treatments-xml/data/8B/F8/55/8BF85597960F59559C2FD2442E216CE1.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1014.60426" ID-GBIF-Dataset="97cd82f3-4a3a-43ca-a68f-92a5a2074d07" ID-PMC="PMC7875960" ID-Pensoft-Pub="1313-2970-1014-1" ID-Pensoft-UUID="62961664CAED5F15B9C8A9990D7D388D" ID-PubMed="33603534" ModsDocID="1313-2970-1014-1" checkinTime="1612410909014" checkinUser="pensoft" docAuthor="Achterberg, Cornelis van" docDate="2021" docId="8BF85597960F59559C2FD2442E216CE1" docLanguage="en" docName="ZooKeys 1014: 1-118" docOrigin="ZooKeys 1014" docPubDate="2021-02-03" docSource="http://dx.doi.org/10.3897/zookeys.1014.60426" docTitle="Phanerotoma hellyeri Achterberg 2021, sp. nov." docType="treatment" docVersion="4" id="62961664CAED5F15B9C8A9990D7D388D" lastPageNumber="1" masterDocId="62961664CAED5F15B9C8A9990D7D388D" masterDocTitle="The tribe Phanerotomini (Hymenoptera, Braconidae, Cheloninae) of the Arabian Peninsula, with special reference to the United Arab Emirates and Yemen" masterLastPageNumber="118" masterPageNumber="1" pageNumber="1" updateTime="1668149759257" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The tribe Phanerotomini (Hymenoptera, Braconidae, Cheloninae) of the Arabian Peninsula, with special reference to the United Arab Emirates and Yemen</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Achterberg, Cornelis van</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6495-4853</mods:nameIdentifier>
<mods:affiliation>Naturalis Biodiversity Center, Postbus 9517, 2300 RA Leiden, the Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">kees@vanachterberg.org</mods:nameIdentifier>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-03</mods:number>
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<mods:detail type="volume">
<mods:number>1014</mods:number>
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<mods:start>1</mods:start>
<mods:end>118</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1014.60426</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1014.60426</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1014-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">62961664CAED5F15B9C8A9990D7D388D</mods:identifier>
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<treatment ID-GBIF-Taxon="177209562" LSID="urn:lsid:zoobank.org:act:A84BE796-CCB6-4FDB-8BE7-A46C5DD97503" httpUri="http://treatment.plazi.org/id/8BF85597960F59559C2FD2442E216CE1" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/A84BE796-CCB6-4FDB-8BE7-A46C5DD97503" authority="Achterberg, 2021" authorityName="Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Phanerotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanerotoma hellyeri" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hellyeri" status="sp. nov.">Phanerotoma hellyeri</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 152–155" captionStartId="F23" captionText="Figures 152 - 155. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype (but 153 - 155 of ♂, paratype) 152 habitus lateral 153 antenna 154 middle femur and tibia lateral 155 hind femur and tibia lateral." figureDoi="10.3897/zookeys..60426.figures152-155" httpUri="https://binary.pensoft.net/fig/504646" pageId="0" pageNumber="1">Figs 152-155</figureCitation>
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">, 156-166</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
, ♀ (RMNH), &quot; 
<emphasis bold="true" pageId="0" pageNumber="1">United Arab Emirates</emphasis>
, Sharjah Desert Park (2517), light trap, 20.x.-8.xi.2005, 
<geoCoordinate degrees="25" direction="north" minutes="17" orientation="latitude" precision="925" value="25.283333">25°17'N</geoCoordinate>
