13172A28E3B9FF2FFF75FDE3B9C3FDDA" box="[151,255,582,607]" captionStart="FIGURE 183" captionStartId="191.[151,250,1440,1462]" captionTargetBox="[340,1247,195,1408]" captionTargetId="figure@191.[340,1247,195,1408]" captionTargetPageId="191" captionText="FIGURE 183. Madagaster barbata, holotype, frontal view, habitus dorsal, lateral, and ventral." httpUri="https://zenodo.org/record/1041124/files/figure.png" pageId="190" pageNumber="191">Figs. 183</figureCitation>
(habitus), 185 (aedeagus), 301 (map)
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3B9FF2FFF75FD2BBA62FD6A" pageId="190" pageNumber="191" type="materials_examined">
<paragraph id="8B9336ADE3B9FF2FFF75FD2BBA62FD6A" blockId="190.[151,1437,654,2011]" pageId="190" pageNumber="191">
<emphasis id="B958EABFE3B9FF2FFF75FD2BB8FAFD22" bold="true" box="[151,454,654,679]" pageId="190" pageNumber="191">
<typeStatus id="5497880FE3B9FF2FFF75FD2BB9EEFD22" box="[151,210,654,679]" pageId="190" pageNumber="191">Type</typeStatus>
Material.
<typeStatus id="5497880FE3B9FF2FFEBBFD2BB8FAFD22" box="[345,454,654,679]" pageId="190" pageNumber="191" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<collectingRegion id="49E8F84FE3B9FF2FFDCDFD2BBBF2FD22" box="[559,718,654,679]" country="Madagascar" name="Fianarantsoa" pageId="190" pageNumber="191">
<emphasis id="B958EABFE3B9FF2FFDCDFD2BBBF2FD22" bold="true" box="[559,718,654,679]" pageId="190" pageNumber="191">Fianarantsoa</emphasis>
</collectingRegion>
, Andringitra NP: Riambavy R. (female) cascade hygropetric habitat, N -22° 8.761, E:46° 53.4, wet rock surfaces next to high waterfall, elev.
<quantity id="4CD49B48E3B9FF2FFBC1FD16BD47FD4F" box="[1059,1147,691,715]" metricMagnitude="3" metricUnit="m" metricValue="1.739" pageId="190" pageNumber="191" unit="m" value="1739.0">1739 m</quantity>
,
<date id="FF92106DE3B9FF2FFB65FD16BDD2FD4E" box="[1159,1262,690,715]" pageId="190" pageNumber="191" value="2006-05-08">8 v 2006</date>
, Bergsten
<emphasis id="B958EABFE3B9FF2FFA81FD11BCA0FD4E" box="[1379,1436,690,715]" italics="true" pageId="190" pageNumber="191">et al.</emphasis>
(P39) (
<collectionCode id="ED3DAE68E3B9FF2FFF06FD72B802FD6A" box="[228,318,727,751]" pageId="190" pageNumber="191">BMNH</collectionCode>
).
<emphasis id="B958EABFE3B9FF2FFEB3FD73B8EEFD6A" bold="true" box="[337,466,726,751]" pageId="190" pageNumber="191">
<typeStatus id="5497880FE3B9FF2FFEB3FD73B8F1FD6A" box="[337,461,726,751]" pageId="190" pageNumber="191" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same data as
<typeStatus id="5497880FE3B9FF2FFD90FD73BBE8FD6A" box="[626,724,726,751]" pageId="190" pageNumber="191" type="holotype">holotype</typeStatus>
(2
<collectionCode id="ED3DAE68E3B9FF2FFD1BFD72BA6EFD6A" box="[761,850,727,751]" pageId="190" pageNumber="191">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3B9FF2FFF25FD5FBDDCFC42" pageId="190" pageNumber="191" type="diagnosis">
<paragraph id="8B9336ADE3B9FF2FFF25FD5FBDDCFC42" blockId="190.[151,1437,654,2011]" pageId="190" pageNumber="191">
<emphasis id="B958EABFE3B9FF2FFF25FD5FB8E0FC96" bold="true" box="[199,476,762,787]" pageId="190" pageNumber="191">Differential Diagnosis.</emphasis>
Similar in habitus to
<taxonomicName id="4C2C4D2EE3B9FF2FFD1CFD59BAA0FC96" authorityName="Perkins" authorityYear="2017" box="[766,924,764,787]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="procarina">
