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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1131.91235" ID-Pensoft-Pub="1313-2970-1131-173" ID-Pensoft-UUID="2D635B88B2DD5FD6BA7430BFAAC148F1" ID-ZooBank="E6519ACB2945462DBA95BB1B14404D62" ModsDocID="1313-2970-1131-173" checkinTime="1669268533807" checkinUser="pensoft" docAuthor="Li, Yun-Chun" docDate="2022" docId="CE52411FF7CD5528983C7DCCCF7D887E" docLanguage="en" docName="ZooKeys 1131: 173-195" docOrigin="ZooKeys 1131" docPubDate="2022-11-23" docSource="http://dx.doi.org/10.3897/zookeys.1131.91235" docTitle="Tyrannochthonius dongjiensis Li 2022, sp. nov." docType="treatment" docUuid="B395357D-20CA-4B95-8423-9BC09EF468B4" docUuidSource="ZooBank" docVersion="1" id="2D635B88B2DD5FD6BA7430BFAAC148F1" lastPageNumber="173" masterDocId="2D635B88B2DD5FD6BA7430BFAAC148F1" masterDocTitle="Five new troglobitic species of Tyrannochthonius (Arachnida, Pseudoscorpiones, Chthoniidae) from the Yunnan, Guizhou and Sichuan Provinces, China" masterLastPageNumber="195" masterPageNumber="173" pageNumber="173" updateTime="1669268533807" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five new troglobitic species of Tyrannochthonius (Arachnida, Pseudoscorpiones, Chthoniidae) from the Yunnan, Guizhou and Sichuan Provinces, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Yun-Chun</mods:namePart>
<mods:affiliation>College of Life Science, China West Normal University, Nanchong, Sichuan 637009, China</mods:affiliation>
<mods:nameIdentifier type="email">liyunchun2260@126.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-23</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1131</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>173</mods:start>
<mods:end>195</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1131.91235</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1131.91235</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1131-173</mods:identifier>
<mods:identifier type="ZooBank">E6519ACB2945462DBA95BB1B14404D62</mods:identifier>
<mods:identifier type="Pensoft-UUID">2D635B88B2DD5FD6BA7430BFAAC148F1</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:B395357D-20CA-4B95-8423-9BC09EF468B4" httpUri="http://treatment.plazi.org/id/CE52411FF7CD5528983C7DCCCF7D887E" lastPageNumber="173" pageId="0" pageNumber="173">
<subSubSection pageId="0" pageNumber="173" type="nomenclature">
<paragraph pageId="0" pageNumber="173">
<taxonomicName LSID="https://zoobank.org/B395357D-20CA-4B95-8423-9BC09EF468B4" authority="Li, 2022" authorityName="Li" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius dongjiensis" order="Pseudoscorpiones" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="dongjiensis" status="sp. nov.">Tyrannochthonius dongjiensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="173">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="173" type="description">
<paragraph pageId="0" pageNumber="173">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">Figs 1</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A, B Tyrannochthonius dongjiensis sp. nov., dorsal views A holotype male B paratype female C, D T. huaerensis sp. nov., dorsal views C holotype male D paratype female. Scale bar: 1.00 mm (A-D)." figureDoi="10.3897/zookeys.1131.91235.figure6" httpUri="https://binary.pensoft.net/fig/773437" pageId="0" pageNumber="173">, 6A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="173" type="type material">
<paragraph pageId="0" pageNumber="173">Type material.</paragraph>
<paragraph pageId="0" pageNumber="173">
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Holotype</emphasis>
male: China, Guizhou Province, Luodian County, Dongjia Town, Dongjia Village, Nameless Cave,
<geoCoordinate degrees="25" direction="north" minutes="38.53" orientation="latitude" precision="9" value="25.642166">25°38.53'N</geoCoordinate>
,
<geoCoordinate degrees="106" direction="east" minutes="54.67" orientation="longitude" precision="9" value="106.91116">106°54.67'E</geoCoordinate>
, 869 m a.s.l., 7 October 2019, Yun-Chun Li leg., in MCWNU (Ar-Ps-GZ-0055).