, 
<geoCoordinate degrees="55" direction="east" minutes="42" orientation="longitude" precision="925" value="55.7">55°42'E</geoCoordinate>
, A. v. Harten, 
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: 2♀, 3♂: Same data as holotype; 3♀: Idem, 29.iii.-6.iv.2005; 2♀, 1♂: Idem, 6-13.iv.2005; 1♀, 2♂: Idem, 13-23.iv.2005; 1♀: Idem, 25.ii.-25.iii.2006; 1♀: Idem, 10.xi.2004; 1♀: Idem, 30.iv.-7.v.2005; 2♀: Idem, 23-30.iv.2005; 1♀: Idem, 21-29.iii.2005; 1♀: Idem, 1.ii.-14.iii.2005; 2♀, 1♂: &quot;United Arab Emirates, SSW of ad-Dhaid (6154), light tr[ap], 24-30.v.2006, 
<geoCoordinate degrees="25" direction="north" minutes="09" orientation="latitude" precision="925" value="25.15">25°09'N</geoCoordinate>
, 
<geoCoordinate degrees="55" direction="east" minutes="48" orientation="longitude" precision="925" value="55.8">55°48'E</geoCoordinate>
, A. v. Harten, 
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 2♀: &quot;United Arab Emirates, NARC near Sweihan (1410), light trap, 1.ii.-14.iii.2005, 
<geoCoordinate degrees="24" direction="north" minutes="24" orientation="latitude" precision="925" value="24.4">24°24'N</geoCoordinate>
, 
<geoCoordinate degrees="55" direction="east" minutes="26" orientation="longitude" precision="925" value="55.433334">55°26'E</geoCoordinate>
, A. v. Harten, 
<normalizedToken originalValue="RMNH’05”">RMNH'05&quot;</normalizedToken>
; 6♀, 2♂: Idem, 28.iii.-2.iv.2005; 24♀, 2♂: Idem, 9-20.iv.2005; 22♀, 3♂: Idem, 20-30.iv.2005; 1♂: Idem, 14-28.iii.2005; 1♀: &quot;United Arab Emirates, al-Ajban (6426), Malaise tr[ap], 25.v.-26.vi.2006, 
<geoCoordinate degrees="24" direction="north" minutes="36" orientation="latitude" precision="925" value="24.6">24°36'N</geoCoordinate>
, 
<geoCoordinate degrees="55" direction="east" minutes="01" orientation="longitude" precision="925" value="55.016666">55°01'E</geoCoordinate>
, A. v. Harten, 
<normalizedToken originalValue="RMNH’07”">RMNH'07&quot;</normalizedToken>
; 3♀, 1♂: Idem, 22.x.-9.xi.2005; 3♀, 3♂: Idem, 7-28.xii.2006; 6♀: Idem, 17.iv.-29.v.2006; 3♀, 1♂: Idem, 27.v.-26.vi.2006; 1♂: Idem, 17.x.-9.xi.2005; 1♂: Idem, 12-19.vi.2006; 2♂: &quot;United Arab Emirates, Fujairah (1314), light tr[ap], 13-19.iv.2005, 
<geoCoordinate degrees="25" direction="north" minutes="08" orientation="latitude" precision="925" value="25.133333">25°08'N</geoCoordinate>
, 
<geoCoordinate degrees="56" direction="east" minutes="21" orientation="longitude" precision="925" value="56.35">56°21'E</geoCoordinate>
, A. v. Harten, 
<normalizedToken originalValue="RMNH’05”">RMNH'05&quot;</normalizedToken>
; 1♂: Idem, 2-13.v.2005; 6♂: Idem, 2.v.-5.vi.2005; 1♀, 1♂: Idem, 19.iv.-2.v.2005; &quot;; 1♀: Idem, 5-24.iii.2005; 2♀, 2♂: &quot;United Arab Emirates, Sharjah (2279), light trap, 30.vi.-21.vii.2005, 
<geoCoordinate degrees="25" direction="north" minutes="17" orientation="latitude" precision="925" value="25.283333">25°17'N</geoCoordinate>
, 
<geoCoordinate degrees="55" direction="east" minutes="42" orientation="longitude" precision="925" value="55.7">55°42'E</geoCoordinate>
, A. v. Harten, 
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 1♂: &quot;United Arab Emirates, Sharjah x Khor Kalba (6308-6311), light trap, [
<geoCoordinate degrees="24" direction="north" minutes="59" orientation="latitude" precision="925" value="24.983334">24°59'N</geoCoordinate>
, 
<geoCoordinate degrees="56" direction="east" minutes="09" orientation="longitude" precision="925" value="56.15">56°09'E</geoCoordinate>
,] 31.v.-7.vi.2006, A. v. Harten, 