<emphasis id="B958EABFE3B9FF2FFD1CFD59BAA0FC96" box="[766,924,764,787]" italics="true" pageId="190" pageNumber="191">M. procarina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D2EE3B9FF2FFC04FD59BD41FC96" authorityName="Perkins" authorityYear="2017" box="[998,1149,762,787]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B958EABFE3B9FF2FFC04FD59BD41FC96" box="[998,1149,762,787]" italics="true" pageId="190" pageNumber="191">M. bergsteni</emphasis>
</taxonomicName>
(
<figureCitation id="13172A28E3B9FF2FFB77FD5FBC36FC96" box="[1173,1290,762,787]" captionStart="FIGURE 179" captionStartId="187.[151,250,1464,1486]" captionText="FIGURE 179. Madagaster bergsteni, holotype, habitus dorsal, lateral, and ventral." pageId="190" pageNumber="191">Figs. 179</figureCitation>
,
<figureCitation id="13172A28E3B9FF2FFAC2FD5EBC70FC96" box="[1312,1356,763,787]" captionStart="FIGURE 180" captionStartId="188.[151,250,1601,1623]" captionText="FIGURE 180. Madagaster procarina, holotype, habitus dorsal, lateral, and ventral." pageId="190" pageNumber="191">180</figureCitation>
,
<figureCitation id="13172A28E3B9FF2FFA80FD5EBCACFC96" box="[1378,1424,763,787]" captionStart="FIGURE 183" captionStartId="191.[151,250,1440,1462]" captionTargetBox="[340,1247,195,1408]" captionTargetId="figure@191.[340,1247,195,1408]" captionTargetPageId="191" captionText="FIGURE 183. Madagaster barbata, holotype, frontal view, habitus dorsal, lateral, and ventral." httpUri="https://zenodo.org/record/1041124/files/figure.png" pageId="190" pageNumber="191">183</figureCitation>
). Differentiated from both species by the larger size (ca.
<quantity id="4CD49B48E3B9FF2FFCE8FCBABA4CFCB2" box="[778,880,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="190" pageNumber="191" unit="mm" value="2.4">2.40 mm</quantity>
) and, in males, by the strong ridge on the genae, the mesotibiae markedly widened and with fringe of setae on the anterior margin, and the metatibiae, which are arcuate toward the midline. The pronotal foveae are deeper than in the compared species. The very complex male genitalia of the three species share a hook-shaped process on the ventral surface of the main piece, but the parameres and processes of the distal part of the main piece differ markedly (
<figureCitation id="13172A28E3B9FF2FFC15FC0BBDA6FC42" box="[1015,1178,942,967]" captionStart="FIGURES 181 182" captionStartId="189.[151,264,1764,1787]" captionText="FIGURES 181 182. Madagaster, aedeagi of holotypes." pageId="190" pageNumber="191">Figs. 181, 182</figureCitation>
,
<figureCitation id="13172A28E3B9FF2FFB44FC0BBDE8FC42" box="[1190,1236,942,967]" captionStart="FIGURES 185 186" captionStartId="193.[151,264,1869,1892]" captionText="FIGURES 185 186. Madagaster, aedeagi of holotypes." pageId="190" pageNumber="191">185</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3B9FF2FFF25FC77B84BF832" pageId="190" pageNumber="191" type="description">
<paragraph id="8B9336ADE3B9FF2FFF25FC77B841FBB7" blockId="190.[151,1437,654,2011]" pageId="190" pageNumber="191">
<emphasis id="B958EABFE3B9FF2FFF25FC77B86DFC6E" bold="true" box="[199,337,978,1003]" pageId="190" pageNumber="191">Description</emphasis>
. Size:
<typeStatus id="5497880FE3B9FF2FFE41FC77BB39FC6E" box="[419,517,978,1003]" pageId="190" pageNumber="191" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.40/1.06; head width 0.61; pronotum 0.60/0.89, PA 0.66, PB 0.69; elytra 1.43/1.06. Dorsum and venter black, legs dark brown, ocelli light brown to testaceous.