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Paratypes</emphasis>
: 2 males, 4 females, collected with the holotype, in MCWNU (Ar-Ps-GZ-0008); 5 males, 2 females, Guizhou Province, Pingtang County, Tangbian Town, Baima Cave,
<geoCoordinate degrees="25" direction="north" minutes="40" orientation="latitude" precision="1" seconds="6.13" value="25.66837">25°40'6.13&quot;N</geoCoordinate>
,
<geoCoordinate degrees="106" direction="east" minutes="45" orientation="longitude" precision="1" seconds="53.89" value="106.76497">106°45'53.89&quot;E</geoCoordinate>
, 870 m a.s.l., 6 October 2019, Yun-Chun Li leg., in MCWNU (Ar-Ps-GZ-0010).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="173" type="diagnosis">
<paragraph pageId="0" pageNumber="173">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="173">
Troglobiont habitus. This new species is distinguished from other members of the genus
<taxonomicName authorityName="J. C. Chamberlin" authorityYear="1929" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius" order="Pseudoscorpionida" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="173">Tyrannochthonius</emphasis>
</taxonomicName>
by the following combination of characters: carapace without eyes or eyespots, anterior margin with six setae; epistome absent; rallum composed of six blades; tergites I-IV with two setae; apex of coxa I with long and rounded anteromedial process, near the apex with a seta; chelal hand dorsal surface with chemosensory setae; fixed chelal finger with 24 or 25 teeth, movable chelal finger with 27-29 retrorse teeth. Pedipalpal femur (♂) 7.58-7.63
<normalizedToken originalValue="×">x</normalizedToken>
, (♀) 7.36-7.42
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad, length (♂) 0.91-0.95 mm, (♀) 1.03-1.07 mm; chela (♂) 7.88-7.90
<normalizedToken originalValue="×">x</normalizedToken>
, (♀) 7.06-7.10 longer than deep, length (♂) 1.25-1.28 mm, (♀) 1.20-1.24 mm; ratio movable chelal finger/chelal hand (♂) 1.86-1.90
<normalizedToken originalValue="×">x</normalizedToken>
, (♀) 1.88-1.93
<normalizedToken originalValue="×">x</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="173" type="etymology">
<paragraph pageId="0" pageNumber="173">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="173">Latinized adjective, derived from the village of Dongjia, located near the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="173" type="description">
<paragraph pageId="0" pageNumber="173">Description.</paragraph>
<paragraph pageId="0" pageNumber="173">
<emphasis bold="true" pageId="0" pageNumber="173">Adult male</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A, B Tyrannochthonius dongjiensis sp. nov., dorsal views A holotype male B paratype female C, D T. huaerensis sp. nov., dorsal views C holotype male D paratype female. Scale bar: 1.00 mm (A-D)." figureDoi="10.3897/zookeys.1131.91235.figure6" httpUri="https://binary.pensoft.net/fig/773437" pageId="0" pageNumber="173">6A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="173">
Pale yellow-orange, chelicera slightly darker, soft parts pale (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A, B Tyrannochthonius dongjiensis sp. nov., dorsal views A holotype male B paratype female C, D T. huaerensis sp. nov., dorsal views C holotype male D paratype female. Scale bar: 1.00 mm (A-D)." figureDoi="10.3897/zookeys.1131.91235.figure6" httpUri="https://binary.pensoft.net/fig/773437" pageId="0" pageNumber="173">6A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="173">
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Carapace</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1A</figureCitation>
): 1.26-1.30
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad, no eyes or eyespots; epistome absent; carapace surface smooth, lateral margins distinctly constricted posteriorly. With 18 setae arranged 6: 4: 4: 2: 2, anterolateral setae much shorter than others.
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Coxae</emphasis>
: manducatory process pointed, with two distal setae, one long and the other slightly shorter. Pedipalpal coxa with three setae, coxa I 3, II 4, III 5, IV 5; intercoxal tubercle absent. Apex of coxa I with long and rounded anteromedial process, near the apex with a seta (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1E</figureCitation>
); coxae II with nine terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for ~
<normalizedToken originalValue="½">1/2</normalizedToken>
their length (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1D</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Chelicera</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1B</figureCitation>
): 1.82-1.85
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad, hand with five setae and one lyrifissure dorsally, movable finger with one submedial seta. Cheliceral hand with moderate hispid granulation dorsally. Fixed finger with eight or nine teeth, distal one largest, decreasing in size proximally; movable finger with 12 or 13 teeth; galea absent. Serrula exterior with 22-25 blades. Rallum composed of six blades (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1C</figureCitation>