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 1♂: Idem, 16-31.i.2006; 1♂: &quot;United Arab Emirates, Hatta (6398), at light, 17-24.viii.2006, 
<geoCoordinate degrees="24" direction="north" minutes="49" orientation="latitude" precision="925" value="24.816668">24°49'N</geoCoordinate>
, 
<geoCoordinate degrees="56" direction="east" minutes="07" orientation="longitude" precision="925" value="56.116665">56°07'E</geoCoordinate>
, A. v. Harten, 
<normalizedToken originalValue="RMNH’06”">RMNH'06&quot;</normalizedToken>
; 2♀: Idem, 21.vi.-19.vii.2006; 1♀: &quot; 
<emphasis bold="true" pageId="0" pageNumber="1">Yemen</emphasis>
(no. 3111), 
<normalizedToken originalValue="Ta’izz">Ta'izz</normalizedToken>
, light trap, 26-28.v.1998, A. van Harten &amp; Ahmad Ahwad, 
<normalizedToken originalValue="RMNH’98”">RMNH'98&quot;</normalizedToken>
; 1♀: Idem, ix.2000; 1♀: Idem, vii.2002; 1♀: &quot;Yemen (6141), Al Kowd, light trap, 16-20.viii.2001, A. van Harten &amp; S. Al Haruri, 
<normalizedToken originalValue="RMNH’02”">RMNH'02&quot;</normalizedToken>
; 1♀: Idem, 27-31.vii.2001; 1♀: &quot;Yemen (7501), Al Kadan, light trap, i.2003, A. v. Harten &amp; T. Abdul-Haq, 
<normalizedToken originalValue="RMNH’03”">RMNH'03&quot;</normalizedToken>
; 2♀: Idem, v.2002; 1♀: Idem, x.2001; 1♀: &quot;Yemen (6158), Al Lahima, 17.ix.-14.xi.2001, Mal[aise] trap, A. v. Harten, 
<normalizedToken originalValue="RMNH’02”">RMNH'02&quot;</normalizedToken>
.
</paragraph>
<caption doi="10.3897/zookeys..60426.figures152-155" httpUri="https://binary.pensoft.net/fig/504646" pageId="0" pageNumber="1" start="Figures 152–155" startId="F23">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 152-155.</emphasis>
<taxonomicName authorityName="Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Phanerotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanerotoma hellyeri" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hellyeri">
<emphasis italics="true" pageId="0" pageNumber="1">Phanerotoma hellyeri</emphasis>
</taxonomicName>
van Achterberg, sp. nov., ♀, holotype (but 
<emphasis bold="true" pageId="0" pageNumber="1">153-155</emphasis>
of ♂, paratype) 
<emphasis bold="true" pageId="0" pageNumber="1">152</emphasis>
habitus lateral 
<emphasis bold="true" pageId="0" pageNumber="1">153</emphasis>
antenna 
<emphasis bold="true" pageId="0" pageNumber="1">154</emphasis>
middle femur and tibia lateral 
<emphasis bold="true" pageId="0" pageNumber="1">155</emphasis>
hind femur and tibia lateral.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Robust species with twelfth-16th (counted from apex of antenna) antennal segments of ♀ widened (compared to more basal segments) and ventrally flattened, 13th segment from apex of antenna of ♀ as long as wide (Figs 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">165</figureCitation>
, 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">166</figureCitation>
) and eighth-tenth apical segments moniliform, stocky, matt or slightly shiny (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">166</figureCitation>
); stemmaticum yellow, more or less infuscate subapically; eye 2.0-2.1 
<normalizedToken originalValue="×">x</normalizedToken>
as wide as median width of temple in lateral view (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">164</figureCitation>
); POL of ♀ slightly less than width of posterior ocellus; frons with median carina anteriorly; second submarginal cell somewhat longer than in 
<taxonomicName family="Braconidae" kingdom="Animalia" lsidName="P. mesocellata" order="Hymenoptera" pageId="0" pageNumber="1" rank="species" species="mesocellata">