</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF25FB9BBA07FB62" blockId="190.[151,1437,654,2011]" pageId="190" pageNumber="191">Entire dorsum dull, with very small, closely spaced tubercles, each with a short recumbent seta. Labrum bilobed, sparsely granulate and with sparse small setae except with longer setae along anterior margin. Labrum originating at anterior of slightly overhanging margin of clypeus. Labroclypeal suture moderately deep. Short, deep sulcus in front of each ocellus. Frons with deep concavity between ocelli. Maxillary palpi ca. as along as antenna. Antenna with penultimate antennomere larger than ultimate.</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF25FB56B838FAF2" blockId="190.[151,1437,654,2011]" pageId="190" pageNumber="191">Pronotum with two narrowly separated midlongitudinal fovea, anterior ca. twice as long as posterior; each side with two admedian and two sublateral foveae; posteroadmedian fovea oblique, deep; transverse fovea along base; pronotum slightly cordiform, lateral margin rounded except emarginate near base; postocular emarginations deep and wide.</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF25FA27BBC3FA67" blockId="190.[151,1437,654,2011]" pageId="190" pageNumber="191">Elytral suture costate in posterior. Each elytron with four non-interrupted costae; first costa (from suture) slightly curved in ca. middle ½ of length. Each costa with unilinear row of granules. Areas between costae with four irregular rows of granules. Elytral apices truncate.</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF25FA4ABAD6F9CA" blockId="190.[151,1437,654,2011]" pageId="190" pageNumber="191">Mentum and postmentum densely micropunctulate, sparsely pubescent; Genae micropunctulate, dull, concave behind cardo, with very strong, high posterior ridge, obliquely oriented on each side of midline. Subocular groove for antenna moderately deep and wide. Posterior margin of eye not carinate.</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF25F9FEB882F912" blockId="190.[151,1437,654,2011]" pageId="190" pageNumber="191">Prosternum with very low, almost absent, midlongitudinal carina in basal ½, carina stronger on intercoxal process, enlarged apically.</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF25F907BBEFF8A2" blockId="190.[151,1437,654,2011]" pageId="190" pageNumber="191">
Mesoventrite without tubercle on intercoxal process. Metaventrite densely micropunctulate and with very short and very dense vestiture; median depression oval, moderately deep (
<figureCitation id="13172A28E3B9FF2FFC43F963BD39F95A" box="[929,1029,1734,1759]" captionStart="FIGURE 183" captionStartId="191.[151,250,1440,1462]" captionTargetBox="[340,1247,195,1408]" captionTargetId="figure@191.[340,1247,195,1408]" captionTargetPageId="191" captionText="FIGURE 183. Madagaster barbata, holotype, frontal view, habitus dorsal, lateral, and ventral." httpUri="https://zenodo.org/record/1041124/files/figure.png" pageId="190" pageNumber="191">Fig. 183</figureCitation>
). First four abdominal ventrites and anterior margin of fifth with very dense, recumbent pubescence (plastron?). Last ventrite (male only) oval, flat, very slightly concave, and with sparse pubescence.
</paragraph>
<paragraph id="8B9336ADE3B9FF2FFF25F897B84BF832" blockId="190.[151,1437,654,2011]" pageId="190" pageNumber="191">
Protibia slightly widened apically. Mesotrochanter with small, sharply pointed tubercle. Mesotibia markedly widened and with fringe of setae on anterior margin. Hind legs rather elongate. Metatibia slender, slightly arcuate (away from midline), apex slightly thickened and with brush of setae. Female (
<quantity id="4CD49B48E3B9FF2FFBFAF8DEBDBEF817" box="[1048,1154,1915,1939]" metricMagnitude="-3" metricUnit="m" metricValue="2.09" pageId="190" pageNumber="191" unit="mm" value="2.09">2.09 mm</quantity>
; n=1) smaller than male. Female legs simple.
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3B9FF2FFF25F867BA78F85E" box="[199,836,1986,2011]" pageId="190" pageNumber="191" type="etymology">
<paragraph id="8B9336ADE3B9FF2FFF25F867BA78F85E" blockId="190.[151,1437,654,2011]" box="[199,836,1986,2011]" pageId="190" pageNumber="191">
<emphasis id="B958EABFE3B9FF2FFF25F867B872F85E" bold="true" box="[199,334,1986,2011]" pageId="190" pageNumber="191">Etymology.</emphasis>
Named in reference to the hairy mouthparts.
</paragraph>
</subSubSection>
</treatment>
</document>