), distal blade weakly recumbent basally, with fine barbules and set apart from the other blades, the latter tightly grouped and with long pinnae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Pedipalp</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1H-J</figureCitation>
): all setae acuminate. Trochanter 1.56-1.61
<normalizedToken originalValue="×">x</normalizedToken>
, femur 7.58-7.63
<normalizedToken originalValue="×">x</normalizedToken>
, patella 2.73-2.76
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad and with three lyrifissures (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1H</figureCitation>
). Femur 2.22-2.31
<normalizedToken originalValue="×">x</normalizedToken>
longer than patella. Chela 7.88-7.90
<normalizedToken originalValue="×">x</normalizedToken>
, hand 2.63-2.66
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep; movable chelal finger 1.86-1.90
<normalizedToken originalValue="×">x</normalizedToken>
longer than hand. Chelal hand dorsal surface with a single row of five chemosensory setae between
<emphasis italics="true" pageId="0" pageNumber="173">esb</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="173">ib</emphasis>
/
<emphasis italics="true" pageId="0" pageNumber="173">isb</emphasis>
trichobothria; distal paraxial seta of hand not enlarged. Fingers straight in dorsal view (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1J</figureCitation>
). Fixed finger with 24 or 25 teeth, middle ones larger than those at both ends; movable finger with 27-29 retrorse teeth (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1I</figureCitation>
). Venom apparatus absent. Fixed chelal finger with eight trichobothria and movable finger with four,
<emphasis italics="true" pageId="0" pageNumber="173">ib</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="173">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis italics="true" pageId="0" pageNumber="173">eb</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="173">esb</emphasis>
, and
<emphasis italics="true" pageId="0" pageNumber="173">ist</emphasis>
forming a straight oblique row at base of fixed chelal finger;
<emphasis italics="true" pageId="0" pageNumber="173">it</emphasis>
slightly distal to
<emphasis italics="true" pageId="0" pageNumber="173">est</emphasis>
, situated subdistally;
<emphasis italics="true" pageId="0" pageNumber="173">et</emphasis>
slightly nearer to tip of fixed finger;
<emphasis italics="true" pageId="0" pageNumber="173">dx</emphasis>
situated distal to
<emphasis italics="true" pageId="0" pageNumber="173">et</emphasis>
;
<emphasis italics="true" pageId="0" pageNumber="173">sb</emphasis>
half-way between
<emphasis italics="true" pageId="0" pageNumber="173">st</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="173">b</emphasis>
;
<emphasis italics="true" pageId="0" pageNumber="173">b</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="173">t</emphasis>
situated subdistally,
<emphasis italics="true" pageId="0" pageNumber="173">t</emphasis>
situated at same level as
<emphasis italics="true" pageId="0" pageNumber="173">est.</emphasis>
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Opisthosoma</emphasis>
: tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy (I-XII): 2: 2: 2: 2: 3-4: 4: 4: 4: 4: 2: 0; sternal chaetotaxy (IV-XII): 12: 7: 7: 7: 7: 7: 7: 0: 2. Anterior genital operculum with ten setae, genital opening slit-like, with 14 or 15 setae on the right side and 18 on the left (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1K</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Legs</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1F, G</figureCitation>
): leg I: trochanter 1.00-1.03
<normalizedToken originalValue="×">x</normalizedToken>
, femur 7.63-7.66
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep and 1.91-1.97
<normalizedToken originalValue="×">x</normalizedToken>
longer than patella; patella 4.57-4.59
<normalizedToken originalValue="×">x</normalizedToken>
, tibia 4.33-4.37
<normalizedToken originalValue="×">x</normalizedToken>
, tarsus 13.40-13.44
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep. Leg IV: trochanter 0.88-0.96
<normalizedToken originalValue="×">x</normalizedToken>
, femoropatella 3.83-3.87
<normalizedToken originalValue="×">x</normalizedToken>
, tibia 6.89-6.92
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep, basitarsus 3.75-3.80
<normalizedToken originalValue="×longer">xlonger</normalizedToken>
than deep, with a basal tactile seta (TS = 0.24-0.25), telotarsus 15.2-15.5
<normalizedToken originalValue="×longer">xlonger</normalizedToken>
than deep and 2.53-2.55
<normalizedToken originalValue="×">x</normalizedToken>
longer than basitarsus, with a tactile seta near base (TS = 0.23-0.24). Arolia on legs I and IV shorter than claws.