<emphasis italics="true" pageId="0" pageNumber="1">P. mesocellata</emphasis>
</taxonomicName>
; parastigma usually largely yellow, rarely infuscate; vein 1-M (as usually parastigma) slightly darker than yellow M+CU1 of fore wing; blister of middle tibia medium-sized; hind femur and tibia rather stout (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">160</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1" start="Figures 156–166" startId="F24">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 156-166.</emphasis>
<taxonomicName authorityName="Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Phanerotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanerotoma hellyeri" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hellyeri">
<emphasis italics="true" pageId="0" pageNumber="1">Phanerotoma hellyeri</emphasis>
</taxonomicName>
van Achterberg, sp. nov., ♀, holotype 
<emphasis bold="true" pageId="0" pageNumber="1">156</emphasis>
fore wing 
<emphasis bold="true" pageId="0" pageNumber="1">157</emphasis>
mesosoma dorsal 
<emphasis bold="true" pageId="0" pageNumber="1">158</emphasis>
first-third metasomal tergites dorsal 
<emphasis bold="true" pageId="0" pageNumber="1">159</emphasis>
metasoma lateral 
<emphasis bold="true" pageId="0" pageNumber="1">160</emphasis>
hind leg lateral 
<emphasis bold="true" pageId="0" pageNumber="1">161</emphasis>
mandible ventral 
<emphasis bold="true" pageId="0" pageNumber="1">162</emphasis>
head dorsal 
<emphasis bold="true" pageId="0" pageNumber="1">163</emphasis>
head anterior 
<emphasis bold="true" pageId="0" pageNumber="1">164</emphasis>
head lateral 
<emphasis bold="true" pageId="0" pageNumber="1">165</emphasis>
antenna lateral 
<emphasis bold="true" pageId="0" pageNumber="1">166</emphasis>
apical half of antenna lateral.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Female, holotype, length of body (excluding ovipositor) 4.2 mm; antenna 3.0 mm; fore wing 3.1 mm; visible part of ovipositor sheath 0.3 mm (0.1 mm erect setose).</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Width 1.6 
<normalizedToken originalValue="×">x</normalizedToken>
median length in anterior view and part of head above eye in lateral view 0.2 
<normalizedToken originalValue="×">x</normalizedToken>
height of eye (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">164</figureCitation>
); antenna with 23 cylindrical segments and as long as fore wing, 10 apical antennal segments small, rather serrate and moniliform (Figs 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">165</figureCitation>
, 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">166</figureCitation>
), with short bristles apically and apical segment with rather long spine, third, fourth and penultimate segments 3.0, 2.6 and 1.3 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide in lateral view, respectively; area of stemmaticum rugulose; OOL: diameter of posterior ocellus: POL = 20: 10: 9; length of eye 3.8 
<normalizedToken originalValue="×">x</normalizedToken>
temple in dorsal view (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">162</figureCitation>
); frons with median carina (V-shaped dorsally: Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">162</figureCitation>
), smooth antero-medially, posteriorly with curved rugae and finely rugose laterally; vertex rugose and with satin sheen, but posteriorly transversely rugulose; temple rugose but coriaceous near eye, convex, parallel-sided in lateral view and with satin sheen, directly narrowed behind eyes; face transversely rugose laterally, rugulose and with obsolescent median bump and rather shiny; clypeus smooth (except punctulation), shiny and as wide as minimum width of face, intertentorial distance 3.8 
<normalizedToken originalValue="×">x</normalizedToken>
minimum width between clypeus and eye (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">163</figureCitation>
), long erect setose and with 3 distinct blunt teeth medio-ventrally (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">161</figureCitation>
); eye large, strongly convex and in lateral view twice wider than temple (measured medially; Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">164</figureCitation>
), in anterior view its height 0.9 
<normalizedToken originalValue="×">x</normalizedToken>
minimum width of face (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">163</figureCitation>
); upper condyle of mandible above lower level of eyes (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">163</figureCitation>
); malar space rugulose, with satin sheen and 0.5 
<normalizedToken originalValue="×">x</normalizedToken>
as long as basal width of mandible; lower tooth of mandible half as long as apical tooth, robust (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">161</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
(Figs 
<figureCitation captionStart="Figures 152–155" captionStartId="F23" captionText="Figures 152 - 155. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype (but 153 - 155 of ♂, paratype) 152 habitus lateral 153 antenna 154 middle femur and tibia lateral 155 hind femur and tibia lateral." figureDoi="10.3897/zookeys..60426.figures152-155" httpUri="https://binary.pensoft.net/fig/504646" pageId="0" pageNumber="1">152</figureCitation>
, 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">157</figureCitation>
). Length1.4 
<normalizedToken originalValue="×">x</normalizedToken>
its width in lateral view; side of pronotum reticulate-punctate; posteriorly propleuron bulging near central groove; mesosternum smooth and shiny; mesoscutum densely reticulate-rugose, with satin sheen, notauli anteriorly impressed; scutellum flat, finely punctate-rugose; scutellar sulcus medium-sized, with twelve carinae (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">157</figureCitation>
); metanotum with median carina and medio-posterior tooth, its posterior border finely serrate; propodeum coarsely reticulate-rugose, dorsal face short, without transverse or median carinae, and latero-posteriorly weakly tuberculate. 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Wings</emphasis>
.
</emphasis>
Fore wing 2.7 
<normalizedToken originalValue="×">x</normalizedToken>
longer than its maximum width; 1-R1 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
as long as pterostigma; distance between wing apex and marginal cell apex 0.25 
<normalizedToken originalValue="×">x</normalizedToken>
length of 1-R1; r issued far beyond middle of pterostigma and 0.2 
<normalizedToken originalValue="×">x</normalizedToken>
3-SR; 2-SR distinctly curved and subparallel with posterior margin of pterostigma (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">156</figureCitation>
); SR1 curved; m-cu interstitial; parastigma large; 1-CU1 0.3 
<normalizedToken originalValue="×">x</normalizedToken>
as long as vein 2-CU1, cu-a 1.3 
<normalizedToken originalValue="×">x</normalizedToken>
1-CU1, strongly inclivous; r:3-SR:SR1 = 5:22:51; 2-SR:3-SR:r-m = 27:22:7; r-m reclivous; 2-M oblique and slightly curved (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">156</figureCitation>
). Hind wing: M+CU:1-M:1r-m = 21:19:10. 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Legs</emphasis>
.