</paragraph>
<caption doi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="173">
<emphasis bold="true" pageId="0" pageNumber="173">Figure 1.</emphasis>
<taxonomicName authorityName="Li" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius dongjiensis" order="Pseudoscorpionida" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="dongjiensis">
<emphasis italics="true" pageId="0" pageNumber="173">Tyrannochthonius dongjiensis</emphasis>
</taxonomicName>
sp. nov., holotype male (
<emphasis bold="true" pageId="0" pageNumber="173">A-K</emphasis>
) and paratype female (
<emphasis bold="true" pageId="0" pageNumber="173">L</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="173">A</emphasis>
carapace
<emphasis bold="true" pageId="0" pageNumber="173">B</emphasis>
left chelicera
<emphasis bold="true" pageId="0" pageNumber="173">C</emphasis>
rallum of left chelicera
<emphasis bold="true" pageId="0" pageNumber="173">D</emphasis>
coxal spines
<emphasis bold="true" pageId="0" pageNumber="173">E</emphasis>
process of left coxa I, ventral view
<emphasis bold="true" pageId="0" pageNumber="173">F</emphasis>
left leg I, lateral view
<emphasis bold="true" pageId="0" pageNumber="173">G</emphasis>
left leg IV, lateral view
<emphasis bold="true" pageId="0" pageNumber="173">H</emphasis>
palp (minus chela)
<emphasis bold="true" pageId="0" pageNumber="173">I</emphasis>
chela, retrolateral view
<emphasis bold="true" pageId="0" pageNumber="173">J</emphasis>
chela, dorsal view
<emphasis bold="true" pageId="0" pageNumber="173">K</emphasis>
male genital area
<emphasis bold="true" pageId="0" pageNumber="173">L</emphasis>
female genital area. Scale bars: 0.20 mm
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="173">
<emphasis bold="true" pageId="0" pageNumber="173">Adult female</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A, B Tyrannochthonius dongjiensis sp. nov., dorsal views A holotype male B paratype female C, D T. huaerensis sp. nov., dorsal views C holotype male D paratype female. Scale bar: 1.00 mm (A-D)." figureDoi="10.3897/zookeys.1131.91235.figure6" httpUri="https://binary.pensoft.net/fig/773437" pageId="0" pageNumber="173">6B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="173">Mostly the same as the holotype with the differences listed below.</paragraph>
<paragraph pageId="0" pageNumber="173">
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Carapace</emphasis>
: slightly longer than broad (1.13-1.15
<normalizedToken originalValue="×">x</normalizedToken>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Chelicera</emphasis>
: 2.30-2.33
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad.
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Pedipalp</emphasis>
: trochanter 1.80-1.86
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad, femur 7.36-7.42
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad, patella 2.63-2.70
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad, femur 2.45-2.49
<normalizedToken originalValue="×">x</normalizedToken>
longer than patella. Chela 7.06-7.10
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep, hand 2.41-2.46
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep; movable finger 1.88-1.93
<normalizedToken originalValue="×">x</normalizedToken>
longer than hand.
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Opisthosoma</emphasis>
: tergal chaetotaxy (I-XII): 2: 2: 2: 2: 4: 4: 4: 4: 4: 2: 2: 0; sternal chaetotaxy (IV-XII): 14: 12: 8: 7: 7: 9: 7: 0: 2. Anterior genital operculum with 9 + 14 setae on posterior margin (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tyrannochthonius dongjiensis sp. nov., holotype male (A-K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm" figureDoi="10.3897/zookeys.1131.91235.figure1" httpUri="https://binary.pensoft.net/fig/773432" pageId="0" pageNumber="173">1L</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="173">
<emphasis bold="true" italics="true" pageId="0" pageNumber="173">Dimensions</emphasis>
(mm, length/width or, in the case of the legs, chela, and chelal hand, length/depth).
</paragraph>
<paragraph pageId="0" pageNumber="173">Males (females in parentheses): body length 2.24-2.30 (2.49-2.56). Carapace 0.54-0.57/0.43-0.45 (0.53-0.55/0.47-0.48). Pedipalp: trochanter 0.25-0.28/0.16-0.18 (0.27-0.29/0.15-0.17), femur 0.91-0.95/0.12-0.14 (1.03-1.07/0.14-0.16), patella 0.41-0.44/0.15-0.17 (0.42-0.44/0.16-0.18), hand 0.42-0.45/0.16-0.17 (0.41-0.44/0.17-0.18), length of movable chelal finger 0.78-0.80 (0.77-0.79), chela 1.25-1.28/0.16-0.17 (1.20-1.24/0.17-0.18). Chelicera: 0.51-0.53/0.28-0.29 (0.53-0.55/0.23-0.24). Leg I: trochanter 0.15-0.17/0.15-0.16 (0.15-0.17/0.14-0.16), femur 0.61-0.64/0.08-0.09 (0.62-0.65/0.08-0.09), patella 0.32-0.35/0.07-0.08 (0.31-0.34/0.07-0.08), tibia 0.26-0.27/0.06-0.07 (0.27-0.29/0.06-0.07), tarsus 0.67-0.69/0.05-0.06 (0.66-0.68/0.05-0.06). Leg IV: trochanter 0.15-0.17/0.17-0.18 (0.17-0.19/0.14-0.16), femoropatella 0.88-0.92/0.23-0.25 (0.90-0.93/0.22-0.24), tibia 0.62-0.65/0.09-0.10 (0.63-0.65/0.09-0.10), basitarsus 0.30-0.32/0.08-0.09 (0.29-0.31/0.08-0.09), telotarsus 0.76-0.79/0.05-0.06 (0.79-0.82/0.06-0.07).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="173" type="distribution">
<paragraph pageId="0" pageNumber="173">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="173">China (Guizhou).</paragraph>
</subSubSection>
</treatment>
</document>