</emphasis>
Hind femur matt, 3.2 
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide and robust; hind tibia swollen (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">161</figureCitation>
); middle tibia with distinct ivory blister; inner spur of middle tibia 0.5 
<normalizedToken originalValue="×">x</normalizedToken>
its basitarsus; hind coxa mostly smooth, but dorsally partly superficially granulate and rather shiny.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Metasoma</emphasis>
(Figs 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">158</figureCitation>
, 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">159</figureCitation>
). Oval in dorsal view, 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide and 1.1 
<normalizedToken originalValue="×">x</normalizedToken>
as long as mesosoma; first and second tergites densely and coarsely longitudinally rugose; second metasomal suture rather wide and slightly curved; third tergite 1.6 
<normalizedToken originalValue="×">x</normalizedToken>
longer than second tergite and laterally curved, convex medially, rounded posteriorly in dorsal view (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">158</figureCitation>
), obtuse posteriorly in lateral view (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">159</figureCitation>
), densely reticulate-rugose and with satin sheen (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">158</figureCitation>
), lateral lamella narrow, wide latero-apically and medio-apically; ovipositor sheath narrow, apically somewhat widened and darkened (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">159</figureCitation>
), its visible and setose part 0.1 
<normalizedToken originalValue="×">x</normalizedToken>
as long as fore wing and 0.2 
<normalizedToken originalValue="×">x</normalizedToken>
metasomal carapace, and with erect setae; hypopygium of ♀ with short widely triangular and up curved apical protuberance (Fig. 
<figureCitation captionStart="Figures 156–166" captionStartId="F24" captionText="Figures 156 - 166. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype 156 fore wing 157 mesosoma dorsal 158 first-third metasomal tergites dorsal 159 metasoma lateral 160 hind leg lateral 161 mandible ventral 162 head dorsal 163 head anterior 164 head lateral 165 antenna lateral 166 apical half of antenna lateral." figureDoi="10.3897/zookeys..60426.figures156-166" httpUri="https://binary.pensoft.net/fig/504647" pageId="0" pageNumber="1">159</figureCitation>
) and with short setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Pale brownish yellow (including stemmaticum); apex of antenna, hind tibia apically and subbasally and apex of ovipositor sheath rather brown; parastigma and vein 1-M yellow; clypeus, palpi, tegulae, remainder of legs, mesoscutum medio-posteriorly, first and second tergites and metasoma baso-ventrally pale yellowish or ivory; pterostigma (but basally and apically pale yellowish) and most veins brown; wing membrane slightly brownish below pterostigma.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="male">
<paragraph pageId="0" pageNumber="1">Male.</paragraph>
<paragraph pageId="0" pageNumber="1">
Similar to female (including hind femur and tibia: Fig. 
<figureCitation captionStart="Figures 152–155" captionStartId="F23" captionText="Figures 152 - 155. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype (but 153 - 155 of ♂, paratype) 152 habitus lateral 153 antenna 154 middle femur and tibia lateral 155 hind femur and tibia lateral." figureDoi="10.3897/zookeys..60426.figures152-155" httpUri="https://binary.pensoft.net/fig/504646" pageId="0" pageNumber="1">155</figureCitation>
), but antenna slenderer (Fig. 
<figureCitation captionStart="Figures 152–155" captionStartId="F23" captionText="Figures 152 - 155. Phanerotoma hellyeri van Achterberg, sp. nov., ♀, holotype (but 153 - 155 of ♂, paratype) 152 habitus lateral 153 antenna 154 middle femur and tibia lateral 155 hind femur and tibia lateral." figureDoi="10.3897/zookeys..60426.figures152-155" httpUri="https://binary.pensoft.net/fig/504646" pageId="0" pageNumber="1">153</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variations.</paragraph>
<paragraph pageId="0" pageNumber="1">Length of fore wing of ♀ 2.4-3.7 mm, of ♂ 2.2-3.2 mm; vein 2-SR usually evenly curved, but sometimes distinctly bent and parallel with posterior margin of pterostigma; vein 1-M of fore wing and parastigma pale yellowish, but sometimes more or less brown; pterostigma partly dark brown, largely or entirely pale yellowish; third tergite brown to rather dark brown; stemmaticum brownish yellow, only rarely darkened; apical half of antenna brownish yellow or dark brown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">United Arab Emirates, Yemen.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The new species is named after Peter Hellyer for his life-long research on the archaeology and ecology of the United Arab Emirates and his support of the series &quot;Arthropod Fauna of the UAE&quot;.</paragraph>
</subSubSection>
</treatment>
</document>